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Brotula clarkae Hubbs, 1944
PACIFIC BEARDED BROTULA
Life   Vertebrata   Fish   Ophidiidae   Brotula

Brotula clarkae
© Copyright Ross Robertson, 2006 · 12
Brotula clarkae

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Brotula clarkae
© Copyright Ross Robertson, 2006 · 12
Brotula clarkae
Brotula clarkae
© Copyright Ross Robertson, 2006 · 12
Brotula clarkae
español

Overview
Main identification features
  • adult: red-brown
  • juvenile: eye-stripe + large spots
Dorsal rays 108-112; anal rays 78-89; pectoral rays 27-28; pelvic fins under middle of head, each consisting of a pair of filaments that are joined basally; three short barbels on each side of snout and three barbels on each side of lower jaw; dorsal and anal fins confluent with pointed tail fin.

Generally reddish brown; median fins dusky dark brown to blackish; juveniles (under about 20 cm) with dark stripe on head behind eye and numerous, large dark brown spots on body.

Size: attains 98 cm.

Inhabits rocky reefs and adjacent sand bottoms.

Depth: 1-650 m .

Southern Baja and the central Gulf of California to Peru.


Attributes
Abundance: Common.
Cites: Not listed.
Climate Zone: Northern Subtropical (Cortez Province + Sinaloan Gap); Northern Tropical (Mexican Province to Nicaragua + Revillagigedos); Equatorial (Costa Rica to Ecuador + Galapagos, Clipperton, Cocos, Malpelo); South Temperate (Peruvian Province ).
Depth Range Max: 650 m.
Depth Range Min: 1 m.
Diet: mobile benthic crustacea (shrimps/crabs); bony fishes.
Eastern Pacific Range: Northern limit=32; Southern limit=-6; Western limit=-115; Eastern limit=-78; Latitudinal range=38; Longitudinal range=37.
Egg Type: Pelagic; Pelagic larva.
Feeding Group: Carnivore.
FishBase Habitat: Bentho-Pelagic.
Global Endemism: Tropical Eastern Pacific (TEP) endemic; East Pacific endemic; All species.
Habitat: Sand & gravel; Soft bottom (mud, sand,gravel, beach, estuary & mangrove); Reef associated (reef + edges-water column & soft bottom); Reef (rock &/or coral); Rocks; Reef and soft bottom; Water column.
Inshore Offshore: Inshore; Inshore Only.
IUCN Red List: Not evaluated / Listed.
Length Max: 98 cm.
Regional Endemism: Continent; Continent only; TEP endemic; Continental TEP endemic; 3 provinces (Cortez + Mexican + Panamic) endemic; All species.
Residency: Resident.
Salinity: Marine; Marine Only.
Water Column Position: Bottom; Near Bottom; Bottom + water column;


Names
Scientific source:

Links to other sites

References
  • Acero, A. and Franke, R., 2001., Peces del parque nacional natural Gorgona. En: Barrios, L. M. y M. Lopéz-Victoria (Eds.). Gorgona marina: Contribución al conocimiento de una isla única., INVEMAR, Serie Publicaciones Especiales No. 7:123-131.
  • Béarez, P., 1996., Lista de los Peces Marinos del Ecuador Continental., Revista de Biologia Tropical, 44:731-741.
  • Findley, L.T., Hendrickx, M.E., Brusca, R.C., van der Heiden, A.M., Hastings, P.A., Torre, J., 2003., Diversidad de la Macrofauna Marina del Golfo de California, Mexico., CD-ROM versión 1.0. Projecto de la Macrofauna del Golfo .  Derechos reservados de los autores y Conservación Internacional.
  • Franke , R. and Acero P., A., 1995., Las Especies Colombianas del Genero Brotula (Pisces: Ophidiidae)., Caldasia, 17:597-602.
  • García-Rodríguez , F. J. and Aurioles-Gamboa, D., 1997., Contribución al conocimiento de la diversidad íctica en la Bahía de La Paz por … lobo marino de California, Zalophus californianus californianus. En Urbán Ramírez, J. y M. Ramírez Rodríguez (Eds.). La Bahía de La Paz investigación y conservación., Universidad Autónoma de Baja California Sur:151-162.
  • Hildebrand, S.F., 1946., A descriptive catalog of the shore fishes of Peru., Bull. U.S. Nat. Mus., 189:1-530.
  • Hubbs, C.L., 1944., Species of the circumtropical fish genus Brotula., Copeia, 1944(3):162-178.
  • Jimenez-Prado, P., Béarez, P., 2004., Peces marinos del Ecuador continental / Marine fishes of continental Ecuador., SIMBIOE/NAZCA/IFEA tomo 1 y 2.
  • Lopez , M. I. and Bussing, W. A., 1982., Lista provisional de los peces marinos de la Costa Rica., Revista de Biologia Tropical, 30(1):5-26.
  • Love, M.S., Mecklenburg, C.W., Mecklenburg, T.A., Thorsteinson, L.K., 2005., es of the West Coast and Alaska: a checklist of North Pacific and Artic Ocena species from Baja California to the Alaska-Yukon border., U.S. Department of the Interior, U.S. Geological Survey, Biological Resources Division, 288pp.
  • Rubio, E.A., 1988., Estudio taxonomico de la ictiofauna acompañante del camaron en areas costeras del Pacifico de Colombia., Memorias del VI Seminario Nacional de las Ciencias del Mar. Comisión Colombiana de Oceanografía. Bogota, Colombia., :169-183.
  • Van der Heiden , A. M. and Findley, L. T., 1988., Lista de los peces marinos del sur de Sinaloa, México., Anales del Centro de Ciencias del Mar y Limnologia de la Universidad Autonoma Nacional de Mexico, 15:209-224.

