D I S C O V E R    L I F E   
Bee Hunt! Odonata Lepidoptera 
  HomeAll Living ThingsIDnature guidesGlobal mapperAlbumsLabelsSearch
  AboutNewsEventsResearchEducationProjectsStudy sitesHelp


Tetraloniella wilmattae (Cockerell, 1917)
Xenoglossodes wilmattae Cockerell, 1917; Xenoglossodes pallidicauda Cockerell, 1934

Life   Insecta   Hymenoptera   Apoidea   Apidae   Tetraloniella
Subgenus: Tetraloniella


Click on map for details about points.

IDnature guide

Overview
Tetraloniella wilmattae (Cockerell), new combination

Xenoglossodes wilmattae Cockerell, 1917, in Cockerell, W.P., 1917, Jour. New York Ent. Soc., 25:191. Xenoglossodes pallidicauda Cockerell, 1934, American Mus. Nov. No. 697, p. 11. (new synonymy).

This small pale species is very similar to T. eriocarpi. The female of wilmattae differs from that of eriocarpi by having the labrum entirely or mostly black, the clypeal pale macula narrow (usually one-fourth of clypeus along midline) and without a pointed median extension posteromedially, and the metasomal terga 2–4 apical areas usually with relatively narrowly hyaline margins. The male of wilmattae can be immediately recognized by the form of the sixth metasomal sternum. In wilmattae the lateral teeth are somewhat smaller and the carina leaving the apical edge of each tooth proceeds medially forming a straight or convex line (the bulge facing apically). In addition, the males of wilmattae usually have the last three and one-half flagellar segments dark below and the vestiture is usually white instead of ochraceous.

FEMALE. Measurements and Ratios.- N = 20; length, 7.5–9.0 mm; width, 2.5–3.5 mm; wing length, M = 2.26±0.062 mm; hooks in hamulus, M = 10.35±0.167; flagellar segment 1/2, M = 1.42±0.029.

Integumental Color.- Integument piceous except as follows: clypeus with narrow apical band of yellow usually equaling onefourth or less of clypeus, in all specimens posterior margin slightly concave medially; mandible with basal yellow maculae (occasionally reduced); labrum dark with small mediobasal pale spot usually equal to one-third or less of labral area; tegulae hyaline, yellow to orange; terga 1–4 with apical areas hyaline, colorless to yellow in apical half, red to brown basally; distitarsi red; wing membranes hyaline, veins dark brown to reddish brown except costal vein usually yellow or orange.

Structure.- Clypeus, oculoclypeal distance, supraclypeal area as in eriocarpi. Face above antennal fossae as in eriocarpi but punctures usually separated by half a puncture width or less. Vertex and genal area as in eriocarpi. Galea as in eriocarpi, maxillary segmental ratio about as 1.0:0.7:0.7:0.5:0.3. Flagellar segments as in eriocarpi. Mesoscutum as in eriocarpi but anterior third with punctures separated mostly by half a puncture width. Propodeum sculptured as in eriocarpi but dorsal area punctures not elongate. Mesepisterna and metasomal terga sculptured as in eriocarpi. Pygidial plate and metasomal sterna as in eriocarpi.

Vestiture.- White to pale ochraceous, often darker or bright ochraceous on thoracic dorsum and vertex. Metasomal terga with vestiture as in eriocarpi but tergum 3 as well as tergum 2 with interband zone of short, suberect to erect hairs not hiding surface, a few highly plumose hairs may be scattered across the median fourth of tergum 3. Sternal hairs and scopal hairs as in eriocarpi.

MALE. Measurements and Ratios.- N = 20; length, 7–10 mm; width, 2.5–8.0; wing length, M = 2.39±0.098 mm; hooks in hamulus, M = 9.45±0.170; flagellar segment 2/1, M = 11.04±0.251.

Integumental Color.- Integument black except as follows: clypeus, base of mandibles, and labrum entirely yellow, flagellum red below and dark red above except last three and one half segments dark reddish brown below and dark brown above (in some individuals fewer than three and one half segments are darkened); wing membranes hyaline, colorless, veins dark brown to dark reddish brown except costal vein pale; terga with apical areas hyaline, often narrowly infuscated posteriorly; sterna piceous; distitarsi pale, tibial spurs pale ochraceous.

Structure.- Head sculptured as in female; maxillary palpus 5-segmented, in ratio of about 1.0:0.8:0.8:0.3:0.5; antennae long, flagellum round in cross-section, last five or six segments moderately crenulate. Mesoscutum with posteromedian area punctures sparse, separated mostly by two puncture widths or more, apical third with punctures separated by half a puncture width or slightly more, surface shiny. Metasomal terga sculptured as in eriocarpi; pygidial plate narrow, base not broader or only slightly broader than apex (unless worn), with weak subapical lateral notches. Tergum 7 with lateral arm of postgradulus carinate, without tooth. Sternum 7 with lateral carina forming small lateral teeth, end of each tooth with a carina extending mesally ending short of midline, carina straight from tooth to midline or forming a convex curve facing apically. Terminalia (Figs. 40– 41) essentially as in T. eriocarpi.

