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Trichoptera

Caddisflies

Links

Kinds

Overview
Trichoptera consists of about 7000 species of insects known as "caddisflies" that are found in freshwater environments. Trichopterans are distinguished from other insects by their characterisitically hairy wings and long antennae. Some are plant eaters and others are meat eaters. Like moths, caddisflies are strongly attracted to light. They are ecologically important because they feed on debris, cleaning out the freshwater systems in which they live. Also, they are an important food source for fish, and they are often used as bait by fishermen. -- (Britannica.com)

Click here for Caddisfly checklist in the Great Smoky Mountains.


Identification
Trichoptera are small to medium sized insects that are similar to moths in appearance. The wings are membranous, hairy (and sometimes have scales), and are held over the body in a rooflike fashion when at rest. The antennae are long and slender. Only a few trichopterans are brightly colored; most have dull coloration. -- (Borror et al., 1989)

Phylogeny

Taxonomic Category Scientific Name Common Name
Phylum Arthropoda Arthropods
Class Insecta Insects
Order Trichoptera Caddisfly


Photographs

Trichopteran "Arctopsyche"

Copyright Skip Hodges.

Trichopteran "Helicopsyche" cases

Copyright Skip Hodges.

Species: Limnephilus lunatus

Copyright The Virtual Field Guide.

Species: Limnephilus lunatus

Copyright The Virtual Field Guide.


Geographic distribution
Caddisflies are worldwide in distribution and, like most insects, they are concentrated in warm rather than cold areas.

Natural history
Caddisfly larvae are very similar to caterpillars in appearance. Their abdominal segments bear gills, making caddisflies aquatic insects. The larvae occur in ponds, lakes, stream, and other bodies of water. Some are case makers, others build nets when submerged, and a some are free-living. Case making larvae use silk to fuse together leaves, twigs, sand, and pebbles to make cases in which they live. They feed on plants. Net making larvae spend most of their time near underwater silk nets and feed off materials that are trapped in the nets. The free-living larvae are predaceous.

The larvae fasten their cases to objects in the water when they have completed their growth and pupate in the cases. The fully developed pupa chews its way out of the case, emerges from the water, and crawls onto some object like a rock or stick. Then it developes into an adult.

Adult caddislflies are generally weak fliers. The eggs are laid in masses or strings of several hundred, and whether laid in or near the water, the female usually attaches them to some sort of object like a stone. The young hatch within a few days of deposition. Interestingly, caddisflies spend most of their lives as larvae, maturing primarily in order to reproduce. Larvae take about a year to develop into adults, but the adults only live about a month.

Caddisflies are ecologically important mainly because they occupy a special niche in the food chain. They are preyed upon by fish and other aquatic organisms. -- (Borror et al., 1989)


How to encounter
Caddisflies are usually found near water. Check near ponds and streams, and even the underside of bridges. Sweeping in vegetation in these areas is a good way to collect them. Lights are probably the best way to collect them, and blue lights seem to be more attractive than other colors. -- (Borror et al., 1989)

Links to other sites
Caddisflies are usually found near water. Check near ponds and streams, and even the underside of bridges. Sweeping in vegetation in these areas is a good way to collect them. Lights are probably the best way to collect them, and blue lights seem to be more attractive than other colors. -- (Borror et al., 1989)

References
  • Borror, Donald J., Norman F. Johnson, and Charles A. Triplehorn. An Introduction to the Study of Insects. Philadelphia: Saunders College Publishing. 1989. 6th ed.

Acknowledgements
Stephen Boyd, Scentific Illustration Major, University of Georgia, Athens.

Thanks to Sabina Gupta, Denise Lim, and Dr. John Pickering for technical and web support in developing this page.

Special Thanks to: The Virtual Field Guide, Skip Hodges, Clemson University, and David Walker.



Following modified from University of Guelph
   
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Order - TRICHOPTERA
(Greek, thrix = hair; pteron = wing)
Common Name: caddisflies
Distribution: Cosmopolitan

Description
Caddisflies are very closely related to butterflies and moths, and they look a lot like small moths with hairy wings held roof-like over the body.

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Following modified from NC State University
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TRICHOPTERA


Caddisflies

The name Trichoptera, derived from the Greek words " trichos " meaning hair and " ptera " meaning wings, refers to the long, silky hairs that cover most of the body and wings.
Immature and adult stages in the life cycle of a caddisfly (Limnophilus sp.) are depicted on this 1999 stamp from Poland


Classification

Life History & Ecology

Distribution

Physical Features

Economic Importance

Major Families

Fact File

Hot Links






Life History & Ecology:

The order Trichoptera (caddisflies) is another likely descendant of the Mecopteran lineage. Adults are mostly nocturnal, weak-flying insects that are often attracted to lights.   During the day, they hide in cool, moist environments such as the vegetation along river banks.   The body and wings are clothed with long silky hairs (setae) -- a distinctive characteristic of the order.   In flight, the hind wings are coupled to the front wings by specially curved hairs.   At rest the wings are held tent-like over the abdomen.   Many caddisflies have reduced or vestigal mouthparts.   Few species have actually been observed feeding, and most adults are relatively short-lived.

