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Overview |
Main identification features
- adult: red-brown
- juvenile: eye-stripe + large spots
Dorsal rays 108-112; anal rays 78-89; pectoral rays 27-28; pelvic fins under middle of head, each consisting of a pair of filaments that are joined basally; three short barbels on each side of snout and three barbels on each side of lower jaw; dorsal and anal fins confluent with pointed tail fin.
Generally reddish brown; median fins dusky dark brown to blackish; juveniles (under about 20 cm) with dark stripe on head behind eye and numerous, large dark brown spots on body.
Size: attains 98 cm.
Inhabits rocky reefs and adjacent sand bottoms.
Depth: 1-650 m .
Southern Baja and the central Gulf of California to Peru.
Attributes
Abundance: Common.
Cites: Not listed.
Climate Zone: Northern Subtropical (Cortez Province + Sinaloan Gap); Northern Tropical (Mexican Province to Nicaragua + Revillagigedos); Equatorial (Costa Rica to Ecuador + Galapagos, Clipperton, Cocos, Malpelo); South Temperate (Peruvian Province ).
Depth Range Max: 650 m.
Depth Range Min: 1 m.
Diet: mobile benthic crustacea (shrimps/crabs); bony fishes.
Eastern Pacific Range: Northern limit=32; Southern limit=-6; Western limit=-115; Eastern limit=-78; Latitudinal range=38; Longitudinal range=37.
Egg Type: Pelagic; Pelagic larva.
Feeding Group: Carnivore.
FishBase Habitat: Bentho-Pelagic.
Global Endemism: Tropical Eastern Pacific (TEP) endemic; East Pacific endemic; All species.
Habitat: Sand & gravel; Soft bottom (mud, sand,gravel, beach, estuary & mangrove); Reef associated (reef + edges-water column & soft bottom); Reef (rock &/or coral); Rocks; Reef and soft bottom; Water column.
Inshore Offshore: Inshore; Inshore Only.
IUCN Red List: Not evaluated / Listed.
Length Max: 98 cm.
Regional Endemism: Continent; Continent only; TEP endemic; Continental TEP endemic; 3 provinces (Cortez + Mexican + Panamic) endemic; All species.
Residency: Resident.
Salinity: Marine; Marine Only.
Water Column Position: Bottom; Near Bottom; Bottom + water column;
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Names |
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Links to other sites |
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References |
- Acero, A. and Franke, R., 2001., Peces del parque nacional natural Gorgona. En: Barrios, L. M. y M. Lopéz-Victoria (Eds.). Gorgona marina: Contribución al conocimiento de una isla única., INVEMAR, Serie Publicaciones Especiales No. 7:123-131.
- Béarez, P., 1996., Lista de los Peces Marinos del Ecuador Continental., Revista de Biologia Tropical, 44:731-741.
- Findley, L.T., Hendrickx, M.E., Brusca, R.C., van der Heiden, A.M., Hastings, P.A., Torre, J., 2003., Diversidad de la Macrofauna Marina del Golfo de California, Mexico., CD-ROM versión 1.0. Projecto de la Macrofauna del Golfo . Derechos reservados de los autores y Conservación Internacional.
- Franke , R. and Acero P., A., 1995., Las Especies Colombianas del Genero Brotula (Pisces: Ophidiidae)., Caldasia, 17:597-602.
- García-Rodríguez , F. J. and Aurioles-Gamboa, D., 1997., Contribución al conocimiento de la diversidad íctica en la Bahía de La Paz por … lobo marino de California, Zalophus californianus californianus. En Urbán Ramírez, J. y M. Ramírez Rodríguez (Eds.). La Bahía de La Paz investigación y conservación., Universidad Autónoma de Baja California Sur:151-162.
- Hildebrand, S.F., 1946., A descriptive catalog of the shore fishes of Peru., Bull. U.S. Nat. Mus., 189:1-530.
- Hubbs, C.L., 1944., Species of the circumtropical fish genus Brotula., Copeia, 1944(3):162-178.
- Jimenez-Prado, P., Béarez, P., 2004., Peces marinos del Ecuador continental / Marine fishes of continental Ecuador., SIMBIOE/NAZCA/IFEA tomo 1 y 2.
- Lopez , M. I. and Bussing, W. A., 1982., Lista provisional de los peces marinos de la Costa Rica., Revista de Biologia Tropical, 30(1):5-26.
- Love, M.S., Mecklenburg, C.W., Mecklenburg, T.A., Thorsteinson, L.K., 2005., es of the West Coast and Alaska: a checklist of North Pacific and Artic Ocena species from Baja California to the Alaska-Yukon border., U.S. Department of the Interior, U.S. Geological Survey, Biological Resources Division, 288pp.
- Rubio, E.A., 1988., Estudio taxonomico de la ictiofauna acompañante del camaron en areas costeras del Pacifico de Colombia., Memorias del VI Seminario Nacional de las Ciencias del Mar. Comisión Colombiana de Oceanografía. Bogota, Colombia., :169-183.
