Index

Subfamily 1 Scarabaeinae

TRIBE 1 SCARABAEINI

Genus 1	Deltochilum Eschscholtz 1822 gibbosum gibbosum (Fabricius 1775)
Genus 2	Canthon Hoffmannsegg 1817 chalcites (Haldeman 1843) depressipennis LeConte 1859 pilularius (Linnaeus 1758) probus (Germar 1824) vigilans LeConte 1858 viridis viridis (Beauvois 1805)
Genus 3	Melanocanthon Halffter 1958 bispinatus (Robinson 1941)
Genus 4	Pseudocanthon Bates 1887 perplexus (LeConte 1847)

TRIBE 2 COPRINI

Genus 1	Ateuchus Weber 1801 histeroides Weber 1801 lecontei (Harold 1868)
Genus 2	Dichotomius Hope 1838 carolinus (Linnaeus 1767)
Genus 3	Copris Geoffroy 1762 fricator (Fabricius 1787) minutus (Drury 1770)
Genus 4	Phanaeus MacLeay 1819 igneus MacLeay 1819 triangularis triangularis (Say 1823) vindex MacLeay 1819

TRIBE 3 ONTHOPHAGINI

Genus 1

Onthophagus Latreille 1802 concinnus Laporte 1840 gazella (Fabricius 1787) hecate hecate (Panzer 1794) oklahomensis Brown 1927 orpheus orpheus (Panzer 1794) pennsylvanicus Harold 1871 striatulus striatulus (Beauvois 1809) subaeneus (Beauvois 1811) taurus (Schreber 1759) tuberculifrons Harold 1871

Index

Subfamily 2 Aphodiinae

TRIBE 1 AEGIALIINI

Genus 1

Aegialia Latreille 1807 blanchardi Horn 1887 cartwrighti Stebnicka 1977 humeralis Brown 1931

TRIBE 2 APHODIINI

Genus 1	Aphodius Illiger 1798 badipes Melsheimer 1845 bicolor Say 1823 brimleyi Cartwright 1939 campestris Blatchley 1912 crassulus Horn 1870 distinctus (Muller 1776) erraticus (Linnaeus 1758) femoralis Say 1823 fimetarius (Linnaeus 1758) granarius (Linnaeus 1767) haemorrhoidalis (Linnaeus 1758) lentus Horn 1870 leopardus Horn 1870 lividus (Olivier 1789) lodingi Cartwright 1957 lutulentus Haldeman 1843 nigrita Fabricius 1801 pseudolividus Balthasar 1941 robinsoni Cartwright 1939 rubeolus Beauvois 1805 rusicola Melsheimer 1845 serval Say 1837 stercorosus Melsheimer 1845 stupidus Horn 1870 terminalis Say 1823 vittatus Say 1825
Genus 2	Dialytes Harold 1869 striatulus (Say 1825) truncatus (Melsheimer 1846) ulkei Horn 1875
Genus 3	Dialytellus Brown 1929 dialytoides (Fall 1907) humeralis (LeConte 1878)

TRIBE 3 EUPARIINI

Genus 1	Pseudataenius Brown 1927 contortus Cartwright 1974
Genus 2	Aphotaenius Cartwright 1952 carolinus (Van Dyke 1928)
Genus 3	Saprosites Redtenbacher 1858 ventralis (Horn 1887)
Genus 4	Ataenius Harold 1867 abditus (Haldeman 1848) alternatus (Melsheimer 1844) apicalis Hinton 1937 brevis Fall 1930 cylindrus Horn 1871 erratus Fall 1930 exiguus Brown 1932 fattigi Cartwright 1948 figurator Harold 1874 gracilis (Melsheimer 1844) imbricatus (Melsheimer 1844) insculptus Horn 1878 miamii Cartwright 1934 ovatulus Horn 1871 picinus Harold 1867 platensis (Blanchard 1846) rhyticephalus (Chevrolat 1864) saxatilis Cartwright 1944 simulator Harold 1868 spretulus (Haldeman 1848) strigatus (Say 1823) wenzelii Horn 1887
Genus 5	Martineziella Chalumeau 1986 dutertrei (Chalumeau 1983)

TRIBE 4 PSAMMODIINI

Genus 1	Neopsammodius Rakovic 1986 interruptus (Say 1835)
Genus 2	Platytomus Mulsant 1842 atlanticus (Cartwright 1948) caelicollis (Cartwright 1948) longulus (Cartwright 1948) notialis (Cartwright 1948)

Index

Subfamily 3 Ochodaeinae

Genus 1

Ochodaeus Saint-Fargeau and Serville 1828 frontalis LeConte 1863 musculus (Say 1835)

