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© Copyright source/photographer
· 5
Andrena sayi, m, face, St. Marys Co, MD |
© Copyright source/photographer
· 5
Andrena sayi, m, side, St. Marys Co, MD |
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© Rebekah Andrus Nelson
· 1
Andrena sayi, female, face |
© Copyright source/photographer
· 1
Andrena sayi, male, sternal plates 7 and 8 |
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Overview |
Reprinted with permission of the American Entomological Society from:
Bouseman, J. K., LaBerge, W. E. 1978. A revision of the bees of the genus Andrena of the Western Hemisphere. Part IX. Subgenus Melandrena. Transactions of the American Entomological Society 104: 275-390.
Please report text errors to: leah at discoverlife dot org.
Although this is a large conspicuous bee occupying a relatively large area of southeastern North America, it is relatively rare in collections. Both sexes of sayi can be distinguished from other members of this subgenus by the distinct but interrupted pale tergal bands and by the propodeal dorsal enclosure being relatively abruptly declivous to the posterior surface. The female lacks the labral median crista and sulci and the male has a relatively weak clypeal beard.
FEMALE. MEASUREMENTS AND RATIOS. — N = 20; length, 12-15 mm; width, 3.0-4.5 mm; wing length, M = 5.22 ± 0.111 mm; FL/FW, M = 0.93 ± 0.004; FOVL/FOVW, M = 3.14 ± 0.042.
INTEGUMENTAL COLOR. — Black except as follows: mandible with basal half and often base rufescent; flagellar segments 3-10 dark reddish-brown below; tegula translucent, reddish-brown; wing membranes moderately infumate, yellow to brown at tips, veins reddish-brown, subcosta darker; terga hyaline apically, clear to reddish; sterna with apical areas hyaline, usually clear, basal areas usually rufescent; distitarsi and often hind basitarsi dark rufescent.
STRUCTURE. — Antennal scape length equals first two and one-half flagellar segments or slightly less; flagellar segment 1 about as long as segments 2 plus 3; segment 2 quadrate, shorter than 3; segment 3 shorter than 4; segments 4-9 about one and one-fourth times as long as broad. Eyes each almost four times as long as broad, inner margins diverging slightly toward mandibles. Mandible, galea and stripes as in carlini, but stripes punctate throughout. Maxillary palpus as in carlini but segmental ratio about as 0.9:1.0:0.9:0.9:0.9:0.9. Labial palpus as in carlini but ratio about as 1.0:0.6:0.6:0.6. Labral process large, usually entire, trapezoidal, shiny, rarely with weak apical emargination; labrum below process punctate except margin, without strong cristae or sulci. Clypeus with large round punctures irregularly separated by half to almost one puncture width, with narrow, usually complete, median, impunctate line; surface shiny except near base. Supraclypeal area and face above antennal fossae as in carlini but face with longitudinal rugulae finer. Facial fovea as in carlini. Vertex as in carlini. Genal area in profile only slightly broader than eye (about as 7:6), sculptured as in carlini.
Thoracic form and sculpturing as in carlini except as follows: mesoscutum with anterior half and peripherally with punctures separated mostly by half a puncture width, posteromedially punctures separated largely by two to three puncture widths, surface finely tessellate; scutellum with punctures sparse medially, finely tessellate to reticularly shagreened, often moderately shiny; propodeal dorsal enclosure finely punctatorugose throughout, abruptly declivous to posterior surface; dorsal and posterior surfaces outside of enclosure with fine crowded punctures and fine tessellation dulling surfaces; lateral surface propodeum without vermiculate rugulae posteriorly. Wing venation as in carlini but second submarginal cell usually narrower along anterior margin than along posterior margin.
Metasomal tergum 1 with basal area distinctly punctate, punctures separated largely by two puncture widths; apical area punctures separated by half to one puncture width; surface shiny to moderately shiny, with reticular shagreening. Tergum 2 with basal area punctures separated largely by one to two puncture widths, apical area with apical rim impunctate, otherwise with small punctures slightly denser than in basal area; surface moderately dulled to shiny, reticular shagreening fine. Terga 3 and 4 similar to 2 but progressively more sparsely punctate and more densely shagreened. Pygidial plate broad, with apex blunt or rounded. Sterna 2-5 with narrow apical areas impunctate and impunctate at extreme bases, otherwise relatively coarsely punctate, surfaces reticularly shagreened.
VESTITURE. — White to pale ochraceous except as follows: terga 5 and 6 with long hairs golden brown to yellow; inner surfaces hind tarsi yellow; basitibial plate and just below reddish-brown to pale brown. Thoracic dorsum with hairs relatively short and sparse, scarcely hiding surface. Metasomal terga 2-4 with distinct apical pale fasciae, interrupted medially usually by about one-third width of tergum, each lateral segment tapering toward midline. tibial scopal hairs while; pollen-collecting hairs normal for subgenus.
MALE. MEASUREMENTS AND RATIOS. — N = 20; length, 12-14 mm; width, 2.5-3.0 mm; wing length, M = 4.46 ± 0.223 mm; FL/FW, M = 0.90 ± 0.007; FS1/FS2, M = 1.00 ± 0.012.
