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Superfamily APOIDEA
Family ANDRENIDAE

This is a large family of pollen-collecting bees which is present on all the continents except
Australia. Nearly all of the species excavate their nests in the ground and most are solitary
although a few species in each of the two subfamilies live in colonies. Their intrafloral relation-
ships are varied and range from numerous examples of oligolecty in varying degrees to broad
patterns of polylecty.

Even though both subfamilies are well represented in America north of Mexico by a great
many species, the majority of our fauna consists of the chiefly Holarctic genus Andrena and the
Nearctic genus Perdita.

Taxonomy: Rozen, 1951. Kans. Ent. Soc, Jour. 24: 142-150, 17 figs, (male genitalia).
—Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 86-330, figs. 1, 2, 4, 15-79, tables 3-9
(eastern U. S. spp.). —Yager and Rozen, 1966. Amer. Mus. Novitates 2265: 1-13, 18 figs,
(pupae).

Subfamily ANDRENINAE

Taxonomy: Yager and Rozen, 1966. Amer. Mus. Novitates 2265: 5-6 (pupae).

Genus ANDRENA Fabricius

Since studies of the relationships of the numerous subgenera within the genus Andrena are
incomplete, the subgenera are arranged alphabetically in this catalog. Those species currently
not assigned to any subgenus are placed after the recognized subgenera.

Taxonomy: Cockerell, 1898. N. Mex. Univ., Bui. 1: 48 (N. Mex. spp.) (reprint: Denison Univ.
Sci. Labs., Bui. 11: 48). —Cockerell, 1899. Ent. News. 10: 253 (misc.). —Cockerell, 1901.
Ent. News 12: 74 (misc.). —Cockerell, 1901. N. Y. Ent. Soc, Jour. 9: 132 (N. J. spp.).
—Robertson, 1902. Amer. Ent. Soc, Trans. 28: 187 (111. spp.). — Bruner, 1903. Amer. Ent.
Soc, Trans. 29: 240 (N. Amer. spp.). — Viereck, 1904. Canad. Ent. 36: 157, 189, 221 (Pacific
Northwest spp.). —Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 429-437 (Colo. spp.).
—Viereck, 1907. Ent. News 18: 280 (Conn. spp.). —Cockerell, 1907. Colo. Univ., Studies 4:
244 (Colo. spp.). —Cockerell, 1914. Ann. and Mag. Nat. Hist. (8) 14: 10 (Colo. spp.).
—Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc 48: 1 (Rocky Mts. spp.). —Viereck,
1917 (1916). Conn. State Geol. and Nat. Hist. Survey, Bui. 22: 709-720 (Conn. spp.).
—Viereck, 1924. Canad. Ent. 56: 20 (subgenera). —Cockerell, 1930. Pan-Pacific Ent. 7: 7
(misc.). —Cockerell, 1931. Amer. Mus. Novitates 458: 1-20 (Rocky Mts. spp.). —Cockerell,
1932. Canad. Ent. 64: 155-158 {Micrandrena). —Cockerell, 1932. Pan-Pacific Ent. 8:
173-177 (misc. Calif, spp.). -Cockerell, 1932. Canad. Ent. 64: 285-288 {sladeni group).
— Atwood, 1934. Canad. Jour. Res. 10 (2): 206-214, figs. 31-91 (Nova Scotia spp.).
—Cockerell, 1937. Amer. Mus. Novitates 948: 15 {Diandrena, pars). —Linsley, 1938. Calif.
Acad. Sci., Proc. (4) 23: 263-282 (black spp.). — Lanham, 1941. Ent. Soc. Amer., Ann. 34:
702-713 (Colo. spp.). —Lanham, 1949. Pan-Pacific Ent. 25: 33-35 (females of carlini group).
— Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 183-238 (subgenera). — Timberlake, 1951. U. S.
Natl. Mus., Proc. 101: 373-414 (misc Calif, spp.). —Linsley and MacSwain, 1955.
Pan-Pacific Ent. 31: 163-172 (N. Amer. spp. of Melandrena). —Linsley and MacSwain,
1956. Pan-Pacific Ent. 32: 111-121 {Oenothera visiting spp.). —Mitchell, 1960. N. C. Agr.




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Expt. Sta. Tech. Bui. 141: 86-257, figs. 2, 4, 6-57, tables 3-7 (eastern U. S. spp.). — Linsley
and MacSwain, 1961. Pan-Pacific Ent. 37: 31-41 (Diandrena spp. associated with flowers
of Oenothera). —Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 117-130
(O^iagraudrena spp. associated with the flowers of Clarkia and Oenothera). —Linsley and
MacSwain, 1962. Pan-Pacific Ent. 38: 49-52 (N. spp. of Onagrandrena). —Linsley and
MacSwain, 1963. Pan-Pacific Ent. 39: 189-198 (N. spp. of Onagrandrena). — Knerer and
Atwood, 1964. Ent. Soc. Ontario, Proc. 94: 41-56 (Ontario spp.). — LaBerge, 1964. Nebr.
Univ. State Mus., Bui. 4: 279-316 (N. Amer. subgenera). — Ribble, 1967. Nebr. Univ. State
Mus., Bui. 6: 27-42, figs. 1-5, 1 table, 1 map (subg. Larandrena). —LaBerge, 1967. Nebr.
Univ. State Mus., Bui. 7: 1-316, 340 figs., 4 tables, 14 maps (subg. Callandreyia).
— Warncke, 1968. Coimbra Univ. Est. Mus. Zool., Mem. 307: 1-110 (west. Palaearctic
subgenera). —Ribble, 1968. Kans. Ent. Soc, Jour. 41: 220-236, 15 figs., 1 map, 1 table (subg.
Belandrena). —Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 237-394, 124 figs., 18 maps
(subgenera Derandrena and M icrandrena). —Thorp, 1969. Calif. Univ. Pubs. Ent. 52:
1-46, 88 figs., 5 tables, 13 maps (subg. Diandrena). —LaBerge, 1969. Amer. Ent. Soc,
Trans. 95: 1-47, 38 figs., 3 tables (subgenera Aporandrena, Charitandrena, Plastandrena).
—LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 543-605, 75 figs., 5 maps
(subg. Tylandrena). —Warncke, 1970. Bayerischen Ent. Nachr. 19: 28-32 (lectotypic
designations). —LaBerge, 1971. Pan-Pacific Ent. 47: 47-57, 15 figs. (subg. Nemandrena).
—LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 441-520, 68 figs., 1 table (subgenera
Rhaphandrena, Scrapteropsis, Xiphandrena). —LaBerge and Ribble, 1972. Amer. Ent.
Soc, Trans. 98: 27-358, 76 figs, (subgenera Geissandrena, Gonandrena, Parandrena,
Pelicandrena). —LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 235-371, 69 figs., 2 tables, 15
maps (subg. Trachandrena). —Ribble, 1974. Amer. Ent. Soc, Trans. 100: 101-189, 99 figs.,
5 tables (subg. Scaphandrena). —LaBerge and Ribble, 1975. Amer. Ent. Soc, Trans. 101:
371-446, 74 figs. (subg. Euandrena). —Donovan, 1977. Calif. Univ. Pubs. Ent. 81: 1-107,
130 figs., 17 maps (subg. Cnemidandrena; received too late for inclusion in this catalog).

— LaBerge, 1977. Amer. Ent. Soc, Trans. 103: 1-143, 128 figs., 3 tables (subgenera
Dasyandrena, Euandrena, Oxyandrena, Psammandrena, Rhacandrena, Thysandrena;
received too late for inclusion in this catalog).

Biology: Graenicher, 1905. Wis. Acad. Sci., Arts and Letters 15: 89-97 (floral relationships).

— Linsley, 1937. Brooklyn Ent. Soc, Bui. 32: 125-127 (occurrence of double broods in N.
Amer. spp.). —Linsley, MacSwain and Smith, 1955. Pan-Pacific Ent. 31: 173-185 (floral
relationships of Me land rena). —Linsley and MacSwain, 1959. Calif. Univ. Pubs. Ent. 16:
1-46, 9 plates, 6 textfigs. (Ranunculus visiting spp.). — Linsley, MacSwain and Raven,
1963. Calif. Univ. Pubs. Ent. 33: 1-58, 6 plates, 6 textfigs. (comparative behavior of spp.
visiting flowers of Onagraceae in the Colorado Desert and Great Basin). —Linsley,
MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 59-98, 3 plates, 17 tables
(comparative behavior of spp. visiting flowers of Onagraceae in the Mojave Desert).
—Stephen, 1966. Kans. Ent. Soc, Jour. 39: 42-50, 2 figs, (comparative bionomics).

— Youssef and Bohart, 1968. Kans. Ent. Soc, Jour. 41: 442-444 (literature references to
nesting habits). — Cruden, 1972. Madrono 21: 505-515, 1 fig., 2 tables {Nemophila visiting
spp.). —Cruden, 1972. Evolution 26: 373-389, 8 figs., 9 tables (oligolectic spp. of Nemophila
menziesii, evolution of oligolectic bees). — MacSwain, Raven and Thorp, 1973. Calif. Univ.
Pubs. Ent. 70: 1-80, 3 plates, 10 textfigs., 18 tables (comparative behavior of Clarkia
visiting spp.). —Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71:
1-68, 6 plates, 15 textfigs., 10 tables (comparative behavior oi Camissonia and Oenothera
visiting spp.). —Davis and LaBerge, 1975. 111. Nat. Hist. Survey, Biol. Notes 95: 2
(literature references to nesting habits).

Genus ANDRENA Subgenus ANDRENA Fabricius

Andrena Fabricius, 1775. Systema Ent., p. 376.

Type-species: Apis helvola Linnaeus. Desig. by Viereck, 1912.
Anthrena Illiger, 1801. Mag. Insektenk. 1: 127. Emend.
Anthocharessa Gistel, 1850. Isis (Encycl. Ztschr.) 6: 82.

Type-species: Apis helvola Linnaeus. Autobasic; proposed unnecessarily for
Andrena Fabricius which Gistel considered to be preoccupied by Anthrenus
Geoffroy, 1764.




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Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 205 (tax. characters). —Mitchell, 1960.

N. C. Agr. Expt. Sta. Tech. Bui. 141: 104-110, figs. 15-16, 20-22, table 3 (eastern U. S. spp.).

—Lanham, 1965. Kans. Ent. Soc, Jour. 38: 198-201 (Salix visiting spp. of the Colorado

Front Range).

Andrena advarians Viereck
B. C.

Andrena advarians Viereck, 1904. Canad. Ent. 36: 192, 224. ♀, ♂.

Andrena albihirta (Ashmead)
Rocky Mts. and Pacific Coast. Pollen: Collects pollen from Salix,
including S. laevigata, S. lasiolepis.

Cilissa albihirta Ashmead, 1890. Colo. Biol. Assoc, Bui. 1: 5. ♀.

Andrena bebbiana Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 39. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 8 (tax. characters). —Lanham, 1941.
Ent. Soc. Amer. Ann. 34: 705, 708. (tax characters, as bebbiana) female, male (key).
— Lanham, 1965. Kans. Ent. Soc, Jour. 38: 199-200 (tax. status, synonymy).

Biology: Lanham, 1974. Kans. Ent. Soc, Jour. 47: 373-377, 1 fig., 1 table (effect of melanism

on adult behavior, floral relationships, stylopization).

Andrena albosellata Cockerell
Wyo., Colo. Predator: Philanthus crabroniformis Sm.

Andrena albosellata Cockerell, 1931. Amer. Mus. Novitates 458: 7. ♀.

Andrena asmi Viereck
Wash.

Andreyia asmi Viereck, 1904. Canad. Ent. 36: 192, 225. ♀, ♂.

Andrena banffensis Viereck
Alta.

Andrena (Andrena) banffensis Viereck, 1924. Canad. Ent. 56: 32. ♀.

Andrena bella Viereck
B. C.

Andrena {Andrena) bella Viereck, 1924. Canad. Ent. 56: 22. ♀.

Andrena birtwelli Cockerell
N. Mex., Colo., Calif. Pollen: Unknown, but visits flowers of Antennaria,
Eriogonum, Potentilla, Solidago.
Andrena birtwelli Cockerell, 1901. Psyche 9: 283. ♀.
Andrena birtwelli subatrata Cockerell, 1931. Amer. Mus. Novitates 458: 8. ♀.

Taxonomy: Viereck, 1903. Amer. Ent. Soc, Trans. 29: 54. 6. —Lanham, 1941, Ent. Soc.
Amer., Ann. 34: 705. 9 (key).

Andrena buckelli Viereck
B. C.

Ayidrena (Andrena) buckelli Viereck, 1924. Canad. Ent. 56: 22. ♀, ♂.

Andrena carrikeri Viereck and Cockerell
Nebr.

Aiidrena carrikeri Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 14. ♂.

Andrena ceanothifloris Linsley
South. Calif. Pollen: Possibly an oligolege of Ceanothus including C.
cordulatus, C. integerrinius, C. leucodemiis, but visits other flowers including
Arctostaphylos patula, Eriodictyon californicum, Horkelia, Ribes.
Andrena ceanothifloris Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 270. ♀, ♂.

Andrena clarkella (Kirby)
Holarctic; Alaska to Ont., Que., Mass. and Maine, south to Colo., Minn, and
N. Y.; Europe. Pollen: Apparently an oligolege of Salix.
Melitta Clarkella Kirby, 1802. Monog. Apum Angliae, v. 2, p. 130. ♀.

Taxonomy: Sladen, 1919. Canad. Ent. 51: 124. 9, 6. —Mitchell, 1960. N. C. Agr. Expt. Sta.

Tech. Bui. 141: 104-105, figs. 15-16, 21, table 3 (redescription). —Lanham, 1963. Pan-Pacific

Ent. 40: 9 (geogr. records). —Lanham, 1965. Kans. Ent. Soc, Jour. 38: 199 (floral and

geogr. records).

Andrena clypeoporaria Viereck
Wash., Oreg.

Andrena clypeoporaria Viereck, 1904. Canad. Ent. 36: 192. ♀.

Andrena cristata Viereck
Nev.

Andrena (Andrena) cristata Viereck, 1917. Acad. Nat. Sci. Phila., Proc. 68: 556. ♀.

Andrena diversicolor Viereck
Alta.

Andrena (Andrena) diversicolor Viereck, 1924. Canad. Ent. 56: 76. ♀.

Andrena edwardsi Viereck
Idaho.

Andrena edwardsi Viereck, 1916. Amer. Mus. Nat. Hist., Bui. 35: 731. ♀.

Andrena edwiniae Cockerell
Colo.

Andrena edwiniae Cockerell, 1906. Ann. and Mag. Nat. Hist. (7) 17: 310. ♀.




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Biology: Clements and Long, 1923. Carnegie Inst. Wash., Pub. 326: 249 (ecology).

Andrena enigmatica Viereck and Cockerell
Nebr.

Andrena enigmatica Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 33. ♂.

Andrena erecta Viereck
B. C. to Calif. Pollen: Unknown, but visits flowers of Chamaebatia foliolosa,
Clarkia dudleyana, Erigeron glaiicus, Paeonia browni, Phacelia imbricata, Rubus,
Thysanocarpus curvipes.
Andrena (Andrena) erecta Viereck, 1924. Canad. Ent. 56: 28. ♀, ♂.

Andrena excellens Viereck
B. C.

Andrena (Andrena) excellens Viereck. 1924. Canad. Ent. 56: 76. ♀.

Andrena frigida Smith
Alaska to N. S., south to Va., 111., Wis., Minn., Wash, and Oreg. Pollen:
Unknown, but visits flowers of Conius, Pninus, Salix discolor.
Andrena frigida Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 115. ♀.
Andrena moesta Smith, 1879. Desc. New Species Hym. Brit. Mus. p. 54. ♀, ♂.
Andrena cockerelli Graenicher, 1903. Canad. Ent. 35: 163. ♀, ♂.
Andrena hirticeps Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 116. ♂. Preocc.
Andrena hirtignatha Linsley, 1951. In Muesebeck, Krombein and Townes, U. S. Dept.
Agr., Agr. Monog. 2: 1068. N. name.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 149. 9 (type). —Cockerell, 1906.
Psyche 13: 33 (type). — Atwood, 1934. Canad. Jour. Res. 10: 207, 209. 9, S (key). —Morice
and Cockerell, 1901. Canad. Ent. 33: 149 (type of hirticeps). —Morice and Cockerell, 1901.
Canad. Ent. 33: 153 (type of moesta). —Cockerell. 1906. Psyche 13: 26 (type of hirticeps)
—Cockerell, 1906. Psyche 13: 34 (type of ynoesta). —Viereck, 1907. Ent. News 18: 283,
285. 9, 6 (key, as cockerelli). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
105-106, figs. 15-16, table 3 (redescription, synonymy).

Andrena fulvicrista Viereck
B. C.

Andrena {Andrena) fulvicrista Viereck, 1924. Canad. Ent. 56: 30. ♀, ♂.

Andrena harveyi Viereck
B. C, Oreg.

Andrena harveyi Viereck, 1904. Canad. Ent. 36: 192, 194. ♀, ♂.

Andrena hemileuca Viereck
Wash.

Andrena hemileuca Viereck, 1904. Canad. Ent. 36: 192, 193. ♀, ♂.

Andrena hitei Cockerell
Colo.

Andrena hitei Cockerell, 1907. Ann. and Mag. Nat. Hist. (7) 19: 369. ♀.

Taxonomy: Nelson, 1930. Ent. News 41: 322, fig. 9,6. — Lanham, 1941. Ent. See. Amer.,
Ann. 34: 705. 9 (key).

Andrena impuncta Kirby
Maine, N. Y.

Andrena impuncta Kirby, 1837. Faun. Bor.-Amer., v. 4, p. 268. ♀.

Andrena irana Cockerell
Colo. Parasite: Nomada morrisoni Cress., A^. vallesina Ckll.?

Andrena irana Cockerell, 1931. Ann. and Mag. Nat. Hist. (10) 3: 392. ♀.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 265. 9 (key). -Lanham 1941. Ent.
Soc. Amer., Ann. 34: 705. 9 (key).

Biology: Hicks, 1934. Colo. Univ., Studies 21: 267 (nest, parasite).

Andrena laminibucca Viereck and Cockerell
Colo.

Andrena laminibucca Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 37. ♂.

Andrena lewisii Cockerell
Colo. Pollen: Unknown, but visits flowers oi Rubus.

Andrena lewisii Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 435. ♀.

Taxonomy: Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 57. 9 (var.). —Cockerell,
1927. Ent. Soc. Amer., Ann. 20: 397. ♀.

Biology: Clements and Long, 1923. Carnegie Inst. Wash., Pub. 336: 249 (ecology).

Andrena lillooetensis Viereck
B. C.

Andrena {Andrena) lillooetensis Viereck, 1924. Canad. Ent. 56: 237. ♂.

Andrena lummiorum Viereck
B. C.

Andrena {Andrena) lummiorum Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 564. ♀.




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Andrena macoupinensis Robertson
111., Mass., N. C. Pollen: Unknown, but visits flowers of Prumis,
Salix, Vicia.
Andrena macoupinensis Robertson, 1900. Acad. Sci. St. Louis, Trans. 10: 48. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 191. 9 (key). —Mitchell, 1962. N. C.
Agr. Expt. Sta. Tech. Bui. 141: 106, figs. 15-16, 21, table 3 (redescription).

Andrena magnifica Viereck
B. C.

Andre7ia (Andrena) magnifica Viereck, 1924. Canad. Ent. 56: 77. ♀.

Andrena mandibularis Robertson
N. S. and Ont., south to Ga., west to 111. and Minn. Pollen: Unknown,
but visits flowers of Amelanchier, Claytonia, Comus, Crataegus, Hepatica, Kahnia,
Mains, Pninus, Pyrus, Rhamnus, Rhus, Salix, Staphylea, Uvularia, Viburnum,
Zanthoxylum.
Andrena mandibularis Robertson, 1892. Amer. Nat. 26: 272. ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 191, 192. 9, 6 (key). — Atwood,
1934. Canad. Jour. Res. 10: 209. 6 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui.
141: 106-107, figs. 15, 22, table 3 (redescription).

Biology: Salt, 1927. Jour. Expt. Zool. 48: 252 (stylopization).

Andrena mesoleuca Cockerell
Calif.

Andrena mesoleuca Cockerell, 1924. Pan-Pacific Ent. 1: 60. ♀.

Andrena metea Cockerell
Wyo.

Andrena metea Cockerell, 1924. Ent. News 35: 350. ♂.

Andrena milwaukeensis Graenicher
N. S. to Ga., west to Minn. Pollen: Unknown, but visits flowers of
Aruncus, Comus, Ilex, Pyrus malus, Rubus, Viburnum.
Andrena milwaukeensis Graenicher, 1903. Canad. Ent. 35: 164. ♀, ♂.

Taxonomy: Viereck, 1907. Ent. News 18: 283, 285. 9, 6 (key). —Atwood, 1934. Canad. Jour.
Res. 10: 207, 209. 9, S (key). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
107-108, figs. 15-16, 20, table 3 (redescription).

Biology: Brittain, 1933. Canad. Dept. Agr., Bui. 162: 94, figs, (ecology).

Andrena moesticolor Viereck and Cockerell
Colo.

Andrena moesticolor Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 43: 31. ♀, ♂.

Andrena monogonoparia Viereck
Nev.

Andrena (Andrena) monogonoparia Viereck, 1917. Acad. Nat. Sci. Phila., Proc. 68: 566. ♂.

Andrena nitidarum Viereck
Alta.

Andrena (Andrena) nitidarum Viereck, 1924. Canad. Ent. 56: 78. ♀.

Andrena nodosa Viereck
Alta.

Andrena (Andrena) nodosa Viereck, 1924. Canad. Ent. 56: 238. ♀.

Andrena paenefulva Viereck and Cockerell
Colo.

Andrena paenefulva Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 51. ♀.

Andrena pallida Viereck
Alta.

Andrena (Andrena) pallida Viereck, 1924. Canad. Ent. 56: 78. ♀.

Andrena palpalis Timberlake
South. Calif. Pollen: Unknown, but visits flowers of Calandrinia

menziesii, Cryptantha barbigera, Larrea tridentata, Phacelia affinis, P. distans, Salix
gooddingii.

Andrena (Andrena) palpalis Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 400. ♀, ♂.

Andrena perarmata Cockerell
B. C. to Calif. Pollen: Oligolege of Salix including S. lasiolepis, but visits
other flowers for nectar including Baccharis.

Andrena perar^nata Cockerell, 1898. Entomologist 31: 88. ♂.

Taxonomy: Cockerell, 1911. Ann. and Mag. Nat. Hist. (8) 7: 232. 6. — Lanham, 1941. Ent. Soc.
Amer., Ann. 34: 705, 708. 9, 6 (key). —Lanham, 1965. Kans. Ent. Soc, Jour. 38: 199-200
(tax. status, geogr. range).

Andrena pyrrhacita
***authority mismatch
mosina Cockerell. Colo. Pollen: Oligolege of Salix.

Andrena pyrrhacita var. mosina Cockerell, 1908. Ann. and Mag. Nat. Hist. (8) 2: 330. ♀.

Taxonomy: Lanham, 1965. Kans. Ent. Soc, Jour. 38: 201 (tax. status).




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Andrena pyrrhacita
***authority mismatch
pyrrhacita Cockerell. B. C. to Colo., Utah, Calif. Pollen: Oligolege of Salix.
Aiidrena pyrrhacita Cockerell, 1917. Ann. and Mag. Nat. Hist. (7) 19: 536. ♀.
Andrena pyrrhacita var. coloradensia Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc.
48:39. ♀.

Ta.xonomy: Lanham, 1965. Kans. Ent. Soc, Jour. 38: 200-201 (tax. status).