Vestiture.- White to pale ochraceous, occasionally brighter on thoracic dorsum and vertex; metasomal tergum 1 with apical pale pubescent band reduced to lateral fasciae each equal to one-fourth or less of width of tergum, never complete or nearly complete; metasomal terga as in eriocarpi but terga 3 and 4 and often 5 with area between basal and apical pale bands often with simple hairs not hiding surface as in tergum 2. Sternal hairs as in eriocarpi.

Type Material.- The holotype female of Tetraloniella wilmattae (CAS No. 15,555) was collected at Point Isabel (near Brownsville), Texas, in April 1917, from flowers of a small composite, by W.P. Cockerell. The holotype female of pallidicauda (AMNH) was collected in Bexar Co., Texas, by H.B. Parks. The holotype female has the clypeal yellow, macula reduced to two small yellow apicolateral maculae.

Distribution.- Tetraloniella wilmattae ranges from Texas south into northeastern México (Fig. 4). It has been taken from March 27 through December 24 but chiefly in April and May. In addition to the type specimen mentioned above, 105 females and 98 males were collected from localities listed below (the type locality is included).

México.- COAHUILA: Piedras Negras (192 km. S). DURANGO: El Salto. NUEVO LEON: Linares (10 mi. S and 12 mi. W); Vallecillo. TAMAULIPAS: El Limon; San Fernando (at the Río Conchos and 63 mi. N); Santander Jiménez.

United States.- TEXAS: Austin, Travis Co.; Ben Bolt, Jim Wells Co.; Bentzen-Rio Grande St. Park; Bexar Co., Brackettville; Brownsville (Del Mar); Cotula; Eastland County; Edinburg; Eldorado (10 mi. S); Fedor, Lee Co.; Fort Stockton (25 and 31 mi. SE), Pecos Co.; Harper; Hidalgo; Johnson City; Kerrville; Kingsville; Laredo; Lee County; Lopeno; Ozona; Palmetto St. Park, Gonzales Co.; Pharr (10 mi. SW); Point Isabel (near Brownsville); Sabinal; San Benito; Sonora; Southmost, Cameron Co.; Starr Co.; Stonewall; Streeter, Mason Co.; West (3 mi. W), McLennan Co.

Flower Records.- T. wilmattae is probably an oligolege of composites as is its close relative T. eriocarpi, and has been collected from flowers of the plants listed below.

Actinea sp.; Agastache breviflora; Callirhoe sp.; Coreopsis sp.; Gaillardia sp., G. suavis; Helenium latifolium, H. microcephalum; Medicago minima; Ratibida columnaris; Rudbeckia sp.; Tetragonotheca ludoviciana.


Identification
Extracted from Wallace E. LaBerge. 2001. Revision of the Bees of the Genus Tetraloniella in the New World (Hymenoptera: Apidae) (November 2001)

