13.
Campylopus schmidii
(Müller Hal.) A. Jaeger, Thätigk. St. Gallischen Ber. Naturwiss. Ges. 1870-1871: 439. 1872.
Dicranum schmidii
Müller Hal., Bot. Zeitung (Berlin) 11: 37. 1853;
Campylopus aureus
Bosch & Sande La Coste
Plants
2-5 cm, yellowish green, stiff, evenly foliate, the perichaetia in comal tufts.
Leaves
5-6 mm, erect-patent when wet, appressed when dry, from oblong base gradually contracted to a long subula, ending in a straight, hyaline, serrate tip; alar cells hardly differentiated; basal laminal cells thin-walled, rectangular, hyaline; distal laminal cells chlorophyllose, oval to narrow or elongate oval, incrassate; costa filling
1
/2-
2
/3 of leaf width, in transverse section showing adaxial hyalocysts as large as the median deuter cells, and abaxial groups of stereids, ribbed abaxially.
Specialized asexual reproduction
by deciduous buds produced in the distalmost part of the stem.
Sporophytes
not known from North America.
Soil in open pine, cedar and cypress forests; 80-200 m; Calif., Oreg.; Mexico; Asia (China, s India, Sri Lanka, Java, Sulawesi, Borneo, Taiwan); Africa (c Africa); Indian Ocean Islands (Madagascar); Pacific Islands (Hawaii); n Australia.
Campylopus schmidii
does not fruit in North America, where apparently only female plants exist. The range of this species is mainly southeastern Asia. From there it extends south to Queensland, west to Madagascar and Central Africa, east to Hawaii, California, Oregon, and Mexico. In California it is known from two localities, in Oregon from one, and in Mexico from one, which suggests that the occurrence of
C. schmidii
may result from occasional long distance dispersal events across the Pacific Ocean; it may not be native in North America. The first collection was made in California in 1933. Plants of
C. schmidii
resemble
C. pilifer
but are distinguished by elongate-oval rather than oval distal laminal cells and costa smooth at the abaxial surface and not with lamellae 3-4 cells high as in
C. pilifer
.
Campylopus introflexus
has lamellae 2 cells high and a similar areolation as
C. pilifer
but is distinguished in the field by reflexed hairpoints. All three species are more or less vicariant sister species,
C. introflexus
in the subantarctic to subtropical parts of the southern hemisphere,
C. pilifer
in tropical India, Africa, and South America and from there extending to southeastern North America and southwestern Europe, and
C. schmidii
mainly in southeastern Asia.