Andrena revelstokensis Viereck
B. C.

Andrena (Andrena) revelstokensis Viereck, 1924. Canad. Ent. 56: 239. ♀.

Andrena rhodotricha Linsley
Calif.

Andre7ia rhodotricha Linsley, 1939. Pan-Pacific Ent. 15: 157. ♀, ♂.

Taxonomy: MacSwain, 1945. Pan-Pacific Ent. 21: 134 (nesting habits).

Andrena ribesina Cockerell
Colo. Pollen: Unknown, but visits flowers of Ribes.

Andrena ribesina Cockerell, 1906 Amer. Mus. Nat. Hist., Bui. 22: 433. ♀.
Andrena jacobaea Cockerell, 1915. Ann. and Mag. Nat. Hist. (8) 15: 267. ♀.

Taxonomy: Linsley, 1951. In Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr.
Monog. 2: 1079 (synonymy).

Andrena ribifloris Viereck and Cockerell
Colo. Pollen: Unknown, but visits flowers of Ribes.
Andrena ribifloris Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 32. c? ( 9
misdet.).

Andrena saccata Viereck
Wash, to Calif. Pollen: Unknown, but visits flowers of Eriogonum.
Atidrena saccata Viereck, 1904. Canad. Ent. 36: 192, 195. 9, s'.

Taxonomy: Cockerell, 1937. Amer. Mus. Novitates 899: 3 (tax. characters).

Andrena salicicola Viereck and Cockerell
Colo. Pollen: Unknown, but visits flowers of Salix.
Andrena salicicola Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 52. ♀.

Andrena semifulva Viereck
Calif.

Andrena (Andreiia) semifulva Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 575. ♀.

Andrena singularis Viereck
B. C.

Ajtdrena {Andrena) singularis Viereck, 1924. Canad. Ent. 56: 80. ♀.

Andrena thaspii Graenicher
N. S. to N. J. west to Ind., Wis. and Minn. Pollen: Unknown, but visits

flowers oi Angelica atropurpurea, Brassica, Chrysanthemum, Diervilla, Ledum, Pyrus
mains, Raphanus, Rosa, Rubus, Solidago, Stellaria, Taraxacum, Thaspium, trifoliatum
aureum, Trifolium, Vaccinium.
Andrena thaspii Graenicher, 1903. Canad. Ent. 35: 162. ♀, ♂.

Taxonomy: Cockerell, 1912. Ann. and Mag. Nat. Hist. (8) 9: 382. 9 (var.). — Atwood, 1934.
Canad. Jour. Res. 10: 208, 209. 9, 6 (key). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141: 108-109, table 3 (redescription).

Andrena thaspiiformis Viereck
Calif. Pollen: Oligolege of Salix including S. gooddingii, S. hindsiana, S.
laevigata, S. lasiolepis, S. nigra, but visits other flowers presumably for nectar
including Sainbuciis caenilea. Spiraea prunifolia.
Andrena {Andrena) perezana thaspiiformis Viereck, 1917. Amer. Ent. Soc, Trans. 43: 386.
9.

Andrena tincta Viereck
Colo.

Andrena (Andrena) tincta Viereck, 1917. Amer. Ent. Soc, Trans. 43: 387. ♀.

Andrena topazana Cockerell
Colo., Idaho, Wyo.

A7idre7ia topazana Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 434. ♀.

Taxonomy: Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc 48: 44, 45. 9. —Cockerell,
1927. Ent. Soc. Amer., Ann. 20: 397. ♀.

Andrena tridens Robertson
New England States, south to Tenn. and Ga., west to Mich, and 111. Pollen:
Unknown, but visits flowers of Azalea, Dentaria, Hepatica, Hydrangea, Rhododendron,
Rhus.
Andrena tridens Robertson, 1902. Amer. Ent. Soc, Trans. 28: 192. ♂.
Andrena comelli Viereck, 1907. Ent. News 18: 282. ♀.




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Taxonomy: Mitchell, 1958. In Krombein, U. S. Dept. Agr., Agr. Monog. 2, First Suppl. pp.
213, 216 (synonymy). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 109-110, figs.
15-16, 20-21, table 3 (redescription).
Washington! Cockerell. B. C. to Wash, and Colo.

Andrena washingtoni Cockerell, 1901. Psyche 9: 284. ♀, ♂.

Taxonomy: Viereck, 1904. Canad. Ent. 36: 192, 195. 9, S (key).

Biology: Clements and Long, 1923. Carnegie Inst. Wash., Pub. 336: 249 (ecology).

Genus ANDRENA Subgenus APORANDRENA Lanham

Andrena subg. Aporandrena Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 201.

Type-species: A^idrena coactipostica Viereck. Monotypic and orig. desig.

Revision: LaBerge, 1969. Amer. Ent. Soc, Trans. 95: 34-40, figs. 29-33 (N. Amer. spp.).

Andrena coactipostica Viereck
Calif. Pollen: Polylectic, apparently prefers flowers of the families

Papaveraceae and Boraginaceae in that order, but visits a variety of flowers including
Calandrinia menziesii, Capsella bursa- jMstoris, Cryptantha interruedia, Encelia
farinosa, Eschscholzia californica, Lasthenia gracilis, Phacelia distayis, Plagiobothrys
californicus, P. nothofulvus, Platystemon califomicus.
Andrena (Andrena) coactipostica Viereck, 1917. Amer. Ent. See, Trans. 43: 372. ♀.

Genus ANDRENA Subgenus BELANDRENA Ribbic

Andrena subg. Belandrena Ribble, 1968. Kans. Ent. Soc, Jour. 41: 221.
Type-species: Andrena nemophilae Ribble. Orig. desig.

The species of this subgenus exhibit a strong preference for the flowers of the families
Hydrophyllaceae and Malvaceae.

Revision: Ribble, 1968. Kans. Ent. Soc, Jour. 41: 220-236, 15 figs., 1 map, 1 table (N. Amer.
spp.).

Biology: Cruden, 1972. Madrono 21: 505-515, 1 fig., 2 tables (floral relationships). — Cruden,
1972. Evolution 26: 373-389, 8figs., 9 tables (floral relationships).

Andrena nemophilae Ribble
Calif. Parasite: Caloglyphus sp., Meloe sp. Pollen: Oligolectic on flowers of
Nemopliila including A'^. maculata, N. menziesii atonieria, N. m. integrifolia, N. m.
menziesii, N. pulchella, but visits other flowers for nectar including Arctostaphylos,
Ceanothus, Gilia, Montia, Pholistorna atiritum.
Andrena (Belandrena) nemophilae Ribble, 1968. Kans. Ent. Soc, Jour. 41: 223, figs. 1-5,
map 1, table 1. ♀, ♂.

Andrena sagittagalea Ribble
Tex. Parasite: Vidia sp. Pollen: Apparently oligolectic on flowers of
Phacelia, but also visits flowers of Salix.
Andrena (Belandrena) sagittagalea Ribble, 1968. Kans. Ent. Soc, Jour. 41: 230, figs. 6-10,
map 1, table 1. ♀, ♂.

Andrena sphaeralceae Linsley
Ariz, west to south. Calif, and south. Nev. Pollen: Apparently oligolectic
on flowers of Sphaeralcea including S. ambigua, S. laxa, S. orcutti, S. subhastata, but
visits other flowers presumably for nectar including Koeberlinia, Phacelia distans.
Andrena (Opandrena) sphaeralceae Linsley, 1939. Pan-Pacific Ent. 15: 160. ♀, ♂.

Genus ANDRENA Subgenus CALLANDRENA Cockerell

Callandrena Cockerell, 1898. Amer. Ent. Soc, Trans. 25: 186.

Type-species: Panurgus manifestus Fox. Monotypic.
Pterandrena Robertson, 1902. Amer. Ent. Soc, Trans. 28: 193.

Type-species: Andrena pulchella Robertson. Orig. desig.

Most species of this subgenus obtain pollen from the flowers of the Compositae.

Revision: LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 1-316, 340 figs., 4 tables, 14 maps
(N. Amer. spp.).




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Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 198 (tax. characters). — LaBerge, 1964.

Nebr. Univ. State Mus., Bui. 4: 294 (tax. characters).

Andrena accepta Viereck
N. J. to S. C, west to Alta., Oreg., east. Calif. (Big Pine), and Ariz.; Mexico

(Chihuahua). Parasite: Myopaf or Zodion obliquefasciatiim (Macq.). Pollen: Oligolege of
Compositae, especially Heliantlius including H. angustifolius, H. ayi7iuus, H.
divarkatus, H. giganteus, H. grosseserratus, H. maxiyiiilUani, H. petiolaris, H.
salicifolius, H. scabberimus, H. strumosus, H. tuberosus, but visits other flowers
including Aster ericoides villosiis, Bidens aristosa. Cassia chamaecrista,
Chrysothamnus, Coreopsis tripferis, Gaillardia pulchella, Grindelia squarrosa,
Gutierrezia sarotlirae, Haplopappits, Machaeranthera, Medicago sativa, Prionopsis
ciliata, Silphium integrifolium, S. perfoliatum, S. speciosum, Solidago nemoralis,
Verbesina occidentalis, V. oreophila.

Andrena pulchella Robertson, 1891. Amer. Ent. Soc, Trans. 18: 57. ♀. Preocc.

Andrena accepta Viereck, 1916. Biol. Soc. Wash., Proc. 29: 127. N. name.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 193, 194. 9, 6 (key). —Mitchell,
1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 137-138, figs. 24-26, 28, table 4 (redescription).
— Rozen, 1973. N. Y. Ent. Soc, Jour. 81: 58-61, figs. 3-10 (mature larva).

Biology: Rozen, 1973. N. Y. Ent. Soc, Jour. 81: 54-58, 2 figs, (nest architecture, life history,
parasite).

Andrena afimbriata LaBerge
Tex. Pollen: Unknown, but visits flowers of Berlandiera, Polygala alba,

Pyrrhopappus carolinianus.
Andrena (Callandrena) afimbriata LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 165,

figs. 169-172. 9, 6:

Andrena aliciae Robertson
N. Y. to Ga., west to Minn. Nebr. and Kans. Pollen: Oligolege of

Compositae, especially Helianthus including H. angustifolius, H. divaricatus, H.

giganteus, H. microcephalus, H. strumosus, H. tuberosus, but visits other flowers

including Aster azurella, Bidens aristosa, B. chrysanthemoides, B. laevis, B.

trichosperma, Cucurbita, Heliopsis helianthoides, Rudbeckia fulgida, R. laciniata, R.

lanceolata, R. triloba, Silphiu7n perfoliatum, Solidago occidentale.
Andrena aliciae Robertson, 1891. Amer. Ent. Soc, Trans. 18: 57. ♀, ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 193. 9, 6 (key). —Mitchell, 1960. N.
C. Agr. Expt. Sta. Tech. Bui. 141: 138-139, figs. 24-25, table 4 (redescription).

Andrena aliciarum Cockerell
N. Mex., Ariz.; Mexico (Sonora). Pollen: Unknown, but visits flowers of
Eriogonnm, Pedis papposa.
Andrena aliciarum Cockerell, 1897. Entomologist 30: 138. ♀.

Taxonomy: Cockerell, 1899. Ent. News 10: 254. ♂.

Andrena aridis LaBerge
Colo., Nebr., N. Mex., Tex. Pollen: Unknown, but visits flowers of
Chrysothamnus nauseosus, Gutierrezia microcephala, G. sarothrae.
Andrena (Callandrena) aridis LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 249, figs.
277-281. ♀, ♂.

Andrena asteris Robertson
Que. to Ga., west to Man., N. Dak., Nebr., and Mo. Pollen: Oligolege of

Compositae, especially flowers of Aster including A. ericoides, A. paniculatus, but visits
other flowers including Echinacea, Eupatorium altissimuni. Polygonum
pennsylvanicum, Solidago nemoralis, S. puberula, S. rigida, S. ulmifolia. Predator:
Philanthus solivagus Say.
Andrena asteris Robertson, 1891. Amer. Ent. Soc, Trans. 18: 56. 9, cJ. '

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 194. 9, 6 (key). —Mitchell, 1960. N.
C. Agr. Expt. Sta. Tech. Bui. 141: 139-140, figs. 24-26, 28, table 4 (redescription).

Biology: Salt, 1927. Expt. Zool., Jour. 48: 251 (stylopization). — Britton, 1937. Conn. (State)
Agr. Expt. Sta., Bui. 408: 261 (economic).

Andrena asteroides Mitchell
N. J. and Pa., south to S. C. and Miss. Pollen: Apparently an oligolege of
Aster including A. ericoides, A. duynosus, A. paniculatus, but visits other flowers
including Chrysopsis, Haplopappus.
Ayidrena (Pterandrena) asteroides Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 140,
figs. 24,25. ♀, ♂.




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Andrena auripes LaBerge
Ariz. (Graham Mts.); Mexico (Distrito Federal and Mexico).

Ajidrena (Callandrena) atiripes LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 204, figs.
219-223. ♀, ♂.

Andrena balsamorhizae LaBerge
Nev. (Lake Mead, Overton Beach and Rodgers Springs). Pollen:
Unknown, but visits flowers oi Balsamorliiza.
Andrena (Callandrena) balsamorhizae LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 252,
figs. 282-286. ♀, ♂.

Andrena barberi Cockerell
N. Mex. (Eagle Creek and Ruidoso Forks, White Mts.); Mexico (Durango,
Guadalajara, Guanajuato, Hidalgo, Mexico, Puebla, Tlaxcala and Vera Cruz). Pollen:
Apparently an oligolege of Compositae, including Bidens triplinervia var. macrantha,
Heterotkeca, Rudbeckia laciniata, Solidago trinervata, but visits other flowers including
Argemone, Primus.
Andrena barberi Cockerell, 1898. Ann. and Mag. Nat. Hist. (7) 2: 448. ♀, ♂.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 14. 9 (key).

Andrena beameri LaBerge
Ark., Kans. Pollen: Unknown, but visits flowers of Coreopsis grandiflora.
Echinacea angustifolia, Helenium nudiflora.
Andrena (Callandrena) beameri LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 135. ♀.

Andrena berkeleyi Viereck and Cockerell
Colo., Okla., Tex. Pollen: Unknown, but visits flowers of
Actinea, Engelmannia pinnatifida.
Andrena berkeleyi Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 19. ♂.

Taxonomy: Cockerell, 1933. Pan-Pacific Ent. 9: 153. ♂.

Andrena biscutellata Viereck
Colo., Kans., N. Mex., Tex. Pollen: Unknown, but visits flowers of
Gaillardia, Monarda.
Andrena (Ptilandrena) biscutellata Viereck, 1917. Amer. Ent. Soc, Trans. 43: 393. ♀.

Andrena braccata Viereck
New England States to Va. Pollen: Apparently an oligolege of Solidago
including S. altissiyna, S. nevioralis, but also visits flowers oi Aster.
Andrena braccata Viereck, 1907. Ent. News 18: 284, 286. ♀, ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 141-142, figs. 24-25, table 4
(redescription).

Biology: Clements and Long, 1923. Carnegie Inst. Wash., Pub. 336: 249 (ecology).

Andrena bullata LaBerge
Tex. (Brazos and Lee Counties). Pollen: Unknown, but visits flowers of
Gutierrezia texana, Heterotheca subaxillaris.
Andrena (Callandrena) bullata LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 154, figs.
154-158. ♀, ♂.

Andrena calvata LaBerge
Ariz. (22 miles west of Springerville). Pollen: Unknown, but visits flowers of
Viguiera annua.
Andrena (Callandrena) calvata LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 74, figs.
60-64. ♂.

Andrena crawfordi Viereck
Kans., Okla., Tex. Pollen: Oligolege of Compositae, tribe Cichoriae, visits
flowers of Berlandiera, Coreopsis, Engelmannia bipinnatifida, Lindheimeria texana,
Polygala alba, Pyii-lwpappus caroliniamis, P. geiseri, P. gra}>diflora, P. multicaulis,
Serinea oppositifolia, Sitilias.
Andrena crawfordi Viereck, 1909. Ent. Soc. Wash., Proc. 11: 143. ♀, ♂.

Andrena duplicata Mitchell
N. J., N. C, Ohio, 111., Mo. Pollen: Apparently an oligolege of Compositae,
visits flowers of Bidens coronata, Helianthus divaricatus, H. grosseserratus, Solidago.
A7idrena (Pterandrena) duplicata Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 144,
figs. 24-26, 28. ♀, ♂.

Andrena fulvipennis Smith
N. J. to Fla. on coastal plain, also east. Tex. Pollen: Apparently an oligolege
of Compositae, visits flowers of Aster, Chrysopsis mariana, Coreopsis, Haplopappus,
Heterotheca latifolia, H. subaxillaris, Gutierrezia texaiia.
Andrena fulvipennis Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 117. ♀.

Taxonomy: Viereck, 1902. Ent. News 13: 237. 9. —Mitchell, 1960. N. C. Agr. E.xpt. Sta. Tech.
Bui. 141: 145-146, figs. 24-25, table 4 (redescription).




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Andrena gardineri Cockerell
Md. to Ga., west to S. Dak., Colo, and Tex. Pollen: Apparently an oligolege
of early summer flowering Compositae, especially Senecio including S. aureus, S.
pauperculus var. balsaniitae, S. sviallii, S. multilobatus, S. triangularis, but visits other
flowers including Aster, Astragalus striatus, Baileya ynultiradiata, Crepis runcinata,
Geraea, Hynienoxys glabra, Lesqnerella ovalifolia, Penstemon, Rubus.

Andrena (Pterandrena) gardineri Cockerell, 1906. Ann. and Mag. Nat. Hist. (7) 17: 307. ♀.

Andrena ashmeadi Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 45. ♀.

Andrena (Opandrena) lamellicauda Cockerell, 1925. Ann. and Mag. Nat. Hist. (9) 16: 629.
6.

Andrena campbelli Cockerell, 1933. Pan-Pacific Ent. 9: 153. ♂.

Taxonomy: Lanham, 1941. Ent. Soc. Amer., Ann. 34: 711 (tax. characters). — Lanham, 1951.
In Linsley In Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1066
(synonymy). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 146-147, figs. 24-26,
table 4 (redescription).

Biology: Clements and Long, 1923. Carnegie Inst. Wash., Pub. 366: 249 (ecology).
haynesi Viereck and Cockerell. Alta. and N. Dak., south to Tex. Pollen: Oligolege of

Helianthus, mostly H. petiolaris, but also visits other flowers presumably for nectar
including Campanula, Echinacea pallida, Soiidago rigida.
Andrena haynesi Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 26. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 14. 9 (key).

Andrena helianthi Robertson
Que. south to Va., west to Alta., Oreg., Calif, and Ariz. Pollen: Oligolege
of Compositae, apparently preferring the flowers of Helianthus including H. annuus, H.
coloradinus, H. coronatus, H. divaricatus, H. giganteus, H. grosseserratus, H.
inaximilliani, H. petiolaris, H. rigidus, H. subrhomboideus, H. tuberosus, but also visits
other flowers including Aster novaeangliae, Bidens aristosa, Chrysothamnus nauseosus,
Cirsiuvi undulatum, Cleome semdata, Gilia, Gutierrezia sarothrae, Medicago sativa,
Rudbeckia laciniata, Silphium perfoliatum, Soiidago canadensis, S. rigida, Verbesina
encelioides, V. oreophila.

Andre7ia helianthi Robertson, 1891. Amer. Ent. Soc, Trans. 18: 55. ♀, ♂.

Andrena nitidior Cockerell, 1900. Ann. and Mag. Nat. Hist. (7) 5: 406. ♀.

Andrena graenicheri Cockerell, 1902. Ann. and Mag. Nat. Hist. (7) 9: 104. ♀.

Andrena lincolnella Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 46. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 194. 9,6. —Mitchell, 1960. N. C.
Agr. Expt. Sta. Tech. Bui. 141: 147-148, figs. 24-25 (redescription). — LaBerge, 1967. Nebr.
Univ. State Mus., Bui. 7: 94-101, figs. 8, 94-98, table 3 (redescription, synonymy).

Biology: Hicks, 1926. Colo. Univ., Studies 15: 222 (nesting habits).

Andrena helianthiformis Viereck and Cockerell
Mont, and N. Dak., south to Kans. Pollen: Apparently
an oligolege of Echinacea including E. angustifolia, E. pallida, but also visits other
flowers including Amorpha canescens, Gaillardia, Heliopsis helianthioides, Melilotus
officinalis.
Andrena helianthiformis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 26. ♀.

Andrena humeralis LaBerge
Tex. Pollen: Unknown, but visits flowers oi Aster, Baccharis.

Andrena (Callandrena) humeralis LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 76. ♀.
6.
ignota LaBerge. S. C. (Florence).

Andrena {Callandrena) ignota LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 83. ♀.

Andrena irrasus LaBerge
111., Nebr., Kans., Wyo., N. Mex. Pollen: Unknown, but visits flowers of
Amphiachyris dracunculoides, Aster, Chrysothamnus.
Andrena (Callandrena) irrasus LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 131, figs.
129-133. ♀, ♂.

Andrena isocomae Timberlake
Calif. Pollen: Oligolege of Compositae, visits flowers oi Chrysothamnus,
Eriogonum, Gaillardia, Gutierrezia californica, G. lucida, G. sarothrae, Haplopappus
bemardinus, H. vemonioides, Heterotheca grandiflora, Senecio douglasii, Soiidago.
Andrena (Pterandrena) isocomae Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 378. ♀, ♂.




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Andrena krigiana Robertson
Conn, to Fla., west to Minn., Okla. and Tex. Pollen: Oligolege of Krigia

including K. biflora, K. Virginia, but visits other flowers presumably for nectar including
Conuis, Hieraceum venosum, Ranunculus acris.

Andrena krigiana Robertson, 1901. Canad. Ent. 33: 229. ♀, ♂.

Andrena (Ptilandrena) parakrigiana Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
157. ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 194. 9,6. — Viereck, 1917 (1916).
Conn. State Geol. and Nat. Hist. Survey, Bui. 22: 777. 9,6. — LaBerge, 1967. Nebr. Univ.
State Mus., Bui. 7: 157-162, figs. 12, 159-163 (redescription, synonymy).

Andrena levipes LaBerge
Oreg., Calif. Pollen: Collects pollen from the flowers of Lasthe7iia

chrysostoma and Linanthus aureus, but visits other flowers including Arenaria,
Coreopsis bigelovii, Cryptantha, Gilia, Phacelia distans.
Andrena (Callandrena) levipes LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 289, figs.
336-340. ♀, ♂.

Andrena melliventris Cresson
Kans. to Tex., Ariz.? Pollen: Unknown, but visits flowers of Aster

tanacetifolium, Coreopsis cardaminefolia, Gaillardia pulchella, Monarda punctata.
Andrena melliventris Cresson, 1872. Amer. Ent. Soc, Trans. 4: 257. ♀.

Andrena monticola LaBerge
Ariz. (Chiricahua Mts.); Mexico (Chihuahua, Durango, Guanajuato and
Mexico). Pollen: Unknown, but visits flowers of Aster, Cirsium, Heliopsis, Solidago.
Andrena (.Callandrena) monticola LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 125, figs.
119-123. ♀, ♂.

Andrena neomexicana LaBerge
Ariz., N. Mex. Pollen: Unknown, but visits flowers of Baileya
pleniradiata, Erigeron.
Andrena {Callandrena) neomexicana LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 246,
figs. 272-276. ♀, ♂.

Andrena ofella LaBerge
Tex., N. Mex., Ariz. Pollen: Unknown, but visits flowers of Aplopappus
gracilis, Chrysothamnus, Gntierrezia microcephala, G. sarothrae.
Andrena (Callandrena) ofella LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 267, figs.
307-311. ♀, ♂.

