Reprinted by permission of the Regents of the University of California from:
Donovan, B. J. 1977. A revision of North American bees of the subgenus Cnemidandrena (Hymenoptera: Andrenidae). University of California Publications in Entomology: 1-107.
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A. ramaleyi may be separated from other species of the Citrinihirta group by the vestiture, white or nearly white with the wings nearly hyaline, the metasomal terga with abundant vestiture, and the antennae yellow at least ventrally.
FEMALE. MEASUREMENTS AND RATIOS. — -N = 11; length, 7.0-8.2 mm; width, 2.4-2.8 mm; wing length, M = 2.29 ± 0.104 mm; FL/FW, M = 1.0 ± 0.017; FOVL/FOVW, M = 3.12 ± 0.237.
INTEGUMENTAL COLOR. — Black except as follows: antenna ventrally with apex of flagellar segments 1 and 2 yellow, remaining flagellar segments bright yellow, with only a narrow dorsal area black to flagellar segment 7; mandible with apical half red; tibial spurs white tinged with yellow; tarsi except basitarsi and basal half of tibial claws bright yellow, apical half of tibial claws red; tegula hyaline light brown; wing veins and pterostigma moderately yellow-brown; wing membrane very faintly clouded apically, remainder hyaline; metasomal terga with apical, depressed area hyaline; tergum 1 with lateral reflexed area hyaline, pygidial plate brown-black apically, red-yellow basally; sterna widely hyaline apically.
STRUCTURE. — Scape equal to flagellar segments 1 to 3 plus one-third of flagellar segment 4; flagellar segment 1 slightly longer than flagellar segments 2 plus 3; flagellar segment 3 equal to flagellar segment 4. Eye more than 4 times longer than wide, with inner margin diverging moderately above except for upper end slightly incurved. Malar space 8 times wider than long. Mandible in repose extends for one-fourth its length or slightly more beyond middle of labrum, with notch of mandible tooth slightly more than one-third distance between middle of labrum and mandible apex from middle of labrum. Galea weakly to moderately tessellate, dorsal surface one-third wider than lateral surface. Maxillary palpus with apical segment extending beyond apex of galea, segmental ratio 0.7:0.7:0.6 :0.5:0.5:0.6. Labial palpus with basal segment laterally compressed and curved, apical segment attached subapically, segmental ratio 1.0:0.6:0.4:0.6. Labral process slightly reflexed apically, slightly more than twice as wide as long, apical emargination depth one-fourth length of labral process. Clypeus moderately protuberant, extending below eyes for two-fifths its length, with large punctures except for small, impunctate, ventromedian area; punctures separated by 1 diameter or less medially, punctures smaller, closer laterally and dorsally, very light shagreening on narrow upper periphery, remainder shiny. Supraclypeal area with small, irregular punctures and very light shagreening. Genal area wider than eye with medium-sized punctures irregularly spaced, third nearest eye shiny, remainder with moderate shagreening. Vertex equal to 1 ocellar diameter with medium-sized to large deep punctures closely spaced, with light shagreening, nearly shiny. Supra-antennal area with irregular, longitudinal rugulae and small to medium-sized punctures separated by 1 to 2 diameters. Facial fovea slightly more than 3 times longer than wide, wider above than below, rounded above and below, reaching above to a line above median ocellus, reaching below to a line equidistant between antennal fossa and clypeus.
Pronotum with humeral angle well developed, humeral ridge very well developed, humeral ridge not depressed where reached by indistinct, pronotal suture, pronotum dorsally with medium-sized punctures separated by 1 to 2 diameters, posterior margin of pronotum moderately shagreened, remainder shiny. Scutum anteriorly with medium-sized punctures separated by 1 to 2 diameters, dorsal area with very few, scattered punctures, very shiny. Scutellum almost impunctate, smooth, shiny. Metanotum with very few punctures, shagreened laterally, almost smooth and shiny medially. Propodeal enclosure with a broad, median, longitudinal, moderately tessellate depression, tessellation roughened anteriorly; dorsoposterior surface with nearly large punctures separated by 2 to 4 diameters, with light shagreening; corbicular area with widely spaced, medium-sized punctures in anterior third, smooth, shiny. Mesepisternum with large punctures separated by 1 diameter, with light shagreening in anterior two-thirds, remainder shiny. Metepisternum impunctate, smooth, shiny. Mesobasitarsus widest medially, more than 3 times longer than wide, slightly wider than metabasitarsus. Metabasitarsus widest medially in basal third, 4 times longer than wide, slightly more than two-thirds as wide as metatibia. Metatibia widest apically, 3.3 times longer than wide. Wing with pterostigma 4 times longer than wide, vein 1st m-cu meets second submarginal cell one-third distance from apex of cell. Metasomal tergum 1 with medium-sized punctures except for very small, median, impunctate area and depressed, apical margins, punctures separated by 1 to 2 diameters, lightly shagreened; terga 2 to 4 similar except punctured throughout with punctures closer on apical margins of discs; tergum 5 with very large, deep punctures separated by less than 1 diameter, moderately shagreened. Pygidial plate rounded apically, median area and wide lateral areas raised, posterior half weakly reflexed, smooth, anterior half of median area with small, contiguous punctures. Sternum 1 with very small, median emargination, with medium-sized punctures separated by 1 diameter except for anterior median area and impunctate hyaline apex.