- Van der Heiden , A. M. and Findley, L. T., 1988., Lista de los peces marinos del sur de Sinaloa, México., Anales del Centro de Ciencias del Mar y Limnologia de la Universidad Autonoma Nacional de Mexico, 15:209-224.
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Acknowledgements |
I thank Ashley MacDonald and John Pickering, University of Georgia, for technical support in building this page.
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http://www.fishbase.org/Summary/speciesSummary.php?genusname=Brotula&speciesname=clarkae ---> http://192.134.151.83/Summary/speciesSummary.php?genusname=Brotula&speciesname=clarkae http://192.134.151.83/Summary/speciesSummary.php?genusname=Brotula&speciesname=clarkae ---> https://fishbase.mnhn.fr/Summary/speciesSummary.php?genusname=Brotula&speciesname=clarkae https://fishbase.mnhn.fr/Summary/speciesSummary.php?genusname=Brotula&speciesname=clarkae ---> https://fishbase.mnhn.fr/summary/Brotula-clarkae.html
Brotula clarkae, Pacific bearded brotula : fisheries
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Brotula clarkae
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Teleostei (teleosts) >
Ophidiiformes
(Cusk eels) >
Ophidiidae
(Cusk-eels) > Brotulinae
Etymology:
Brotula:
Latin, brotula, -ae = little, bud, shoot (Ref.
45335
)
.
More on author:
Hubbs
.
Environment: milieu / climate zone / depth range / distribution range
Ecology
Marine; benthopelagic; depth range ? - 650 m (Ref.
34024
), usually 40 - 75 m (Ref.
36413
). Tropical; 25°N - 6°S
Eastern Central Pacific: Gulf of California to Peru.
Length at first maturity / Size / Weight / Age
Maturity: L
m
75.4
  range ? - ? cm
Max length : 115 cm TL male/unsexed; (Ref.
55763
); common length : 45.0 cm TL male/unsexed; (Ref.
55763
)
Short description
Morphology
|
Morphometrics
Dorsal
spines
(total): 0;
Dorsal
soft rays
(total): 108-118;
Anal
spines
: 0;
Anal
soft rays
: 78 - 89;
Vertebrae
: 55 - 56. Body completely covered with small, imbricate, cycloid scales; barbels present on snout (6) and chin (6); branchiostegal rays 8; median basibranchial tooth patch absent; developed gill rakers on first arch 4 or fewer; pelvic fin rays inserted at about level of preopercle, well behind eye (Ref.
34024
).
Adults benthopelagic to 650 m depth, juveniles common on reefs (Ref.
34024
). At depths of about 40-75 m, adults are found deep on mud and broken shell bottom (Ref.
36413
). Smaller silvery specimens taken far out at sea in the epipelagic (Ref.
34024
). Caught occasionally (Ref.
34024
). Oviparous, with late post-flexion and transformation stage larvae found in midwater (Ref.
36413
). Oval, pelagic eggs float in a gelatinous mass (Ref.
205
).
Life cycle and mating behavior
Maturity
|
Reproduction
|
Spawning
|
Eggs
|
Fecundity
|
Larvae
Oviparous, with no specialized copulatory organ (Ref.
36413
).
Nielsen, J.G., D.M. Cohen, D.F. Markle and C.R. Robins
, 1999. Ophidiiform fishes of the world (Order Ophidiiformes). An annotated and illustrated catalogue of pearlfishes, cusk-eels, brotulas and other ophidiiform fishes known to date. FAO Fish. Synop. 125(18):178p. Rome: FAO. (Ref.
34024
)
IUCN Red List Status (Ref.
130435
)
Data deficient (DD)
; Date assessed:
23 May 2007
CITES
Not Evaluated
Not Evaluated
Threat to humans
Harmless
Human uses
Fisheries: minor commercial
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Zoological Record
Estimates based on models
Preferred temperature (Ref.
123201
): 16 - 25.2, mean 19.4 °C (based on 24 cells).
Phylogenetic diversity index (Ref.
82804
): PD
50
= 0.5156 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00380 (0.00200 - 0.00722), b=3.12 (2.94 - 3.30), in cm total length, based on LWR estimates for this species & (Sub)family-body (Ref.
93245
).
Trophic level (Ref.
69278
): 4.1 ±0.77 se; based on food items.
Resilience (Ref.
120179
): Medium, minimum population doubling time 1.4 - 4.4 years (Preliminary K or Fecundity.).
Prior r = 0.38, 95% CL = 0.25 - 0.57, Based on 1 stock assessment.
Fishing Vulnerability (Ref.
59153
): Moderate to high vulnerability (45 of 100).
Price category (Ref.
80766
):
Medium
.
Nutrients (Ref.
124155
): Calcium = 56.5 [31.2, 137.3] mg/100g; Iron = 0.759 [0.396, 1.366] mg/100g; Protein = 16.2 [13.7, 18.7] %; Omega3 = 0.202 [0.105, 0.413] g/100g; Selenium = 151 [68, 350] μg/100g; VitaminA = 11.9 [3.5, 39.7] μg/100g; Zinc = 0.728 [0.481, 1.126] mg/100g (wet weight);
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Updated: 2024-04-19 19:37:01 gmt
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