Andrena pecosana
***authority mismatch
Cockerell. N. Mex., Colo., Utah, Ariz.; Mexico (Chihuahua, Hidalgo, Jalisco and

Zacatecas). Pollen: Unknown, but presumably an oligolege of Compositae, visits flowers
of Grindelia, Gutierrezia sarothrae, Helianthus, Heliopsis Viguiera annua.
Andrena pecosana Cockerell, 1913. Ann. and Mag. Nat. Hist. (8) 12: 104. ♀.
Andrena townsendi Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 49. ♀.
Andrena colletoides Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 27. ♂.

Taxonomy: LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 112-116, figs. 9, 84-88
(redescription, synonymy, floral relationships).

Andrena pectidis (Cockerell)
N. Mex., Tex. Pollen: Unknown, but visits flowers of Pedis papposa.
Panurgus pectidis Cockerell, 1897. Amer. Ent. Soc, Trans. 24: 148. ♀.

Taxonomy: Cockerell and Porter, 1899. Ann. and Mag. Nat. Hist. (7) 4: 418 (tax. characters).

Andrena perpunctata LaBerge
Ariz. (Chiricahua Mts.); Mexico (Guerrero, Mexico and Morelos). Pollen:
Unknown, but visits flowers of Heterotheca subaxillaris.
Andrena (Callandrena) perpunctata LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 222,
figs. 243-247. ♀, ♂.

Andrena placata Mitchell
Que. and Ont., south to N. C. west to Minn, and 111. Pollen: Oligolege of
Solidago, but visits other flowers including Aster lateriflorum, A. macrophyllum,
Eupatorium serotinum, Fagopyrum esculentum. This species evidently hybridizes
occasionally throughout its range with the closely related A. simplex Smith.
Andrena (Pterandrena) placata Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 148,
figs. 24, 25. ♂.

Andrena reflexa Cresson
Tex.; Mexico (Chihuahua, Jalisco and Michoacan). Pollen: Unknown, but visits
flowers of C rota laria long irost rata.
Andrena reflexa Cresson, 1872. Amer. Ent. Soc, Trans. 4: 256. ♂.
Andrena permitis Cresson, 1872. Amer. Ent. Soc, Trans. 4: 257. ♀.




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Taxonomy: LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 70-74, figs. 55-59 (redescription,
synonymy).

Andrena rudbeckiae Robertson
N. C. to Mich., Wis., Minn, and Colo., south to Tex. in the Great Plains
States. Pollen: Oligolege of Compositae, especially Ratibida including R. Columbians, R.
columnifera, R. pinnata and Rudbeckia including R. hirta, R. lanceolata, R. triloba, but
visits other flowers including Centaurea aynericana, Chrysopsis, Coreopsis, Echinacea
angustifolia, Gaillardia pulchella, Lepachys, Verbesiyia helianthioides.
Andrena rudbeckiae Robertson, 1891. Amer. Ent. Soc, Trans. 18: 56. ♀, ♂.

Taxonomy: Cockerell, 1899. Ent. News 10: 255. 6 (key). —Robertson, 1902. Amer. Ent. Soc,

Trans. 28: 194. 9, S (key). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 149, 151,

figs. 25-28, table 4 (redescription). —LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 90-94,

figs. 7 (redescription).

Andrena senticulosa LaBerge
Tex. Pollen: Unknown, but visits flowers of Berlandiera, Engelmannia
bipinnatifida, Pyrrhopappus, Serinia oppositifolia.

Andrena (Callandrena) senticulosa LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 177,
189-193. ♀, ♂.

Andrena simplex Smith
N. H. to N. C, west to Minn., Nebr. and Kans. Pollen: Oligolege of Compositae,
especially Aster and Solidago, visits flowers of Am phiachyris dracunculoides, Aster
anomalus, A. azureus, A. commutatus, A. ericoides, A. e. vilosus, A. laterifolius, A.
multiflorus, A. novaeangeliae, A. paniculatus, A. praelatus, Boltonia asteroides,
Eupatorium perfoliatum, Euthamia graminifolia, Gnaphalium polycephahim,
Grindelia, Helianthus tuberosus, Polygonuyn hydropiperoides, P. scandens, Solidago
altissima, S. canadensis, S. graminifolia, S. lanceolata, S. nemoralis, S. rigida, S.
serotina. Predator: Philanthus solivagus Say. This species evidently hybridizes
occasionally throughout its range with the closely related A. placata Mitchell.

A7idre7ia simplex Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 114. ♀.

Andrena solidagijiis Robertson, 1893. Amer. Ent. Soc, Trans. 18: 55. ♀, ♂.

Andrena radmitricha Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc 48: 51. ♀.

Andrena {Piilandrena) determinata Viereck, 1917. Amer. Ent. Soc, Trans. 43: 394. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 124 (type). —Cockerell, 1906. Psyche
13: 8 (type). -LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 137-143, figs. 10, 134-138,
table 4 (redescription, synonymy, floral relationships).

Andrena simulata Smith
Ariz., N. Mex., Colo.; Mexico. Pollen: Unknown, but visits flowers of Aster,
Viguiera annua.
Andrena simulata Smith, 1879. Descr. n. spp. Hymen. Brit. Mus., p. 52. ♀.
Andrena aureocincta Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 88. ♀, ♂.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 48. 6 (as aureocincta).
—LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 189-193, figs. 204-208 (redescription,
synonymy).

Andrena sitiliae Viereck
Okla., Tex. Pollen: Possibly an oligolege of Pyrrhopappus including P.
caroli7iianus, but also visits Coreopsis tinctoria.
Andrena sitiliae Viereck, 1909. Ent. Soc. Wash., Proc. 11: 144. ♀.

Andrena sonorensis LaBerge
Ariz. (Show Low), N. Mex. (Carizozo). Pollen: Unknown, but visits
flowers oi Gutierrezia microcephala.
Andrena (Callandrena) sonorensis LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 110,
figs. 99-103. ♀, ♂.

Andrena tegularis LaBerge
Ariz. (Chiricahua and Santa Rita Mts.). Pollen: Unknown, but visits flowers
of Heliopsis.
Andrena (Callandrena) tegularis LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 195, figs.
209-213. ♀, ♂.

Andrena texana Cresson
Tex. (Roosevelt and Victoria); Mexico (San Luis Potosi). Pollen: Unknown, but
visits flowers of Aynphiachyris, Aster.
Andrena texana Cresson, 1872. Amer. Ent. Soc, Trans. 4: 258. ♂.

Andrena tonkaworum Viereck
Nebr., Kans., Colo., Tex., N. Mex. Pollen: Possibly an oligolege of
Engelmannia.
Andrena {Ptilandrena) tonkaworum Viereck, 1917. Amer. Ent. Soc, Trans. 43: 396. ♀.




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Andrena trimaculata LaBerge
Ariz. (Canelo); Mexico (Durango).

Ayidrena (Callandrena) trimaculata LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 68,
figs. 50-54. ♀, ♂.

Andrena utahensis LaBerge
Utah, Colo., Ariz. Pollen: Unknown but visits flowers of Geraea.

Andrena (Callandreiia) utahensis LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 258, figs.
292-296. ♀, ♂.

Andrena verbesinae Viereck and Cockerell
Tex. (Cotulla and San Antonio). Pollen: Unknown, but visits
flowers of Verbesina encelioides.
Andrena verbesinae Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 1. ♀, ♂.

Andrena verecunda Cresson
Tex. Pollen: Unknown, but visits flowers of Engelmannia bipinnatifida,
Gaillardia, Pyrrhopappus carolinianus, P. grandiflora.
Andrena verecunda Cresson, 1872. Amer. Ent. Soc, Trans. 4: 257. ♀.

Andrena vulpicolor Cockerell
Wyo., south to N. Mex., west to Oreg. and Calif. Pollen: Apparently an

oligolege of Chrysothammis including C. iiauseosus, C. n. albicaulis, C. n. coyisimilis, C.
parryi, C. viscidiflonis pumilis, C. v. stenophyllus, but also visits flowers of Erigeron
neomexicana.
Andrena vulpicolor Cockerell, 1897. Ann. and Mag. Nat. Hist. (6) 20: 512. ♀.
Andrena nubilipennis Viereck, 1904. Canad. Ent. 36: 193. ♀.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 49. 9 (key). —Cockerell, 1931.
Amer. Mus. Novitates 458: 13. 9 (key). —LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7:
106-110, figs. 6, 79-83 (redescription, synonymy).

Genus ANDRENA Subgenus CHAULANDRENA LaBerge

Andrena subg. Ckaulandrena LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 314.
Type-species: A7idre7ia porterae Cockerell. Orig. desig.

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 284, 288, 314-315 (tax. characters).

Andrena porterae Cockerell
Sierra Nevada, Cascade and Rocky Mts. to N. Mex. Parasite: Stylops sp.
Pollen: Apparently oligolectic, presumably gathers pollen from the flowers o{ Ribes.
Andrena porterae Cockerell, 1900. Ann. and Mag. Nat. Hist. (7) 5: 401. ♀.
Ayidrejia leptanthi Viereck and Cockerell, 1904. Ann. and Mag. Nat. Hist. (7) 14: 27. ♂.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 264, 267. ♀, ♂.

Biology: Linsley, 1937. Pan-Pacific Ent. 13: 157 (stylopization).

Genus ANDRENA Subgenus CNEMIDANDRENA Hedicke

Andrena subg. Cnemidandrena Hedicke, 1933. Berlin Zool. Mus., Mitt. 19: 212.
Type-species: Melitta nigriceps Kirby. Orig. desig.

Revision: Donovan, 1977. Calif. Univ. Pubs. Ent. 81: 1-107, 130 figs., 17 maps (N. Amer. spp.;
received too late for inclusion in this catalog).

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 211 (tax. characters). —LaBerge, 1964.

Nebr. Univ. State Mus., Bui. 4: 303 (tax. characters).

Andrena antonitonis Viereck and Cockerell
Colo.

Andrena antonitonis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 56. ♀.

Andrena apacheorum Cockerell
Rocky Mts. to N. Mex., Ariz.

Andrena apacheorum Cockerell, 1897. Entomologist 30: 306. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 14, 15. 9, cJ (key).

Biology: Clements and Long, 1923. Carnegie Inst. Wash., Pub. 336: 249 (ecology). —Hicks,
1926. Univ. Colo., Studies 15: 221 (habits).

Andrena autumnalis Viereck and Cockerell
Nebr.

Andrena autumnalis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 44. ♀.

Andrena beulahensis Viereck
N. Mex.

Andrena beulahensis Viereck, 1903. Amer. Ent. Soc, Trans. 29: 53. ♀.




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Andrena canadensis
***authority mismatch
canadensis Dalla Torre. N. S. to N. Y. and N. J., west to Minn. Pollen: Unknown,

but visits flowers of Aster, Epilobium, Solidago.
Ayidrena simulafa Provancher, 1888. Addit. Corr. Faune Ent. Canada, Hym., p. 313. ♀, ♂.

Preocc.
Andrena canadensis Dalla Torre, 1896. Cat. Hym., v. 10, p. 107. N. name.
Andrena (Pterandrena) persnnilis Graenicher, 1904. Ent. News 15: 66. ♀.

Taxonomy: Viereck, 1907. Ent. News 18: 282, 286. 9, 6 (key). — Atwood, 1934. Canad. Jour.
Res. 10: 207, 209, fig. 9, S (key). -Timberlake, 1951. In Muesebeck, Krombein and
Townes, U. S. Dept. Agr., Agr. Monog. 2: 1059 (synonymy). —Mitchell, 1960. N. C. Agr.
Expt. Sta. Tech. Bui. 141: 172, figs. 30-31, 33, table 4 (redescription).

Biology: Salt, 1927. Jour. Expt. Zool. 48: 245 (stylopization).

Andrena canadensis
***authority mismatch
oslarella Viereck and Cockerell. Colo., N. Mex.

Andrena canadensis oslarella Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 28.

6.

Biology: Clements and Long, 1923. Carnegie Inst. Wash. Pub. 336: 249 (ecology).

Andrena chromotricha Cockerell
N. Mex., Wis.

A7idrena chromotricha Cockerell, 1899. Entomologist 32: 128. ♀.
Andrena clypeonitens Cockerell, 1902. Canad. Ent. 34: 47. ♀.
Ayidrena truncata Viereck, 1903. Amer. Ent. Soc, Trans. 29: 53. ♀.

Taxonomy: Graenicher, 1910. Canad. Ent. 42: 160. 6. —Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 172-173, table 4 (redescription of female).

Andrena citrinihirta Viereck
Calif. Pollen: Unknown, but visits flowers of Chrysothamnus,
Heterotheca grandiflora, Solidago californica.
Andrena (.Andrena) citrinihirta Viereck, 1917. Amer. Ent. Soc, Trans. 43: 371. ♀.

Andrena colletina Cockerell
B. C. to Calif., east to Colo. Pollen: Unknown, but visits flowers of

Achillea millefolium, Chrysothamnus nauseosus consimilis, C. viscidiflonis typicus,
Haplopappus hloomeri var. angustatns.
Andrena colletina Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 454. ♂.

Taxonomy: Cockerell, 1909. Ann. and Mag. Nat. Hist. (8) 5: 260. 9. —Cockerell, 1931. Amer.
Mus. Novitates 458: 14, 15. 9, d (key).
Columbiana Viereck. B. C. to Calif., N. Mex. Pollen: Unknown, but visits flowers of Achillea
lanulosa, Erigeron coulteri, E. pygmaeus, Microseris nutans, Senecio aureus var.
subnudus, S. canus, S. integirrimus, S. scorzonella.
Andrena {Andrena) columbiana Viereck, 1917. Amer. Ent. Soc, Trans. 43: 374. ♀.

Andrena costillensis Viereck and Cockerell
Colo., N. Mex.

Andrena costillensis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 50. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 14. ♀, ♂.

Andrena davidsoni Viereck and Cockerell
Calif.

Andrena davidsoni Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 52. ♀.

Taxonomy: Cockerell, 1930. Pan-Pacific Ent. 7: 8. 9 (key).

Andrena hirticincta Provancher
Alta. to N. S., south to Ga. Pollen: Unknown, but visits flowers of
Aster, E pilobium, Solidago.
Andrena fimbriata Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 116. ♀, ♂. Preocc.
Andrena hirticincta Provancher, 1888. Addit. Corr. Faune Ent. Canada, Hym., p. 308. ♀.

i.
Anthrena americana Dalla Torre, 1896. Cat. Hym., v. 10, p. 102. N. name.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 149 (type of fimbriata). —Cockerell,
1906. Psyche 13: 9 (types o{ fimbriata). —Atwood, 1934. Canad. Jour. Res. 10 (2): 207, 209,
fig. 9, 6 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 173-174, figs. 30-31,
33-34, table 4 (redescription).

Biology: Salt, 1927. Jour. Expt. Zool. 48: 233, figs, (stylopization).

Andrena indecisa Cockerell
B. C, Alta., Mont.

Andrena costillensis indecisa Cockerell, 1937. Canad. Ent. 69: 34. ♀.




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Andrena mentzeliae Cockerell
Colo., N. Mex. Pollen: Unknown, but visits flowers of Mentzelia.
Andrena ynentzeliae Cockerell, 1897. Entomologist 30: 307. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 14, 15. 9, 6. — Lanham, 1941. Ent.
Soc. Amer., Ann. 34: 706, 708. 9, 6 (key).

Andrena nubecula Smith
East. Canada., south to Ga., west to B. C. Pollen: Unknown, but visits flowers
of Aster, Euthamia, Solidago. Predator: Philayithus solivagus Say.
Andrena nubecula Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 117. ♀.
Andrena nubecula tristicomis Cockerell, 1931. Canad. Ent. 63: 22. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 150 (type). —Robertson, 1902. Amer.
Ent. Soc, Trans. 28: 191, 192. 9, S (key). —Cockerell, 1906. Psyche 13: 9 (type).
—Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 174-175, figs. 31, 33-34, table 4
(redescription).

Andrena pacta Viereck
N. Mex.

Andreyia pacta Viereck, 1903. Amer. Ent. Soc, Trans. 43: 54. ♀, ♂.

Andrena peckhami Cockerell
Minn, to Vt., south to N. C. Pollen: Unknown, but visits flowers of
Baptisia, Solidago.
Andreyia peckhami Cockerell, 1902. Ann. and Mag. Nat. Hist. (7) 9: 105. ♀.

Taxonomy: Graenicher, 1910. Canad. Ent. 42: 159. 9,6. — LaBerge and Kibble, 1972. Amer.
Ent. Soc, Trans. 98: 272 (subgeneric position).

Andrena ramaleyi Cockerell
Colo.

Andreyia ramaleyi Cockerell, 1931. Ann. and Mag. Nat. Hist. (10) 7: 346. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 14 (key).

Andrena robervalensis Mitchell
Que., Mich., Minn. Pollen: Unknown, but visits flowers of Melilotus
alba.
Andrena {^Cnemidandrena) robervalensis Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui.
141: 175. ♀.

Andrena scutellinitens Viereck
Oreg., Calif., Nev. Pollen: Unknown, but visits flowers of

Chrysothamnus nauseosus speciosus, C. viscidiflorus typicus, Haplopappus bloomeri
var. ayigustatus, Oxypolis occidentalis, Solidago elongata.
Andrena {Andrena) scutellinitens Viereck, 1916. Acad. Nat. Sci. Phila, Proc. 68: 573. ♀, ♂.

Taxonomy: LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 294 (subgeneric position).

Andrena segregans Cockerell
N. Mex.

Andrena segregans Cockerell, 1900. Ann. and Mag. Nat. Hist. (7) 5: 404. ♀, ♂.

Andrena surda Cockerell
Wyo., Colo., Oreg., Calif., N. Mex. Pollen: Unknown, but visits flowers of
Chrysothamnus nauseosus, C. viscidiflorus typicus, Erigeron pygmaeus,
Sphenosciadium capitellatum.
Andreyia hirticiyicta var. stirda Cockerell, 1910. Ann. and Mag. Nat. Hist. (8) 5: 264. ♂.
Ayidreyia pertarda Cockerell, 1916. Entomologist 49: 157. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 13, 14. 9, cJ (key). —Lanham, 1941.
Ent. Soc. Amer., Ann. 34: 706, 708. 9, 6 (key). — Timberlake, 1951. In Linsley In
Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1082 (synonymy).

Andrena xanthigera Cockerell
N. Mex.

Ayidreyia xayithigera Cockerell, 1900. Ann. and Mag. Nat. Hist. (7) 5: 402. ♀.
Ayidreyia albovirgata Cockerell, 1900. Ann. and Mag. Nat. Hist. (7) 5: 403. ♀.

Taxonomy: Cockerell, 1931. Canad. Ent. 63: 22. 9. —Cockerell, 1931. Amer. Mus. Novitates
458: 19. ♀.

Genus ANDRENA Subgenus CONANDRENA Viereck

Ayidrena subg. C oyiayidreyia Viereck, 1924. Canad. Ent. 56: 20.

Type-species: Ayidreyia bradleyi Viereck. Monotypic and orig. desig.

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 206 (tax. characters). —LaBerge, 1964.
Nebr. Univ. State Mus., Bui. 4: 284, 288, 300 (tax. characters).




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Andrena bradleyi Viereck
Colo, to N. S., south to Ga. Pollen: Unknown, but visits flowers of Pru^ius,
Pyrus, Rliododendron, Ribes, Vaccinimn.
Andrena bradleyi Viereck, 1907. Ent. News 18: 285, 286. ♀, ♂.
Andrena saccharina Cockerell and Rohwer, 1907. Ann. and Mag. Nat. Hist. (4) 20: 128. ♂.

Taxonomy: Atwood, 1934. Canad. Jour. Res. 10: 206, 208, figs. 9, S (key). — Timberlake,
1951. In Linsley In Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2:
1059 (synonymy). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 226-228, figs. 40,
46-47, 49-50 (redescription).

Biology: Salt, 1927. Jour. Expt. Zool. 48: 249 (stylopization).
Carolina Viereck. Minn, to N. S., south to Tenn. and Ga. Pollen: Unknown, but visits flowers of
Azalea, Claytonia, Dentaria, Ledum, Pyrus, Rhodora, Vaccinium.
Andrena Carolina Viereck, 1909. Ent. News 20: 126. ♀.

Taxonomy: Atwood, 1934. Canad. Jour. Res. 10: 208, 209. 9, d. —Mitchell, 1960. N. C. Agr.
Expt. Sta. Tech. Bui. 141: 228-229, figs. 40, 45-46, 49 (redescription, flower records).

Andrena cheyennorum Viereck and Cockerell
Nebr., Wyo., Idaho.

Andrena cheyennorum Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 20. ♀, ♂.
Andrena angustifrons Cockerell, 1934. Pan-Pacific Ent. 9: 155. ♀.

Taxonomy: Timberlake, 1951. In Linsley hi Muesebeck, Krombein and Townes, U. S. Dept.
Agr., Agr. Monog. 2: 1061 (synonymy).

Andrena durangoensis Viereck and Cockerell
Colo, to N. S., south to W. Va. Ecology: Unknown, but
visits flowers Ilex, Rubus.
Andrena durangoensis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 36. ♂.
Andrena (Andrena) media Viereck, 1922. Boston Soc. Nat. Hist., Occas. Papers 5: 41. ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 229-230, figs. 40, 45, 46
(redescription, synonymy).

Andrena rufosignata Cockerell
Minn, to N. B. and New England states.
Andrena rufosignata Cockerell, 1902. Canad. Ent. 34: 46. ♀.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 230, fig. 49 (redescription,
subgeneric assignment). — Knerer and Atwood, 1963. Canad. Ent. 95: 585, figs. 5, 7. ♂.

Genus ANDRENA Subgenus DACTYLANDRENA Viereck

Andrena subg. Dactylandrena Viereck, 1924. Canad. Ent. 56: 20.

Type-species: Andrena (Dactylandrena) maura Viereck. Monotypic and orig. desig.

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 204 (tax. characters). — LaBerge, 1964.
Nebr. Univ. State Mus., Bui. 4: 285, 288, 299 (tax. characters).

Andrena berberidis Cockerell
Colo. Pollen: Unknown, but visits flowers of Berberis.

Andrena milwaukeensis var. berberidis Cockerell, 1905. Canad. Ent. 37: 371. ♀.

Taxonomy: Cockerell, 1914. Ann. and Mag. Nat. Hist. (8) 13: 9. 6. —Lanham, 1941. Ent. Soc.
Amer., Ann. 34: 705, 707. 9, i (key).

Andrena caliginosa Viereck
Calif. Pollen: Unknown, but possibly an oligolege oi Ribes.

Andrena (Andrena) caliginosa Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 552. ♀.
Andrena (Dactylandrena) maura Viereck, 1924. Canad. Ent. 56: 31. ♂ (♀ misdet.).

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 267. 9, 6 (key).

Andrena submaura Linsley
South. Calif. (The Gavilan and Sandbergs). Pollen: Unknown, but possibly
gathers pollen from Ribes indecorum.
Andrena submaura Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 269. ♂, ♀.

Genus ANDRENA Subgenus DERANDRENA Ribble

Andrena subg. Derandrena Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 333.
Type-species: Andrena vandykei Cockerell. Orig. desig.

Revision: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 333-367, figs. 90-124, maps 11-18,
tables 12-14 (N. Amer. spp.).




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Andrena arctostaphylae Ribble
Calif. Pollen: Apparently an oligolege of Arctostaphylos including A.
Crustacea, A. glauca, A. mariposa, A. pungens.
Andrena (Derandrena) arctostaphylae Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 344,
figs. 95-99, map 12. ♀, ♂.

