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Engraulis mordax Girard, 1854
CALIFORNIAN ANCHOVY
Northern Anchovy

Life   Vertebrata   Fish   Engraulidae   Engraulis

Engraulis mordax
© Copyright Photographer/SFTEP, 2002 · 0
Engraulis mordax

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Engraulis mordax
www.fao.org Copyright Michel Lamboeuf · 0
Engraulis mordax
español

Overview
Main identification features
  • top jaw reaches onto preoperculum
  • lower jaw tip under nostril
  • anal fin origin under en dorsal fin dorsal fin
Slender, elongate, rounded; snout long and sharply pointed; top jaw medium length, reaching onto preopercle, tip sharply pointed; tip of bottom jaw reaching under nostril; no large canine teeth; membrane between gill covers not expanded at rear; third gill arch without rakers; 37-45 rakers on lower limb of first gill arch; dorsal fin origin in mid-body; anal fin base short, 19-23 rays, origin under or behind last dorsal ray; pectoral short, not reaching pelvics.

Silver stripe along flank in young fish only.

Size: 25 cm.

Habitat: marine, coastal pelagic, usually within 30km of shore.


Depth: 0-310 m.

Temperate; British Colombia to the tip of Baja, most of the western and the central eastern Gulf of California.


Attributes
Abundance: Common.
Cites: Not listed.
Climate Zone: North Temperate (Californian Province &/or Northern Gulf of California); Northern Subtropical (Cortez Province + Sinaloan Gap).
Depth Range Max: 310 m.
Depth Range Min: 0 m.
Diet: zooplankton; phytoplankton; pelagic fish larvae; pelagic fish eggs.
Eastern Pacific Range: Northern limit=51; Southern limit=24; Western limit=-128; Eastern limit=-110; Latitudinal range=27; Longitudinal range=18.
Egg Type: Pelagic; Pelagic larva.
Feeding Group: Planktivore.
FishBase Habitat: Pelagic.
Global Endemism: TEP non-endemic; East Pacific endemic; All species.
Habitat: Water column.
Inshore Offshore: Inshore; Inshore Only.
IUCN Red List: Not evaluated / Listed.
Length Max: 25 cm.
Regional Endemism: Continent; Temperate Eastern Pacific, primarily; Tropical Eastern Pacific (TEP) non-endemic; California province, primarily; Continent only; All species.
Residency: Vagrant.
Salinity: Marine; Marine Only.
Water Column Position: Near Surface; Surface; Water column only;


Names
Scientific source:

Links to other sites

References
  • De la Cruz , J. , Galvan , F. , Abitia , L. A. , Rodriguez , J. and Gutierrez, F. J., 1994., Lista sistematica de los peces marinos de Bahia Magdalena, Baja California Sur (Mexico). Systematic List of marine fishes from Bahia Magdalena, Baja California Sur (Mexico)., Ciencias Marinas, 20:17-31.
  • Eschmeyer , W. N. , Herald , E. S. and Hamman, H., 1983., A field guide to Pacific coast fishes of North America from the Gulf of Alaska to Baja California. Peterson Field Guide Ser. 28., Houghton Mifflin:336pp.
  • Findley, L.T., Hendrickx, M.E., Brusca, R.C., van der Heiden, A.M., Hastings, P.A., Torre, J., 2003., Diversidad de la Macrofauna Marina del Golfo de California, Mexico., CD-ROM versión 1.0. Projecto de la Macrofauna del Golfo .  Derechos reservados de los autores y Conservación Internacional.
  • Fischer , W. , Krup , F. , Schneider , W. , Sommer , C. , Carpenter , K. E. and Niem, V. H., 1995., Guia FAO para la Identificacion de Especies de para los fines de la Pesca. Pacifico Centro-Oriental. Volumen II. Vertebrados - Parte 1., FAO2:647-1200.
  • Fowler, H.W., 1944., Results of the Fifth George Vanderbilt Expedition (1941) (Bahamas, Caribbean sea, Panama, Galapagos Archipelago and Mexican Pacific Islands). The Fishes., Acad. Nat. Sci. Philadel., Monographs, 6:57-529.
  • Galván-Magaña, F., Gutiérrez-Sánchez, F., Abitia-Cárdenas, L.A., Rodríguez-Romero, J., 2000., The distribution and affinities of the shore fishes of the Baja California Sur lagoons. In Aquatic Ecosystems of Mexico: Status and Scope. Eds. M. Manuwar, S.G. Lawrence, I.F. Manuwar & D.F. Malley. Ecovision World Monograph Series., Backhuys Publishers:383-398.
  • Girard, C., 1854., Descriptions of new fishes, collected by Dr. A. L. Heermann, naturalist attached to the survey of the Pacific Railroad Route, under Lieut. R. S. Williamson, U. S. A., Proc. Acad. Nat. Sci. Phila., 7:129-140.
  • Grant , W. S. and Bowen, B. W., 1998., Shallow population histories in deep evolutionary lineages of marine fishes: Insights from sardines and anchovies and lessons for conservation., Journal of Heredity, 89:415-426.
  • Love, M.S., Mecklenburg, C.W., Mecklenburg, T.A., Thorsteinson, L.K., 2005., es of the West Coast and Alaska: a checklist of North Pacific and Artic Ocena species from Baja California to the Alaska-Yukon border., U.S. Department of the Interior, U.S. Geological Survey, Biological Resources Division, 288pp.
  • Osburn , R. C. and Nichols, J. T., 1916., Shore Fishes Collected by the 'Albatross' Expedition in Lower California with Descriptions of New Species., Bull. Amer. Mus. Nat. Hist., 35:139-181.
  • Whitehead , P.J.P. , Nelson , G.J. and Wongratana, T., 1985., Clupeoid fishes of the world (Suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, anchovies and wolf-herrings. Part 2. Engraulididae. FAO species catalogue. Vol. 7., FAO Fish. Synop. No 125, 125(2):305-579.