Index

Subfamily 4 Hybosorinae

Genus 1

Hybosorus MacLeay 1819 illigeri Reiche 1853

Index

Subfamily 5 Geotrupinae

TRIBE 1 BOLBOCERATINI

Genus 1	Bolboceras Kirby 1819 alabamensis (Wallis 1929) cornigerus Melsheimer 1844 darlingtoni (Wallis 1928) liebecki (Wallis 1928) thoracicornis (Wallis 1928)
Genus 2	Bolbocerosoma Schaeffer 1906 farctum (Fabricius 1775) tumefactum (Beauvois 1805)
Genus 3	Eucanthus Westwood 1848 impressus Howden 1964 lazarus (Fabricius 1775) subtropicus Howden 1955

TRIBE 2 GEOTRUPINI

Genus 1

Geotrupes Latreille 1796 balyi Jekel 1865 blackburnii blackburnii (Fabricius 1781) egeriei Germar 1824 hornii Blanchard 1888 semiopacus Jekel 1865 splendidus (Fabricius 1775) ulkei Blanchard 1888

Index

Subfamily 6 Glaphyrinae

Genus 1

Lichnanthe Burmeister 1844 vulpina (Hentz 1827)

Index

Subfamily 7 Ceratocanthinae

Genus 1	Ceratocanthus White 1842 aeneus (MacLeay 1819)
Genus 2	Germarostes Paulian 1982 aphodioides (Illiger 1800) globosus (Say 1835)

Index

Subfamily 8 Troginae

Genus 1	Trox Fabricius 1775 aequalis Say 1831 affinis Robinson 1940 capillaris Say 1823 foveicollis Harold 1872 hamatus Robinson 1940 scaber (Linnaeus 1767) sordidus LeConte 1854 terrestris Say 1825 tuberculatus (DeGeer 1774) unistriatus Beauvois 1805 variolatus Melsheimer 1846
Genus 2	Omorgus Erichson 1847 asper LeConte 1854 monachus (Herbst 1790) scabrosus (Beauvois 1805) suberosus (Fabricius 1775) tytus (Robinson 1941)

Index

Subfamily 9 Melolonthinae

TRIBE 1 SERICINI

Genus 1

Serica MacLeay 1819 alabama Dawson 1967 aspera Dawson 1922 atracapilla (Kirby 1837) blatchleyi Dawson 1932 carolina Dawson 1920 delicata Dawson 1922 floridana Dawson 1967 georgiana georgiana Leng 1911 imitans Chapin 1931 intermixta Blatchley 1910 iricolor (Say 1824) loxia Dawson 1920 mystaca Dawson 1922 opposita Dawson 1921 parallela Casey 1884 sericea (Illiger 1802) trociformis Burmeister 1855 vespertina Gyllenhal 1817

Genus 2

Maladera Mulsant and Rey 1871 castanea (Arrow 1913)

TRIBE 2 MELOLONTHINI

Genus 1	Diplotaxis Kirby 1837 bidentata LeConte 1856 frondicola (Say 1825) harperi Blanchard 1850 liberta (Germar 1824) punctatorugosa Blanchard 1850 sordida (Say 1825) subcostata Blanchard 1850
Genus 2	Phyllophaga Harris 1827 aemula (Horn 1887) anxia (LeConte 1850) apicata Reinhard 1939 arkansana (Schaeffer 1906) barda (Horn 1887) calceata (LeConte 1856) clemens (Horn 1887) clypeata (Horn 1887) corrosa (LeConte 1856) crenulata (Froelich 1792) cupuliformis Langston 1924 debilis (LeConte 1856) diffinis (Blanchard 1850) dispar (Burmeister 1855) drakii (Kirby 1837) ephilida (Say 1825) fervida (Fabricius 1775) forsteri (Burmeister 1855) foxii Davis 1920 fraterna Harris 1842 fusca (Froelich 1792) futilis (LeConte 1850) georgiana (Horn 1885) glaberrima (Blanchard 1850) gracilis (Burmeister 1855) gracilis var. angulata Glasgow 1925 hirsuta (Knoch 1801) hirticula (Knoch 1801) ilicis (Knoch 1801) impar Davis 1920 implicita (Horn 1887) knochii (Schoenherrr and Gyllenhal 1817) latifrons (LeConte 1856) longispina (Smith 1889) lota Luginbill 1928 luctuosa (Horn 1887) marginalis (Horn 1856) mariana Fall 1929 micans (Knoch 1801) omani Sanderson 1937 parvidens (LeConte 1856) paternoi Glasgow 1925 perlonga Davis 1920 postrema (Horn 1887) prununculina (Burmeister 1855) quercus (Knoch 1801) soror Davis 1920 subpruinosa (Casey 1884) tristis (Fabricius 1781) ulkei (Smith 1889) uniformis (Blanchard 1850) vehemens (Horn 1887) vilifrons (LeConte 1856)
Genus 3	Polyphylla Harris 1841 comes Casey 1914