Andrena californiensis Ribble
Calif. Pollen: Unknown, most often collected on flowers of Salix

including S. argophylla, S. lasiolepis, but visits a wide variety of flowers including
Arctostaphylos pungens, Cryptantha intermedia, Descourainia sophia, Eriodictyon
caiifornicuni, Gilia multicaulis, Lasthenia chrysostoma, Lomatimn dasycarpmn,
Orthocarpus, Plagiobothrys nothofulvus, Quercus, Rhamnus crocea, Rhus trilobata,
Ribes indecorum. Salvia columbariae. Sisymbrium irio, Tamarix gallica, Trifolium.
Andrena (Derandrena) califomiensis Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 348,
figs. 100-104, map 13, table 13. ♀, ♂.

Andrena hermosa Ribble
Calif. (Central Valley). Pollen: Unknown, but possibly oligolectic on
Lasthenia.
Andrena (Derandrena) hermosa Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 360, figs.
115-119, map 18. ♀, ♂.

Andrena murietae Ribble
Calif. (11 miles northwest of California Hot Springs). A. penutiani Ribble
may be the male.
Andrena (Derandrena) murietae Ribble, 1968. Nebr. Univ. State Mus. Bui. 8: 346. ♀.

Andrena penutiani Ribble
Cent. Calif. Pollen: Unknown, but visits flowers of Lasthenia. A. murietae
Ribble may be the female.
Andrena (Derandrena) penutiani Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 352, map
15. ♂.

Andrena timberlakei Cockerell
Calif. (San Benito and Tulare Cos., south to Riverside Co.). Pollen:
Based upon the mouth parts of the female, this species evidently collects pollen from
the flowers of Cryptantha including C. intermedia, but visits other flowers presumably
for nectar including Descourainia sophia, Encelia farinosa, Lomatium dasycarpum,
Plagiobothrys califomicus, P. nothofulvus, Rhamnus crocea.
A7idrena timberlakei Cockerell, 1929. Ann. and Mag. Nat. Hist. (10) 4: 300. ♀, ♂.

Taxonomy: Cockerell, 1931. Canad. Ent. 64: 158. ♀, ♂.

Andrena vandykei Cockerell
South. Oreg. (12 miles southwest of Keno), Calif.; Mexico (Baja California).
Pollen: Unknown, most often collected on flowers of Arctostaphylos including A.
Crustacea, A. glauca, A. mariposa, A. pungens as well as Ceanothus including C.
arboreus, C. cuneatus, C.foliosus, but visits other flowers including Baccharis,
Phoradenron villosum, Rhamnus crocea, Rhus diversiloba, R. ovata, Salix, Tamarix,
Xylococcus bicolor.
Andrena (Micrandrena) solutula Cockerell, 1936. Pan-Pacific Ent. 12: 150. ♀.
Andrena (Micrandrena) vandykei Cockerell, 1936. Pan-Pacific Ent. 12: 151. ♂.

Andrena viridissima Ribble
Calif.; Mexico (Baja California). Pollen: Unknown, but visits flowers of
Brassica, Lasthenia tenella, Nemophila raceinosa, Phacelia ciliata, P. distaiis, P.
douglasii, Pholisma racemosa. Ranunculus canus.
Andrena (Derandrena) viridissima Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 360, figs.
115-119, map 18. ♀, ♂.

Andrena ziziaeformis Cockerell
Mass. to Ga., Tenn., 111. Pollen: Unknown, but visits flowers of
Potentilla including P. canadensis, Rubus, Waldsteinia.
Andrena ziziaeformis Cockerell, 1908. Canad. Ent. 40: 234. ♀.

Taxonomy: Cockerell, 1932. Canad. Ent. 64: 157. 9 (key). -Mitchell, 1960. N. C. Agr. Expt.
Sta. Tech. Bui. 141: 169-171, figs. 29-32 (redescription).

Genus ANDRENA Subgenus DIANDRENA Cockerell

Diandrena Cockerell, 1903. Psyche 10: 75.

Type-species: Panurgus chalybaeus Cresson. Orig. desig.

Revision: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 1-146, 88 figs, 5 tables, 13 maps (N. Amer.
spp., includes comparative ecological information).




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Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 219-220 (tax. characters). — Linsley and
MacSwain, 1961. Pan-Pacific Ent. 37: 31-41 (Camissonia visiting spp.). — LaBerge, 1964.
Nebr. Univ. State Mus., Bui. 4: 283, 287, 309-310 (tax. characters). -Linsley, MacSwain,
Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 13-26, figs. 6-11 (Camissonia visiting
spp. of Calif, and Baja Calif.).

Biology: Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 74-75, pis. 1-3 (floral
relationships of Mojave Desert spp.). —Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 14-43,
124-128, figs. 1-2, tables 1-5 (nest architecture, life history, comparative ecology, parasites,
predators). —Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 13-26,
47-50, tables 1-2 (life history, floral relationships of Camissonia visiting spp.).
abiegata (Cockerell). Idaho, Wyo., Colo., Utah Calif. Pollen: Oligolectic on ligulate Compositae,
most likely collects pollen from early morning-opening flowers of the subtribe
Microseridinae including Agoseris glauca, Microseris nutans, Taraxacum officinale, but
visits other flowers including Camissonia subacaulis, Cardaria draba, Lomatium
dissectum.
Diandrena abiegata Cockerell, 1922. Amer. Mus. Novitates 40: 1. ♀.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 32-33 (tax. characters).
—Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 93-95, figs. 1, 29-30, 56, map 7 (redescription).

Andrena agoseridis Thorp
Calif. Pollen: Collects pollen from ligulate Compositae, primarily from

Agoseris heterophylla, Anisocoma acaulis, but visits other flowers including Calycoseris
parryi, Camissonia campestris campestris, C. pallida pallida, Descourainia sophia,
Erodium cicutarium, Euphorbia albomarginata, Hesperochiron, Lasthenia
chrysostoma, Layia glandulosa, Malacothrix califomica, M. glabrata, Phacelia ciliata,
Rafinesquia neomexicana. Sisymbrium, Tamarix gallica.
Andreiia (Diandrena) agoseridis Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 116, figs. 1, 49-50,
56, map 12. ♀, ♂.

Andrena anatolis Linsley and MacSwain
South. Cahf.; Mexico (Baja California). Pollen: Collects pollen
from morning-opening flowers of Camissonia including C. bistorta, C. campestris, C.
cheirantkifolia suffruticosa, but visits other flowers including Calandrinia ciliata,
Camissonia crassifolia, Ceanothus crassifolius, Cryptantha intermedia, PotentiUa,
Sitanion for nectar.
Andreyia (Diandrena) anatolis Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 36. ♀, ♂.

Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 98-99, figs. 1, 33-34, 56, map 8. 9, d
(redescription).

Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 99, fig. 1 (floral relationships). —Linsley,
MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 21-22, 49, fig. 9 (floral
relationships).

Andrena apasta Linsley and MacSwain
Calif. Pollen: Collects pollen from morning-opening flowers of
Camissonia, principally from C. campestris, but occasionally from C. ovata in
northernmost localities; also visits other flowers including Agoseris heterophylla,
Brassica, Descourainia sophia, Layia, Monolopia lanceolata for nectar.
Andrena (Diandrena) apasta Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 34. ♀.

Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 118-119, figs. 1, 51-52, 56, 68, 80, map 10
(redescription).

Biology: Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 75, fig. 11 (nest,
floral relationships). —Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 99 (floral relationships).
—Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 26, 50, fig. 11
(floral relationships).

Andrena blennospermatis Thorp
Calif. Pollen: Collects pollen from flowers of Blennosperma including
B. bakeri, B. nanum.
Andrena (Diandrena) blennospermatis Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 81, figs. 1,
19-20,56,63,75. ♀, ♂.

Andrena chalybaea (Cresson)
Calif. (Central coast, from Mendocino to San Luis Obispo Counties).
Parasite: Nomada sp. Pollen: Collects pollen only from the early morning-opening




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flowers of Camissonia ovata, but visits other flowers for nectar including
Blennosperma bakeri, Brassica, Cryptantha, Ranunculus caUfomicus.
Panurgus chalybaeus Cresson, 1878. Amer. Ent. Soc, Trans. 7: 61. ♀, ♂.

Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 57-61, figs. 1, 3-4, 55-58, 69, 85-86, map 1
(redescription, post defecating larva).

Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 5-14, 59, fig. 2 (nest architecture, floral
relationships, parasite). — Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs.
Ent. 71: 15-17, 48, fig. 6, pi. 1 (nest, floral relationships, mating).

Andrena chalybioides (Viereck)
Wash., Oreg., Calif. Parasite: Myopa rubida (Bigot)? Pollen: Collects
pollen almost exclusively from ligulate Compositae in the early morning, primarily from
Agoseris including ^4. heterophylla, occasionally from introduced weedy species of
Cichorieae including Hypochoeris radicata, Taraxacum officinale, but also visits other
flowers for nectar including Blennosperma, Ceanothus, Convolvulus, Daucus, Lasthenia
chrysostoma, Layia platyglossa, Linum, Ranunculus orthorhynchus.

Parandrena chalybioides Viereck, 1904. Canad. Ent. 36: 229. ♀, ♂.

Andrena {Parandrena) perchalybea Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 591. ♀.

Diandrena purdyi Cockerell, 1936. Pan-Pacific Ent. 12: 155. ♂.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 31, 33 (synonymy). —Thorp,
1969. Calif. Univ. Pubs. Ent. 52: 83-85, figs. 1, 21-22, 56 (redescription).

Biology: MacSwain and Bohart, 1947. Pan-Pacific Ent. 23: 30 (parasite). —Thorp, 1969. Calif.
Univ. Pubs. Ent. 52: 34, 85 (floral relationships, parasite).

Andrena chlorosoma Linsley and MacSwain
Calif. (Central Coast Ranges, from Mendocino County to
Ventura County). Pollen: Collects pollen from early morning opening ligulate
Compositae including Agoseris grandiflora, A. heterophylla as well as from introduced
weedy species of Cichorieae including Hypochoens radicata, but visits these and other
flowers for nectar including Camissoyiia ovata, Hemizonia, Lasthenia chrysostoma,
Layia chrysanthemoides. Ranunculus californicus. Taraxacum officinale.
Andrena {Diandrena) subchalybea chlorosoma Linsley and MacSwain, 1961. Pan-Pacific
Ent. 37:32. ♀, ♂.

Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 107-111, figs. 1, 41-44, 56 (redescription).

Biology: Linsley and MacSwain, 1959. Calif. Univ. Pubs. Ent. 16: 18, 30, 31 (floral
relationships, as purdyi). —Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 110, 111 (floral
relationships).

Andrena cuneilabris Viereck
Oreg., Calif. Parasite: Myopa perplexa Camras, M. rubida (Bigot), Stylops
pacifica Bohart? Pollen: Collects pollen and nectar chiefly from Ranunculus including
R. californicus, R. occidentalis, R. orthorhynchus, occasionally collects pollen from
Platyglossa californicus during periods when Ranunculus pollen is scarce or absent;
visits other flowers for nectar including Amsinckia, Agoseris grandiflora, A.
heterophylla, Blennosperma bakeri, Brassica, Calandrinia ciliata, Camissonia ovata, C.
tanacetifolia, Geum macrophyllum, Lasthenia, Lomatium, Wyethia.
Andrena {Parandrena) cuneilabris Viereck, 1926. Calif. Acad. Sci., Proc. (4) 15: 400. ♀.

Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 64-67, figs. 1, 7-8, 56, 59, 70, 81-82, map 2
(redescription).

Biology: MacSwain and Bohart, 1946. Pan-Pacific Ent. 23: 30 (parasite). —Camras and Hurd,

1957. Calif. Ins. Survey, Bui. 6: 39 (parasite). —Linsley and MacSwain, 1959. Calif. Univ.

Pubs. Ent. 16: 2-31, figs. 4, 6 (floral relationships, parasites). —Thorp, 1969. Calif. Univ.

Pubs. Ent. 52: 34, 66-67 (floral relationships, parasites).

Andrena cyanosoma (Cockerell)
Calif.; Mexico (Baja California). Pollen: Collects pollen exclusively

from the flowers of Camissonia including C. bistorta, C. campestris, C. cheiranthifolia
suffruticosa, C. dentata, C. sierrae sierrae, C. spiralis, but visits these and other flowers
for nectar including Cryptantha intermedia, Lasthenia aristata, L. coronaria, Rhus
trilobata, Salix.

Diandrena cyanosoma Cockerell, 1916. Pomona Jour. Ent. Zool. 8: 49. ♀, ♂.




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Andrena (Parandrena) austrocalifomica Viereck, 1917. Acad. Nat. Sci. Phila., Proc. 68:

587. ♀.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 36 (synonymy). —Thorp, 1969.
Calif. Univ. Pubs. Ent. 52: 99-102, figs. 1, 35-36, 56, 65, 77, map 9 (redescription).

Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 101 (floral relationships). —Linsley,

MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 23-24, fig. 10 (floral

relationships).

Andrena eothina Linsley and MacSwain
Calif. (Merced County to Ventura County). Pollen: Collects
pollen principally from Camissonia campestris and C. sierrae sieirrae and also from C.
boothii decorticans, C. contorta, C. graciliflora, but visits these and other flowers for
nectar including Agoseris heteropkylla, Amsinckia, Encelia virginiensis actoni,
Lasthenia chrysostoma, Layia, Monolopia lanceolata, Tropidocarpum gracile.

Andrena (DiandreJia) eothina linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 39. ♀, ♂.

Andrena (Diandrena) anatolis mahdina Linsley and MacSwain, 1961. Pan-Pacific Ent. 37:
38. ♀, ♂.

Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 95-97, figs. 1, 31-32, 56, map 8
(redescription, synonymy).

Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 97 (floral relationships). —Linsley,
MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 22-23, fig. 9 (nest
architecture, floral relationships).

Andrena evoluta Linsley and MacSwain
Wash., Idaho and Wyo., south to northeast. Calif., Nev. and
northern Ariz. (Grand Canyon). Pollen: Collects pollen primarily from microseridine
Compositae including Agoseris glauca, A. g. var. monticola, Microseris nutans and
locally obtains pollen from some of the crepidine Compositae such as Crepis
occidentalis, the onagraceous Camissonia including C. pallida, C. tanacetifolia, and the
introduced weed Taraxacum officinale; visits other flowers for nectar including
Ranunculus califomicus.
Andrena {Diandrena) evoluta Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 33. ♀.

Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 85-88, figs. 1, 23-24, 56, map 6
(redescription).

Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 40-41 (floral
relationships). —Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 88 (floral relationships).

Andrena foxii Cockerell
Calif. Pollen: Collects pollen exclusively from flowers of Camissonia, chiefly C.
campestris, but also from C. contorta, C. dentata parishii and C. pallida pallida when
C. campestris is seasonally or locally rare or absent; visits these and other flowers for
nectar including Coreopsis califomica, Cryptantha, Hemizonia, Isomeris arborea,
Lupinus, Malacothrix.
Andrena foxii Cockerell, 1898. Amer. Ent. Soc, Trans. 25: 188. ♀, ♂.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 130 (subgeneric position).
—Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 67-69, figs. 1, 9-10, 56, 60, 71, 87, map 2
(redescription).

Biology: Linsley and MacSwain, 1956. Pan-Pacific Ent. 32: 120-121 (floral relationships).

— Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 75 (floral relationships).

—Thorp 1969. Calif. Univ. Pubs. Ent. 52: 69 (floral relationships). —Linsley, MacSwain,

Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 18-20, 48-49, fig. 7, pi. 1 (floral

relationships).

Andrena gnaphalii (Cockerell)
South. Calif. (Catalina Island and Transverse and Peninsular Ranges).
Pollen: Collects pollen apparently only from early morning opening flowers of Agoseris
including A. heteropkylla, A. retrorsa, but visits these and other flowers for nectar
including Anisocoma acaulis, Ceanothus integerrimus, Descourainia sophia,
Gnaphalium bicolor, Lasthenia chrysostoma, Potentilla, Sisymbrium, Sonchus
oleraceus.

Diandrena gnaphalii Cockerell, 1938. Ann. and Mag. Nat. Hist. (11) 2: 148. ♂.




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Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 91-93, figs. 1, 27-28, 56, map. 13

(redescription).
Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 92-93 (floral relationships).

Andrena lewisorum Thorp
Calif. Pollen: Collects pollen from flowers of the genus Clarkia including C.
breweri, C. concinna, C. cylindrica, C. douglasiana, C. dudleyana, C. unbricata, C.
purpurea, C. speciosa, C. unguiculata, C. williamsonii, C. xantiana, but visits these and
other flowers for nectar including Baccharis, Cryptantha flaccida, Datura, Enogonum
fasciculatum, Hemizonia kelloggii, Lonicera interrupta. Mini ul us, Ranuncuhis
occidentalis, Rhamnus californica.
Andrena (Diandrena) lewisorum Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 72, figs. 1, 13-14,
56, 61, 73, map 3. ♀, ♂.

Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 73-74 (floral relationships). — MacSwain,
Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 70: 10-13, fig. 1 (floral relationships).

Andrena macswaini Linsley
Calif. (San Joaquin Valley). Pollen: Collects pollen from the flowers of
Camissonia campestris campestris and C. sierrae sierrae, but visits these and other
flowers for nectar including Camissonia boothii decorticans, Erodium, Isomeris
arborea, Malacothrix californica.
Andrena (Diandrena) macswaini Linsley, 1960. Pan-Pacific Ent. 36: 97. ♀, ♂.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 36 (tax characters). —Thorp,
1969. Calif. Univ. Pubs. Ent. 52: 102-105, figs. 1, 37-38, 56, 66, 78, map 10 (redescription).

Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 105 (floral relationships). —Linsley,
MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 24-26, 50, pi. 2, fig. 11 (nest
architecture, floral relationships).

Andrena malacothricidis Thorp
South. Calif.; Mexico (Baja California). Pollen: Collects pollen from
stephanomerine Compositae, primarily Malacothrix including M. californica, M.
glabrafa, but visits other flowers including Anisocoma acaulis, Calochortus,
Camissonia campestris campestris. Convolvulus malacophyllus, Cryptantha,
Descourainia sophia. Euphorbia albomarginata, Lasthenia chrysostoma, Linanthus
aureus, Phacelia ciliata.
Andrena (Diandrena) malacothricidis Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 113, figs. 1,
47-48, 56, map 11. ♀, ♂.

Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 115-116 (floral relationships).

Andrena nothocalaidis (Cockerell)
B. C. to north. Calif., east to Wyo. and Colo. Pollen: Collects pollen
from flowers of ligulate Compositae, primarily Microseridinae including Agoseris glauca,
Microseris nutans, Nothocalais cuspidata, Taraxacum officinale, but visits other
flowers for nectar including Antenyiaria, Camissonia tanacetifolia tanacetifolia, Crepis
occidentalis, Erigeron, Lepidium, Mimulus.
Diandrena nothocalaidis Cockerell, 1905. Biol. Soc. Wash., Proc. 18: 183. ♀, ♂.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 33-35 (tax. characters and
status). —Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 105-107, figs. 1, 39-40, 56, map 11
(redescription).

Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 107 (floral relationships).

Andrena olivacea Viereck
Ariz, to south. Nev. and south. Calif. Pollen: Collects pollen almost

exclusively from the flowers of stephanomerine Compositae, primarily Anisocoma
acaulis and Malacothrix including M. californica, M. glabrata, but visits other flowers
including Agoseris heterophylla. Astragalus, Baileya, Calycoseris wrightii, Camissonia
campestris. Coreopsis bigelovii, C. californica, Cryptantha barbigera, Encelia farinosa,
Fallugia paradoxa, Geraea canescens, Hemizonia, Hyptis emoryi, Layia glandulosa,
Phacelia, Rafinesquia neomexicana. Predator: Lestomyia sp.
Andrena (Parandrena) olivacea Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 590. ♀, ♂.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 35 (tax. characters). —Thorp,
1969. CaHf. Univ. Pubs. Ent. 52: 88-91, figs. 1, 25-26, 56, 64, 76, map 7 (redescription).

Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 91 (floral relationships).




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Andrena parachalybea Viereck
South. Calif.; Mexico (Baja California). Pollen: Collects pollen from
Cainissonia including C. bistorta, C. campestris cavipestris, C. cheirantliifolia
suffruticosa, C. dentate, C. pallida pallida, but visits these and other flowers for nectar
including Agoseris heterophylla, Astragalus, Baccharis virninea, Ceanothus crassifolius,
Coreopsis bigelovii, C. califomica, Cryptantha intermedia, Descourainia sophia,
Erodium cicutarium, Haplopappus linearifolius, Eriophyllum confertiflornm,
Lasthenia, Layia glandulosa, L. platyglossa campestris, Lomatium dasycarpum,
Monolopia lanceolata, Plagiobothrys califomicus, Rhus trilobata, Salix laevigata, S.
lasiolepis, Sisymbrium irio, Tamarix.
Andrena (Parandrena) parachalybea Viereck, 1917. Amer. Ent. Soc, Trans. 43: 391. ♀.

Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 61-63, figs. 1, 5-6, 56, map 1
(redescription).

Biology: Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 74-75 (floral
relationships). —Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71:
17-18, 48, fig. 6 (floral relationships).

Andrena puthua (Cockerell)
Calif.; Mexico (Baja California). Parasite: Stylops sp. Pollen: Collects pollen
from Lasthenia including L. aristata, L. chrysostoma, L. coronaria, L. debilis, L.
gracilis, L. minor, L. tenella, but visits these and other flowers for nectar including
Calandrinia ciliata, Chaenactis glabriuscula, Cryptayitha, Eschscholzia califomica,
Layia platyglossa, Rhamnus crocea.

Diandrena puthua Cockerell, 1910. Ann. and Mag. Nat. Hist. (8) 5: 24. cJ (9 misdet.).

Diandrena beatula Cockerell, 1916. Pomona Jour. Ent. Zool. 8: 48. ♀.

Diandrena clariventris Cockerell, 1916. Pomona Jour. Ent. Zool. 8: 50. ♂.

Diandrena sanctorum Cockerell, 1941. San Diego Soc. Nat. Hist., Trans. 9: 346. ♀.

Taxonomy: Timberlake, 1951. In Linsley hi Muesebeck, Krombein and Townes, U. S. Dept.
Agr., Agr. Monog. 2: 1078 (synonymy). —Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 74-78,
figs. 1, 15-16, 56, map 4 (redescription, synonymy).

Biology: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 34, 77 (floral relationships, parasite).
sperryi (Cockerell). Calif. Pollen: Collects pollen from the flowers of Caynissonia including C.
campestris, C. clavaeforrnis, C. pallida pallida, but visits these and other flowers for
nectar including Anisocoma acaulis. Coreopsis califomica, C. bigelovii, Malacothrix
glabrata. Predator: Lestomyia sp.
Diandrena sperryi Cockerell, 1937. Amer. Mus. Novitates 948: 14. ♀.

Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 69-72, figs. 1, 11-12, 56, 72, map 3
(redescription).

Biology: Linsley, MacSwain and Thorp, 1964. Calif. Univ. Pubs. Ent. 33: 74, table 3 (floral
relationships). —Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 33, 72 (floral relationships,
predator). —Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 20, 49,
fig. 8 (floral relationships).

Andrena subapasta Thorp
Calif. Pollen: Apparently collects pollen primarily from Arenaria califomica,
but visits other flowers including Lasthenia, Orthocarpus erianthus.
Andrena (Diandrena) subapasta Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 120, figs. 1, 53-54,
56, map 13. ♀, ♂.

