Reprinted with permission of the American Entomological Society from:
LaBerge, W. E. 1971b. A revision of the bees of the genus Andrena of the Western Hemisphere. Part IV. Scrapteropsis, Xiphandrena, and Rhaphandrena. Transactions of the American Entomological Society 97: 441-520.
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This small species is known from only a few specimens from New Jersey and Pennsylvania. A. daeckei can be easily recognized by the form of the mouthparts in both sexes. The galeae are opaque, the surfaces dulled by coarse, granular tessellation, and covered by short blunt erect hairs, and the maxillary and labial palpi are reduced in length. Of the Scrapteropsis only A. kalmiae has mouthparts of this type in addition to daeckei and these two Andrena are closely related. Atwood (1934), Viereck (1907) and Mitchell (i960) all missed seeing these mouthpart characters or did not realize their uniqueness and thus had some difficulty in separated daeckei from other species of Scrapteropsis (or Trachandrena).
A. daeckei is very close to A. kalmiae and whether or not these two forms should remain as distinct species or be considered as well-marked races of one species cannot be decided at present. They are here kept as distinct species until such time as additional specimens provide us with more conclusive evidence. A. daeckei can be separated from kalmiae by the shorter vertex (at least in females), the sparser mesoscutal punctures and the longer apical areas of terga 2 - 4 of the female.
FEMALE. MEASUREMENTS AND RATIOS. — N = 6: length, 7-8 mm; width, 3 mm; wing length, M = 2.51 ± 0.187 mm; FL/FW, M = 1.08 ± 0.006; FOVL/FOVW, M = 3.30 ± 0.112.
INTEGUMENTAL COLOR. — Black except as follows: Mandible with apical half to almost entirely rufescent; flagellar segments 3-10 reddish-brown below; wing membranes hyaline, slightly infumate. veins dark brown; terga 1-4 with extremely narrow apical margins hyaline, apical areas slightly rufescent; sterna hyaline apically; distitarsi rufescent.
STRUCTURE. — Antennae as in imitatrix except flagellar segment 3 shorter than 4 and quadrate. Eyes each about four times as long as broad, inner margins parallel. Malar space and mandible as in imitatrix. Galea broad. surface opaque, dulled by dense, coarse, granular tessellation. Maxillary palpus short, extended forward surpasses galea by little more than last segment. segmental ratio about 0.9: 1.0: 0.6: 0.6: 0.3: 0.4. Labial palpus short. especially last 3 segments reduced, ratio about 1.0: 0.4: 0.3: 0.2. Labral process trapezoidal, weakly emarginate apically: labrum apically without transverse sulcus. punctate. Clypeus rounded, rather long. densely punctate, punctures round, separated mostly by less than half a puncture width. without median impunctate line. surface shiny. Supraclypeal area with minute crowded punctures scarcely dulling surface. Genal area broader than eye in profile (about as 3: 4), with small but distinct punctures and fine reticular shagreening dulling surface, without rugulae: under surface of head without rugulae, with scattered punctures. Vertex short, above lateral ocellus equal to distinctly less than one ocellar diameter, surface dulled by small punctures and tessellation. Face above antennal fossae with well-formed longitudinal rugae and sparse interrugal punctures. Facial fovea extremely long, broad.
Pronotum as in imitatrix but punctures more distinct. Mesoscutum with punctures small, extremely sparse, posteromedially separated by five puncture widths or more, in anterior fourth rather irregularly spaced but separated by half to three puncture widths; surface dulled by fine reticular shagreening. Scutellum sparsely punctate except peripherally, punctures larger than on mesoscutum, surface shiny. Metascutum as in imitatrix. Propodeum as in imitatrix but dorsolateral and posterior surfaces coarsely rugatulopunctate and lateral (corbicular) surface rugulate at least in posterior half to three-fifths. Wing venation as in imitatrix but pterostigma smaller, only slightly broader than from inner margin prestigma to wing margin. Hind femur without ventral carinae; hind tibia narrow, broadest at apex.
Metasomal tergum 1 with basal area punctures separated mostly by two to four puncture widths, apical area punctures separated mostly by half to one puncture width, surface shiny, slightly shagreened if at all. Tergum 2 with apical area distinctly longer than one-third length of tergum; basal area punctures as in tergum 1, apical area punctures slightly sparser than on tergum 1 especially basad, surfaces moderately dulled by distinct reticular shagreening. Terga 3 and 4 similar to 2 but punctures progressively sparser and shagreening denser. Pygidial plate and sterna as in imitatrix.
VESTITURE. — Generally white except thoracic dorsum and vertex pale ochraceous. Form and disposition of hairs as imitatrix except as follows: galeae and maxillary palpi with abundant, extremely short, erect hairs; propodeal corbicula without plumose hairs anteriorly, internal hairs simple; terga 2-4 with apical pale fasciae absent or weak, usually broadly interrupted medially if present.