Acknowledgements

I thank Ashley MacDonald and John Pickering, University of Georgia, for technical support in building this page.


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http://www.fishbase.org/Summary/speciesSummary.php?genusname=Engraulis&speciesname=mordax ---> http://192.134.151.83/Summary/speciesSummary.php?genusname=Engraulis&speciesname=mordax
http://192.134.151.83/Summary/speciesSummary.php?genusname=Engraulis&speciesname=mordax ---> https://fishbase.mnhn.fr/Summary/speciesSummary.php?genusname=Engraulis&speciesname=mordax
https://fishbase.mnhn.fr/Summary/speciesSummary.php?genusname=Engraulis&speciesname=mordax ---> https://fishbase.mnhn.fr/summary/Engraulis-mordax.html
Engraulis mordax, Californian anchovy : fisheries, bait
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Common name (e.g. trout)

Genus + Species (e.g. Gadus morhua)

Engraulis mordax Girard , 1854

Californian anchovy Add your observation in Fish Watcher
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Engraulis mordax   AquaMaps   Data sources: GBIF OBIS
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Pictures | Google image Image of Engraulis mordax (Californian anchovy) Engraulis mordax
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Classification / Names Common names | Synonyms | Catalog of Fishes( genus , species ) | ITIS | CoL | WoRMS | Cloffa

Teleostei (teleosts) > Clupeiformes (Herrings) > Engraulidae (Anchovies) > Engraulinae
Etymology: Engraulis: Greek, eggraulis, -eos = anchovy (Ref. 45335 ) ;   mordax: Genus name from the Greek 'engraulis' meaning anchovy; species name from the Latin 'mordax' meaning biting (Ref. 27436 ) .
More on author: Girard .

Environment: milieu / climate zone / depth range / distribution range Ecology

Marine; pelagic-neritic; depth range 0 - 310 m (Ref. 96339 ), usually ? - 219 m (Ref. 54433 ). Subtropical; 51°N - 21°N, 131°W - 108°W (Ref. 54433 )

Distribution Countries | FAO areas | Ecosystems | Occurrences | Point map | Introductions | Faunafri

Northeast Pacific: northern Vancouver Island south to Cape San Lucas, Baja California, Mexico. Two subspecies recognized: Engraulis mordax mordax from British Columbia to Baja California and Engraulis mordax nanus in Bays of California.

Length at first maturity / Size / Weight / Age

Maturity: L m 9.6   range ? - ? cm
Max length : 24.8 cm SL male/unsexed; (Ref. 27436 ); common length : 15.0 cm TL male/unsexed; (Ref. 9988 ); max. published weight: 68.00 g (Ref. 56527 ); max. reported age: 7 years (Ref. 6884 )

Short description Morphology | Morphometrics

Dorsal spines (total): 0; Dorsal soft rays (total): 14-19; Anal spines : 0; Anal soft rays : 19 - 26; Vertebrae : 43 - 47. Snout quite sharply pointed; maxilla moderate, tip sharply pointed, reaching to or almost to hind border of pre-operculum, projecting well beyond tip of second supra-maxilla; tip of lower jaw below nostril. gill rakers slender, long; absent on hind face of third epibranchial. Anal fin origin under about base of last dorsal fin ray. A silver stripe along flank, disappearing with age.