Andrena subchalybea Viereck
Calif.; Mexico (Baja California). Parasite: Dalmannia picta Will. Pollen:
Collects pollen primarily from ligulate Compositae, especially Agoseris heterophylla;
visits flowers of Am sinckia, Brassica, Camissonia campestris campestris. Coreopsis
califomica, Crepis vesicaria, Descourainia sophia, Erodium, Euphorbia albomarginata,
Haplopappus Linearifolius, Hesperochiron califomicus, Lasthenia chrysostoma, Layia
chrysanthemoides, Layia platyglossa, Limnanthes douglasii. Madia gracilis,
Malacothrix, Microseris douglasii, M. linearifolia, Mimulus, Monolopia lanceolata,
Phacelia ciliata, Platystemon califomicus. Ranunculus califomicus. Sisymbrium,
Sonchus oleraceus.
Andrena (Parandrena) subchalybea Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 593. ♀.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 32 (tax. status). —Thorp, 1969.
Calif. Univ. Pubs. Ent. 52: 111-113, figs. 1, 45-46, 56, 67, 79, 83-84, map 12 (redescription).




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Biology: Bohart, 1938. Pan-Pacific Ent. 14: 132 (parasite). —Thorp, 1969. Calif. Univ. Pubs.
Ent. 52: 34, 113 (floral relationships, parasite).

Andrena submoesta Viereck
Calif. Parasite: Stylops sp. Pollen: Collects pollen from the flowers of

Lasthe7iia including L. chrysostoina, L. coronaria, L. gracilis, L. minor, L. tenella, but
visits these and other flowers for nectar including Haplopappus, Hypochoeris radicata,
Layia, Monolopia lanceolata, Tamarix.
Andrena (Parandrena) submoesta Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 594. ♀.
Diandrena marinensis Cockerell, 1936. Pan-Pacific Ent. 12: 158. ♂.

Taxonomy: Timberlake, 1951. In Linsley, In Muesebeck, Krombein and Townes, U. S. Dept.
Agr., Agr. Monog. 2: 1082 (synonymy). —Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 78-81,
figs. 1, 17-18, 56, 62, 74, map 5 (redescription).

Biology: Thorp, 1969. Calif Univ. Pubs. Ent. 52: 34, 79-80 (floral relationships, parasite).

Genus ANDRENA Subgenus EREMANDRENA LaBerge

Andrena subg. Eremandrena LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 295.
Type-species: Pterandrena pallidiscopa Viereck. Monotypic and orig. desig.

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 283, 293, 295-296 (tax. characters).

Andrena pallidiscopa
***authority mismatch
pallidiscopa (Viereck). Calif., Oreg., Colo. Parasite: Myopa rubida (Bigot). Pollen:
Unknown, but visits flowers of Lomatiiim, Sanicula.
Pterandrena pallidiscopa Viereck, 1904. Canad. Ent. 36: 227. ♀.
Pterandrena nudimedicomis Viereck, 1904. Canad. Ent. 36: 227. ♂.

Taxonomy: Timberlake and Cockerell, 1933. Pan-Pacific Ent. 9: 28 (tax. characters).

Biology: Bohart, 1941. Pan-Pacific Ent. 17: 95 (parasite).

Andrena pallidiscopa
***authority mismatch
trifasciata Timberlake and Cockerell. South. Calif. Parasite: Myopa rubida
(Bigot). Pollen: Unknown, but visits flowers of Calandrinia menziesii, Lasthenia
chrysostoma, Lomatium dasycarpum, L. utriculatum, Rhus trilobata, Sanicula
bipinnatifida.
Andrena pallidiscopa trifasciata Timberlake and Cockerell, 1933. Pan-Pacific Ent. 9: 28.
9.

Genus ANDRENA Subgenus EUANDRENA Hedicke

Andrena subg. Eiiandrena Hedicke, 1933. Berlin Zool. Mus., Mitt. 19: 212.

Type-species: Andrena bicolor Fabricius. Orig. desig.
Andrena subg. Xanthandrena Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 218.

Type-species: Andrena auricoma Smith. Orig. desig.
Andrena subg. Geandrena LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 313.

Type-species: Ayidrena caerulea Smith. Orig. desig.

Revision: LaBerge and Kibble, 1975. Amer. Ent. Soc, Trans. 101: 371-446, 74 figs. (N. Amer.

spp.).

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 309 (tax. characters, as

Xanthandrena). —LaBerge, 1977. Amer. Ent. Soc, Trans. 103: 107-135, figs, (revised key

to included spp., synonymy; received too late for inclusion in this catalog).

Andrena algida Smith
N. S. to Alta. and N. W. T., south to N. J., N. Y., Mich., Minn., S. Dak., N. Mex.
and Ariz. Pollen: Apparently polylectic with some preference for the flowers of Salix
including S. brachycarpae, but visits other flowers including Comus mas, Fragaria,
Pulsatilla hirsutissima, Ranunculus iesticulatus, Ribes vallicola. Taraxacum
officinale, Thlaspi arvense.

Andrena algida Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 116. ♀, ♂.

Andrena fragiliformis Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 435. ♀.

Andrena femaldiella Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 34. ♂.

Andrena brachycarpae Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 55. ♀.

Andrena {Andrena) fuscisignata Viereck, 1917. Amer. Ent. Soc, Trans. 45: 379. ♀, ♂.

Andrena {Andrena) albisigna Viereck, 1922. Boston Soc. Nat. Hist., Occas. Papers 5: 38.
9.




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Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 150 (type). — Cockerell, 1906. Psyche
13: 7. 9 (type). -Atwood, 1934. Canad. Jour. Res. 10: 207, 209, figs. 9,6. -Mitchell, 1960.
N. C. Agr. Expt. Sta. Tech. Bui. 141: 218-219, figs. 42-44 (redescription). — LaBerge and
Ribble, 1975. Amer. Ent. Soc, Trans. 101: 396-399, figs. 3, 25-29 (redescription, synonymy).

Andrena astragali Viereck and Cockerell
Wash, to Calif, east to Utah, Wyo. and Nebr. Pollen:

Unknown, although frequently collected from the flowers of Zygadenus fremontii, but
visits other flowers including Agoseris glauca, Allium haematochiton, Astragalus,
Cardaria draba var. repens, Cliamaebatia foliolosa, Conius, Eriogomim, Oxytropis,
Salix, Taraxacum officinale, Tofieldia occideyitalis.
Andrena astragali Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 46. ♀.
Andrena zygadeni Cockerell, 1932. Pan-Pacific Ent. 8: 174. ♀, ♂.

Andrena auricoma Smith
B. C. to south. Calif., east to Wyo., Colo, and south to Ariz. Pollen: Polylectic,
most frequently collected at the flowers oi Ranunculus, Descourainia, Salix and
Potentilla in that order, but visits a great variety of flowers including Baccliaris,
Barbarea orthoceras, Brassica 7iigra, Camissonia, Ceanothus, CItainaebatia foliolosa,
Conius califomicus, Crataegus, Cryptantha intermedia, C. muricata, Daucus carota,
Dendromecon rigida, Descourainia sophia, Eriodictyon califomicum, Eriogonum
marifolium, Eriopbyllum obovatum, Euphorbia albomarginata, Fragaria califoniica,
Fntillaria lanceolata. Geranium, Gilia capitata, Hackelia floribunda, H. patens,
Lasthenia chrysostoma, Layia, Linanthus montanus, Lomatiuvi dissectum, L.
triternatum, Lupinus bicolor, Malacothamnus arcuatus, Malus, Mentzelia, Mirabilis
laevis, Montia perfoliata, Nemophila integrifolia, Ortbocarpus erianthus, Phacelia
cicutaria, P. distans, P. hispida, P. ramosissiyna, P. tanacetifolia, Plagiobothrys
califomicus, P. tenellus, Potentilla glandulosa, Pnaius ilicifolia, P. melanocarpa, P.
subcordata, Pyracantha, Quercus agrifolia, Ra7iu7iculus califomicus, R. occidentalis,
Rhannius crocea, Rhus diversiloba, R. trilobata, Rubus ursiyius, Salix laevigata, S.
lasiolepis, Salvia columbariae, Sanicula crassicaula, S. nevadensis, Saxifraga
integrifolia, Scrophularia califomica, Sedum, Sisymbriinn irio, Sisyrinchium bellum,
Stachys, Stenotopsis linearifolius, Tamarix gallica. Taraxacum officinale,
Thysanocarpus curvipes.
Andrena auricoma Smith, 1879. Descr. New Species Hym. Brit. Mus., p. 56. ♂.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 155. 6 (notes on type). —Cockerell,

1906. Psyche 13: 8 (notes on type). —Viereck, 1904. Canad. Ent. 36: 193. 9, 6 (key).

—Lanham, 1941. Ent. Soc. Amer., Ann. 34: 706, 708. 9, 6 (key).

Andrena caerulea Smith
B. C. (Vancouver Is.) to Calif., east to Idaho, Nev., and Utah (Mendon).

Parasite: Myopa riibida (Bigot), Myopa sp., Nomada obscurella Fowler, N. opacella
Timberlake, Stylops pacifica Bohart. Pollen: Oligolege of Ranunculus, especially R.
califomicus, but visits other species of Ranimcuius including R. alsijuaefolius, R.
occidentalis, R. o. dissectus, R. orthorhynchus as well as the flowers of Arctostaphylos,
Astragalus, Brassica, Ceanothus including C. cuneatus, Cryptantha, Fagus, Gilia,
Ipomoea, Lasthenia chrysostoma, Lomatium, Lysichiton kayntschatcense, Mimulus,
Nemophila, Phacelia, Polygonum bistortoides, Rhamnus, Ribes, Rubus, Salix, Seiiecio,
Sanicula 7ievade7isis, Sisyrinchium bellmn, Taraxacmn officinale. Predator: Xysticus.

Andre7ia caerulea Smith, 1879. Descr. New Species Hym. Brit. Mus., p. 55. ♀.

Andre7ia coerulea var. territa Cockerell, 1898. Entomologist 31: 89. ♀.

Pterandre7ia erigenoides Viereck. 1904. Canad. Ent. 36: 227. ♀.

Ptera7idre7ia crypta Viereck, 1904. Canad. Ent. 36: 227. ♀.

Pterandre7ia acrypta Viereck, 1904. Canad. Ent. 36: 227. ♀.

Ptera7idrena co7)iplexa Viereck, 1904. Canad. Ent. 36: 227. ♀.

Andrena (Ptilandre7ia) fra7icisca Viereck, 1917. Acad. Nat. Sci. Phila., Proc.'68: 595. ♀.

A7idre7ia supervirens Cockerell, 1924. Pan-Pacific Ent. 1: 64. ♀.

A7idre7ia supervire^is var. auresceiis Cockerell, 1924. Pan-Pacific Ent. 1: 64. ♀.

A7idrena (A7idre7ia) i7mo7ninata Viereck, 1926. Calif. Acad. Sci., Proc. (4) 15: 404. ♂.

A7idre7ia (Ptilandre7ia) tristis Linsley, 1951. In Muesebeck, Krombein and Townes, U. S.
Dept. Agr., Agr. Monog. 2: 1062. Nomen nudum.




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Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 154 (type). — Cockerell, 1906. Psyche
13: 9 (type). -Viereck, 1904. Canad. Ent. 36: 227. 9 (key, as territa). —Cockerell, 1932.
Canad. Ent. 64: 287. 9 (as complexa). — Michener, 1953. Kans. Univ. Sci. Bui. 35: 1034,
figs. 82, 83, 85 (larva, as complexa). — LaBerge and Ribble, 1975. Amer. Ent. Soc, Trans.
101: 406-415, figs. 6, 35-39 (redescription, synonymy).

Biology: Bohart, 1936. Pan-Pacific Ent. 12: 9 (parasite). — Bohart, 1941. Pan-Pacific Ent. 17:
95 (parasite). —Bohart, 1941. Calif. Univ. Pubs. Ent. 7: 91 (parasite). — MacSwain and
Bohart, 1947. Pan-Pacific Ent. 23: 30 (parasite). —MacSwain, 1949. Pan-Pacific Ent. 25:
89-90 (parasite). — Linsley and MacSwain, 1955. Wasmann Jour. Biol. 13: 253 (parasite).
— Linsley and MacSwain, 1957. Calif. Univ. Pubs. Ent. 11: 395 (parasite). —Linsley and
MacSwain, 1959. Calif. Univ. Pubs. Ent. 16: 5-13, text figs. 1-5, plate figs, lb, 3c and d
(nest construction, life history, overwintering, emergence, behavior, sex ratio, mating,
floral relationships, parasites).

Andrena chlorura Cockerell
Wash, to Calif., east to Utah and Airz.; Mexico (Baja California). Pollen:
Polylectic, visits a wide variety of flowers including Acer macrophyllum, Arbutus
menziesii, Arctostapliylos bicolor, A. ci^ustacea, A. drupacea, A. glandulosa, A. glauca,
A. mariposa, A. patula, Berberis pinnata, Ceanothus arboreus, C. cordulatus, C.
crassifolius, C. cuneatus, C. greggii, C. integerrimus, C. leucodermis, C. palmeri, C.
soredatus, C. velutinus, C. verrucosa, Cercocarpus betulifolius, Claytonia spathulata,
Cryptantha, Dentaria califoniica, Fragai'ia califomica, Gilia, Lasthenia chrysostoma,
Mains, Paeonia brownii, Petasites palmata, Prunus subcordata, Quercus, Ranunculus,
Rhamnus califomica, R. crocea, R. ilicifolia, Rhus iyitegrifolia, R. ovata, R. trilobata,
Ribes indecorum, Salix lasiolepis. Salvia mellifera, Tamarix.
Andrena chlorura Cockerell, 1916. Pomona Jour. Ent. Zool. 8: 48. ♀.
Andrena {Andre7ia) complicata Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 554. ♀.
Andrena ablusula Cockerell, 1936. Pan-Pacific Ent. 12: 135. ♀.
Andrena Clementina Timberlake, 1941. South. Calif. Acad. Sci., Bui. 39: 193. ♀.

Taxonomy: Timberlake, 1951. In Linsley, In Muesebeck, Krombein and Townes, U. S. Dept.
Agr., Agr. Monog. 2: 1061 (synonymy). —LaBerge and Ribble, 1975. Amer. Ent. Soc,
Trans. 101: 402-406, figs. 5, 30-34 (redescription, synonymy).

Andrena dissimulans Timberlake
Calif. Pollen: Evidently an oligolege oi Lasthenia including L.
aristata, L. chrysostoma, L. gracilis, L. tenella, but also visits the flowers of
Calandrinia menziesii, Salvia.
Andrena {Thysandrena) dissimulans Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 408. ♀.

6.
Andrejia (Thysandrena) blandula Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 410. ♀.

Taxonomy: LaBerge and Ribble, 1975. Amer. Ent. Soc, Trans. 101: 426-429, figs. 3, 55-59
(redescription, synonymy).

Andrena hamulata LaBerge and Ribble
Calif. Pollen: Unknown, but visits flowers of Salix.

Andrena (E uandrena) hamulata LaBerge and Ribble, 1975. Amer. Ent. Soc, Trans. 101:
399, figs. 3, 20-24. ♀, ♂.

Andrena lawrencei Viereck and Cockerell
B. C. to Calif., east to Alta., Idaho, Wyo., Utah and Nebr.
Pollen: Unknown, but visits flowers of Balsamorhiza sagittata, Carnaria draba,
Helianthus nuttallii. Phlox hoodii, Silphium, Wyethia mollis.
Andrena lawrencei Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 15. ♀.
Andreyia (lomelissa) extensa Viereck, 1924. Canad. Ent. 56: 240. ♀, ♂.

Taxonomy: Cockerell, 1932. Canad. Ent. 64: 286. 9 (key, as extensa). —LaBerge and Ribble,
1975. Amer. Ent. Soc, Trans. 101: 386-390, figs. 3, 18-19 (redescription, synonymy).

Andrena misella Timberlake
Calif. Pollen: Unknown, but visits flowers of Am.sinckia eastwoodae,
Anagallis arve7isis, Brassica, Calandrinia ciliata, Lasthenia, Montia perfoliata,
Phacelia, Ranunculus.
Andrena misella Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 410. ♀.

Andrena nigrihirta (Ashmead)
Alaska and transcont. south. Canada, south to Calif., Nev., Utah, Colo.,
N. Dak., Minn., Wis., 111., Mich., N. Y. and Conn. Pollen: Apparently polylectic, visits
flowers oi Angelica, Dentaria califomica, Eriogonum, Eriophyllum confertiflorum,




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Eryfhronium grandiflorum , Fragaria, Fritillaria puniila, Lonkera, Lotus, Montia
perfoliata, Peiasifes palmata, Pulsatilla hirsutissima, Ranunculus, Rhododendron
occidentale, Rosa, Rubus, Sviilacina stellata, Synthiridis plantagium.

Cillissa nigrihirta Ashmead, 1890. Colo. Biol. Assoc, Bui. 1: 6. ♂.

Andrena longihirtiscopa Viereck, 1904. Canad. Ent. 36: 191. ♀.

Andrena decussata Viereck, 1904. Canad. Ent. 36: 193, 194. ♀, ♂.

Andrena decnssatula Viereck, 1904. Canad. Ent. 36: 193. ♀.

Andrena synthiridis Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 436. ♀.

Andrena novae-angliae Viereck, 1907. Ent. News 18: 283. ♀.

Aiidrena nivaloides Graenicher, 1911. Pub. Mus. Milwaukee, Bui. 1: 235. ♀.

Andrena discolor Viereck, 1916. Amer. Mus. Nat. Hist., Bui. 35: 370. ♀.

Andrena (Andrena) tumida Viereck, 1922. Boston Soc. Nat. Hist., Occas. Papers 5: 35. ♀.
S.

Andrena {Andrena) vancouvereyisis Viereck, 1924. Canad. Ent. 56: 80. ♂.

Andrena (Andrena) nigrovaria Viereck, 1924. Canad. Ent. 56: 237. ♀.

Andrena (Andrena) marina Viereck, 1926. Calif. Acad. Sci., Proc. (4) 15: 405. ♂.

Andrena ripariella Cockerell, 1936. Pan-Pacific Ent. 12: 148. ♂.

A7idre7ia (Thysandrena) crenata Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 221,
figs. 44, 51. ♂.

Taxonomy: Atwood, 1934. Canad. Jour. Res. 10: 207. 9 (key, as novaeangliae). — Lanham,

1941. Ent. Soc. Amer., Ann. 34: 705. 9, 6 (key). — LaBerge and Kibble, 1975. Amer. Ent.

Soc, Trans. 101: 380-386, figs. 2, 8-12 (redescription, synonymy).

Andrena nigrocaerulea Cockerell
B. C. to Calif., east to Wyo. and Colo. Parasite: Stylops pacifica
Bohart. Pollen: Polylectic, visits a wide variety of flowers including Allium, Arbutus
menziesii, Arctostaphylos nevadensis, Arenaria kingii, Barbarea orthoceras, B. vulgaris,
Berberis aquilifolium, Brassica, Cakile edentula califomica, Caiyptridium umbellatum,
Ceanothus integerrinuis, Cercocarpus betuloides, C hamaebatia foliolosa, Cryptantha
intermedia, Daucus carota, Descourainia sophia, Doronicum, Eriodictyon califomicum,
Eriogonum lobbii, E. marifolium, Erodium botrys. Erysimum asperum, Fragaria
califomica, Gilia capitata, G. niulticaulis, G. tricolor, Hackelia patens, Hesperochiron
californicus, Horkelia tillingi, Lappula, Linanthus androsaceus, L. parviflorus,
Lithophragma affinis, Lomatium dissectum, Lonicera, Lotus, Mertensia, Microseris
nutans, Mimulus guttatus, Montia perfoliata, M. sibirica, Nemophila integrifolia, N.
spatulata, Oenothera, Penstemon cyananthus, Phacelia humilis, P. linearis, P. pusilla,
P. tanacetifolia, Plectritis macrocera, Potentilla glandulosa, Prtmus virginiana, P.
subcordata, Pyracantha, Quercus agrifolia. Ranunculus californicus, Raphanus sativa,
Rhamnus californicus, R. crocea, Rubus, Salix, Sanicula nevadensis, Senecio canus, S.
integerrimus, Sisyrinchium bellum, Spraguea umbellata, Taraxacum officinale, T.
vulgare, Trifolium repens.

Andrena nigrocaerulea Cockerell, 1897. Entomologist 30: 309. ♀, ♂.

A7idrena seattlensis Viereck, 1904. Canad. Ent. 36: 191, 195. ♀, ♂.

Andrena epileuca Cockerell, 1924. Pan-Pacific Ent. 1: 62. ♀, ♂.

Andrena lustraiis Cockerell, 1924. Pan-Pacific Ent. 1: 63. ♀.

Taxonomy: Cockerell, 1932. Canad. Ent. 64: 286 (key). —Cockerell, 1932. Canad. Ent. 64: 286.
9 (variation, as lustrans). —Cockerell, 1932. Canad. Ent. 64: 286. 9 (key, as epileuca).
—LaBerge and Kibble, 1975. Amer. Ent. Soc, Trans. 101: 390-396, figs. 4, 13-17
(redescription, synonymy).

Biology: Linsley and MacSwain, 1957. Calif. Univ. Pubs. Ent. 11: 399 (stylopization).
— Linsley and MacSwain, 1959. Calif. Univ. Pubs. Ent. 16: 19 (floral relationships,
stylopization).

Andrena penemisella
***authority mismatch
LaBerge and Kibble. Calif, and Oreg. (Cayuse and The Dalles). Pollen: Unknown,
but visits flowers of Calandrinia, Cryptantha, Erodium, Montia perfoliata.
Andrena (Euandrena) penemisella LaBerge and Kibble, 1975. Amer. Ent. Soc, Trans. 101:
422. ♀.

Andrena suavis Timberlake
Oreg. (Klamath Co.), Calif. Parasite: Nomada obliquella Timberlake, A^.
opacellus Timberlake, Stylops pacifica Bohart. Pollen: Oligolectic, apparently collects




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pollen only from the flowers of Ranunculus califominis, but visits other flowers
presumably for nectar including Astragalus, Calandrinia menziesii, Ceanothus,
Lastlienia chrysostoma, Lomatium utriculatum, Montia perfoliata, Oenothera ovata,
Orthocarpus densiflorus, Platystemon califomicus, Rhus trilobata, Wyethia
angustifolia. Predator: Xysticus sp.
Andrena (Ptilandrena) suavis Timberlake, 1938. Pan-Pacific Ent. 14: 24. ♀, ♂.

Biology: Bohart, 1941. Calif. Univ. Pubs. Ent. 7: 128 (parasite). — MacSwain, 1949.
Pan-Pacific Ent. 25: 89 (stylopization). — Linsley and MacSwain, 1955. Wasmann Jour. Biol.
13: 253 (parasite). —Linsley and MacSwain, 1957. Calif. Univ. Pubs. Ent. 11: 395-422
(parasite, stylopization). —Linsley and MacSwain, 1959. Calif. Univ. Pubs. Ent. 16: 13-16,
plate 3, figs, a and b (nest architecture, life history, overwintering, emergence, sex ratio,
floral relationships, parasites, predator).

Andrena subdepressa Timberlake
Calif. Pollen: Unknown, but visits flowers of Brassica, Dentaria
califoruica, Ranuyiculus califomicus, Salix lasiolepis.
Andrena {Jhysandrena) subdepressa Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 407. ♀.
6.

Biology: Linsley and MacSwain, 1959. Calif. Univ. Pubs. Ent. 16: 19 (floral relationships).

Andrena venata LaBerge and Ribble
Calif. (Sonora Pass). Pollen: Unknown, but visits flowers of
Ranunculus.
Andrena (Euandrena) venata LaBerge and Ribble, 1975. Amer. Ent. Soc, Trans. 101: 429,
figs. 60-64. ♀, ♂.