This small pale species is very similar to T. eriocarpi. The female of wilmattae differs from that of eriocarpi by having the labrum entirely or mostly black, the clypeal pale macula narrow (usually one-fourth of clypeus along midline) and without a pointed median extension posteromedially, and the metasomal terga 2–4 apical areas usually with relatively narrowly hyaline margins. The male of wilmattae can be immediately recognized by the form of the sixth metasomal sternum. In wilmattae the lateral teeth are somewhat smaller and the carina leaving the apical edge of each tooth proceeds medially forming a straight or convex line (the bulge facing apically). In addition, the males of wilmattae usually have the last three and one-half flagellar segments dark below and the vestiture is usually white instead of ochraceous.
FEMALE. Measurements and Ratios.- N = 20; length, 7.5–9.0 mm; width, 2.5–3.5 mm; wing length, M = 2.26±0.062 mm; hooks in hamulus, M = 10.35±0.167; flagellar segment 1/2, M = 1.42±0.029.
Integumental Color.- Integument piceous except as follows: clypeus with narrow apical band of yellow usually equaling onefourth or less of clypeus, in all specimens posterior margin slightly concave medially; mandible with basal yellow maculae (occasionally reduced); labrum dark with small mediobasal pale spot usually equal to one-third or less of labral area; tegulae hyaline, yellow to orange; terga 1–4 with apical areas hyaline, colorless to yellow in apical half, red to brown basally; distitarsi red; wing membranes hyaline, veins dark brown to reddish brown except costal vein usually yellow or orange.
Structure.- Clypeus, oculoclypeal distance, supraclypeal area as in eriocarpi. Face above antennal fossae as in eriocarpi but punctures usually separated by half a puncture width or less. Vertex and genal area as in eriocarpi. Galea as in eriocarpi, maxillary segmental ratio about as 1.0:0.7:0.7:0.5:0.3. Flagellar segments as in eriocarpi. Mesoscutum as in eriocarpi but anterior third with punctures separated mostly by half a puncture width. Propodeum sculptured as in eriocarpi but dorsal area punctures not elongate. Mesepisterna and metasomal terga sculptured as in eriocarpi. Pygidial plate and metasomal sterna as in eriocarpi.
Vestiture.- White to pale ochraceous, often darker or bright ochraceous on thoracic dorsum and vertex. Metasomal terga with vestiture as in eriocarpi but tergum 3 as well as tergum 2 with interband zone of short, suberect to erect hairs not hiding surface, a few highly plumose hairs may be scattered across the median fourth of tergum 3. Sternal hairs and scopal hairs as in eriocarpi.
MALE. Measurements and Ratios.- N = 20; length, 7–10 mm; width, 2.5–8.0; wing length, M = 2.39±0.098 mm; hooks in hamulus, M = 9.45±0.170; flagellar segment 2/1, M = 11.04±0.251.
Integumental Color.- Integument black except as follows: clypeus, base of mandibles, and labrum entirely yellow, flagellum red below and dark red above except last three and one half segments dark reddish brown below and dark brown above (in some individuals fewer than three and one half segments are darkened); wing membranes hyaline, colorless, veins dark brown to dark reddish brown except costal vein pale; terga with apical areas hyaline, often narrowly infuscated posteriorly; sterna piceous; distitarsi pale, tibial spurs pale ochraceous. Structure.- Head sculptured as in female; maxillary palpus 5-segmented, in ratio of about 1.0:0.8:0.8:0.3:0.5; antennae long, flagellum round in cross-section, last five or six segments moderately crenulate. Mesoscutum with posteromedian area punctures sparse, separated mostly by two puncture widths or more, apical third with punctures separated by half a puncture width or slightly more, surface shiny. Metasomal terga sculptured as in eriocarpi; pygidial plate narrow, base not broader or only slightly broader than apex (unless worn), with weak subapical lateral notches. Tergum 7 with lateral arm of postgradulus carinate, without tooth. Sternum 7 with lateral carina forming small lateral teeth, end of each tooth with a carina extending mesally ending short of midline, carina straight from tooth to midline or forming a convex curve facing apically. Terminalia (Figs. 40– 41) essentially as in T. eriocarpi.
Vestiture.- White to pale ochraceous, occasionally brighter on thoracic dorsum and vertex; metasomal tergum 1 with apical pale pubescent band reduced to lateral fasciae each equal to one-fourth or less of width of tergum, never complete or nearly complete; metasomal terga as in eriocarpi but terga 3 and 4 and often 5 with area between basal and apical pale bands often with simple hairs not hiding surface as in tergum 2. Sternal hairs as in eriocarpi.
Type Material.- The holotype female of Tetraloniella wilmattae (CAS No. 15,555) was collected at Point Isabel (near Brownsville), Texas, in April 1917, from flowers of a small composite, by W.P. Cockerell. The holotype female of pallidicauda (AMNH) was collected in Bexar Co., Texas, by H.B. Parks. The holotype female has the clypeal yellow, macula reduced to two small yellow apicolateral maculae.
Distribution.- Tetraloniella wilmattae ranges from Texas south into northeastern México (Fig. 4). It has been taken from March 27 through December 24 but chiefly in April and May. In addition to the type specimen mentioned above, 105 females and 98 males were collected from localities listed below (the type locality is included).
México.- COAHUILA: Piedras Negras (192 km. S). DURANGO: El Salto. NUEVO LEON: Linares (10 mi. S and 12 mi. W); Vallecillo. TAMAULIPAS: El Limon; San Fernando (at the Río Conchos and 63 mi. N); Santander Jiménez. United States.- TEXAS: Austin, Travis Co.; Ben Bolt, Jim Wells Co.; Bentzen-Rio Grande St. Park; Bexar Co., Brackettville; Brownsville (Del Mar); Cotula; Eastland County; Edinburg; Eldorado (10 mi. S); Fedor, Lee Co.; Fort Stockton (25 and 31 mi. SE), Pecos Co.; Harper; Hidalgo; Johnson City; Kerrville; Kingsville; Laredo; Lee County; Lopeno; Ozona; Palmetto St. Park, Gonzales Co.; Pharr (10 mi. SW); Point Isabel (near Brownsville); Sabinal; San Benito; Sonora; Southmost, Cameron Co.; Starr Co.; Stonewall; Streeter, Mason Co.; West (3 mi. W), McLennan Co.
Flower Records.- T. wilmattae is probably an oligolege of composites as is its close relative T. eriocarpi, and has been collected from flowers of the plants listed below. Actinea sp.; Agastache breviflora; Callirhoe sp.; Coreopsis sp.; Gaillardia sp., G. suavis; Helenium latifolium, H. microcephalum; Medicago minima; Ratibida columnaris; Rudbeckia sp.; Tetragonotheca ludoviciana.


Names
Scientific source:

Supported by
go to Discover Life's Facebook group

Updated: 2017-11-18 20:26:41 gmt
Discover Life | Top
© Designed by The Polistes Corporation