INTEGUMENTAL COLOR. — Black except as follows: mandibles with apical third or more and extreme base rufescent; flagellar segments 2-11 dark reddish-brown to brown below; tegula translucent, ferrugineous; wing membranes hyaline, moderately infumate especially apically, yellowish, veins red to reddish-brown; metasomal terga with apical areas hyaline, colorless apically to yellow basally; sterna 2-5 with apical areas colorless to yellow, basal areas rufescent; tarsi and often tip of hind tibia rufescent; tibial spurs testaceous.
STRUCTURE. — Antennal scape length equals first two and one-third flagellar segments or slightly more; flagellar segment 1 about as long as segment 2, shorter than 3; segments 3-10 about one and one-half times as long as broad or slightly shorter. Eyes each almost four times as long as broad, inner margins diverging slightly toward mandibles. Mandible moderate in length, in repose surpassing midlabrum by about one-fourth length of mandible, bidentate. galea as in carlini. Maxillary palpus as in carlini but segmental ratio about as in female. Labial palpus as in carlini but ratio about as 1.0:0.7:0.6:0.7. Labral process shiny, trapezoidal, weakly emarginate apically; labrum apical to process shiny, with minute punctures. Clypeus with large round punctures separated mostly by less than half a puncture width, without median impunctate line but occasionally small subapical median area with punctures sparser, shiny except basally where shagreened. Supraclypeal area and face above antennal fossae as in carlini. Vertex above lateral ocellus equals about one and one-half ocellar diameters, dulled by punctures and dense tessellation. Genal area in profile somewhat broader than eye (about as 6:5), sculptured as in carlini.
Thoracic form and sculpturing as in female but dorsal and posterior surface of propodeum more coarsely punctate and lateral surface punctate and tessellate except row of several short weak rugulae along lower margin in posterior third. Wing venation as in female.
Metasomal terga 1-5 punctate as in female terga 1-4 but punctures sparser; tergum 2 with basal area punctures separated mostly by two to three puncture widths, apical area punctures smaller and less distinct; surfaces shiny to moderately shiny, reticular shagreening weak. Tergum 7 with narrow shiny pygidial area, apex rounded or acute. Sterna 2-5 as in carlini but punctures more abundant and surface usually moderately dulled by reticulotransverse shagreening. Sternum 6 with apical margin turned down, shallowly emarginate; surface punctate, without triangular area.
Terminalia as in figures 39-43; note penis valves with narrow tips and sternum 8 not modified as in carlini in lateral view.
VESTITURE. — White to pale ochraceous except as follows: dorsum of thorax often yellow; tergal segments 6 and 7 often with golden-brown to yellow hairs; inner surfaces hind tarsi pale yellow. Clypeal beard weak; thoracic dorsum with hairs long but relatively sparse; terga 2-5 with interrupted pale apical bands as in female terga 2-4; sterna without fimbriae; sternum 6 with abundant moderately long to short hairs directed mostly rearward.
Reprinted with permission from: Mitchell, T.B. 1960. Bees of the Eastern United States. North Carolina Agricultural Experiment Station Technical Bulletin No. 141.
FEMALE. — Length 13 mm.; face slightly longer than distance between eyes above; eyes nearly parallel; foveae broad, occupying above most of area between eyes and ocelli, covered with pale yellowish tomentum; vertex broad, lateral ocelli separated from its hind margin by about twice their diameter; cheeks slightly broader than eyes; clypeus rather strongly protuberant, protruding about one-half below suborbital line, smooth and shining, with rather fine but deep and distinct, evenly distributed punctures; malar space linear; process of labrum broad, entire, truncate, apex about equal to each oblique side; basal segment of flagellum about equal to 2nd and 3rd combined, median segments somewhat longer than broad; vertex and cheeks smooth but rather dull, with minute, rather evenly distributed punctures, area below ocelli finely striate; pubescence of head and thorax rather short and thin, entirely pale ochraceous, more dense and elongate laterad of antennae and on pleura; scutum and scutellum smooth but rather dull, punctures fine but quite distinct, close and more obscure on scutum anteriorly, becoming rather sparse in center posteriorly, scattered and irregular on scutellum; pleura dull, more or less granular, with very fine and indefinite, close punctures; dorsal area of propodeum finely granular or subrugose; propodeal corbicula well developed, dorsal hairs not extremely long, with a dense anterior fringe, and numerous hairs on interior; trochanteral floccus imperfect; hind tibial scopa and pubescence of legs in general ochraceous; hind tibial spurs relatively straight; wings lightly infuscated, somewhat darker toward apex, veins and stigma testaceous; 2nd submarginal considerably shorter than 3rd, stigma more ferruginous; tegulae piceous; abdominal terga somewhat shining between deep and distinct but rather close and fine punctures, these more sparse on basal tergum; apical margins of segments 2 and 3 rather narrowly and shallowly impressed, these areas well punctured; discal pubescence very short, erect, entirely pale ochraceous, apical fasciae evident laterally but not well developed, apical fimbria ochraceous.