Genus ANDRENA Subgenus GEISSANDRENA LaBerge

Andrena subg. Geissandrena LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 302.
Type-species: Andreiia trevoris Cockerell. Monotypic and orig. desig.

Revision: LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 302-307, figs. 1, 36-40.

Andrena trevoris Cockerell
B. C. and Alta., south to Calif., south to Idaho, Utah and ?Ariz. (Huachuca
Mts. and Nogales). Pollen: Unknown, but visits flowers of Aesculus, Cirsium, Lonicera,
Rosa, Symphoricarpos.
Andrena trevoris Cockerell, 1897. Entomologist 30: 306. ♂.
Andrena semipolita Viereck, 1904. Canad. Ent. 36: 192. ♀.
Andrena ricardonis Cockerell, 1916. Canad. Ent. 48: 272. ♂.

Taxonomy: Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 380. ♀.

Genus ANDRENA Subgenus GONANDRENA Viereck

Andrena subg. Gonandrena Viereck, 1917. Amer. Ent. Soc, Trans. 43: 390.

Type-species: Andrena (Gonandrena) persiinulata Viereck. Monotypic.
Andrena subg. Tropandrena Viereck, 1924. Canad. Ent. 56; 21.

Type-species: Andrena fragilis Smith. Monotypic and orig. desig.

Revision: LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 271-302, figs. 1-4, 10-35 (N.
Amer. spp.).

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 210 (tax. characters). —LaBerge, 1964.
Nebr. Univ. State Mus., Bui. 4: 302 (tax. characters).

Andrena avulsa LaBerge and Ribble
Tex. Pollen: Unknown, but visits flowers of Parkinsonia,
Phacelia.
Andrena {Gonandrena) avulsa LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 298,
figs. 4, 26-30. ♀, ♂.

Andrena flocculosa LaBerge and Ribble
Wash., Calif. (Gold Lake and Meadow Valley), Nev. (Lamoille).
Andrena (Gonandrena) flocculosa LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98:
296, figs. 4, 24-25. ♀, ♂.

Andrena fragilis Smith
Que. to Ga., west to Wis., Iowa and Kans. Pollen: Apparently an oligolege of

Comus including C. amonum, C. paniculata, but visits other flowers including Aruncus,




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Castanea pumila, Ceanothus, Ciciita maculata, Cirsium, Hydrangea, Pastinaca sativa,

Ptelea, Salix, Svida, Trifolium, Viburnum dentatum, V. molle.
Andrena fragilis Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 115. ♂.
Andrena laticeps Provancher, 1888. Addit. Corr. Fauna Ent. Canada, Hym., p. 307. ♂.

Preocc.
Andrena provanckeri Dalla Torre, 1896. Cat. Hym. v. 10, p. 147. N. name.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 149 (type of fragilis). — Cockerell,
1906. Psyche 13: 7 (type of fragilis). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
231-232, figs. 46-49, table 6 (redescription). -LaBerge and Ribble, 1972. Amer. Ent. Soc,
Trans. 98: 278-284, figs. 2, 10-14 (redescription, synonymy).
Integra Smith. Que. and Ont, south to N. C, west to Minn, and Kans. Pollen: Possibly an

oligolege of Cortius including C. stolonifera, but visits other flowers including Achillea,
Aninais, Barbarea vulgare, Crataegus, Cryptotaenia, Melilotus alba, M. officinalis,
Pastinaca, Ptelea, Rhus, Viburnum, Vitis.
Andrena integra Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 114. ♀.
Andrena lineata Provancher, 1888. Addit. Corr. Faune Ent. Canada, Hym., p. 309. ♀.
Andre7ia (Gonandrena) lucifera Cockerell, 1932. Canad. Ent. 64: 155. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 124 (type of integra). —Cockerell,

1906. Psyche 13: 6 (type of integra). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:

232-233, figs. 46-49, table 6 (redescription). —LaBerge and Ribble, 1972. Amer. Ent. Soc,

Trans. 98: 284-287, figs. 2, 15-19 (redescription, synonymy).

Andrena nigrifrons (Cresson)
Ont. and N. H. south to N. C, west to Man., Minn., Nebr., Kans. and Tex.
Pollen: Possibly polylectic, although seems to prefer pollen of Comus including C.
asperifolia, C. paniculata, C. racemosa, C. sericea, C. stolonifera, but visits other
flowers including Achillea, Barbarea vulgaris, Ceanothus americanus, Cryptotaenia
canadensis, Erigeron philadelphus, Hieraceum, Lotus comiculatus, Melilotits alba, M.
officinalis, Ptelea trifoliata, Rhus glabra, Rubus, Sambucus americanus. Spiraea
aruncus, S. vanhouttei, Taraxacum, Toxicodendron, Viburnum dentatum, Zizia aurea.

Panurgus nigrifrons Cresson, 1878. Amer. Ent. Soc, Trans. 7: 62. ♂.

Andrena platyparia Robertson, 1895. Amer. Ent. Soc, Trans. 22: 119. ♀, ♂.

Andrena (Andrena) barbarica Viereck, 1917. Amer. Ent. Soc, Trans. 43: 369. ♀.

Andrena (Gonandrena) monroensis Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
233. ♀.

Taxonomy: LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 287-292, figs. 3, 20-21
(redescription, synonymy).

Andrena persimulata Viereck
N. S. to N. W. T. and Alta., south to Conn., N. Y., Mich., Wis., Minn.,

Nebr., Colo, and Utah. Pollen: Possibly an oligolege of Comus including C. asperifolia,
C. stolonifera, but visits other flowers including Angelica atropurpurea, Crataegus,
Deutzia gracilis, Fragaria, Lomatium dissectum. Polygonum convolvulus, Pnimis,
Spiraea, Taraxacum officinalis. Viburnum.
Andrena (Gonandrena) persimulata Viereck, 1917. Amer. Ent. Soc, Trans. 43: 390. ♀.

Taxonomy: Atwood, 1934. Canad. Jour. Res. 10: 206, 209. 9, 6 (key). —Mitchell, 1960. N. C.
Agr. Expt. Sta. Tech. Bui. 141: 235-236, figs. 45-46, 48, table 6 (redescription). —LaBerge
and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 292-296, figs. 4, 22-23 (redescription).

Genus ANDRENA Subgenus HESPERANDRENA Timberlake

Andrena subg. H esperandrena Timberlake, 1949. hi Lanham, Calif. Univ. Pubs. Ent. 8:
208.

Type-species: Andrena escondida Cockerell. Orig. desig.

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 283, 293, 301 (tax. characters).

Andrena baeriae Timberlake
Calif. Pollen: Apparently an oligolege of the flowers of Lasthenia

including L. aristata, L. chrysostoma, L. gracilis, but visits other flowers including
Eriophyllum co7ifertiflorum, Layia platyglossa, Lessingia germanorum., Senecio
calif omica.
Andrena baeriae Timberlake, 1941. South. Calif. Acad. Sci., Bui. 39: 194. ♀, ♂.




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Andrena duboisi Timberlake
North. Calif. Pollen: Unknown, but visits flowers oi Lasthenia
chrysostoma, Layia platyglossa.
Andrejia (Hesperandrena) duboisi Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 390. ♀, ♂.

Andrena escondida Cockerell
South. Calif.; Mexico (Baja California). Pollen: Unknown, but visits
flowers of Encelia, Layia platyglossa.
Andrena escondida Cockerell, 1938. Ann. and Mag. Nat. Hist. (11) 2: 146. ♂.

Andrena lativentris Timberlake
Calif. Pollen: Unknown, but visits flowers of Amsinckia intermedia.

Coreopsis maritima, Hemizonia pungens, Lasthenia chrysostoma, L. gracilis, L. tenella,
Layia platyglossa.
Andrena (Hesper andrena) lativentris Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 388. ♀.
6.

Andrena limnanthis Timberlake
Calif. Pollen: Possibly an oligolege of Limnanthes including L.

douglasii, but visits other flowers including Chamaehatia foliolosa, Layia platyglossa,
Raphanus sativus.
Andrena (H esperandrena) limnanthis Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 387.
9,

Genus ANDRENA Subgenus lOMELISSA Robertson

lomelissa Robertson, 1900. Acad. Sci. St. Louis, Trans. 10: 50.
Type-species: Andrena violae Robertson. Monotypic.

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 207 (tax. characters). — LaBerge, 1964.
Nebr. Univ. State Mus., Bui. 4: 284, 288, 300 (tax. characters).

Andrena violae Robertson
N. Y. to N. C, west to 111. and Colo. Pollen: Obtains pollen solely from the
flowers of Viola, but visits other flowers for nectar including Cardamine, Ellisia,
Oxalis, Potentilla.
Andrena violae Robertson, 1891. Amer. Ent. Soc, Trans. 18: 53. ♀, ♂.
Andrena davisi Viereck, 1907. Ent. News 18: 283. ♀.

Taxonomy: Cockerell, 1914. Ann. and Mag. Nat. Hist. (8) 14: 10. 9 (key). —Lanham, 1941.
Ent. Soc. Amer., Ann. 34: 706, 707. 9, 6 (key). —Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 252-254, figs. 54-56 (redescription, synonymy).

Genus ANDRENA Subgenus LARANDRENA LaBerge

Andrena subg. Larandrena LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 304.
Type-species: A^idrena miserabilis Cresson. Orig. desig.

Revision: Ribble, 1967. Nebr. Univ. State Mus., Bui. 6: 27-42, figs. 1-5, 1 table, 1 map.

Taxonomy: LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 308 (tax. status).

Andrena miserabilis Cresson
Transcont. south. Canada and U. S., south to Fla., Ala., Miss., Tex., N.
Mex., Utah and Calif.; Mexico (Tamaulipas). Parasite: Leucophora obtusa (Zett.),
Nomada sp., Stylops bipunctata Pierce, S. oklahomae Pierce. Pollen: Polylectic,
principally obtains pollen from the flowers of Rosaceae and Salicaceae; visitation
records include Amelanchier, Antennaria, Arabis, Aronia, Berberis, Brassica, Capsella,
Cardamine, Ceanothus, Cercis, Cercocarpus, Claytonia, Comandra, Convolvulus,
Comus, Crataegus, Cydonia, Dentaria, Erigenia, Exochorda, Fragaria, Hepatica,
Heracleum, Ilex, Isopyrum, Mains, Pri(7ius, Ptelea, Pyracayitha, Pyrus, Ranmiculus,
Rhamnus, Rhus, Rubus, Salix, Solidago, Sorbaria, Spiraea, Stachys, Staphylea,
Stellaria, Taraxacum, Viburnum, Viola, Zanthoxylum.

Andrena clypeata Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 115. ♀. Preocc.

Andrena miserabilis Cresson, 1872. Amer. Ent. Soc, Trans. 4: 259. 9 (cJ misdet.).

Andrena bipunctata Cresson, 1872. Amer. Ent. Soc, Trans. 4: 259. ♂.

Andrena flavoclypeata Smith, 1879. Descr. New Species Hym. Brit. Mus., p. 54. ♂.

Andrena scutellaris Robertson, 1893. Amer. Ent. Soc, Trans. 10: 148. ♀. Preocc.

Anthrena clypeolata Dalla Torre, 1896. Cat. Hym., v. 10, p. 133. N. name.

Anthrena scutellata Dalla Torre, 1896. Cat. Hym., v. 10, p. 151. N. name.

Andrena pennsylvanicola Viereck, 1907. Ent. News 18: 284. ♀.

Andrena pronitens Cockerell, 1930. Ann. and Mag. Nat. Hist. (10) 5: 114. ♀.




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Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 124, 152 (types oi clypeata,
flavoclypeata). —Robertson, 1902. Amer. Ent. Soc, Trans. 28: 193. 9, 6 (key).
— Cockerell, 1906. Psyche 13: 9, 36, (types of clypeata, flavoclypeata). — Atwood, 1934.
Canad. Jour. Res. 10 (2): 208, 209, fig. 9, 6 (key). — Lanham and Timberlake, 1951. In
Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1058 (synonymy).
—Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 95, 160, fig. 29, table 4
(redescription). — Ribble, 1967. Nebr. Univ. State Mus., Bui. 6: 27-42, 5 figs., 1 table, 1 map
(redescription, synonymy).

Biology: Michener and Rettenmeyer, 1956. Kans. Univ. Sci. Bui. 37: 679-681, fig. 21 (nest
architecture, life history, parasites, as bipunctata). —Ribble, 1967. Nebr. Univ. State
Mus., Bui. 6: 37, table 1 (floral relationships).

Genus ANDRENA Subgenus LEUCANDRENA Hedickc

Andrena subg. Leucandrena Hedicke, 1933. Berlin Zool. Mus., Mitt. 19: 215.
Type-species: Apis sericea Christ. Orig. desig.

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 220 (tax. characters). — LaBerge, 1964.
Nebr. Univ. State Mus., Bui. 4: 285, 290, 310 (tax. characters).

Andrena bifurcata Mitchell
Mich., Ohio.

Andrena (^Leucandrena) bifurcata Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 211,
figs. 40-42. ♂.

Andrena chippewaensis Mitchell
Mich., N. B.

A^idrena (^^Leucandrena) chippewaensis Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui.
141: 212, fig. 43. ♀.

Taxonomy: Knerer and Atwood, 1963. Canad. Ent. 95: 584, figs. 2, 4. ♂.

Andrena electrica Casad and Cockerell
N. Mex.

Andrena electrica Casad and Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 89. ♂, ♀.

Taxonomy: Cockerell, 1898. Denison Univ Sci. Labs., Bui. 11: 49. 9, 6 (key).

Andrena erythronii Robertson
Que. to Mich., 111. and Kans., south to Maine and Mass. Pollen: Primarily
collects pollen oi Erythronium including E. mesochoreum, but also collects pollen from
the flowers of Mains, Prunus, Quercus, Taraxacum; visits other flowers presumably
for nectar including Amelancliier, Claytonia, Erigenia, Hepatica, Salix.
Andrena erythronii Robertson, 1891. Amer. Ent. Soc, Trans. 18: 53. ♀, ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 190, 193. 9, S (key). —Michener,
1953. Kans. Univ. Sci. Bui. 35: 1035, figs. 88-91 (larva). -Mitchell, 1960. N. C. Agr. Expt.
Sta. Tech. Bui. 141: 212-213, figs. 40-42 (redescription). —Yager and Rozen, 1966. Amer.
Mus. Novitates 2265: 5 (pupa).

Biology: Michener and Rettenmeyer, 1956. Kans. Univ. Sci. Bui. 37: 645-679, figs. 1-20 (nest
architecture, life history, floral relationships, mating, number of females per nest,
immature stages).
lupin! Cockerell. Calif. Pollen: Unknown, but visits flowers of Arctostaphylos glandulosa, A.
glauca, Ceanothus cuneatus, C. greggii, C. integerrimus, C. verrucosus. Clematis
lasiantha, Erodium botrys, Eschscholzia califomica, Lasthenia chrysostoma, Lathyms
alfeldii, Lupinus, Phacelia.
Ayidrena lupini Cockerell, 1936. Pan-Pacific Ent. 12: 142. ♀.

Andrena mariposorum Viereck
Calif.

Andrena (Andrena) mariposorum Viereck, 1917. Amer. Ent. Soc, Trans. 43: 382. ♀.

Andrena parnassiae Cockerell
Mich., Wis., Vt. Pollen: Unknown, but visits flowers of Pamassia.
Andrena paniassiae Cockerell, 1902. Ann. and Mag. Nat. Hist. (7) 9: 105. ♀.

Taxonomy: Graenicher, 1904. Ent. News 15: 66. 9,6. —Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 213-214, figs. 40-43 (redescription).

Andrena perezana Viereck and Cockerell
Nebr.

Andrena perezana Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 47. ♀.




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Andrena picta Mitchell
N. Y. (Ithaca).

Andreiia (Leucandrena) picta Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 214. ♀.

Andrena placida
***authority mismatch
placida Smith. Alaska to N. S., south to Va., Ohio, 111., Mich., Minn, and Calif. Parasite:
Meloe opacus LeC, M. strigulosus Mann., Stylops sp.?. Pollen: Polylectic, analyzed pollen
stores reveal that a given female, with one exception, provisions her cells with only one
kind of pollen which in California consisted of predominantly pollen of Eschscholzia
caiifomica, while others stored pollen of a ligulate Compositae (Agoseris^), legume
(Lupinus^), and a crucifer (Raphaiius^); visitation records for other flowers include
Acer, Amelanchier, Fragaria, Layia camosa, Prunus, Pyrus malus, Taraxacum,
Viburnum.

Andrena placida Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 112. ♀.

Andrena macilenta Provancher, 1888. Addit. Corr. Faune Ent. Canada, Hym., p. 313. ♀, ♂.

Andrena macgillivrayi Cockerell, 1897. Entomologist 30: 308. ♀.

Andrena salicacea Robertson, 1900. Acad. Sci. St. Louis, Trans. 10: 48. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 151 (type of placida). —Cockerell,
1906. Psyche 13: 7 (type of placida). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
214-216, figs. 40-42, table 6 (redescription). —Thorp and Stage, 1968. Ent. Soc. Amer., Ann.
61: 1583-1584, figs. 3-9. (immature stages, chromosome number).

Biology: Clements and Long, 1923. Carnegie Inst. Wash., Pub. 336: 249 (ecology). —Thorp
and Stage, 1968. Ent. Soc. Amer., Ann. 61: 1580-1586, 9 figs., table (nest architecture, life
history, floral realtionships, parasites).

Andrena placida
***authority mismatch
sapellonis Cockerell. Colo., N. Mex.

Andreyia sapellonis Cockerell, 1900. Ann. and Mag. Nat. Hist. (7) 5: 19. ♀.

Andrena recta Mitchell
Minn, to N. Y. south to N. C. Pollen: Unknown, but visits flowers oi Helianthus
anniius. Predator: Philanthus sp.
Andrena (Leucandrena) recta Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 217, fig.
43. ♀.

Andrena seavillensis Mitchell
N. J. (South Seaville).

Andrena {? Leucandrena) seavillensis Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
217, fig. 43. ♀.

Andrena trapezoidina Viereck and Cockerell
Nebr.

Andrena trapezoidina Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 58. ♀.

Genus ANDRENA Subgenus MELANDRENA Perez

Melandrena Perez, 1890. Soc. Linn. Bordeaux, Actes 44: 170.

Type-species: Andrena niorio Brulle. Desig. by Hedicke, 1933.
Andrena subg. Gymnandrena Hedicke, 1933. Berlin Zool. Mus., Mitt. 19: 213.

Type-species: Apis thoracica Fabricius. Orig. desig.
Andrena subg. Cryptandrena Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 222. Preocc.

Type-species: Andrena carlini Cockerell. Orig. desig.
Andrena subg. Bythandrena Lanham, 1950. Ent. News 61: 140. N. name.

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 310, 311 (tax. characters, as
Gymnandrena and Melandrena. —LaBerge, 1971. Pan-Pacific Ent. 47: 47 (synonymy).

Andrena bisignata Mitchell
Mass. to N. C, west to Minn. Pollen: Unknown, but visits flowers of
Erythronium americanum. Ilex verticillata, Prunus serotina, Pyrus malus, Salix
sericea. Viburnum acerifolium.
Andrena (Gymnandrena) bisignata Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
122, figs. 20, 22, 23. ♀, ♂.

Andrena bryanti Cockerell
Alta.

Andrena bryanti Cockerell, 1938. Canad. Ent. 70: 5. ♀.

Andrena carlini
***authority mismatch
carlini Cockerell. N. S. to Ga., west to Minn., Colo., Wyo. and Oreg.?, B. C? Pollen:
Apparently polylectic, visits flowers oi Amelanchier, Anemonella, Arabis, Brassica,
Cercis, Chaerophyllum, Claytonia, Cornus, Dentaria, Dicentra, Erigenia, Erythronium,
Hepatica, Heracleum, Hydrophyllum, Isopyrum, Malus, Polemonium, Prunus, Pyrus,




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Rhus, Ribes, Rubus, Salix, Sanguinaria, Sassafras, Smilacina, Solidago, Thaspium,
Tragana, Trifolium, Uvularia, Vacchiiuni, Viburnum, Vicia.
Andrena carlini Cockerell, 1901. Canad. Ent. 23: 150, 153. ♀.

Taxonomy: Atwood, 1934. Canad. Jour. Res. 10: 207, 209. 9, 6 (key). — Lanham, 1941. Ent.
Soc. Amer., Ann. 34: 705, 708. 9, S (key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui.
141: 111-112, figs. 17, 19, table 3 (redescription).

Biology: Atwood, 1933. Canad. Jour. Res. 9: 456 (floral relationships). — Brittain, 1933. Canad.
Dept. Agr., Bui. 162: 94, figs, (floral relationships).

Andrena carlini
***authority mismatch
neorhodura Mitchell. Mass., Pa., Mich.

Andrena (Bythandrena) carlini neorhodura Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141: 113. ♂.

Andrena carliniformis Viereck and Cockerell
B. C. to Calif., east to Wyo., Colo, and Utah. Pollen:

Unknown, but visits flowers ot Agastache urticifolia. Allium, Calochortus, Ceanothus,
C hamaebatia foliolosa, Eriogonum, Euphorbia crenulata, Gilia, Horkelia, Lappula,
Lotus nevadensis, Lupinus bicolor, Phacelia Plagiobothrys nothofulvus, Potentilla,
Prunus demissa. Ranunculus califomicus, Rhanuius crocea, R. califomica, Salix,
Salvia columbariae. Taraxacum officinale, Vicia, Wyethia.

Andrena carliniformis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 25. ♀.

Andrena carolinensis Mitchell
N. C. (Raleigh). Pollen: Unknown, but visits flowers of Vicia
caroliniana.

Andrena {Gymnandrena) carolinensis Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
123, fig. 20. ♀.

Andrena cerasifolii Cockerell
Calif., Ariz., Utah, N. Mex. Pollen: Polylectic, visits flowers of

Arctostaphylos drupacea, A. glauca, A. mariposa, A. patula, Brassica arve^isis, B.
campestris, Ceanothus cuneatus, C. greggii, C. pahneri, Cirsium califomicum, Clarkia
biloba, C. nitens, C. unguiculata, Cryptantha intermedia, Emmenanthe penduliflora,
Eriogonum fasciculatum, Erodium, Erysimum aspenim, Heteromeles arbutifolia,
Horkelia bolanderi, Hyptis emoryi, Peyistemon spectabilis, Phacelia brachyloba, P.
cicutaria, Rhamnus califomicus, R. crocea, Rhus trilobata, Ribes indecorum, Salix
hindsiana, S. lasiolepis. Sisymbrium irio, Swertia parryi, Trichostema parryi,
Vibumtim.

Andreiia cerasifolii Cockerell, 1896. Ent. Monthly Mag. (2) 7: 220. ♂.

Andrena mimetica Cockerell, 1903. Ann. and Mag. Nat. Hist. (7) 12: 412. ♀, ♂.

Andrena mimetica war.falli Cockerell, 1907. Ann. and Mag. Nat. Hist. (7) 19: 536.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 48. S (key). —Cockerell, 1924.
Pan-Pacific Ent. 1: 58. 6. — Timberlake, 1951. hi Muesebeck, Krombein and Townes, U. S.
Dept. Agr., Agr. Monog. 2: 1060 (synonymy).

Biology: Linsley, 1937. Brooklyn Ent. Soc, Bui. 32: 125 (floral relationships, number of
generations). — MacSwain, Raven, Thorp, 1973. Calif. Univ. Pubs. Ent. 70: 46 (floral
relationships).