TRIBE 3 MACRODACTYLINI

Genus 1	Dichelonyx Harris 1827 albicollis (Burmeister 1855) diluta (Fall 1901) fuscula (LeConte 1856) linearis (Gyllenhal 1817) subvittata (LeConte 1856)
Genus 2	Macrodactylus Latreille 1825 angustatus (Beauvois 1805) subspinosus (Fabricius 1775)

TRIBE 4 HOPLIINI

Genus 1

Hoplia Illiger 1803 modesta Haldeman 1843 mucorea (Germar 1824) trifasciata Say 1825 trivialis Harold 1869

Index

Subfamily 10 Rutelinae

TRIBE 1 ANOMALINI

Genus 1	Anomala Samouelle 1819 binotata (Gyllenhal 1817) flavipennis flavipennis Burmeister 1844 innuba (Fabricius 1787) lucicola (Fabricius 1798) marginata (Fabricius 1792) mendica Casey 1915 minuta Burmeister 1844 oblivia Horn 1884 orientalis (Waterhouse 1875) parvula Burmeister 1844 semilivida LeConte 1878 umbra Casey 1915 undulata Melsheimer 1845
Genus 2	Strigoderma Burmeister 1844 arbicola (Fabricius 1792) pygmaea (Fabricius 1798)
Genus 3	Popillia Saint-Fargeau and Serville 1828 japonica Newman 1841

TRIBE 2 RUTELINI

Genus 1	Parastasia Westwood 1841 brevipes (LeConte 1856)
Genus 2	Pelidnota MacLeay 1819 punctata (Linnaeus 1758)
Genus 3	Cotalpa Burmeister 1844 lanigera (Linnaeus 1758)

Index

Subfamily 11 Dynastinae

TRIBE 1 CYCLOCEPHALINI

Genus 1	Dyscinetus Harold 1869 morator (Fabricius 1789)
Genus 2	Cyclocephala Latreille 1829 borealis Arrow 1911 lurida Bland 1863 parallela (Casey 1915) puberula LeConte 1863 seditiosa LeConte 1863

TRIBE 2 PENTODONTINI

Genus 1	Ligyrus Burmeister 1847 gibbosus gibbosus (DeGeer 1774) neglectus (LeConte 1847) relictus (Say 1825) subtropicus Blatchley 1922
Genus 2	Aphonus LeConte 1856 castaneus (Melsheimer 1846) tridentatus (Say 1823) variolosus (LeConte 1847)
Genus 3	Euetheola Bates 1888 humilis rugiceps (LeConte 1856)

TRIBE 3 ORYCTINI

Genus 1	Xyloryctes Hope 1837 jamaicensis (Drury 1773)
Genus 2	Strategus Kirby 1828 aloeus (Linnaeus 1758) antaeus (Drury 1773)

TRIBE 4 DYNASTINI

Genus 1

Dynastes MacLeay 1819 tityus (Linnaeus 1763)

TRIBE 5 PHILEURINI

Genus 1

Phileurus Latreille 1807 truncatus (Beauvois 1807) valgus valgus (Linnaeus 1758)

Index

Subfamily 12 Cetoniinae

TRIBE 1 GYMNETINI

Genus 1

Cotini Burmeister 1842 nitida (Linnaeus 1764)

TRIBE 2 CETONIINI

Genus 1	Stephanucha Burmeister 1842 areata (Fabricius 1775)
Genus 2	Euphoria Burmeister 1842 fulgida (Fabricius 1775) herbacea (Olivier 1790) inda (Linnaeus 1758) sepulcralis (Fabricius 1801)

TRIBE 3 CREMASTOCHEILINI

Genus 1	Psilocnemis Burmeister 1842 leucosticta Burmeister 1842
Genus 2	Cremastocheilus Knoch 1801 canaliculatus Kirby 1827 castaneae Knoch 1801 harrisii Kirby 1827 variolosus Kirby 1826

Index

Subfamily 13 Trichiinae

TRIBE 1 OSMODERMINI

Genus 1

Osmoderma Saint-Fargeau and Serville 1828 eremicola (Knoch 1801) scabra (Beauvois 1805)

TRIBE 2 TRICHIINI

Genus 1	Gnorimella Casey 1915 maculosa (Knoch 1801)
Genus 2	Trigonopeltastes Burmeister 1840 delta (Forster 1771)
Genus 3	Trichiotinus Casey 1915 affinis (Gory and Percheron 1833) bibens (Fabricius 1775) lunulatus (Fabricius 1775) piger (Fabricius 1775)

Index

Subfamily 14 Valginae

Genus 1

Valgus Scriba 1790 canaliculatus (Fabricius 1775) seticollis (Beauvois 1805)

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