MALE. — Length 10-11 mm.; face somewhat longer than distance between eyes; eyes subparallel; vertex broad, lateral ocelli separated from its hind margin by about twice their diameter; cheeks somewhat broader than eyes, rounded posteriorly (as in obscuripennis, fig. 21); clypeus rather strongly protuberant, protruding nearly one-half below suborbital line, smooth and shining, with rather close and fine, evenly distributed punctures; malar space equal to about one-third width of mandible at base; process of labrum subtriangular, quite large with truncate tip; mandibles slender, with a subapical inner tooth, overlapping nearly one-half; basal segment of flagellum longer than segment 2, but shorter than 2 and 3 combined; pubescence of head rather short and thin, entirely pale ochraceous; vertex and cheeks smooth but rather dull, with very fine punctures, these becoming quite close on cheeks below; pubescence of thorax entirely pale ochraceous, almost whitish, more ochraceous on dorsum; scutum and scutellum smooth but rather dull, punctures fine, rather close but obscure anteriorly, becoming sparse on scutum posteriorly and very fine and vague on scutellum; pleura smooth but dull, with very fine vague punctures; dorsal area of propodeum with a few fine basal striations; legs black, apical tarsal segments somewhat reddened, pubescence entirely pale; wings lightly infuscated, veins and stigma testaceous, 2nd submarginal slightly shorter than 3rd; tegulae testaceous-hyaline, somewhat darker anteriorly; abdominal terga smooth, somewhat shining, punctures fine, rather evenly distributed, well separated but not sparse, impressed apical margins hardly one-third of median length of segments 2-4, these becoming hyaline along apical rim, pubescence very short and inconspicuous, entirely pale, fasciae barely evident toward sides; sternum 7 with a deep, triangular, median emargination; sternum 8 slender and elongate apically, with a slight, subapical, ventral protuberance; penis valves of genital armature slender and elongate, the tip markedly attenuated, gonocoxites only very slightly dilated apically, the gonocoxal lobes not appreciably produced (armature similar to that of commoda, fig. 22).
DISTRIBUTION. — Tennessee and North Carolina, north to Minnesota, Michigan, New York, New Brunswick and the New England states; March to June.
FLOWER RECORDS. — Amelanchier, Aronia, Brassica, Cercis, Crataegus, Malus, Padus, Prunus, Pyrus, Rubus, Salix, Stellaria, Taraxacum and Vaccinium.
Described using the synonymous name: Andrena seviensis Mitchell
FEMALE. — Length 9 mm.; eyes parallel; foveae rather broad, occupying about two-thirds of space between eyes and ocelli; vertex very narrow, lateral ocelli separated from its hind margin by a space somewhat less than their diameter; cheeks very slightly broader than eyes; clypeus moderately convex, protruding about one-half below suborbital line, smooth and shining, punctures rather deep and distinct, well separated but not sparse medially, becoming somewhat closer at extreme sides; punctures of cheeks and vertex minute and obscure, surface smooth but rather dull; malar space evident but very short; process of labrum broadly rounded, rather large; basal segment of flagellum about equal to segments 2 and 3 combined, the more apical segments somewhat longer than broad; pubescence of head rather short and thin, entirely whitish, that on thorax somewhat more dense above and around tubercles and on propodeum, rather thin below; scutum and scutellum rather dull but smooth, punctures shallow and rather obscure, rather close on scutum anteriorly, becoming more sparse posteriorly, scutellum somewhat more shining, punctures irregularly scattered; pleura dull and tessellate but rather smooth, with shallow obscure punctures evident anteriorly and beneath; triangle of propodeum finely granular or subrugose, not sharply defined; propodeal corbicula poorly developed, dorsal hairs quite short, without an anterior fringe and with but a few, very fine, scattered hairs on interior; trochanteral floccus imperfect, some of the more apical hairs quite elongated; legs black, pubescence whitish in general, hind tibial scopa of elongate whitish hairs, tibia quite slender; hind tibial spurs yellowish, quite straight; wings subhyaline, veins and stigma testaceous to piceous, 2nd submarginal cell much shorter than 3rd, receiving 1st recurrent near middle; tegulae piceous, becoming somewhat brownish posteriorly; abdominal terga dull, posterior margins of 2nd and 3rd segments very broadly but shallowly impressed, these areas impunctate, basal portions of discs with rather deep and distinct, well separated punctures, but these hardly sparse except on basal tergum, interspaces on 3rd and 4th two or three times their diameter; discal pubescence hardly evident, apparently entirely pale, fasciae not developed, apical fimbria somewhat brownish.
TYPE. — Holotype: Female, Chilhowee Mts., Sevier County, Tenn., May 12, 1951 (A. C. Cole) [Univ. Tenn.]. The poorly developed propodeal corbicula suggests a relationship with the western Oligandrena, but it is so far outside the range of species of this group the writer is inclined to place it in or near Gymnandrena.
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Updated: 2024-04-24 15:48:35 gmt
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