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Andrena wellesleyana Robertson, 1897
Life   Insecta   Hymenoptera   Apoidea   Andrenidae   Andrena
Subgenus: Parandrena

Andrena wellesleyana FEM mm .x ZS PMax
© Copyright Laurence Packer 2014 · 7
Andrena wellesleyana FEM mm .x ZS PMax

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Andrena wellesleyana MALE CFP
© Copyright Laurence Packer 2014 · 7
Andrena wellesleyana MALE CFP
Andrena wellesleyana
© Copyright Micheal Veit 2010 · 6
Andrena wellesleyana

Andrena wellesleyana
© Copyright Micheal Veit 2010 · 6
Andrena wellesleyana
Andrena wellesleyana, f, back, Middlesex Co, MA ---.
© Copyright source/photographer · 5
Andrena wellesleyana, f, back, Middlesex Co, MA ---.

Andrena wellesleyana, f, face, Middlesex Co, MA ---.
© Copyright source/photographer · 5
Andrena wellesleyana, f, face, Middlesex Co, MA ---.
Andrena wellesleyana, f, side, Middlesex Co, MA ---.
© Copyright source/photographer · 5
Andrena wellesleyana, f, side, Middlesex Co, MA ---.

Andrena wellesleyana, m, back, Middlesex Co, MA ---.
© Copyright source/photographer · 5
Andrena wellesleyana, m, back, Middlesex Co, MA ---.
Andrena wellesleyana, m, face, Middlesex Co, MA ---.
© Copyright source/photographer · 5
Andrena wellesleyana, m, face, Middlesex Co, MA ---.

Andrena wellesleyana, m, side, Middlesex Co, MA ---.
© Copyright source/photographer · 5
Andrena wellesleyana, m, side, Middlesex Co, MA ---.
Andrena wellesleyana, figure52d
Mitchell, Bees of the Eastern United States, Vol. I, 1960 · 1
Andrena wellesleyana, figure52d

Andrena wellesleyana, figure55b
Mitchell, Bees of the Eastern United States, Vol. I, 1960 · 1
Andrena wellesleyana, figure55b
Andrena wellesleyana, female, 2 submarginal cells
© USDA Bee Biology and Systematics Laboratory, Logan Utah · 1
Andrena wellesleyana, female, 2 submarginal cells

Andrena wellesleyana, female, hummeral ridge interrupted by suture along dorsoventral ridge2
© USDA Bee Biology and Systematics Laboratory, Logan Utah · 1
Andrena wellesleyana, female, hummeral ridge interrupted by suture along dorsoventral ridge2
Andrena wellesleyana, male, face
© Rebekah Andrus Nelson · 1
Andrena wellesleyana, male, face
Overview
Reprinted with permission of the American Entomological Society from: LaBerge, W. E., Ribble, D. W. 1972. A revision of the bees of the genus Andrena of the Western Hemisphere. Part V: Gonandrena, Geissandrena, Parandrena, Pelicandrena. Transactions of the American Entomological Society 98: 271-358.

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This medium-sized widespread bee is related to andrenoides but not as closely as nida. The female resembles the dark female of andrenoides and was once considered as a race of andrenoides because of this resemblance. However, wellesleyana females are distinctive in the tergal pale bands being composed of longer hairs, the bases of terga 1 and 2 having long, more or less erect or suberect hairs at least medially and by the more protuberant midline of the clypeus. The male of wellesleyana is distinctive because the lateral toothlike angles of sternum 6 are bent forwards so as to be parallel with the surface of the sternum.

FEMALE. MEASUREMENTS AND RATIOS. N = 20; length, 9-11 mm; width, 2.5-3.5 mm; wing length, M = 3.92 0.155 mm; FL/FW, M = 0.95 0.004; FOVL/FOVW, M = 3.45 0.050.

INTEGUMENTAL COLOR. Generally as in dark specimens of andrenoides but wing membranes almost colorless.

STRUCTURE. Antenna as in andrenoides but scape length equal to first three and one-half flagellar segments or slightly more. Eyes and mandible as in andrenoides. Galea as in andrenoides but surface dull, densely shagreened. Labial palpus as in andrenoides but segmental ratio about as 1.0:0.8:0.6:0.8. Maxillary palpus as in andrenoides but ratio about as 1.0:0.9:0.7:0.7:0.7:0.7. Labral process and labrum as in andrenoides but process always sharply pointed apically. Clypeus as in andrenoides but median longitudinal impunctate line strongly protuberant especially near apex where a shiny boss usually developed, entire clypeus appears protuberant in lateral view, midline area shiny, unshagreened, elsewhere dulled by fine reticular shagreening. Supraclypeal area, face, facial fovea and coarse tessellation.

