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Nomada aquilarum Cockerell, 1903
Nomada cockerelli Graenicher, 1911; Nomada (Holonomada) dacotensis Swenk, 1913; Nomada (Nomadita) aquilarum Cockerell, 1903; Nomada (Nomadita) cockerelli Graenicher, 1911; Nomada (Nomadita) dacotensis Swenk, 1913

Life   Insecta   Hymenoptera   Apoidea   Apidae   Nomada
Subgenus: None

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Nomada aquilarum, male, abdomen
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Nomada aquilarum, male, top
Overview
Closely aligned with N. snowii, in the white maculations, but white less extensive
Reprinted with permission from: Mitchell, T.B. 1962 Bees of the Eastern United States. North Carolina Agricultural Experiment Station Technical Bulletin No. 152.


FEMALE�Length 8-9 mm.; lateral ocelli subequally distant from margin of vertex and each other; cheeks about as broad as eyes, posterior margin carinate; shorter side of basal segment of flagellum equal to segment 2 in length, median segments fully as broad as long; mandibles slender and simple apically; wings subhyaline basally, becoming faintly infuscated apically, but with a definitely hyaline area just beyond the discoidal and submarginal cells, with the usual three submarginal cells, 2nd somewhat broader anteriorly than 3rd, veins testaceous to piceous, stigma somewhat more reddish, basal vein interstitial with transverse median; tegulae shining, very sparsely punctate except along inner margin; posterior margin of scutellum very slightly grooved medially; front coxae with short, acute, triangular tubercles that are obscured by the pubescence; apex of hind tibiae with a row of 5 or 6 very short setae; face with a pair of much abbreviated, ivory maculations adjacent to eyes below, head otherwise black; antennal scape reddish, flagellum more reddish-testaceous beneath, piceous above, tubercles and posterior band on scutellum ivory, outer half of tegulae and a narrow median band on pronotal collar maculated in some specimens, thorax otherwise black; femora and trochanters largely piceous, front femora and tibiae yellowish maculated apically, otherwise black, hind tarsi largely piceous, mid and front tarsi more reddish-testaceous, spurs pale yellow; basal abdominal tergum entirely black, terga 2-6 with conspicuous, ivory maculae, interrupted medially on 2-4, very broad laterally and triangularly pointed medially on 2 and 3, entire on 5 but narrowed laterally; abdominal sterna largely black, but 3 and 4 often with narrow, subapical, ivory bands that are more or less widely interrupted medially; punctures of face, vertex and cheeks deep, distinct and quite close, somewhat coarser on vertex and upper half of face, rather fine and very close on cheeks below and on lower half of face; scutum, scutellum and mesopleura rather coarsely and densely rugosopunctate, punctures somewhat more distinct on pleura below and on scutellum, propodeum posteriorly with rather fine, close punctures just below the impunctate triangle, this and the lateral surfaces dull and densely tessellate; basal abdominal tergum with minute, rather close punctures just before the impressed apical margin, these becoming minute and inevident toward the shining base; discs of terga 2-6 with deep, distinct, minute and rather close punctures, apical margins rather narrowly impunctate; pseudopygidium of the usual form; pubescence extremely short and thin, hardly visible, apparently entirely pale.

