Kinds of Scarabaeinae

Checklist of Dung Beetles

Paul Schoolmeester
Belgium

and

Sacha Spector
American Museum of Natural History

Scientific name -- Other names
Copris Geoffrey, 1762
   Copris acutidens Motschulsky, 1860
   Copris afgoi Cambefort, 1992
   Copris agnus Lansberge, 1875
   Copris algol Nguyen-Phung, 1989
   Copris amabilis Kolbe, 1914
   Copris amyntor Klug, 1855
   Copris anceus Olivier, 1789
   Copris andrewesi Waterhouse, 1891
   Copris angusticornis Arrow, 1933
   Copris angusutus Nguyen-Phung and Cambefort, 1986
   Copris anomiopseoides Boucomont, 1924
   Copris antares Ferreira, 1958
   Copris apicepunctatus Balthasar, 1942
   Copris arcturus Gillet, 1907
   Copris ares Zelenka, 1993
   Copris arimotoi Ochi and Araya, 1992
   Copris arizonensis Schaeffer, 1906
   Copris armatus Harold, 1869
   Copris armeniacus Falderman, 1835
   Copris armiger Gillet, 1910
   Copris arrowi Felsche, 1910
   Copris aspericollis Gillet, 1910
   Copris atropolitus Gillet, 1910
   Copris basilewskyi Ferreira, 1962
   Copris basipunctatus Balthasar, 1942
   Copris bellator Chevrolat, 1844
   Copris bengalensis Gillet, 1911
   Copris bidens Kolbe, 1893
   Copris bihamatus Balthasar, 1965
   Copris bootes Klug, 1855
   Copris boucardi Harold, 1869
   Copris bovinus Gillet, 1908
   Copris brachypterus Nomura, 1964
   Copris brittoni Ferreira, 1962
   Copris caelatus Fabricius, 1794
   Copris caliginosus Kohlmann and Cano and Delgado, 2003
   Copris cambeforti Nguyen-Phung, 1988
   Copris camerunus Felsche, 1904
   Copris capensis Waterhouse, 1891
   Copris cariniceps Felsche, 1910
   Copris carinicus Gillet, 1910
   Copris carmelita Fabricius, 1801
   Copris cassius Peringuey, 1901
   Copris celebensis Lansberge, 1886
   Copris colmanti Gillet, 1908
   Copris complexus Nguyen-Phung, 1989
   Copris compressipennis Gillet, 1910
   Copris confucius Harold, 1877
   Copris consobrinus Lansberge, 1886
   Copris coriarius Gillet, 1907
   Copris cornifrons Boheman, 1860
   Copris corniger Sahlberg, 1823
   Copris corpulentus Gillet, 1910
   Copris costaricensis Gahan, 1894
   Copris costaricensis_dolichocerus Matthews, 1961
   Copris cribratus Gillet, 1927
   Copris cribricollis Gillet, 1910
   Copris davisi Nguyen-Phung and Cambefort, 1986
   Copris davisoni Waterhouse, 1891
   Copris decellei Cambefort, 1992
   Copris delicatus Arrow, 1931
   Copris denticulatus Nguyen-Phung, 1988
   Copris diversus Waterhouse, 1891
   Copris doriae Harold, 1877
   Copris draco Arrow, 1906
   Copris dracunculus Ferreira, 1959
   Copris dudleyi Cambefort, 1992
   Copris eburneus Cambefort, 1992
   Copris elpheneroides Felsche, 1910
   Copris elphenor Klug, 1855
   Copris erratus Lansberge, 1886
   Copris erratus_masayukii Ochi and Kon, 2004
   Copris evanidus Klug, 1855
   Copris excisus Waterhouse, 1891
   Copris fallaciosus Gillet, 1907
   Copris fallax Felsche, 1910
   Copris felschei Reitter, 1893
   Copris fidius Olivier, 1789
   Copris formosanus Gillet, 1922
   Copris frankenbergerianus Balthasar, 1958
   Copris freyi Ferreira, 1961
   Copris fricator Fabricius, 1787
   Copris fricator_cartwrighti Robinson, 1941
   Copris fujiokai OchiandKon, 1996
   Copris fukiensis Balthasar, 1952