Pronotum as in andrenoides. Mesoscutum as in andrenoides but surface opaque, dulled by dense, regular tessellation. Scutellum and mesepisternum as in andrenoides. Propodeum as in andrenoides but dorsal enclosure often slightly granular especially basally. Wing venation and tibial spurs as in andrenoides.

Metasomal terga 1-4 with minute punctures separated by 3 to 5 puncture widths, surfaces moderately shiny to dull, reticularly shagreened. Pygidial plate as in andrenoides but internal triangle usually lacking apical ridge connecting to apex of plate or this ridge poorly developed. Sterna as in andrenoides.

VESTITURE. Generally white to pale ochraceous and distributed as in andrenoides except as follows: terga 2-4 with complete pale apical fasciae of long decumbent hairs, occasionally narrowly interrupted medially on tergum 2; tergum 1 with pale apical fascia restricted to small lateral patches (occasionally worn away) usually longer than in andrenoides; 1 and 2 with basal areas with conspicuous, long, suberect, plumose hairs especially medially.

MALE. MEASUREMENTS AND RATIOS. N = 20; length, 8-11 mm; width, 2-3 mm; wing length, 3.73 0.129 mm; FL/FW, M = 0.95 0.006; FS1/FS2, M = 2.00 0.035.

INTEGUMENTAL COLOR. As in andrenoides but mandible lacks basal yellow spot and wing membranes almost colorless.

STRUCTURE. Antennae as in andrenoides but flagellar segment 1 distinctly shorter than segments 2 plus 3 and segment 3 longer than broad. Eyes each about three times as long as broad, inner margins converging slightly towards mandibles. Mandible and galea as in andrenoides but galea dull, reticularly shagreened. Labial palpus as in andrenoides but segmental ratio about as 1.0:0.7:0.6:0.6. Maxillary palpus as in andrenoides but ratio about as 1.0:1.0:0.8:0.8:0.8:0.8. Labral process usually rather sharply triangular, small; labrum apical to process slightly depressed, shiny. Clypeus, supraclypeal area, face and vertex as in andrenoides. Genal area as in andrenoides but punctures denser, surface dulled by fine tessellation.

Pronotum as in andrenoides. Mesoscutum as in andrenoides but surface opaque, dulled by dense tessellation. Scutellum, mesepisternum and propodeum as in andrenoides but dorsal enclosure of propodeum usually granular at least basally. Wing venation and tibial spurs as in andrenoides.

Metasomal terga sculptured as in female but apical areas almost impunctate. Sterna with basal areas with small punctures separated by half to one puncture width except in basal third where sparse and absent in apical area. Sternum 6 with lateral teeth completely reflexed so as to be directed anteriorly parallel to surface, teeth widely separated and apical margin between teeth entire. Terminalia as figured (Figs. 48-52); note the following: terminalia large; gonocoxite with dorsal process rounded apically; penis valves with tips slightly expanded dorsoventrally to form a small downwardly directed knob.

VESTITURE. White to pale ochraceous; distributed as in andrenoides except as follows: terga 1 and 2 with basal areas with abundant long erect to suberect hairs (especially medially on tergum 2); terga 2-5 with apical pale fasciae complete or almost so (unless worn).



Reprinted with permission from: Mitchell, T.B. 1960. Bees of the Eastern United States. North Carolina Agricultural Experiment Station Technical Bulletin No. 141.