Andrena commoda Smith
Southeast. Canad, south to Ga., west to N. Dak. Pollen: Unknown, but visits
flowers of Comus paniculata.
Andrena commoda Smith, 1879. Descr. New Species Hym. Brit. Mus., p. 53. ♀.
Andrena comi Robertson, 1900. Acad. Sci. St. Louis, Trans. 10: 50. ♀.
Aiidrena pyrura Cockerell, 1906. Ann. Mag. and Nat. Hist. (7) 17: 309. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 152 (type). — Coclterell, 1906. Psyche
13: 6. 9 (type). —Viereck, 1907. Ent. News 18: 283, 286. 9, 6 (key). —Cockerell, 1931.
Amer. Mus. Novitates 458: 18, 19. —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
124, figs. 22-23, table 3 (redescription, synonymy).

Andrena confederata Viereck
Ohio to N. J., south to Fla. Pollen: Unknown, but visits flowers of

Amelanchier, Castanea, Crataegus, Fragaria, Ilex, Mains, Padus, Prunus, Pyracantha.
Rubus.
Andrena (A^idreiia) confederata Viereck, 1917. Amer. Ent. Soc, Trans. 43: 375. ♀, ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 125, figs. 22-23, table 3
(redescription).




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Andrena critica Mitchell
N. Y. (Catskill Mts.).

Andrena (Bythandrena) critica Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 113. ♀.

Andrena dunningi Cockerell
Que. to N. C, west to Minn. Pollen: Unknown, but visits flowers of

Claytonia, Gleditsia, Malus, Narcissus, Ribes, Rubiis, Salix, Taraxacum, Viburnum.

Andrena dunningi Cockerell, 1898. Canad. Ent. 30: 103. ♀.

Andreyia vicinifonnis Robertson, 1900. Acad. Sci. St. Louis, Trans. 10: 47. ♀, ♂.

Andrena annae Cockerell, 1931. Canad. Ent. 63: 200. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 190, 192. 9, 6 (key). — Timberlake,
1951. In Linsley, In Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2:
1064 (synonymy). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 113-114, figs.
18-19, table 3 (redescription).

Andrena errans Smith
B. C. to Calif, and N. Mex. Pollen: Unknown, but visits flowers of

Arctosiaphylos mariposa, Ceanothus, Eriogonum marifolium, Lappula, Ranunculus,

Salix.
Andrena ei'rans Smith, 1879. Descr. New Species Hym. Brit. Mus., p. 55. ♀, ♂.
Andrena pluvialis Cockerell, 1901. Canad. Ent. 33: 154. ♀.
Andreiia junonia Viereck, 1904. Canad. Ent. 36: 191. ♀.
Ajidrena compactiscopa Viereck, 1904. Canad. Ent. 36: 191. ♀.
Andrena argentiniae var. trickomelaena Cockerell, 1913. Ann. and Mag. Nat. Hist. (8) 12:

376. ♂, ♀.
A7idre7ia spaldingi Cockerell, 1934. Amer. Mus. Novitates 697: 3. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 153 (type). —Cockerell, 1903. Ann.
and Mag. Nat. Hist. (7) 12: 443. 6 (as pluvialis). —Cockerell, 1906. Psyche 13: 34 (type).
—Timberlake, 1951. In Linsley, In Muesebeck, Krombein and Townes, U. S. Dept. Agr.,
Agr. Monog. 2: 1065 (synonymy).

Andrena flexa Malloch
111. to Tex. Parasite: Nomada sp.

Andrena fie xa Malloch, 1917. Brooklyn Ent. Soc, Bui. 12: 92. ♀.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 126 (redescription).

Biology: Rozen, 1966. Amer. Mus. Novitates 2244: 26 (parasite).

Andrena gabrielsoni Mitchell
Maine, Conn., N. C, Mich.

Andrena (Bythandrena) gabrielsoni Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
115. ♀.

Andrena hilaris Smith
Mich, and Ohio to eastern Canada and New England States, south to Ga. and

Ala. Pollen: Unknown, but visits flowers oi Aronia, Crataegus, Hydrangea, Ilex, Malus,
Prunus, Pyrus, Rubus, Salix, Vaccinium.
A7idre7ia hilaris Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 112. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 151 (type). —Mitchell, 1960. N. C.
Agr. Expt. Sta. Tech. Bui. 141: 126-127, figs. 21-23, table 3 (redescription).

Andrena lupinorum
***authority mismatch
helenae Cockerell. Colo.

Andrena lupinorum helenae Cockerell, 1934. Amer. Mus. Novitates 697: 2. ♀.

Andrena lupinorum
***authority mismatch
lupinorum Cockerell. Wyo., Colo., Nebr. Pollen: Unknown, but visits flowers of
Lupinus.
Andre7ia lupinorum Cockerell, 1906. Ann. and Mag. Nat. Hist. (7) 17: 308. ♀.
Andrena vicina argeiitiniae Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 432. ♀.

Taxonomy: Lanham, 1941. Ent. Soc. Amer., Ann. 34: 705. 9 (key). —Timberlake, 1951. In

Linsley, In Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1070

(synonymy).

Andrena macra Mitchell
N. C. Pollen: Unknown, but visits flowers of Crataegus, Pyracantha, Rubus.

Andrena 7nacra Mitchell, 1951. Elisha Mitchell Sci. Soc, Jour. 67: 246, figs. 1-3. ♀, ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 127-128, figs. 22-23, table 3
(redescription).

Biology: Sivik, 1954. Ent. News 65: 253-255 (nest, supersedure).




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Andrena nigra Provancher
Ariz., south. Calif.; Mexico (Baja California). Pollen: Apparently .an oligolege
of Pkacelia including P. distans, but also visits flowers of Coreopsis maritima,
Cryptantha intermedia, Layia platyglossa, Malacotkrix, Mains, Salix.

Andrena nigra Provancher, 1896. Nat. Canad. 23: 173. ♀.

Andrena griseonigra Cockerell, 1905. Canad. Ent. 37: 371. ♂.

Andrena subtristis Cockerell, 1905. Canad. Ent. 37: 372. ♀.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 266, 279. 9, 6 (key).

Biology: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 280 (floral relationships).

Andrena nigripes Provancher
Calif.

Andrena nigripes Provancher, 1895. Nat. Canad. 22: 173. ♀.

Andrena nivalis Smith
Que. to Ga., west to B. C, south to Calif, and N. Mex. Pollen: Unknown, but
visits flowers of Hydrangea, Rhododeyidron, Rubus, Vagnera.

Andrena nivalis Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 118. ♀, ♂.

Andrena convexa Provancher, 1888. Addit. Corr. Faune Ent. Canada, Hym., p. 311. ♀, ♂.

Andrena semiriifa Cockerell, 1900. Ann. and Mag. Nat. Hist. (7) 5: 407. ♀.

Andrena solidula Viereck, 1904. Canad. Ent. 36: 191, 194. ♀, ♂.

Andrena idahonim Viereck, 1916. Amer. Mus. Nat. Hist., Bui. 35: 732. ♀.

Taxonomy: Morice and Cockerel], 1901. Canad. Ent. 33: 151. 9 (type). —Cockerell, 1906.
Psyche 13: 9 (type). —Cockerell, 1931. Amer. Mus. Novitates 458: 14. 9 (key, as seminifa).
— Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 128-129, figs. 22, 23, table 3
(redescription, synonymy).

Andrena obscuripennis Smith
N. J.(?), N. C, Ga., La.(?). Parasite: Nomada cnidelis Cress.? Pollen:
Unknown, but visits flowers of Batodendron, Crataegus, Ilex, Padus.
Andrena obsmripennis Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 188. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 150. 9 (type). —Cockerell, 1906.

Psyche 13: 36. 9 (type). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 129-131,

figs. 21, 23, table 3 (redescription).

Andrena perimelas Cockerell
Coastal Calif, and Catalina Island Parasite: Nomada edwardsii edwardsii
Cress., Stylops pacifica Bohart, S. vandykei Bohart. Pollen: Apparently polylectic, visits
flowers of Brassica, Coreopsis maritima, Crepis, Limnanthes douglasii,
Mesembryanthevium edule, Prunus, Ranunctilus califomims, Raphanus sativus, Ribes
cereum, Rubus, Salix, Sisymbrium bellum, Zygadenus fremontii.

Andrena perimelas Cockerell, 1905. Canad. Ent. 37: 371. ♀.

Andrena meadowsi Cockerell, 1938. Ann. and Mag. Nat. Hist. (11) 2: 146. ♂.

Biology: Linsley and MacSwain, 1955. Wasmann Jour. Biol. 13: 254-256, 270, pi. 1, fig. 5 (nest,
floral relationships, parasite). —Linsley and MacSwain, 1959. Calif. Univ. Pubs. Ent. 16: 19,
pi. 5 (nest, floral relationships, parasite, stylopization).

Andrena pertristis Cockerell
Calif. Pollen: Unknown, but visits flowers of Brassica, Cryptantha

intermedia, Eriodictyon trichocalyx, Fremontodendron califomica, Lupinus formosus,
Malus, Nemophila integrifolia, Phacelia heteropkylla, Stanleya pinnata.
Andrena pertristis Cockerell, 1905. Canad. Ent. 37: 372. ♀.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 265, 267. 9, 6 (key).

Andrena platyrhina Cockerell
Colo.

Andrena (Pterandrena) platyrhina Cockerell, 1930. N. Y. Ent. Soc, Jour. 37: 446. ♀.

Andrena pruni Robertson
Minn, and 111. to Mass., south to Ga. Pollen: Unknown, but visits flowers of
Anincus, Claytonia, Crataegus, Dentaria, Heracleum, Pastinaca, Polemonium, Prumis,
Ribes, Rubus, Salix, Staphylea, Uvularia, Viburnum, Zanthoxylum.
Andrena pruni Robertson, 1891. Amer. Ent. Soc, Trans. 18: 51. 9, tj.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 191, 192. 9, 6 (key). —Mitchell,
1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 117-118, figs. 19-20, table 3 (redescription).

Andrena pulverulenta Viereck
Oreg.

Andrena pulverulenta Viereck, 1904. Canad. Ent. 36: 190, 195. ♀, ♂.




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Andrena regularis Malloch
B. C. to N. S., in eastern U. S. from Minn, to New England States, south to
Ga. Pollen: Unknown, but visits flowers of Aster, Brassica, Pyrus, Rubus, Syringa,
Vaccinium.
Andrena regularis Malloch, 1917. Brooklyn Ent. Soc, Bui. 12: 91. ♂, ♀.

Taxonomy: Atwood, 1934. Canad. Jour. Res. 10: 207, 209. 9, S (key). —Mitchell, 1960. N. C.
Agr. Expt. Sta. Tech. Bui. 141: 118-119, figs. 18-19, table 3 (redescription).

Andrena sayi Robertson
N. B. to N. C, west to Minn., 111. and Kansas. Pollen: Unknown, but visits
flowers o{ Amelanchier, Aronia, Brassica, Cercis, Crataegus, Malus, Padus, Prunus,
Pyrus, Rubus, Salix, Stellaria, Taraxacum, Vaccinium.
Andrena sayi Robertson, 1891. Amer. Ent. Soc, Trans. 18: 52. ♀, ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 191, 192. 9, 6 (key). —Mitchell,

1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 131-132, fig. 23, table 3 (redescription).

Andrena sevierensis Mitchell
Tenn. (Sevier Co.).

Andrena {^Gymnandrena) seviensis(\) Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
132, fig. 20. ♀.

Andrena {^Gymnandrena) sevierensis Krombein, 1967. In Krombein and Burks, U. S.
Dept. Agr., Agr. Monog. 2, Second Suppl. p. 434. Emend.

Andrena sola Viereck
B. C. to Calif., Ariz.; Mexico (Baja California). Pollen: Unknown, but visits
flowers of Aynsinckia intermedia, Arctostaphylos, Baccharis, Brassica, Ceanothus
crassifolius, Chaenactis glabriuscula. Clematis lasiantha, Cryptantha intermedia,
E7icelia farinosa, Eriodictyon, Gallium nuttallii, Gilia, Layia platyglossa, Melilotus,
Papaver califomica, Plagiobothrys califomicus, Platystemon califomica, Prunus
subcordat2is, Quercus, Ranunculus, Rhamnus califomica, R. crocea, Rhus diversiloba,
R. trilobata, Senecio califomicus, Solanum um.belliferum, Tam.arix.

Andreyia (Andrena) sola Viereck, 1916. Acad. Nat. Sci. Phila., Proc 68: 577. ♀.

Andrena carissima Cockerell, 1924. Pan-Pacific Ent. 1: 58. ♀.

Andreyia microdonta Cockerell, 1924. Pan-Pacific Ent. 1: 61. ♂.

Andrena ensenadensis Cockerell, 1941. San Diego Soc Nat. Hist., Trans. 9: 345. ♀.

Taxonomy: Timberlake, 1951. In Linsley, hi Muesebeck, Krombein and Townes, U. S. Dept.
Agr., Agr. Monog. 2: 1081 (synonymy).

Andrena transnigra Viereck
B. C. to Calif., Colo, and Wyo. Pollen: Unknown, but visits flowers of
Allium, Brassica, Ceanothus integerrimus, C. parviflorus, Chamaebatia foliolosa,
Lappula, Madia ramii. Ranunculus, Ribes, Salix, Taraxacum, Vaccinium, Wyethia.
A7tdre7ia trayisnigra Viereck, 1904. Canad. Ent. 36: 191. ♀.
Andrena cya^iura Cockerell, 1916. Canad. Ent. 48: 252. ♀.
Andrena transnigra paysoni Cockerell, 1924. Ent. News 35: 349. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 18. ♀.

Andrena vicina Smith
N. S. to Ga., west to B. C, Colo, and Oreg. Parasite: Nomada imbricata Sm., N.
obliterata Cress., N. vicina vicina Sm. Pollen: Unknown, but visits flowers of
Amelanchier, Azalea, Caragana, Castanea, Ilex, Malus, Narcissus, Padus, Prunus,
Pyrus, Rhododendron, Rhodura, Rubus, Salix, Scilla, Spiraea, Taraxacum, Trifolium,
Tulipa, Vaccinium, Viburnum.
Andre7ia vicina Smith, 1853. Cat. Hym. Brit. Mus., v. 1 p. 112. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 151 (type). —Cockerell, 1906. Psyche
13: 9. —Atwood, 1934. Canad. Jour. Res. 10: 207, 210, figs. 9, S (key). — Lanham, 1941.
Ent. Soc. Amer., Ann. 34: 705, 708. 9, 6 (key). —Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 132-134, fig. 23, table 3 (redescription).

Biology: Packard, 1869. Guide Study Ins., p. 144 (nesting habits, parasites). —Clements and
Long, 1923. Carnegie Inst. Wash., Pub. 336: 249 (ecology). —Salt, 1927. Jour. Expt. Zool.
48: 251 (stylopization).

Andrena victima Smith
N. S. to Ont. and Conn. Possibly a syn. of A. vicina Smith.
Andreiia victima Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 113. ♀.
Aiidrena desponsa Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 114. ♂.




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Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 123, 124. 9, 6 (types). — Cockerell,
1906. Psyche 13: 36. 9 (type). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
134-135 (stylopization and its possible effect on tax. characters).

Genus ANDRENA Subgenus MICRANDRENA Ashmead

Micrandrena Ashmead, 1899. Amer. Ent. Soc, Trans. 26: 89.

Type-species: Andrena melanochroa Cockerell. Monotypic and orig. desig.
{=Micrandrena pacifica Ashmead).
Andrena subg. Andrenella Hedicke, 1933. Berlin Zool. Mus., Mitt. 19: 210.

Type-species: Melitta minutula Kirby. Orig. desig.

Revision: Kibble, 1968. Nebr. Univ. State Mus., Bui. 8: 242-333, figs. 1-89, maps 1-10, tables
1-11 (N. Amer. spp.).

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 208 (tax. characters). — LaBerge, 1964.

Nebr. Univ. State Mus., Bui. 4: 301 (tax. characters).

Andrena annectens
***authority mismatch
Kibble. Calif. (San Benito Co.). Pollen: Unknown, but visits flowers of Salix.

Andrena (Micrandrena) annectens Kibble, 1968. Nebr. Univ. State Mus., Bui. 8: 323, figs.
80-84. ♀, ♂.

Andrena candidiformis Viereck and Cockerell
B. C. and Wash, to Calif., east to Wyo., Colo, and Ariz.
Pollen: Polylectic, visits a wide variety of flowers including Arctostaphylos patula, A.
pungens, Arenaria douglasii, Baccharis, Ceanoihus cordulatus, C. C7ineatus, C.
divaricatus, C. greggii, C. integerrimus, Chamaebatia foliolosa, Crepis, Cryptantha,
Eriodictyon califomicum, Eriogonum, Heracleum lanatuni, Linuni lewisii, Lomatium
dissectum, L. tritematum, Mains, Montia perfoliata, Nemophila menziesii, Physocarpa
intermedia. Primus subcordata, P. virginiana, Pyrus, Ranunculus califomicus,
Rhaynnus califomicus, Ribes nevadense, Salix foliosus, S. laevigata, S. lasiolepis,
Solanum umbelliferum. Taraxacum, Thysanocarpus curvipes, Trifolium, Veronica
americana, Wyethia.

Andrena candidiformis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 33. ♂.

Andrena nigritarsis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 53. 2.

Andrena semotula Cockerell, 1936. Pan-Pacific Ent. 12: 149. ♂.

Taxonomy: Cockerell, 1936. Pan-Pacific Ent. 12: 150. i. —Lanham, 1941. Ent. Soc. Amer.,
Ann. 34: 706, 708. 9, 6 (key). — Timberlake, 1951. In Linsley, In Muesebeck, Krombein
and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1060 (synonymy). —Kibble, 1968. Nebr.
Univ. State Mus., Bui. 8: 288-293, figs. 49-53, map 4 (redescription, synonymy).

Biology: Kibble, 1968. Nebr. Univ. State Mus., Bui. 8: 291-292, table 6 (floral relationships,

stylopizatiom).

Andrena chlorogaster Viereck
B. C. to Calif., east to Idaho, Utah and Ariz. Parasite: Stylops duboisi
Bohart, Imparipes sp. Pollen: Polylectic, visits a wide variety of flowers including
Alyssum maritimum, Amsinckia, Arctostaphylos Crustacea, A. glauca, A. mariposa, A.
pungens, Baccharis, Brassica, Capsella bursa-pastoris, Ceanothus crassifolius, C.
cuneatus, C. greggii, C. orcuttii, C. soredatus, Cercocarpus betuloides, Collinsia
heterophylla, Cryptantha, Daucus, Descourainia sophia, Eriodictyon califomicum,
Geranium molle, Gilia, Lasthenia chrysostoma, Layia glandulosa, Loynatium
dasycarpum, L. dissectum, L. tritematum, L. utriculatum, Montia perfoliata,
Nemophila menziesii, Phacelia ciliata, P. distans, Pmnns subcordata, Ratiunculus
califomicus, R. canus, Rhaynnus crocea, Rhus diversiloba, Salix gooddiyigii, S.
lasiolepis, S. leucodeyidroides, S. nigra, Saynbucus, Sanicula bipiyniatifida, S.
crassicauiis, S. yievadensis, S. tuberosa, Sedum, Sisyyubriuyn irio, Soyichus, Taniarix,
Taraxacum officiyiale, Trifoliuyn pratense.

Ayidreyia chlorogaster Viereck, 1904. Canad. Ent. 35: 189. ♀.

Ayidreyia subtilicoryiis Viereck, 1926. Pomona Jour. Ent. Zool. 18: 4. ♂.

Ayidreyia {Micrayidrena) nitidicomis Cockerell, 1936. Pan-Pacific Ent. 12: 144. ♂.

Ayidreyia catalinica Cockerell, 1939. Pomona Jour. Ent. Zool. 31: 25. ♀.

Ayidreyia (Micr ayidreyia) chlorogaster nesiotes Timberlake, 1951. U. S. Natl. Mus., Proc.
101: 394. ♂.




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Andrena (Micrandrena) chlorogaster gavilanica Timberlake, 1951. U. S. Natl. Mus., Proc.

101:394. ♂, ♀.
Andrena (Micrandrena) radialis Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 395. ♂, ♀.

Taxonomy: Timberlake, 1951. U. S. Natl. Mus. Proc. 101: 396 (tax. characters, as
nitidicomis). — Ribble, 1965. Kans. Ent. See, Jour. 38: 87, fig. 1 (anomalous specimen).
— Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 279-288, figs. 41-48, map 2 (redescription,
synonymy).

Biology: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 287-288, table 5 (floral relationships,

parasites).

Andrena illinoiensis Robertson
South. Canada (Man. to Alta.) and Ohio, Mich., west to Wash., Oreg.
and south. Calif. (Blythe), south to Tex., N. Mex. and Ariz. Parasite: Stylops brunen
Pierce. Pollen: Polylectic, visits a wide variety of flowers including Amelanchier,
Amorpha fndicosa. Anemone, Antennaria plantaginifolia, Capsella, Cardantine,
Ceanothus ovatus, Comus, Crataegus, Descouraiyiia pinnata, Lepidium, Lomatium
daucifolium, L. foeniculaceum. Nasturtium, Prunus americana, Pyrus, Rhus
aromatica, Ribes missouriense, Rosa, Rubus, Salix amygdaloides, S. babylonica, S.
cordata, S. interior, S. nigra, Senecio, Shepherdia argentea, Sisymhriuyn, Sphaeralcea
coccinea. Spiraea thunbergii. Taraxacum officinale, T. vulgare, Thlaspi arvense, Zizia
aurea.

Andrena illinoiensis Robertson, 1891. Amer. Ent. Soc, Trans. 18: 54. ♀, ♂.

Andrena salicinella Cockerell, 1895. Psyche 7 (sup.): 4. ♀.

Andrena placitae Cockerell, 1903. Ent. News 14: 215. ♂.

Andrena vegana Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 17. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 190, 192. 9, 6 (key). —Cockerell,
1914. Ann. and Mag. Nat. Hist. (8) 14: 10. 9 (key). — Lanham, 1941. Ent. Soc. Amer., Ann.
34: 706, 708. 9, 6 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 162-163,
figs. 29, 31-32, table 4 (redescription). —Ribble, 1968. Nebr. Univ. State Mus., Bui. 8:
299-308, figs. 59-63, map 6 (redescription, synonymy).

Biology: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 306-307, table 8 (floral relationships,
stylopization).

Andrena ishii Ribble
Calif. Pollen: Unknown, but visits flowers of Salix.

Andrena (Micrandrena) ishii Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 319, figs. 70-74,
map 8. ♀, ♂.

Andrena labergei Ribble
Ariz. (Southwestern Research Station), N. Mex. (Fort Wingate). Pollen:
Unknown, but visits flowers of Salix.
Andrena {Micrandrena) labergei Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 321, figs.
75-79, map 8. ♀, ♂.

Andrena lamelliterga Ribble
Md., Pa., Ind., Tenn., 111., Kans. Pollen: Unknown, but visits flowers of
Phacelia including P. dubia, P. ranuncidacea.
Andrena {Micrandrena) lamelliterga Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 272,
figs. 31-35, map A. 9, 6.

Andrena lepidii Ribble
South. Calif.; Mexico (Baja California). Pollen: Unknown, but visits flowers of
Cryptantha, Lasthenia chrysostoma, Lepidium flavuyn.
Andrena (Micrandrena) lepidii Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 262, figs.
16-20, map 3. ♀, ♂.

