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Canis lupus Linnaeus, 1758

Gray Wolf

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Canis lupus
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Canis lupus
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Overview
Wolves range widely across a great variety of open to wooded habitats.

Conservation Biology

  • Special Protection Status

    • Rangewide: The wolf is on the federal Endangered Species List in the 48 contiguous states, but will probably be delisted by 2001.

    • In Park: All plants and animals are protected within Great Smoky Mountains National Park. Collection requires a permit which is usually granted only for research or educational purposes.


Identification
  • Adult total length: 50 - 65 in. (1,270 - 1,670 mm)

  • Tail: 11 - 18 in. (280 - 450 mm)

  • Hind foot: 8 - 10 in. (200 - 250 mm)

  • Weight: 45 - 100 lbs. (21 - 45 kg)

  • Physical Characteristics: The largest wild dog, the gray wolf is usually mottled gray but may be black or white or any grade inbetween. It is distinguished from its nearest relative, the red wolf, by being 10 - 50% larger and by having a broader snout and proportionately shorter ears, and from the coyote by being 50 - 100% larger and having a broader snout and proportionately larger feet.


Names
Scientific source:
      Integrated Taxonomic Information System

Phylogeny
Taxonomic Category Scientific Name Common Name
Phylum Chordata Chordates
Class Mammalia Mammals
Order Carnivora Dogs, Cats, Bears, etc.
Family Canidae Wolves, Foxes

Geographic distribution
The gray wolf was formerly distributed throughout the Northern Hemisphere. However, deliberate extermination has restricted the wolf's current North American range to Canada, Alaska, Montana, northern Minnesota, Wisconsin, and Michigan (including Isle Royale in Lake Superior). Washington, Idaho, Wyoming, and North and South Dakota support sporadic individuals, and Mexico may have a few.

  • Range Maps

    North America

  • Great Smoky Mountains National Park: The first mention of the status of wolves in this area came in 1844 when a letter to a member of the House of Representatives (Lanman, 1849) stated that sheep were destroyed by wolves "which have not yet been entirely eliminated."

    The following account of the history of the gray wolf is taken from Linzey and Linzey (1968): "The gray wolf once occurred in the Great Smoky Mountains in fair numbers but became increasingly less common as more of the land was settled. Buckley (1859) reported that wolves were "troublesome" to the mountain farmers of North Carolina and Tennessee. In 1887, C.H. Merriam ( 1888) noted that wolves "still occur" in the Great Smokies. John Oliver, a former resident of the Park, remembered hearing wolves howling in Cades Cove when he was a boy (1880-90). A resident of Gatlinburg recalled seeing one of these animals that had been caught in a bear trap near the Sugarlands during the 1890's. He also heard two wolves howling near the area that was formerly Chimneys Campground. Brimley (1944) wrote that wolves were "apparently finally exterminated in or about 1890, up to which time they still occurred sparingly in the mountains." Hamnett and Thornton (1953) stated: "In the Mountain Region, wolves existed in the more remote sections until the late 1800's and possibly until the very early 1900's."

    "There have been occasional unconfirmed reports of wolves in the mountains after 1900. The Knoxville News-Sentinel (Brewer, 1964) printed a column indicating that wolves were seen on Mt. LeConte as late as 1925. Ganier (1928) speculated that a few might still be present in the "wilder mountainous sections." An animal that was reported to be a gray wolf was killed near Waynesville, Haywood County, North Carolina, Feburary 27, 1933. Official verification is lacking for all of these reports."


Natural history
  • Reproduction
    After a courtship that may last from days to months, wolves copulate during estrus, which occurs once per year and lasts 5-14 days. The receptive period may be as early as January in low latitudes or as late as April in high latitudes. During copulation the pair remains coupled for as long as 30 minutes, during which ejaculation occurs many times. Gestation lasts 63 days, and litters average six young. The young are born blind and helpless, usually in an underground burrow. The same den may be used year after year ( Mech, 1999).

    The female usually stays near the young for at least three weeks. During this time, the male and other pack members hunt and feed her and the pups. The pups' eyes open between 11 and 15 days of age, and they are weaned when 9 weeks old. Healthy pups join adults in their travels as early as October. In the wild, wolves do not breed until they are 2, 3, or 4 years of age. Both sexes may continue to breed through at least 10 years of age (Mech, 1999).