All caddisfly larvae live in aquatic environments; they may be herbivores, scavengers, or predators.   In most cases, the predatory species are free-living or spin silken structures in the water (webs or tunnels) to entrap prey.   The scavengers and herbivores live within protective "cases" which they build from their own silk and stones, twigs, leaf fragments, or other natural materials.   Case design and construction is distinctive for each family or genus of caddisfly.   The case is usually portable, dragged around like a snail shell as the insect moves, and held in place by a pair of hooked prolegs at the tip of the abdomen.   Most species have thread-like abdominal gills and get oxygen from water that circulates inside the case.   All larval growth and development (including pupation) occurs within the case.






Distribution:

Common worldwide.   Larvae are aquatic and may be abundant in some cool, fresh water habitats.   Adults are less conspicuous, usually nocturnal.
North America
Worldwide
Number of Families 18 43
Number of Species 1,261 >7,000





Classification:

Holometabola
complete development (egg, larva, pupa, adult)





Physical Features:

immature
adult
Immatures Adults
  1. Eruciform (caterpillar-like) body; abdomen usually enclosed in a case made of stones, leaves, twigs, or other natural materials.
  2. Head capsule well-developed with chewing mouthparts
  3. Thread-like abdominal gills usually present in case-makers
  4. One pair of hooked prolegs often present at tip of abdomen
  1. Filiform antennae
  2. Mouthparts reduced or vestigal
  3. Two pairs of wings clothed with long hairs
  4. Wings held tent-like over the abdomen
Examples of caddisfly cases





Economic Importance:

Caddisfly larvae may serve as food for fish and other aquatic vertebrates.   Fishermen often gather them for use as bait for trout and other game fish.   Although a few species have been recorded as pests in rice paddies, most caddisflies have very little economic importance.





Major Families:

  • Hydropsychidae -- Most larvae are filter feeders.   They build silk nets in swift water to snare food particles.   A few species are predatory.

  • Hydroptilidae -- Larvae make purse-like cases of silk, often with small stones attached.

  • Limnephilidae -- Larvae build tubular cases from a variety of natural materials.

  • Phryganeidae -- Larvae construct tubular cases with plant fragments arranged in a spiral orientation.





Fact File:

  • Many species of Trichoptera are very similar in appearance, both as larvae and as adults.   It is often easier to identify a species by the structure of its case than by the features of its body.

  • While still in their pupal case, caddisfly adults have sharp mandibles used for cutting through the pupal case.   Once they emerge, their mandibles degenerate and become nonfunctional.   From this time on they do not feed (or injest food only in liquid form).






Hot Links and Illustrations:





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Last Updated:   8 March 2005
Copyright 2005
John R. Meyer
Department of Entomology
NC State University

Following modified from Lucid via Discover Life
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A caddisfly resting on a leaf

Order - TRICHOPTERA
(Greek, thrix = hair; pteron = wing)
Common Name: caddisflies
Distribution: Cosmopolitan

Description
Adult caddisflies are soft bodied and very much resemble moths. The order Trichoptera is closely related to the order Lepidoptera . The mature insects have large compound eyes with two or three simple eyes (ocelli). Antennae are long and composed of short, straight segments (filiform). Mouthparts are reduced and adults are incapable of consuming solid food. Adults can ingest nectar or water but do not often feed if at all. Two pairs of wings are held at rest in a position which resembles the two sloping sides of a house roof. The wings may be hairy or scaled. Body colour is grey or brown and resting insects are extremely well camouflaged.


Larva
The larval stages are aquatic. Eggs are laid in sticky masses attached to twigs or rocks in the water. On hatching the larva may either build a case in which it lives or remain naked and build a conical net which is used to capture food. Case building larvae are mostly vegetarians and feed on detritus or may graze upon water plants. But others prey on small fresh water animals. Net building larvae are purely carnivorous. Net spinners are restricted to flowing water, but case makers frequent still water. On maturity, the larva pupates either within a cocoon or only as a pupal case protected by silk strands through which water flows. The pupa has free legs and large mandibles with which it chews through the cocoon or silk and swims to the surface. The pupa then splits and is used by the emergent adult as a floating platform.


Members
Caddisflies.


Food
Caddisflies feed on fresh vegetative material or detritus. Small freshwater animal life will also be consumed.

Importance
Caddisflies are very important as a food resource for fish in freshwater streams.

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Updated: 2009-11-22 03:30:03 gmt
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