   Copris furciceps Felsche, 1910
   Copris garambae Cambefort, 1992
   Copris gazellarum Gillet, 1918
   Copris gibbulus Lansberge, 1886
   Copris gilleti Kolbe, 1907
   Copris gladiator Arrow, 1933
   Copris gopheri Hubbard, 1894
   Copris gracilis Waterhouse, 1891
   Copris graueri Gillet, 1910
   Copris halffteri Matthews, 1959
   Copris harrisi Waterhouse, 1891
   Copris hartli Frey, 1975
   Copris hatayamai Ochi and Araya, 1992
   Copris hidakai OchiandKon, 1996
   Copris hispanus Linnaeus, 1764
   Copris hispanus_cavolini Petagna, 1792
   Copris hispanus_persianus Miksic, 1958
   Copris howdeni Matthews and Halffter, 1959
   Copris humilis Gillet, 1908
   Copris hybridus Gillet, 1910
   Copris igualensis Warner, 1990
   Copris illotus Balthasar, 1942
   Copris imitans Felsche, 1910
   Copris incertus Say, 1835
   Copris inemarginatus Blatchley, 1918
   Copris inhalatus Quedenfeldt, 1884
   Copris inhalatus_grandis Nguyen-Phung and Cambefort, 1986
   Copris inhalatus_perturbator Peringuey, 1901
   Copris inhalatus_sanctaeluciae Nguyen-Phung and Cambefort, 1986
   Copris insidiosus Peringuey, 1901
   Copris integer Reiche, 1847
   Copris interioris Kolbe, 1897
   Copris iris Sharp, 1875
   Copris jacchoides Nguyen-Phung and Cambefort, 1987
   Copris jacchus Fabricius, 1775
   Copris jahi Nguyen-Phung and Cambefort, 1986
   Copris jucundus Gillet, 1933
   Copris kasagii Ochi and Kon, 1996
   Copris katangae Cambefort and Nguyen-Phung, 1996
   Copris keralensis Gill, 1986
   Copris kiuchii Masumoto, 1989
   Copris klapperichi Balthasar, 1942
   Copris klugi Harold, 1869
   Copris klugi_sierrensis Matthews, 1961
   Copris laeviceps Harold, 1869
   Copris laevigatus Gillet, 1927
   Copris laiiformis Nguyen-Phung and Cambefort, 1987
   Copris laioides Boucomont, 1932
   Copris laius Harold, 1868
   Copris laius_angusticornis Nguyen-Phung and Cambefort, 1987
   Copris lannathai Hanboonsong and Masumoto and Ochi, 2003
   Copris lecontei Matthews, 1961
   Copris lecontei_isthmiensis Matthews, 1961
   Copris lugubris Boheman, 1858
   Copris lunaris Linnaeus, 1758
   Copris macacus Lansberge, 1886
   Copris macclevei Warner, 1990
   Copris macer Peringuey, 1901
   Copris maesi Ratcliffe, 1998
   Copris magicus Harold, 1881
   Copris martinae Nguyen-Phung, 1988
   Copris martinezi Matthews and Halffter, 1968
   Copris matthewsi Delgado and Kohlmann, 2001
   Copris matthewsi_pacificus Delgado and Kohlmann, 2001
   Copris megaceratoides Waterhouse, 1891
   Copris megasoma Matthews and Halffter, 1959
   Copris mesacanthus Harold, 1878
   Copris mesacanthus_transvaalensis Nguyen-Phung, 1988
   Copris mexicanus Matthews and Halffter, 1959
   Copris mindorensis Ochi, 1992
   Copris minutissimus Kryzhanovskii and Medvedev, 1966
   Copris minutus Drury, 1773
   Copris misellus Peringuey, 1901
   Copris miyakei OchiandKon, 2004
   Copris moechus Leconte, 1854
   Copris moffartsi Gillet, 1907
   Copris mongkhoni Hanboonsong and Masumoto and Ochi, 2003
   Copris montivagus Gillet, 1908
   Copris morphaeus Gillet, 1933
   Copris mourgliai Zelenka, 1993
   Copris muticus Boheman, 1857
   Copris neglectus Moxey, 1963
   Copris nepos Gillet, 1908
   Copris nevinsoni Waterhouse, 1891
   Copris novaki Zelenka, 1992