  • Longevity
    In the wild, wolves can live to be about 16 years of age (Mech, 1999).

  • Terrestrial Ecology
    The wolf is a social animal, usually living in packs of 5-10 members, although packs of up to 36 have been reported. Generally wolf packs contain a set of parents (the "alpha pair") and some of the offspring of their past 2-3 litters.

    Wolves are primarily predators on ungulates, beavers, and hares, although almost any species of animal may be eaten, including any type of domestic animal.

  • Predators and Defense
    Humans are the only enemy of gray wolves.

  • Parasites
    None recorded from the park.


Links to other sites

References

Acknowledgements
  • Text
  • Photographs
    • Roger Barbour

  • Map development
  • Web page design & coding
    • Denise Lim, University of Georgia, Athens
    • John Pickering, University of Georgia, Athens



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Canis lupus

Gray Wolf

Order: Carnivora
Family: Canidae

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Gray Wolf
Credit: Isle Royale National Park
Image of Canis lupus
Canis lupus - typical coloration, standing left; black variant, inset; white variant, standing right; juvenile, center
Click to enlarge. (97 kb)

Gray wolves usually live in packs, led by an "alpha pair." The pack includes some of the alpha pair's offspring and may include some unrelated wolves. A pack's territory can be as large as 13,000 square km. Howling probably helps advertise who "owns" a particular piece of territory. When pups are born, the mother stays near them for the first three weeks, and her mate and others in the pack bring food to her. The pups are weaned when they about nine weeks old. As adults, they may travel as far as 72 km a day with their pack and run as fast as 70 km per hour.

Also known as:
Wolf, Timber Wolf, Tundra Wolf, Lobo, Prairie Wolf, Mexican Wolf, Arctic Wolf

Length:
Average: 1,100 mm males; 1,050 kg, females
Range: 1,000-1,300 mm males; 870-1,170 mm females

Weight:
Average: 55 kg males; 45 kg females
Range: 30-80 kg males; 23-55 kg females

References:

Linnaeus, C., 1758.  Systema Naturae per regna tria naturae, secundum classis, ordines, genera, species cum characteribus, differentiis, synonymis, locis . Tenth Edition. Laurentii Salvii, Stockholm, 1:39, 824 pp.

Links:

Mammal Species of the World

Click here for The American Society of Mammalogists species account

Distribution of Canis lupus

Image of Canis lupus
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Following modified from Global Invasive Species Database
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Following modified from CalPhotos
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CalPhotos     Photo Database

 

Number of matches : 1
Query: SELECT * FROM img WHERE ready=1 and taxon like "Canis lupus signatus%" and (lifeform != "specimen_tag" OR lifeform != "Animal") ORDER BY taxon

Click on the thumbnail to see an enlargement

Canis lupus signatus
Canis lupus signatus
ID: 0000 0000 1005 0826 [detail]
© 2005 Diogo Verissimo

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2010 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT     University of California, Berkeley

Following modified from CalPhotos
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CalPhotos     Photo Database

 

Number of matches : 85
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Click on the thumbnail to see an enlargement

Canis lupus
Canis lupus
Gray Wolf
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Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
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Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
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Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0990 0122 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
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Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
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Gerald and Buff Corsi
© 2002 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0990 0120 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0990 0121 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0987 0063 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0986 0020 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 1335 3153 0668 0126 [detail]
Gerald and Buff Corsi
© 2002 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0990 0118 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0987 0062 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0987 0059 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Alaska timber wolf
ID: 0072 3301 1162 0105 [detail]
Robert Potts
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 1335 3153 0668 0130 [detail]
Gerald and Buff Corsi
© 2002 California Academy of Sciences