VESTITURE. — Light white-yellow throughout. Clypeus with sparse hairs medially. Scutum, scutellum and metanotum almost naked on shiny areas. Propodeal corbicula with scattered, plumose hairs in anterior third. Trochanteral floccus imperfect.
Metasomal terga 1 to 4 with apical fasciae of recumbent hairs obscuring half or more of terga, fascia of tergum 1 less dense with hairs more erect, not wholly obscuring surface, with long, erect hairs on remaining area except for small, median, impunctate area; terga 2 to 4 with shorter, erect hairs medially on discs, long, erect hairs laterally. Tergum 5 almost obscured by dense, reclining hairs. Sternum 1 with long, erect hairs. Sterna 2 to 5 with long, nearly erect, subapical fimbriae in outer thirds, less developed in median thirds; poorly developed apical fringes on lateral margins of apical areas; sterna 2 to 6 with short, erect hairs on punctured areas.
MALE. MEASUREMENTS AND RATIOS. — N = 3; length 7.0-8.0 mm, width 1.9-2.1 mm; wing length 2.20-2.28 mm; FL/FW, 0.91; FS1/FS2, 0.77.
INTEGUMENTAL COLOR. — Black except as follows: antennae bright yellow ventrally beyond pedicel; mandible with about apical third red; tibial spurs as in female; tarsi and basal half of tarsal claws light yellow; apical half of tarsal claws red; tegula and wing veins as in female, wing membrane hyaline; metasomal terga with apex of disc narrowly light yellow, apical area hyaline, tergum 1 with lateral, reflexed area hyaline, sterna hyaline apically.
STRUCTURE. — Scape equal to flagellar segments 1 plus 2 plus half of flagellar segment 3; flagellar segment 1 slightly longer than flagellar segment 2, equal to flagellar segment 3. Eye less than 4 times longer than wide, with inner margin diverging moderately above except for upper end slightly incurved. Malar space and mandible as in female, except mandible without ventrobasal lamella. Galea weakly tessellate, surface nearly shiny. Maxillary palpus as in female except segmental ratio 0.7:0.7:0.5:0.6:0.6:0.7. Labial palpus as in female except basal segment not so strongly compressed laterally, segmental ratio 1.2:0.6:0.6:0.7. Labial process strongly reflexed, twice as wide as long, apical emargination depth one half of reflexed area. Clypeus as in female except punctures smaller, shagreening almost absent. Supraclypeal area with irregular, close punctures, slight shagreening. Genal area produced to a blunt prominence posteriorly opposite midpoint of eye; genal area twice as wide as eye, shiny. Vertex as in female. Supra-antennal area with irregular, longitudinal rugulae with medium-sized punctures separated by 1 diameter; parocular areas similarly punctured but rugulae lacking.
Pronotum with humeral angle and ridge well developed; humeral ridge slightly depressed where crossed by pronotal suture; humeral area lightly shagreened to shiny. Remainder of mesosoma sculptured much as in female.
Metasomal terga 1 to 4 with disc with small punctures 3 to 4 diameters apart, punctures closer on disc of tergum 5, terga 1-5 shiny. Sterna 1-6 as in female. Sternum 7 (fig. 88) with basal apodemes narrowing gradually, apex truncate, with lateral angles with long hairs. Sternum 8 (fig. 89) with apex and neck region a little longer than basal region, apex scarcely expanded, neck with abundant long hairs. Gonoforceps (figs. 85 to 87) in dorsal and ventral view expanded apically, in lateral view scarcely turned down apically. Penis valves in dorsal and ventral view widest medially, in lateral view long, scarcely turned down apically, projecting little below gonoforceps.
VESTITURE. — White or near white throughout. Face with long dense pubescence. Mesosomal pubescence long and dense except less dense on disc of scutum.
Metasomal terga 1-4 with apical fasciae of recumbent hairs almost obscuring half terga; remaining areas of terga 1-4 with long, more widely spaced, erect hairs, hairs shorter from terga 1 to 4. Tergum 5 similar to tergum 4 but hairs not forming a definite apical fascia. Sternum 1 with sparse, long hairs, sterna 2-5 apically with dense, long fringes, remaining areas with shorter, more widely spaced hairs.
VARIATION. — In females the facial foveae vary in width in about the dorsal half. In both sexes the degree of shagreening of an area may vary slightly from one specimen to another.
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