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Superfamily EVANIOIDEA
Family EVANIIDAE

Members of this family are parasitic within the egg cases of cockroaches. Known hosts of the
evaniids found in North America are members of the cockroach families Blattidae and Blatel-
lidae. Species of Evaniidae are most numerous in the tropics.

Revision: Bradley, 1908. Amer. Ent. Soc, Trans. 34: 128-162. -Townes, 1949. U. S. Natl.
Mus., Proc. 99: 525-539.

Taxonomy: Hedicke, 1939. Hym. Cat., pt. 9, 50 p. (world catalog).

Genus EVANIA Fabricius

Ei'a)iia Fabricius, 1775. Systema Ent., p. 345.

Type-species: Ichneutiion appendigaster Linnaeus. Desig. by Curtis, 1829.

According to Townes (1958) Ecaiiia is comprised of larger species which occur mostly in the
Oriental Region.

Taxonomy: Townes, 1958. Ins. of Micronesia, v. 19, no. 2, p. 84.

Evania appendigaster (Linnaeus)
Southeastern Mass., s.e. N. Y., s.e. Pa., D. C, Ga., Fla., Tenn., La.,
Tex., s. N. Mex., s. Ariz.; West Indies and Mexico s. to n. Argentina; Old World
(especially warmer parts). Adventive. Host: Blatta orientalis L.?, Periplaneta
a)iierico)ia (L.). I have found no published rearing records for North America, but
because appendigaater has been reared from P. amevicana in other parts of the world
there is no reason to doubt that it serves as a host in North America. I am not aware of




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records of appeudigaster as a parasite of B. orieyitalis which have been confirmed by
authoritative identification of the adult parasites. Therefore, I do not feel certain that
such records do not in fact apply to Prosevania fuscipes (Illiger). North of a latitude of
approximately 37 degrees in the eastern United States appendigaster apparently ranges
inland only rather short distances from the Atlantic coast. Some of the names cited in
the synonymy below will possibly prove to be synonyms of Prosevania fiiscipes, but
Westwood's (1844) description of a specimen in "the Linnaean cabinet" (presumably the
appemUgaHtev type) seems to indicate that the name appendigaster correctly applies to
the present species.

Ichneinnon appendigaster Linnaeus, 1758. Syst. Nat., ed. 10, p. 566. Sex not indicated.

Ichneumon niger Goeze, 1780. In Degeer, Des ... Degeer Abhandl. zur Gesch. des Insecten,
V. 3, p. 285. Sex not indicated?

Evania laevigata Olivier, 1791. Encycl. Meth., Diet. Ins., v. 6, p. 453. ♀.

Evania iinicolor Say, 1824. //( Keating, Narr. Long's 2nd Exped., v. 2 (App.), p. 320. Sex
not indicated.

Evania Ciibae Guerin-Meneville, 1834. Iconogr. Regne Anim., pi. 65, figs. 1-2. 6. For date
of publication see Cowan (1971, p. 28-29).

Evania Desjardinsii Blanchard, 1840. Hist. Nat. Anim. Artie, Ins., v. 3, p. 299. ♂.

Evania affinis Le Guillou, 1841. Soc. Ent. de France, Ann. 10: 311. cJ. ,

Evania appendigastra Hedqvist, 1973. Ent. Tidskr. 94: 178. Emend.

Taxonomy: Westwood, 1844 (1843). Ent. Soc. London, Trans. 3: 241-242. —Cowan, 1971. Soc.
Bibliog. Nat. Hist., Jour. 6: 18-29 (dating of E. cubae).

Biology: Cameron, 1957. Bui. Ent. Res. 48: 199-209. —Roth and Willis, 1960. Smithsonian
Misc. Coll. 141: 236.

Genus PROSEVANIA Kieffer

Evania subg. Prosevania Kieffer, 1911. Soc. Ent. de France, Ann. 80: 157.

Type-species: Prosevania afra Kieffer. Desig. by Viereck, 1914. Described as a
subgenus, but in the case of afra treated nomenclatorially as a genus.

This is a large genus of the Old World Tropics.

Prosevania fuscipes (Illiger)
n. comb. Northwestern Mass. w. to s. Ohio, s. to n.e. N. C. and Ga.; s.e.
Europe, n.w. Asia, n. Africa. Adventive. Host: Blatta orientalis L., Periplaneta
aniericana (L.). The name punctata (Brulle) has erroneously been treated as the senior
name for this species for many years (since 1919 in North America). The names Evania

Prosevania fuscipes
***authority mismatch
Illiger (1807) and E.flavicomis Curtis (1829) were proposed for E.
appendigaster Linnaeus as misidentified by Panzer (1799), Latreille (1805), and Jurine
(1807). There is little reason to doubt that the species identified as appendigaster by any
of these three authors is the present species, but Panzer's figure in particular leaves no
doubt whatsoever and is, therefore, here selected as lectotype for both the names E.

Prosevania fuscipes
***authority mismatch
Illiger and E.flavicomis Curtis.

Evania fuscipes Illiger, 1807. In Rossi, Fauna Etrusca, ed. 2, v. 2, p. 83. [male].

Evania Jlavicornis Curtis, 1829. Brit. Ent., v. 6, pi. 257. [male].

Evania punctata Brulle, 1833. Exped. Sci. de Moree, v. 3, p. 378. sex not indicated. N. syn.

Evania urbana Bradley, 1908. Amer. Ent. Soc, Trans. 34: 140. ♂, ♀. N. syn.