Canis lupus
Canis lupus
Alaska grey wolf
ID: 0072 3301 1162 0117 [detail]
Robert Potts
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 1335 3153 0668 0122 [detail]
Gerald and Buff Corsi
© 2002 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0986 0037 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 1335 3153 0668 0124 [detail]
Gerald and Buff Corsi
© 2002 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0987 0056 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 1335 3153 0668 0125 [detail]
Gerald and Buff Corsi
© 2002 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0990 0105 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Canis lupus
Canis lupus
Gray Wolf
ID: 0091 3183 0987 0055 [detail]
Gerald and Buff Corsi
© 2001 California Academy of Sciences

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2010 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT     University of California, Berkeley

Following modified from CalPhotos
   Top | See original


CalPhotos     Photo Database

 

Number of matches : 1
Query: SELECT * FROM img WHERE ready=1 and taxon like "Canis lupus lupus%" and (lifeform != "specimen_tag" OR lifeform != "Animal") ORDER BY taxon

Click on the thumbnail to see an enlargement

Canis lupus lupus
Canis lupus lupus
Gray Wolf
ID: 0000 0000 0208 1707 [detail]
© 2008 Maik Dobiey

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2010 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT     University of California, Berkeley

Following modified from CalPhotos
   Top | See original


CalPhotos     Photo Database

 

Number of matches : 3
Query: SELECT * FROM img WHERE ready=1 and taxon like "Canis lupus halstromi%" and (lifeform != "specimen_tag" OR lifeform != "Animal") ORDER BY taxon

Click on the thumbnail to see an enlargement

Canis lupus halstromi
Canis lupus halstromi
New Guinea Singing Dog
ID: 0000 0000 0301 0172 [detail]
© 2001 John White

Canis lupus halstromi
Canis lupus halstromi
New Guinea Singing Dog
ID: 0000 0000 0301 0173 [detail]
© 2001 John White

Canis lupus halstromi
Canis lupus halstromi
New Guinea Singing Dog
ID: 0000 0000 0301 0174 [detail]
© 2001 John White

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2010 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT     University of California, Berkeley

Following modified from CalPhotos
   Top | See original


CalPhotos     Photo Database

 

Number of matches : 2
Query: SELECT * FROM img WHERE ready=1 and taxon like "Canis lupus familiaris%" and (lifeform != "specimen_tag" OR lifeform != "Animal") ORDER BY taxon

Click on the thumbnail to see an enlargement

Canis lupus familiaris
Canis lupus familiaris
Domestic Dog
ID: 0000 0000 1008 0618 [detail]
© 2008 John Dittes

Canis lupus familiaris
Canis lupus familiaris
Domestic Dog
ID: 0000 0000 1008 0619 [detail]
© 2008 John Dittes

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2010 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT     University of California, Berkeley

Following modified from CalPhotos
   Top | See original


CalPhotos     Photo Database

 

Number of matches : 19
Query: SELECT * FROM img WHERE ready=1 and taxon like "Canis lupus baileyi%" and (lifeform != "specimen_tag" OR lifeform != "Animal") ORDER BY taxon

Click on the thumbnail to see an enlargement

Canis lupus baileyi
Canis lupus baileyi
Mexican Gray Wolf
ID: 0000 0000 0601 0072 [detail]
© 2001 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 0903 0145 [detail]
© 2003 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 0204 0056 [detail]
© 2004 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 0204 0060 [detail]
© 2004 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Gray Wolf
ID: 0000 0000 0601 0070 [detail]
© 2001 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Gray Wolf
ID: 0000 0000 0601 0071 [detail]
© 2001 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 0204 0057 [detail]
© 2004 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 0204 0059 [detail]
© 2004 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 0204 0058 [detail]
© 2004 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Gray Wolf
ID: 0000 0000 1004 0219 [detail]
© 2004 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 1204 0165 [detail]
© 2004 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 0205 0178 [detail]
© 2005 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 1205 0226 [detail]
© 2005 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 1205 0227 [detail]
© 2005 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 1205 0228 [detail]
© 2005 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 1006 0383 [detail]
© 2006 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 1006 0384 [detail]
© 2006 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 1006 0385 [detail]
© 2006 John White

Canis lupus baileyi
Canis lupus baileyi
Mexican Wolf
ID: 0000 0000 1006 0386 [detail]
© 2006 John White

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2010 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT     University of California, Berkeley

Updated: 2010-02-10 03:40:29 gmt
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