MALE. MEASUREMENTS AND RATIOS. — N = 2: length. 7 mm; width, 2 mm; wing length, 2.35-2.55 mm; FL/FW, 1.04-1.13; FS1/FS2, 0.89-1.00.
INTEGUMENTAL COLOR. — Generally as in female but terga 2-5 more broadly hyaline apically, apical areas translucent yellow to dark red basad of hyaline margins; tegulae rufescent.
STRUCTURE. — Antennae short, not surpassing scutellum in repose; scape as in imitatrix; flagellar segments as in imitatrix but segment 1 may be as long as 2. Eyes. mandible, malar space and galea as in female. Maxillary palpus as in female but segmental ratio 1.0: 0.9: 0.7: 0.5: 0.3: 0.5. Labial palpus as in female but ratio 1.0: 0.6: 0.3: 0.3. Labral process and labrum as in female. Clypeus as in female but puncture smaller. Supraclypeal area. genal areas, face above antennae and vertex as in female. Facial fovea absent.
Thoracic sculpturing as in female except as follows: propodeum with lateral surfaces coarsely rugatulopunctate (like dorsolateral and posterior surfaces). Wing venation as in female. Hind tibia narrow, not inflated subapically. broadest near apex.
Metasomal tergum 1 with basal area punctures small, sparse, separated mostly by five puncture widths or more. apical area punctures separated mostly by two to three puncture widths, surface shiny. Terga 2-5 similar to tergum 1 but surfaces at least lightly shagreened (shagreening progressively more coarse posteriorly). Sterna much as in imitatrix, sternum 6 flat and emarginate apically.
Terminalia as in figures 37-41; note the following: sternum 7 with only 2 or three minute hairs apically, emargination deep. v-shaped: sternum 8 with apical narrow part broadened basad: gonostyli sparsely haired: gonocoxite dorsal lobe long: penis valve tip blunt.
Venture. — White to cinereous: tergal fasciae absent. Galeae and maxillary and labial palpi with abundant short hairs as in female.
Reprinted with permission from: Mitchell, T.B. 1960. Bees of the Eastern United States. North Carolina Agricultural Experiment Station Technical Bulletin No. 141.
FEMALE. — Length 8 mm.; face somewhat longer than broad; clypeus very slightly convex, projecting nearly one-half below suborbital line, very closely, deeply and rather finely punctate, punctures almost crowded; facial foveae broad, occupying above most of area between eyes and ocelli, only slightly narrowed below, covered with whitish tomentum; space between margin of vertex and lateral ocelli subequal to their diameter; cheeks subequal to eyes in width, rounded posteriorly, somewhat shining, very closely and finely punctate; malar space distinct, but quite short; basal segment of flagellum slightly shorter than 2nd and 3rd combined; process of labrum short and rather broad, somewhat narrowed to the truncate and slightly emarginate tip; pubescence of head, thorax and legs short, whitish-ochraceous, becoming somewhat more yellowish on mid tibiae apically; punctures of scutum fine but rather widely separated anteriorly, becoming quite sparse near center posteriorly, surface somewhat shining, those on scutellum somewhat more coarse and irregular, well separated, the surface shining; pleura rather coarsely rugose above, becoming somewhat more finely so below and posteriorly; dorsal area of propodeum oblique, triangle rather broad, quite coarsely and regularly striate; propodeal corbicula very poorly developed, short, whitish, without an anterior fringe; trochanteral floccus white, short and poorly developed; hind tibiae not much broadened apically, apex only very slightly broader than basitarsi, scopa dense, hairs rather short, simple, whitish, becoming somewhat fuscous toward base posteriorly; front and mid basitarsi slightly narrower than their respective tibiae; 2nd submarginal cell much shorter than 3rd, receiving 1st recurrent near apex; abdominal terga smooth, somewhat shining, quite deeply and rather closely and finely punctate, apical impressed area quite shallow, occupying medially about one-third length of disc, narrowly yellowish-hyaline along rims, discal pubescence very short, thin and erect, entirely pale, forming very thin, whitish, apical fasciae on terga 2-4, this somewhat interrupted on tergum 2 medially, tergum 5 with a brownish-ochraceous, apical fimbria.
DISTRIBUTION. — Maine to New Jersey; May. No detailed description of daeckei has ever been published, as it was established originally merely by including it in a key. Thus the description above, made from a paratype female, is the first. Both sexes were included originally, but there is considerable doubt concerning the identity of the male. Specimens identified as such that have been seen are very similar to kalmiae Atwood or to A. (Mimandrena) imitatrix Cresson. Consequently that sex has been omitted here.
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