Biology     Glossary (e.g. epibenthic)

Usually found in coastal waters within about 30 km from shore, but as far out as 480 km, forming large, tightly packed schools. Enters bays and inlets. Feeds on euphausiids, copepods and decapod larvae, both by random filter-feeding and by 'pecking' at prey. Oviparous, epipelagic batch spawner (Ref. 6882 ). Spawns throughout the year, peaking once (Ref. 6882 ). Processed into fishmeal, used as bait for tuna, occasionally canned (Ref. 9298 ).

Life cycle and mating behavior Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae

Spawns from British Colombia south to Magdalena Bay, Baja California, but most abundantly between Point Conception and Point San Juanico. There are two major spawning areas: 1) off southern California and northern Baja California and 2) off central and southern Baja California. Oviparous (Ref. 35602 ). Spawns either in inlets or offshore, throughout the year but mainly in winter and early spring, depending on hydrological conditions (preferably at 10 to 23.3° C in upper water layers and around 22.00 hours). Spawning occurs at night between 2000 and 0400 hour (Ref. 6882 ).

Main reference Upload your references | References | Coordinator | Collaborators

Whitehead, P.J.P., G.J. Nelson and T. Wongratana , 1988. FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (Suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. FAO Fish. Synop. 125(7/2):305-579. Rome: FAO. (Ref. 189 )

IUCN Red List Status (Ref. 130435 )

  Data deficient (DD)  ; Date assessed: 08 May 2020

CITES

Not Evaluated

CMS (Ref. 116361 )

Not Evaluated

Threat to humans

  Harmless





Human uses

Fisheries: commercial; bait: usually FAO - Fisheries: landings , species profile ; Publication: search | FishSource | Sea Around Us

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Internet sources

AFORO (otoliths) | Alien/Invasive Species database | Aquatic Commons | BHL | Cloffa | BOLDSystems | Websites from users | Check FishWatcher | CISTI | Catalog of Fishes : genus , species | DiscoverLife | ECOTOX | FAO - Fisheries: landings , species profile ; Publication: search | Faunafri | Fishipedia | Fishtrace | GenBank: genome , nucleotide | GloBI | Google Books | Google Scholar | Google | IGFA World Record | MitoFish | OceanAdapt | Otolith Atlas of Taiwan Fishes | Public aquariums | PubMed | Reef Life Survey | Socotra Atlas | Tree of Life | Wikipedia: Go , Search | World Records Freshwater Fishing | Zoobank | Zoological Record

Estimates based on models

Preferred temperature (Ref. 123201 ): 7.6 - 14.9, mean 9 °C (based on 55 cells). Phylogenetic diversity index (Ref. 82804 ):  PD 50 = 0.5020   [Uniqueness, from 0.5 = low to 2.0 = high]. Bayesian length-weight: a=0.00550 (0.00348 - 0.00867), b=3.08 (2.94 - 3.22), in cm total length, based on LWR estimates for this species & Genus-body shape (Ref. 93245 ). Trophic level (Ref. 69278 ):  3.1   ±0.28 se; based on food items. Generation time: 2.3 (1.6 - 2.6) years. Estimated as median ln(3)/K based on 17 growth studies. Resilience (Ref. 120179 ):  Medium, minimum population doubling time 1.4 - 4.4 years (rm=0.36; K=0.2-0.6; tm=1-4; tmax=7; Fec=4000). Prior r = 0.50, 95% CL = 0.33 - 0.75, Based on 2 stock assessments. Fishing Vulnerability (Ref. 59153 ):  Low vulnerability (24 of 100). Climate Vulnerability (Ref. 125649 ):  Low vulnerability (22 of 100). Price category (Ref. 80766 ):   Low . Nutrients (Ref. 124155 ):  Calcium = 159 [69, 355] mg/100g; Iron = 1.23 [0.60, 2.84] mg/100g; Protein = 18.6 [17.4, 19.9] %; Omega3 = 0.841 [0.514, 1.405] g/100g; Selenium = 20.5 [9.8, 42.9] μg/100g; VitaminA = 13.9 [3.2, 56.6] μg/100g; Zinc = 1.01 [0.65, 1.56] mg/100g (wet weight);

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