MALE�Length 3 mm.; lateral ocelli slightly nearer margin of vertex than to each other; cheeks fully as broad as eyes, posterior margin not carinate; antennal scape not much swollen, apex rather deeply excavated and partially enclosing pedicel, basal segment of flagellum slightly longer than segment 2, median segments fully as broad as long; mandibles slender and simple apically; wings subhyaline basally, becoming faintly infuscated apically, but with a definite hyaline area just beyond the discoidal and submarginal cells, with the usual three submarginal cells, 2nd somewhat broader anteriorly than 3rd, veins testaceous to piceous, stigma somewhat more reddish, basal vein interstitial with transverse median; tegulae shining, very sparsely punctate except along inner margin; posterior margin of scutellum slightly grooved medially; front coxae with short but rather robust, triangular spines that are obscured by the pubescence; hind femora unmodified; clypeus and labrum in large part, mandibles at base, and a pair of much abbreviated, lateral maculae, pale yellowish-white or ivory, upper third of clypeus black, head otherwise black; antennal scape yellowish-testaceous anteriorly, flagellum more reddish-testaceous beneath, piceous above; tubercles, outer half of tegulae, a narrow median band on pronotal collar, and a pair of spots on scutellum, ivory; thorax otherwise entirely black; femora and trochanters largely piceous, front femora more yellowish anteriorly, tibiae yellowish at base and apex on outer surface, these narrowly connected, surface otherwise piceous to testaceous, hind tarsi largely piceous, mid and front tarsi more reddish-testaceous, spurs pale yellow; basal abdominal tergum entirely black, terga 2-6 with conspicuous, ivory maculae, interrupted medially on 1-3, very broad laterally and triangularly pointed medially on 2 and 3, entire and of rather uniform width on 5 and 6, on which the apical margins are reddish-hyaline; abdominal sterna largely black, yellowish-hyaline apically, sterna 3-5 with narrow, subapical, ivory bands that are more or less widely interrupted medially; punctures of face, vertex and cheeks deep, distinct and quite close, somewhat coarser and more widely separated on vertex, rather fine and very close on cheeks below and on lower half of face, supraclypeal area rather coarsely rugoso-striate, punctures of maculated area of clypeus more vague and irregular; scutum, scutellum and mesopleura rather coarsely and densely rugoso-punctate, punctures somewhat more distinct on pleura below and on scutellum, propodeum posteriorly with rather fine, shallow, but close punctures just below the impunctate triangle, this and the lateral surfaces dull and densely tessellate; basal abdominal tergum with minute, rather close punctures just before the impressed apical margin, these becoming minute and inevident toward the shining base; discs of terga 2-6 with deep, distinct, rather close and fine punctures, apical margins rather narrowly impunctate and more or less yellowish-hyaline, the punctures slightly coarser on the more apical terga; pubescence extremely short and thin but rather copious over head and thorax, more whitish below, more yellowish above, very fine and inconspicuous on abdomen; median length of pygidial plate about equal to basal width, rather strongly narrowed to the narrowly rounded apex, margins carinate, surface rather shallowly rugoso-punctate; sternum 8 and genital armature similar to placida (figs. 103 & 104).

DISTRIBUTION�Chiefly western, but found in Wisconsin, in July.

FLOWER RECORDS�This species has been recorded on Erigeron, Grindelia, Lactuca, Melilotus and Solidago.



Reprinted from: Broemeling, D.K. 1988 A Revision of the Nomada Subgenus Nomadita of North America (Hymenoptera: Anthophoridae). The Pan-Pacific Entomologist 64; 327


Nomadaaquilarum Cockerell, 1903. Ann. Mag. Nat. Hist. 12:208-209. Holotype, male: "No. 13183, 8 .18, NM, S Fk Eagle Cr, Abt 8000 ft., Coll. Townsend, on fls Erigeron macranthus". Type Depository, United States National Museum.
Nomada cockerelli Graenicher, 1911. Bull. Pub. Mus. Milwaukee 1:221-249. Holotype, male: "No. 37769, Hudson, St. Croix Co., between July 6 and 12,1910". Type Depository, Pub. Mus. Milwaukee.
Nomada dacotensis Swenk, 1913. Nebr. Univ. Studies 12:88. Holotype, female: "2803, Fargo, N. D. Aug. 17,1911,0. A. Stevens, Grindelia squarrosa". Type De�pository, University of Nebraska, Lincoln.


Diagnosis: Differs from placida, timberlakei, and verecunda by the presence of white instead of yellow maculations. Differs from snowii by the lack of ferruginous markings on the legs or thorax and broken transverse median maculations on the abdominal terga. Differs from mutans by the presence of a strong denticle on the pro-coxae, a prominent scutellum which is not flattened or highly polished, and the shelf�like supra-spiracular ridge. It differs from most mutans by having the first tergum all black and several terga with medially interrupted transverse maculations.