FEMALE. Length 10 mm.; clypeus broadly convex, projecting slightly less than one-half below suborbital line, somewhat shining, with a very narrow and imperfect, median, impunctate line, closely and deeply but rather finely punctate on each side; facial foveae rather broad, occupying above most of space between eyes and ocelli (as in nida, fig. 49), covered with pale ochraceous tomentum; space between margin of vertex and lateral ocelli equal to about half their diameter; cheeks slightly broader than eyes, rounded posteriorly, quite densely pubescent, rather dull, punctures minute and obscure, hardly visible; malar space very short; basal segment of flagellum subequal to 2nd and 3rd combined; process of labrum small, rather broadly subtriangular (as in nida, fig. 49); mandibles with a low sub-basal ventral dilation; pubescence of head and thorax pale ochraceous above, becoming more whitish below, that on legs more whitish on the basal segments, becoming somewhat more ochraceous apically; thoracic integument dull, finely tessellate, punctures of scutum very fine and obscure, rather close anteriorly, becoming quite sparse near center posteriorly, those on scutellum very minute and obscure, irregularly scattered, the surface more shining; punctures of pleura minute and obscure, largely hidden by the rather dense pubescence; dorsal area of propodeum rather narrow, oblique, triangle finely tessellate; propodeal corbicula well developed but rather short, with a distinct anterior fringe, pale ochraceous; trochanteral floccus long, dense and copious, whitish; hind tibia rather broad apically, apex nearly twice width of the basitarsi, scopa dense, hairs rather short, obscurely plumose, largely white; front and middle basitarsi slightly narrower than their respective tibiae; wings faintly yellowish, with but two submarginal cells, 2nd submarginal cell receiving 1st recurrent about one-fourth from base, 2nd recurrent at apex; abdominal terga smooth, somewhat shining, very minutely and obscurely punctate, punctures rather close but barely visible, apical impressed area very shallow but rather broad, occupying more than one-third median length of disc, becoming narrowly yellowish-hyaline along rims, discal pubescence very thin and sparse, suberect, entirely pale, terga 2-4 with dense, white, apical fasciae, broadly interrupted medially on tergum 2 and more narrowly so on 3, tergum 5 with a brownish ochraceous, apical fimbria.

MALE. Length 9 mm.; face with conspicuous yellow maculae below, filling area between lateral margins of clypeus and lower end of eye; clypeus broad and flat, projecting but very slightly below suborbital line, yellow except for a pair of small black spots and the dark apical rim, smooth and shining, rather closely and deeply but finely punctate, punctures becoming even closer at extreme sides, space between margin of vertex and lateral ocelli subequal to their diameter; cheeks much broader than eyes, rather flat, subtriangular in outline, with the rounded angle opposite middle of eye, dull, punctures hardly visible beneath rather long, dense, white pubescence; malar space very short; basal segment of flagellum subequal to 2nd and 3rd combined, those beyond 2nd pale ferruginous beneath; process of labrum small, subtriangular; mandibles of moderate length, slender, very slightly bowed, with a distinct, inner, subapical tooth, overlapping about one-third; pubescence of head, thorax and legs entirely whitish, rather long and dense; thoracic integument dull, quite densely tessellate, punctures of scutum exceedingly minute and obscure, well separated, those on scutellum even more obscure, surface densely tessellate; pleura with exceedingly minute and obscure punctures anteriorly, barely visible beneath rather long pubescence; dorsal area of propodeum rather narrow, oblique, triangle finely tessellate, obscurely subrugose toward basal margin; basitarsi slender and elongate, considerably narrower than their respective tibiae; wings with but two submarginal cells, 2nd submarginal receiving 1st and 2nd recurrent veins at subequal distances from base and apex; abdominal terga smooth, somewhat shining, punctures exceedingly minute and obscure, barely visible, apical impressed areas rather narrow, becoming rather broadly yellowish-hyaline along rims, discal pubescence thin but erect, entirely pale, terga 2-4 with thin, white, apical fasciae developed laterally, largely inevident or interrupted medially; lateral angles of sternum 6 very strongly reflexed, the tips projecting anteriorly; apical portion of sternum 8 quite robust, gradually narrowed from base to the subtruncate apex, clothed beneath with extremely short, dense pubescence; penis valves narrowed to a slender, spine-like point apically, broadly dilated and robust basally, not excavated laterally, gonocoxites rather robust and elongate, somewhat dilated apically to form subtriangular, apical lobes which are rather acute at tip, gonocoxal lobes somewhat produced, very broadly rounded.

DISTRIBUTION. Minnesota and Illinois to the New England states; April and May.

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FamilyScientific name @ source (records)
Polycitoridae  Salix sp @ BBSL (5)
Salicaceae  Salix @ AMNH_BEE (13)
_  Withheld @ BBSL (32)

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Updated: 2017-07-22 11:22:23 gmt
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