Taxonomy: Panzer, 1799. Faunae Ins. German., fasc. 62, pi. 12. —Latreille, 1805. Hist. Nat.
Crust. Ins., v. 13, p. 193. —Jurine, 1807. Nouv. Meth. Class. Hym. Dipt., v. 1, p. 85 and pi. 7,
fig. 6. — Bequaert, 1919. Brooklyn Ent. Soc, Bui. 14: 23.

Biology: Edmunds, 1952; 1953. Ohio Jour. Sci. 52: 29-30; 53: 121-122. —Edmunds, 1954. Ent.
Soc. Amer., Ann. 47: 575-592.

Genus EVANIELLA Bradley

Evaniella Bradley, 1905. Canad. Ent. 37: 64.

Type-species: Evaiiia unicolor Say as misidentified by Ashmead (1901). Orig. desig.
According to Bradley (1908), Evaniella seniaeoda Bradley is the species which
Ashmead identified as unicolor.




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This is a large genus of Australian, Nearctic, and Neotropic distribution (particularly the
latter).

Taxonomy: Ashmead, 1901. Canad. Ent. 33: 304. —Bradley, 1908. Amer. Ent. Soc, Trans. 34:
142-147.

Evaniella californica (Ashmead)
East-central Calif.

Evania Califontica Ashmead, 1901. Canad. Ent. 33: 304. ♂.

Taxonomy: Snelling, 1963. Pan-Pacific Ent. 39: 108 (female described).

Evaniella neomexicana (Ashmead)
Southeastern N. Mex., s.e. Calif.

Eraiiia Neotitexicana Ashmead, 1901. Canad. Ent. 33: 304. ♂.

Evaniella semaeoda Bradley
R. I. w. to s. Mich, and n.e. Kans., s. to s. Fla. and s. La.; West Indies.

Evaniella semaeoda Bradley, 1908. Amer. Ent. Soc, Trans. 34: 144. ♂, ♀.

Genus HYPTIA Illiger

Hyptia Illiger, 1807. /*( Rossi, Fauna Etrusca, ed. 2, v. 2, p. 82. Appeared as "Hyptiani" in
accusative case, Hyptia being the nominative equivalent. Fauna Etrusa, ed. 2, v. 2 not
necessarily published prior to volume six of Mag. f. Insectenkunde, but usually treated
as prior in synonymies pertaining to various other generic names.

Hyptia Illiger, 1807. Mag. f. Insektenkunde 6: 192.

Type-species: Evania petiolata Fabricius. Monotypic.

This is a large genus of the New World, with many more Neotropic species than Nearctic
ones. Rearing data are few, but the species appear to be parasites of forest dwelling Blattoidea
such as Parcoblatta spp.

Hyptia femorata Townes
Western Tex.

Hyptia f em orata Townes, 1949. U. S. Natl. Mus., Proc. 99: 537. ♂, ♀.

Hyptia floridana Ashmead
Western N. C. w. to n. La., s. to s. Fla.; Guatemala, Panama.

Hyptia Floridana Ashmead, 1901. Canad. Ent. 33: 303. ♂.

Hyptia harpyoides Bradley
N. H. w. to s.e. Minn., s. to n.w. S. C, n. Miss., and n. Kans. Host:

Parcoblatta uiileriana (Sauss.), P. virginica (Brunn.). This species occurs farther north
than any other evaniid in North America.
Hyptia harpyoides Bradley, 1908. Amer. Ent. Soc, Trans. 34: 151. ♂, ♀.
Hyptia brericalcar var. glabriceps Kieffer, 1910. Soc Ent. de France, Ann. 79: 72. ♂.

Hyptia oblonga Townes
Southwestern Ga., s. Ala., and s.e. Ariz. s. to s. Fla.; s. Mex., Costa Rica.

Hyptia oblonga Townes, 1949. U. S. Natl. Mus., Proc 99: 538. ♂, ♀.

Hyptia reticulata (Say)
Mass. w. to mid Mich., s. to s. Fla., s. La., and s.w. Tex.; s. Mexico.

Brachygaster reticulata Say, 1836. Boston Jour. Nat. Hist. 1: 223. sex not indicated.
Hyptia nyctoides Bradley, 1908. Amer. Ent. Soc, Trans. 34: 159. ♂.
Hyptia prosetetlietra Bradley, 1908. Amer. Ent. Soc, Trans. 34: 160. ♂.

Hyptia thoracica (Blanchard)
Mass., s. Ont, Wis., and n.e. Kans. s. to s. Fla. and s. Tex. Host:
Parcoblatta sp., P. pe)isyU'a)iica (Deg.).
Erania thoracica Blanchard, 1840. Hist. Nat. Anim. Artie, Ins., v. 3, p. 299. Sex not

indicated.
Hyptiam thoracicum Shuckard, 1841. Entomologist 1: 120. 6. Preocc by Blanchard, 1840.
Evania dorsalis Westwood, 1851. Ent. Soc London, Trans. (2) 1: 214. Unnecessary n. name

for E. thoracica Blanchard, there being no E. thoracica Klug or Leach.
Evania brevicalcar Kieffer, 1904. Arkiv for Zool. 1: 541. ♂.
Hyptia mijlacridomanes Bradley, 1908. Amer. Ent. Soc, Trans. 34: 153. ♀.
Hyptia hi'jptiogastris Bradley, 1908. Amer. Ent. Soc, Trans. 34: 160. ♂.
Hyptia texana Bradley, 1908. Amer. Ent. Soc, Trans. 34: 161. ♀.


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