Acknowledgements

I thank Ashley MacDonald and John Pickering, University of Georgia, for technical support in building this page.


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Brotula clarkae, Pacific bearded brotula : fisheries
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Common name (e.g. trout)

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Brotula clarkae Hubbs , 1944

Pacific bearded brotula Add your observation in Fish Watcher
Native range | All suitable habitat | Point map | Year 2050
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Brotula clarkae   AquaMaps   Data sources: GBIF OBIS
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Pictures | Google image Image of Brotula clarkae (Pacific bearded brotula) Brotula clarkae
Picture by Robertson, R.

Classification / Names Common names | Synonyms | Catalog of Fishes( genus , species ) | ITIS | CoL | WoRMS | Cloffa

Teleostei (teleosts) > Ophidiiformes (Cusk eels) > Ophidiidae (Cusk-eels) > Brotulinae
Etymology: Brotula: Latin, brotula, -ae = little, bud, shoot (Ref. 45335 ) .
More on author: Hubbs .

Environment: milieu / climate zone / depth range / distribution range Ecology

Marine; benthopelagic; depth range ? - 650 m (Ref. 34024 ), usually 40 - 75 m (Ref. 36413 ). Tropical; 25°N - 6°S

Distribution Countries | FAO areas | Ecosystems | Occurrences | Point map | Introductions | Faunafri

Eastern Central Pacific: Gulf of California to Peru.

Length at first maturity / Size / Weight / Age

Maturity: L m 75.4   range ? - ? cm
Max length : 115 cm TL male/unsexed; (Ref. 55763 ); common length : 45.0 cm TL male/unsexed; (Ref. 55763 )

Short description Morphology | Morphometrics

Dorsal spines (total): 0; Dorsal soft rays (total): 108-118; Anal spines : 0; Anal soft rays : 78 - 89; Vertebrae : 55 - 56. Body completely covered with small, imbricate, cycloid scales; barbels present on snout (6) and chin (6); branchiostegal rays 8; median basibranchial tooth patch absent; developed gill rakers on first arch 4 or fewer; pelvic fin rays inserted at about level of preopercle, well behind eye (Ref. 34024 ).

Biology     Glossary (e.g. epibenthic)

Adults benthopelagic to 650 m depth, juveniles common on reefs (Ref. 34024 ). At depths of about 40-75 m, adults are found deep on mud and broken shell bottom (Ref. 36413 ). Smaller silvery specimens taken far out at sea in the epipelagic (Ref. 34024 ). Caught occasionally (Ref. 34024 ). Oviparous, with late post-flexion and transformation stage larvae found in midwater (Ref. 36413 ). Oval, pelagic eggs float in a gelatinous mass (Ref. 205 ).

Life cycle and mating behavior Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae

Oviparous, with no specialized copulatory organ (Ref. 36413 ).

Main reference Upload your references | References | Coordinator | Collaborators

Nielsen, J.G., D.M. Cohen, D.F. Markle and C.R. Robins , 1999. Ophidiiform fishes of the world (Order Ophidiiformes). An annotated and illustrated catalogue of pearlfishes, cusk-eels, brotulas and other ophidiiform fishes known to date. FAO Fish. Synop. 125(18):178p. Rome: FAO. (Ref. 34024 )

IUCN Red List Status (Ref. 130435 )

  Data deficient (DD)  ; Date assessed: 23 May 2007

CITES

Not Evaluated

CMS (Ref. 116361 )

Not Evaluated

Threat to humans

  Harmless





Human uses

Fisheries: minor commercial FAO - Publication: search | FishSource | Sea Around Us

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Summary page | Point data | Common names | Photos

Internet sources

AFORO (otoliths) | Aquatic Commons | BHL | Cloffa | BOLDSystems | Websites from users | Check FishWatcher | CISTI | Catalog of Fishes : genus , species | DiscoverLife | ECOTOX | FAO - Publication: search | Faunafri | Fishipedia | Fishtrace | GenBank: genome , nucleotide | GloBI | Google Books | Google Scholar | Google | IGFA World Record | MitoFish | OsteoBase: skull , spine | Otolith Atlas of Taiwan Fishes | PubMed | Reef Life Survey | Socotra Atlas | Tree of Life | Wikipedia: Go , Search | World Records Freshwater Fishing | Zoobank | Zoological Record

Estimates based on models

Preferred temperature (Ref. 123201 ): 16 - 25.2, mean 19.4 °C (based on 24 cells). Phylogenetic diversity index (Ref. 82804 ):  PD 50 = 0.5156   [Uniqueness, from 0.5 = low to 2.0 = high]. Bayesian length-weight: a=0.00380 (0.00200 - 0.00722), b=3.12 (2.94 - 3.30), in cm total length, based on LWR estimates for this species & (Sub)family-body (Ref. 93245 ). Trophic level (Ref. 69278 ):  4.1   ±0.77 se; based on food items. Resilience (Ref. 120179 ):  Medium, minimum population doubling time 1.4 - 4.4 years (Preliminary K or Fecundity.). Prior r = 0.38, 95% CL = 0.25 - 0.57, Based on 1 stock assessment. Fishing Vulnerability (Ref. 59153 ):  Moderate to high vulnerability (45 of 100). Price category (Ref. 80766 ):   Medium . Nutrients (Ref. 124155 ):  Calcium = 56.5 [31.2, 137.3] mg/100g; Iron = 0.759 [0.396, 1.366] mg/100g; Protein = 16.2 [13.7, 18.7] %; Omega3 = 0.202 [0.105, 0.413] g/100g; Selenium = 151 [68, 350] μg/100g; VitaminA = 11.9 [3.5, 39.7] μg/100g; Zinc = 0.728 [0.481, 1.126] mg/100g (wet weight);

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Updated: 2024-04-19 19:37:01 gmt
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