Male: Length 6.3-9.4 mm, forewing length 4.7-6.2 mm, hindwing length 3.8-4.9 mm; scape densely, almost contiguously punctured, IPS shiny, somewhat sha-greened; IOD 0.31 mm, OOD 0.38 mm, MLOD 0.09 mm, MOD 0.17 mm, MOOM 0.39 mm; IPS within ocellar triangle roughened; labrum with broken subapical trans�verse carina; acetabular carina distinctly lamellate; pre-lobar carina sharp, gently sloping; pronotal ridge with abruptly angulate, deeply punctate apex; tegulae very sparsely punctate, glassy; scutellum protruberant, bilobate, coarsely punctate, scuto-scutellar suture deeply depressed, posterior interpunctural surface coarsely, transversely micro-rugose; propodeal sides with very rough IPS, supraspiracular ridge strong, forming distinct flattened shelf above spiracular opening, distinct verti-cal groove posterior to spiracle; metapleuron with ventral half shallowly punctate; short but stout pro-coxal spine rudiment present; hind tibial apex with 5 bristles, pos�terior 2 clear and quite thin; forewing with 3 submarginal cells (left wing of type with incomplete 1st intercubitus), infuscated with clear subapical crescent: COLOR: lower half of clypeus, sides of face along compound eyes halfway to antennal inser�tion, malar space, basal half of mandibles, labrum (in part), pronotal lobes, lateral spots on tegulae, apex of scutellar lobes, apical and basal portions of tibiae, apex of hind-femora, lateral median triangular patches on terga 2-4, complete transverse median bands on terga 5-6, apical lateral crescents on sterna 3-5, creamy-white; an�tennae ferruginous; remainder of body dark brown to black.


Reprinted from: Cockerell,T.A. 1903. New American hymenoptera, mostly of the genus Nomada. Ann. Mag. Nat. Hist. 64(3): 208-209

�Length hardly 6 millim. Black, with cream-coloured markings ; abdomen broad and quite small. Facial quadrangle square, the orbits parallel ; eyes pale grey ; anterior half of clypeus, triangular lateral marks short and blunt above, and broad base of mandibles cream-colour; mandibles simple; labrum dark, very densely punctured ; antennae dark brown above, ferruginous beneath ; third joint a little longer than fourth, paler and less shining than rest of flagellum ; cheeks wholly black ; mesothorax extremely densely punctured ; the cream-coloured tubercles offer the only light marking on thorax ; scute-Hum hardly bilobed ; tegulse testaceous, with a small creamcoloured spot. Wings irregularly stained with brown ; nervures dark ; transverso-medial nervure joining discoidal at its base. Legs black, anterior femora and tibiae in front and anterior tarsi pale ferruginous; middle and hind tibia? at base, spot near apex of middle tibiae, and large wedge-shaped mark on apical portion of hind tibiae cream-colour; hair on inner side of basal joint of hind tarsi very pale golden. Abdomen with shallow punctures ; first segment black, second to fourth with a large wedge-shaped cream-coloured mark on each side, fifth and sixth with bands ; posterior margins of third and following segments reddish ; apical plate not at all emarginate; apical portion of venter with three light bands, the first two very broadly interrupted in the middle. The markings of the hind tibiae and the immaculate metathorax suggest N. verecunda, which, however, is a different thing. Hah. South Fork of Eagle Creek, White Mountains, New Mexico, about 8000 feet, at flowers of Erigeron macranthus, Aug. 18 (G. H. T. Townsend).


Identification
Extracted from: Broemeling K.D., (1988). A Revision of the Nomada Subgenus Nomadita of North America (Hymenoptera: Anthophoridae) Pan-Pacific Entomologist 64(4), 1988, p.321-344

Diagnosis: Differs from placida, timberlakei, and verecunda by the presence of white instead of yellow maculations. Differs from snowii by the lack of ferruginous markings on the legs or thorax and broken transverse median maculations on the abdominal terga. Differs from mutans by the presence of a strong denticle on the pro-coxae, a prominent scutellum which is not flattened or highly polished, and the shelf�like supra-spiracular ridge. It differs from most mutans by having the first tergum all black and several terga with medially interrupted transverse maculations.