   Copris nubilosus Kohlmann and Cano and Delgado, 2003
   Copris numa Lansberge, 1886
   Copris obenbergeri Balthasar, 1933
   Copris obesus Boheman, 1857
   Copris ochus Motschulsky, 1860
   Copris orion Klug, 1835
   Copris orion_caffer Gillet, 1908
   Copris orion_centralis Gillet, 1908
   Copris orion_overlaeti Balthasar, 1961
   Copris orphanus Guerin, 1847
   Copris pauliani Nguyen-Phung and Cambefort, 1986
   Copris pecuarius Lewis, 1884
   Copris pedarioides Lansberge, 1886
   Copris phungae Cambefort, 1992
   Copris phungae_gabonicus Cambefort, 1992
   Copris phylax Gillet, 1908
   Copris potanini Semenov, 1890
   Copris propinquus Felsche, 1910
   Copris pseudobootes Ferreira, 1962
   Copris pseudosinicus Balthasar, 1958
   Copris pueli Mollandin De Boissy, 1905
   Copris punctatus Gillet, 1910
   Copris puncticollis Boheman, 1857
   Copris punctipennis Boucomont, 1914
   Copris punctiventris Waterhouse, 1891
   Copris punctulatus Wiedemann, 1823
   Copris punjabensis Gillet, 1921
   Copris ramosiceps Gillet, 1921
   Copris rebouchei Harold, 1869
   Copris reflexus Fabricius, 1787
   Copris remotus Leconte, 1866
   Copris remotus_dicyrtus Matthews and Halffter, 1959
   Copris renwarti Nguyen-Phung and Cambefort, 1986
   Copris repertus Walker, 1858
   Copris ribbei Lansberge, 1886
   Copris riekoae Ochi and Kon, 2003
   Copris ritsemae Harold, 1875
   Copris ruandanus Balthasar, 1961
   Copris rugosus Gillet, 1908
   Copris ruricola Balthasar, 1933
   Copris sabinus Gillet, 1910
   Copris sacontala Redtenbacher, 1848
   Copris sallei Harold, 1869
   Copris sarpedon Harold, 1868
   Copris saundersi Harold, 1869
   Copris scorpio Zelenka, 1993
   Copris serius Nguyen-Phung, 1987
   Copris sexdentatus Thunberg, 1818
   Copris siamensis Gillet, 1910
   Copris signatus Walker, 1858
   Copris simonettai Cambefort, 1992
   Copris simplex Harold, 1869
   Copris simulator Nguyen-Phung and Cambefort, 1986
   Copris singularis Nguyen-Phung and Cambefort, 1986
   Copris sinicus Hope, 1842
   Copris sinon Olivier, 1789
   Copris sodalis Walker, 1858
   Copris sorex Balthasar, 1942
   Copris sphaeropterus Harold, 1877
   Copris spinator Harold, 1881
   Copris subdolus Balthasar, 1958
   Copris subpunctatus Gillet, 1910
   Copris subsidens Peringuey, 1900
   Copris sumatrensis Gillet, 1921
   Copris surdus Arrow, 1931
   Copris szechouanicus Balthasar, 1958
   Copris tetraodon Gillet, 1910
   Copris tridens Felsche, 1901
   Copris tridentatus Solis and Kohlmann, 2003
   Copris tripartitus Waterhouse, 1875
   Copris truncatus Felsche, 1901
   Copris tullius Olivier, 1789
   Copris typhoeus Gerstaecker, 1884
   Copris uenoi Hanboonsong and Masumoto and Ochi, 2003
   Copris umbilicatus Abeille De Perrin, 1901
   Copris urus Boheman, 1857
   Copris usambaricus Gillet, 1908
   Copris vankhaii Nguyen-Phung, 1988
   Copris vankhaii_genopunctatus Nguyen-Phung, 1988
   Copris victorini Boheman, 1857
   Copris vietnamicus Kabakov, 1994
   Copris vilhenai Ferreira, 1962
   Copris vitalisi Gillet, 1921
   Copris vrydaghi Ferreira, 1962
   Copris wiesei Kolbe, 1914
   Copris youngai Balthasar, 1967

Updated: 2011-01-06 15:25:03

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