Male: Length 6.3-9.4 mm, forewing length 4.7-6.2 mm, hindwing length 3.8-4.9 mm; scape densely, almost contiguously punctured, IPS shiny, somewhat sha-greened; IOD 0.31 mm, OOD 0.38 mm, MLOD 0.09 mm, MOD 0.17 mm, MOOM 0.39 mm; IPS within ocellar triangle roughened; labrum with broken subapical trans�verse carina; acetabular carina distinctly lamellate; pre-lobar carina sharp, gently sloping; pronotal ridge with abruptly angulate, deeply punctate apex; tegulae very sparsely punctate, glassy; scutellum protruberant, bilobate, coarsely punctate, scuto-scutellar suture deeply depressed, posterior interpunctural surface coarsely, transversely micro-rugose; propodeal sides with very rough IPS, supraspiracular ridge strong, forming distinct flattened shelf above spiracular opening, distinct verti�cal groove posterior to spiracle; metapleuron with ventral half shallowly punctate; short but stout pro-coxal spine rudiment present; hind tibial apex with 5 bristles, pos�terior 2 clear and quite thin; forewing with 3 submarginal cells (left wing of type with incomplete 1st intercubitus), infuscated with clear subapical crescent: COLOR: lower half of clypeus, sides of face along compound eyes halfway to antennal inser�tion, malar space, basal half of mandibles, labrum (in part), pronotal lobes, lateral spots on tegulae, apex of scutellar lobes, apical and basal portions of tibiae, apex of hind-femora, lateral median triangular patches on terga 2-4, complete transverse median bands on terga 5-6, apical lateral crescents on sterna 3-5, creamy-white; an-tennae ferruginous; remainder of body dark brown to black.

Female: Length 6.2-8.1 mm, forewing length 5.1-5.6 mm, hindwing length 4.2-4.4 mm; very similar to male.

Discussion: This species is widely distributed along the Rocky Mountains. It has the most strongly developed pro-coxal spines of any of the North American species, but they are not as pronounced as those of the Eurasian species.

Extracted from: Cockerell T.D.A., New American Hymenoptera, mostly of the Genus Nomada pp. 35-37.

FEMALE. —Length about 8 millim. Dark ferruginous, with black markings ; no yellow anywhere. Mandibles simple. Resembles N. Astori, but certainly a different species, readily distinguished by having no yellow at lower corners of face ; having the third antennal joint at least as long as the fourth ; the sides of the dark red abdomen without black marks, and the surface of the abdomen very distinctly punctured, which is not at all the case in Astori. Maxillary palpi with the fourth joint longest, but the third almost as long ; the fifth and sixth very slender the fifth conspicuously shorter than the fourth and the sixth than the fifth. Face with short white hair; occiput with tawny hair ; scape black above apically ; front and region of ocelli suffusedly blackened ; mesothorax with a single broad median black stripe ; scutellum and postscutellum dark red but metathorax black ; tegulse shining coppery red, with very sparse punctures. Wings fairly clear, the broadly darkened apical margin strongly contrasting ; stigma very dark reddish, nervures piceous ; second submarginal cell broad above. Legs red, with little black, but the middle femora at base and the hind femora behind are conspicuously blackened ; hair on inner side of basal joint of hind tarsi pale golden ; a trilobed black mark at base of first abdominal segment.

MALE —Length hardly 6 millim. Black, with cream-coloured markings ; abdomen broad and quite small. Facial quadrangle square, the orbits parallel ; eyes pale grey ; anterior half of clypeus, triangular lateral marks short and blunt above, and broad base of mandibles cream-colour; mandibles simple; labrum dark, very densely punctured ; antennae dark brown above, ferruginous beneath ; third joint a little longer than fourth, paler and less shining than rest of flagellum ; cheeks wholly black ; mesothorax extremely densely punctured ; the cream-coloured tubercles offer the only light marking on thorax ; scuteum hardly bilobed ; tegulse testaceous, with a small creamcoloured spot. Wings irregularly stained with brown ; nervures dark ; transverso-medial nervure joining discoidal at its base. Legs black, anterior femora and tibiae in front and anterior tarsi pale ferruginous; middle and hind tibia? at base, spot near apex of middle tibiae, and large wedge-shaped mark on apical portion of hind tibiae cream-colour; hair on inner side of basal joint of hind tarsi very pale golden. Abdo- men with shallow punctures ; first segment black, second to fourth with a large wedge-shaped cream-coloured mark on each side, fifth and sixth with bands ; posterior margins of third and following segments reddish ; apical plate not at all emarginate; apical portion of venter with three light bands, the first two very broadly interrupted in the middle.


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