Andrena melanochroa Cockerell
N. S. to N. C, west to B. C, Wash., Oreg and Calif. Pollen: Polylectic,
apparently with a preference for pollens of Fragaria and Potentilla, but visits a variety
of flowers including Ceanothus, Cerastium arvense, Comus drummondi, Fragaria
virginiana, Kalmia, Potentilla canadensis. Ranunculus, Rubus, Salix, Taraxacum
officinale, Vaccinium, Waldsteinia.
Andrena melanochroa Cockerell, 1898. Entomologist 31: 89. ♀, ♂.
Micrandrena pacifica Ashmead, 1899. Amer. Ent. Soc, Trans. 43: 54. ♀, ♂.
Andrena fragariana Graenicher, 1904. Ent. News 15: 64. ♀, ♂.
Andrena {Micrandrena) vagans Cockerell, 1932. Canad. Ent. 64: 157. ♀.




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Taxonomy: Cockerell, 1932. Canad. Ent. 64: 157. 9 (key). — Cockerell, 1932. Canad. Ent. 64:
157, 158. 9, 6 (key, a.s fragariana). — Timberlake, 1951. In Linsley, In Muesebeck,
Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1071 (synonymy). —Mitchell, 1960.
N. C. Agr. Expt. Sta. Tech. Bui. 141: 161, figs. 29-32, table 4 (redescription). -Ribble,
1968. Nebr. Univ. State Mus., Bui. 8: 293-299, figs. 54-58, map 5 (redescription, synonymy).

Biology: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 297, table 7 (floral relationshiips,
stylopization).

Andrena micheneri Ribble
Tex. (Brownsville). Pollen: Unknown, but visits flowers of Lesquerella.
Andrena (Micrandrena) micheneri Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 264 figs.
21-25. ♀, ♂.

Andrena microchlora Cockerell
B. C. to Calif., east to N. Dak., Wyo., Colo, and Ariz. Pollen: Polylectic
with some preference for the pollens of the Umbelliferae and to a lesser extent those of
the Cruciferae, Compositae, Caryophyllaceae and Ranunculaceae; visitation records
include Ainsinckin, Areiiaria califoniica, Brassica, Ceanothus cimeatus, Descourainia
sophia. Euphorbia albomarginata, Gilia, Helianthus, Lasthenia chrysostoma, Lomatium
dissectum, L. utriculatum, Musineon, Netnophila, Plectritis macrocera, Pninus
virginiana. Ranunculus califorjiicus, Salix, Sanicula nevadensis, Taraxacum,
Tropidocarpum gracile.

Andrena microchlora Cockerell, 1922. Ann. and Mag. Nat. Hist. (9) 10: 265. ♂, ♀.

Andrena microchlora var. subalia Cockerell, 1936. Pan-Pacific Ent. 12: 143. ♀.

Taxonomy: Lanham, 1941. Ent. Soc. Amer., Ann. 34: 705, 707. 9, 6 (key). —Timberlake,
1951. U. S. Natl. Mus., Proc. 101: 397 (tax. characters and status). —Ribble, 1968. Nebr.
Univ. State Mus., Bui. 8: 274-279, figs. 36-40, map 3 (redescription, synonymy).

Biology: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 277, table 4 (floral relationships,
stylopization).

Andrena neonana Viereck
N. J. to Fla., west to Ark. and Tex. Pollen: Unknown, but visits flowers of
Fragaria virginiana. Ilex, Prumis persica, Pyrus.
Andrena (Scrapter) neoyiana Viereck, 1917. Amer. Ent. Soc, Trans. 43: 400. ♀.

Taxonomy: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 259-262, figs. 11-15, map 3

(redescription).

Andrena nigrae Robertson
Conn, to north. Fla., west to Alta., Wash., Idaho, Utah and Tex. Pollen:

Apparently polylectic, although majority of collections have been made at the flowers of
Salix; visitation records include Alnus, Anemoiie virginiana, Antennaria
plantaginifolia, Arabis laevigata, Barbarea, Capsella, Cotoneaster, Crataegus,
Cymopterus acaulis, Descourainia pinnata, Erigenia bulbosa, Lomatium daucifolium,
L. dissectum, Pninus americana, P. angustifolia, P. serotina, P. virginiana, Pyrus,
Rhus aromatica, Ribes, Rubtis, Salix amygdaloides, S. babylonica, S. cordata, S.
humilis, S. interior, S. nigra, S. sericea. Sassafras albidtim, Sphaeralcea coccinea.
Spiraea, Taraxacum, Thlaspi arvense. Viburnum, Zizia aurea.

Andrena illinoensis var. bicolor Robertson, 1898. Acad. Sci. St. Louis, Trans. 8: 46. ♀.
Preocc.

Andrena nigrae Robertson, 1905. Canad. Ent. 37: 237. N. name.

Andrena salicinellina Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 17. ♂.

Andrena (Andrena) abacta Viereck, 1917. Amer. Ent. Soc, Trans. 43: 365. ♀.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 165. 9. —Ribble, 1968. Nebr.
Univ. State Mus., Bui. 8: 313-319, fig. 69, map 8 (redescription, synonymy).

Biology: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 317, table 10 (floral relationships,
stylopization).

Andrena personata Robertson
Pa. to Ga., west to Nebr., Kans., Okla. and Tex. Pollen: Polylectic, visits
a wide variety of flowers, especially those of the Rosaceae, Salicaceae, Umbelliferae,
and Hydrophyllaceae in that order; visitation records include Alliaria officinalis,
Apocynum cannabinum, Arenaria patula, Ai'uncus, Claytonia virginica, Cornus,
Crataegus crus-galli, C. mollis, Ellisia nyctelea, Erigeron, Euphorbia commutata,
Fragaria, Geranium carolinianum, Lesquerella, Osmorhiza longistylus, Oxalis violacea.




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Pastinaca sativa, Perideridea americana, Phacelia diibia, P. ranunculacea, P. purshii,
Potentilla canadeiisis, Prunus, Ranunculus abortivus, R. septentrionalis, Rubus
flagellaris, Salix nigra, Sanicula marilandica, Sisymbrium, Spiraea, Taenidia
integerri^na, Thaspium barbinode, T. trifoUatum, Viburnum molle.
Andrena personata Robertson, 1897. Acad. Sci. St. Louis, Trans. 7: 336. ♀, ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 193. 9, 3 (key). — Cockerell, 1932.
Canad. Ent. 64: 157, 158. 9, 6 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
165-167, figs. 28, 31-33, table 4 (redescription). — Ribble, 1968. Nebr. Univ. State Mus., Bui.
8: 254-259, figs. 6-10, map 2 (redescription).

Biology: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 257-258, table 2 (floral relationships,
stylopization).

Andrena piperi Viereck
B. C. to Calif., east to Idaho, Colo, and Ariz. (Tucson); Mexico (Baja California).
Pollen: Polylectic, visits a wide variety of flowers, especially those of the Cruciferae;
visitation records include Agoseris auriantiaca, Alyssum, Amsinckia, Barbarea
orthoceras, Brassica alba, B. campestris, B. 7iigra, Capsella bursa-pastoris, Cardaria
draba, C. pubescens, Ceanothus, Cirsium, Cryptantha intermedia, C. muricata,
Desconrainia pinnata, D. sophia, Erysimum repandiim, Lasthenia chrysostoma,
Lepidijim, Lobularia maritima, Lomatium dissectum, Lupinus, Medicago sativa,
Penstemon, Phacelia f rem ontii, Platystemon califomicus. Polygonum, Prunus
subcordata, Ranxinculus califomicus, Raphanus sativus, Ribes, Rubus, Salix lasiolepis,
Salsola kali. Sisymbrium altissimuyn, S. irio, S. pinnatum, Stellaria media, Tamarix,
Taraxacum officinale, Thlaspi arvense, Tropidocarpum gracile.
Andrena piperi Viereck, 1904. Canad. Ent. 35: 189. ♀.

Taxonomy: Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 392. S. —Ribble, 1968. Nebr. Univ.
State Mus., Bui. 8: 248-254, figs. 1-5, map 1 (redescription).

Biology: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 252-254, table 1 (floral relationships,
stylopization),

Andrena salictaria Robertson
Transcont. south. Canada (N. S. to B. C.) and Maine to N. C. west to
Wash., Utah, Colo, and N. Mex. Parasite: Stylops bruneri Pierce. Pollen: Apparently
narrowly polylectic, chiefly Salix including S. bebbiana, S. cordata, S. humilis, but visits
other flowers including Amelanchier, Antennaria, Barbarea, Chamaedaphne, Claytonia,
Cotoneaster, Crataegus, Lomatium, Prunus americana, Rhus, Sassafras, Viburnum
opulus.
Andrena salictaria Robertson, 1905. Canad. Ent. 37: 236. ♀, ♂.

Taxonomy: Atwood, 1934. Canad. Jour. Res. 10: 207, 210. 9, 6 (key). — Lanham, 1941. Ent.
Soc. Amer., Ann. 34: 705, 706, 708. 9, 6 (key). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141: 167-168, figs. 30-32, table 4 (redescription). -Ribble, 1968. Nebr. Univ. State
Mus., Bui. 8: 309-313, figs. 64-68, map 7 (redescription).

Biology: Salt, 1927. Jour. Expt. Zool. 48: 252 (stylopization).

Taxonomy: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 311-312, table 9 (stylopization, floral

relationships).

Andrena ziziae Robertson
New England to Ga., west to Man., N. Dak., S. Dak., Nebr., Kans. and N.
Mex. Parasite: Imparipes sp. Pollen: Polylectic although nearly all floral records are
from Zizia including Z. aptera, Z. aurea, but visits other flowers including Amorpha
fruticosa, Aruncus dioicus, Barbarea vtilgaris, Ceanothus ovatus. Echinacea
angustifolia, Hieracium, Pastinaca sativa, Polytaenia nuttallii, Rudbeckia
amplexicaulis, Sanicula marilandica, Taenidia integerrima, Thaspium barbinode, T.
trifoUatum.

Andrena ziziae Robertson, 1891. Amer. Ent. Soc, Trans. 18: 55. ♀, ♂.

Andrena (Micrandrena) vemalis Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 168,
fig. 29. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 193. 9, 6 (key). —Cockerell, 1932.
Canad. Ent. 64: 157, 158. 9, 6 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:




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168-169, figs. 31-32, table 4 (redescription). -Ribble, 1968. Nebr. Univ. State Mus., Bui. 8:
267-271, figs. 26-30, map 1 (redescription, synonymy).

Biology: Ribble, 1968. Nebr. State Mus., Bui. 8: 270, table 3 (floral relationships, parasite).

Genus ANDRENA Subgenus NEMANDRENA LaBerge

Andrena subg. Nemandrena LaBerge, 1971. Pan-Pacific Ent. 47: 48.
Type-species: Andrena torulosa LaBerge. Orig. desig.

Revision: LaBerge, 1971. Pan-Pacific Ent. 47: 47-57, 15 figs. (N. Amer. spp.).

Biology: Cruden, 1972. Evolution 26: 373-389, 8 figs., 9 tables (floral relationships). — Cruden,
1972. Madrono 21: 505-515, 2 tables (floral relationships).

Andrena crudeni LaBerge
Calif. Pollen: Apparently oligolectic on flowers of Neynophila including N.
viaciilata, N. menziesii menziesii.
Andrena (Nemandrena) crudeni LaBerge, 1971. Pan-Pacific Ent. 47: 54, figs. 6-10. ♀, ♂.

Andrena subnigripes Viereck
Calif. Pollen: Collects pollen from flowers of Nemophila menziesii
menziesii.
Andrena (Andrena) subnigripes Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 581. ♀.

Andrena torulosa LaBerge
Oreg., Calif. Pollen: Apparently oligolectic on flowers o{ Nemophila
including N. menziesii atomeria, N. m. liniflora, N. m. me7iziesii, but visits other
flowers presumably for nectar including Arctostaphylos, Limnanthes douglasii,
Platystemon califomica.
Andrena (Nemandrena) torulosa LaBerge, 1971. Pan-Pacific Ent. 47: 49, figs. 1-5. ♀, ♂.

Genus ANDRENA Subgenus OLIGANDRENA Lanham

AndreJia subg. Oligandrena Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 207.
Type-species: Andrena macrocephala Cockerell. Orig. desig.

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 284, 290, 301 (tax. characters).

Andrena angelesia Timberlake
South. Calif. Pollen: Unknown, but visits flowers of Cryptantha
intermedia, Encelia farinosa, Platystemon califomicus.
Andrena (Oligandrena) angelesia Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 404. ♀, ♂.

Andrena anisochlora Cockerell
Oreg., Calif. Pollen: Unknown, but visits flowers of Amsinckia
douglasiana, Claytonia spathulata, Lasthenia, Montia perfoliata, Nemophila
heterophylla.
Andrena anisochlora Cockerell, 1936. Pan-Pacific Ent. 12: 137. ♀.
Andrena (Micrandrena) dinognatha Timberlake, 1938. Pan-Pacific Ent. 14: 26. ♂.

Taxonomy: Timberlake, 1951. hi Linsley, hi Muesebeck, Krombein and Townes, U. S. Dept.

Agr., Agr. Monog. 2: 1056 (synonymy).

Andrena macrocephala
***authority mismatch
macrocephala Cockerell. Calif, (lowlands); Mexico (Baja California). Parasite:
Stylops timberlakei Bohart. Pollen: Collects pollen from flowers of Nemophila including
N. menziesii, N. integrifolia as well as Pholistoma auritum, but visits other flowers
presumably for nectar including Calandrinia ciliata, Cryptantha intermedia,
Eriogonum fasciculatum, Erodium, Gilia, Orthocarpus densiflonis, Pkacelia distans,
Ranunculus, Rhus trilobata. Sisymbrium irio.

Andrena macrocephala Cockerell, 1916. Ann. and Mag. Nat. Hist. (8) 17: 278. ♂.

Andrena peratra Cockerell, 1916. Pomona Jour. Ent. Zool. 7: 46. ♀.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 281. 9, 6 (synonymy, floral
relationships, parasite).

Biology: Cruden, 1972. Evolution 26: 378-380, fig. 7, tables 3, 4 (floral relationships).
— Cruden, 1972. Madrono 21: 505-515, 1 fig. 2 tables (floral relations'hips).

Andrena macrocephala
***authority mismatch
tetleyi Linsley. South. Calif, (montane). Pollen: Collects pollen from flowers of
Nemophila integrifolia.
Andrena macrocephala tetleyi Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 282. ♀.




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Biology: Cruden, 1972. Evolution 26: 379 (floral relationships). — Cruden, 1972. Madrono 21:
511 (floral relationships).

Andrena nigroclypeata Linsley
Calif. Pollen: Unknown, but visits flowers oi Lasthejiia, Layia

platyglossa, Marah, Phacelia alba, Plafystemon californkus, Pholistoma racemosa,
Raphanus sativa, Tamarix.
Andrena nigroclypeata Linsley, 1939. Pan-Pacific Ent. 15: 155. ♀, ♂.

Genus ANDRENA Subgenus ONAGRANDRENA Linsley and MacSwain

Andrena subg. Onagrandrena Linsley and MacSwain, 1956. Pan-Pacific Ent. 32: 111.
Type-species: Andrena oenotherae Timberlake. Orig. desig.

The species of this subgenus are chiefly dependent upon the pollen and nectar of several
genera within the family Onagraceae, especially Camissonia, Clarkia, and Oenothera. These
bees are largely or entirely dark or black in coloration and their behavior is usually closely
synchronous with the diurnal flowering regimes of their host plants. Thus some of these bees
are matinal species while others are active at the flowers only in the late afternoon and early
evening and still others are active at the flowers both in the early morning and again in the late
afternoon and early evening hours.

Taxonomy: Linsley and MacSwain, 1955. Pan-Pacific Ent. 31: 163-172 (as Melandrena).

— Linsley and MacSwain, 1956. Pan-Pacific Ent. 32-111-114 (tax. characters, included spp.).

— Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 117-130 (Oenothera and Clarkia
visiting spp.). —Linsley and MacSwain, 1962. Pan-Pacific Ent. 38: 49-52 (new spp. from
Calif., Nev., Wyo.). —Linsley and MacSwain, 1963. Pan-Pacific Ent. 39: 189-198 (oenotherae
complex). —Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 5-6 (key to
Colorado Desert spp.). —Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33:
31 (key to Great Basin spp.). —Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent.
33: 64-65 (key to Mojave Desert spp.). — LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4:
284, 287, 290, 292, 315 (tax. characters). —Linsley, MacSwain, Raven and Thorp, 1973.
Calif. Univ. Pubs. Ent. 71: 27 (key to cismontane Calif, and Baja California spp.).

Biology: Linsley, MacSwain and Smith, 1955. Pan-Pacific Ent. 31: 173-185 (nesting habits,
flower relationships). —Linsley and MacSwain, 1956. Pan-Pacific Ent. 32: 114-121 (nesting
habits, floral relationships). —Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent.
33: 1-58, 6 plates, 6 text figs., 19 tables (comparative behavior of Oenothera visiting spp.
on the Colorado Desert and Great Basin). —Linsley, MacSwain and Raven, 1964. Calif.
Univ. Pubs. Ent. 33: 59-98, 3 plates, 17 tables (comparative behavior of Oenothera visiting
spp. on the Mojave Desert). —MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent.
70: 1-80, 3 plates, 10 text figs., 18 tables (comparative behavior of Clarkia visiting spp. in
western U. S.). —Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71:
1-68, 6 plates, 15 text figs., 10 tables (comparative behavior of Camissonia and Oenothera
visiting spp. in cismontane Calif, and Baja California).

Andrena anograe
***authority mismatch
anograe Cockerell. Calif., Colo. Wyo. Pollen: Collects pollen from Oenothera

caespitosa, but visits other onagraceous flowers including Aywgra, Oenothera albicaiilis,
0. caespitosa var. montana, and may collect pollen from these as well as the large white
flowered evening primroses of the subgenus Pachylophis.
Andrena anograe Cockerell, 1901. Canad. Ent. 33: 154. ♀.
Ayidrena micranthophila Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 432. ♀, ♂.

Taxonomy: Cockerell, 1934. Amer. Mus. Novitates 697: 2 (tax. characters). —Linsley and
MacSwain, 1955. Pan-Pacific Ent. 31: 166 (synonymy, geogr. records). —Linsley and
MacSwain, 1961. Pan-Pacific Ent. 37: 126 (changed status). —Linsley, MacSwain, and
Raven, 1963. Calif. Univ. Pubs. Ent. 33: 31 (tax. characters).

Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 39 (floral
relationships).

Andrena anograe
***authority mismatch
knowltoni Linsley and MacSwain. Utah. Pollen: Unknown, but visits flowers of
Camissonia scapoidea scapoidea, Staiileya pinnata.
Andrena (Onagrandrena) anograe knowltoni Linsley and MacSwain, 1961. Pan-Pacific
Ent. 37: 126. ♀.




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Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 39 (floral
relationships).

Andrena bernardina Linsley
Calif. (San Bernardino Mts.). Pollen: Possibly oligolectic on Clarkia, but
visits other flowers for nectar including Potentilla glandulosa.
Andrena bernardina Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 275. ♀, ♂.

Taxonomy: Linsley and MacSwain, 1955. Pan-Pacific Ent. 31: 166 (tax. status). —Linsley and
MacSwain, 1961. Pan-Pacific Ent. 37: 128 (tax. status).

Biology: MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 70: 16, fig. 2, table 6
(floral relationships).

Andrena blaisdelli Cockerell
South. Calif. Pollen: Principal source of pollen is Camissonia campestris,
but also collects pollen from C. bistorta, C. califomica, C. contorta; visits these and
other flowers for nectar including Crxjptantha intermedia, Eriophyllum confertiflorum.
Andrena blaisdelli Cockerell, 1924. Pan-Pacific Ent. 1: 59. ♀.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 266, 267. 9, 6 (key). —Linsley,
MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 27 (key).

Biology: Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 27-29, fig. 12

(floral relationships).

Andrena boronensis Linsley and MacSwain
Calif. (Mojave Desert). Parasite: Stylops sp. Pollen:

Collects pollen almost exclusively from the flowers oi Camissonia campestris, but has
been observed collecting pollen from the flowers of C. clavaefomiis and C. kemensis;
nectar requirements are chiefly satisfied at the flowers of C. campestris, but other
flowers are visited for nectar including Baccharis viminea, Camissonia bistorta,
Cryptantha intermedia, Encelia farinosa, Eriophyllum confertiflorum, Layia
glandulosa, Potentilla, Salix lasiolepis.

Andrena (Onagrandrena) boronensis Linsley and MacSwain, 1962. Pan-Pacific Ent. 38: 49.
9, 6.

Taxonomy: Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 64, 65, 69 (key,
identity).

Biology: Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 69-71, tables 3-4,
7-15 (nest architecture, male behavior, floral relationships, parasite).

Andrena camissoniae Linsley and MacSwain
Calif. (Santa Barbara County). Pollen: Apparently obtains
pollen from the flowers o{ Camissonia campestris.
Andreiia {Onagrandrena) camissoniae Linsley and MacSwain, 1968. Pan-Pacific Ent. 44:
144. ♀.

Taxonomy: Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 27, 29
(key, tax. characters).

Biology: Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 29 (floral
relationship).

Andrena chylismiae Linsley and MacSwain
Calif. (Owens Valley), Nev. Parasite: Nomada sp., Stylops
sp. Pollen: Collects pollen from the flowers of Camissonia clavaeformis cruciformis and
C. c. integrior, but visits these and other flowers for nectar including Haplopappus
acaulis. Sisymbrium altissimum, Stanleya pinnata, Taraxacum officinale.
Andrena (Onagrandrena) chylismiae Linsley and MacSwain, 1961. Pan-Pacific Ent. 37:
121. ♀, ♂.

Taxonomy: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 31 (key).

Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 37-38, fig. 6, tables
16, 17, 19 (floral relationships, parasites).

Andrena convallaria
***authority mismatch
convailaria Linsley and MacSwain. Calif. (Central Valley and south, cent. Coast
Ranges). Pollen: Collects pollen principally from the flowers of Camissonia campestris
although occasionally obtains pollen from the flowers of C. califomica, C.
cheiranthifolia, C. c. obispoensis, C. contorta, C. sierrae; visits these and other flowers




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for nectar including Brassica, Camissonia micrantha, Encelia actoni, Isomeris arborea,
Lasthenia, Lupiniis, Malacothrix.
Andrena (Onagrandrena) convallaria convallaria Linsley and MacSwain, 1963.
Pan-Pacific Ent. 39: 193. ♀, ♂.

Taxonomy: Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 27 (key).

Biology: Linsley, MacSwain, Raven and Thorp, 1973. Univ. Calif. Pubs. Ent. 71: 29-31, fig. 13,
tables 1, 3-8 (floral relationships).

Andrena convallaria
***authority mismatch
subhyalina Linsley and MacSwain. Calif, (west, margin Mojave Desert). Parasite:
Stylops sp. Pollen: Collects pollen almost exclusively from the flowers of Camissonia
campestris although occasionally obtains pollen from the flowers of C. abramsii; visits
these and other flowers for nectar including A7iisco7na acaulis, Camissonia
clavaefonnis clavaeformis, Coreopsis bigelovii, C. califomica, Layia califomica.
Andrena (Oyiagrandrena) convallaria subhyalina Linsley and MacSwain, 1963. Pan-Pacific
Ent. 39: 193. ♀, ♂.

Biology: Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 71-72, tables 3,
11-12, 14 (mating, floral relationships, parasite).

Andrena deserticola Timberlake
Calif. (Mojave Desert). Parasite: Nomada sp., Stylops sp. Pollen:

Collects pollen almost exclusively from the flowers of Camissonia campestris although
on one occasion collected pollen from the flowers of C. kertiensis; visits these and other
flowers for nectar including Baileya multiradiata, Cam.issonia abramsii, C.
clavaeformis, C. bootkii decorticans, Coreopsis bigelovii, C. califomica, Isomeris
arborea.
Andrena deserticola Timberlake, 1937. Pan-Pacific Ent. 12: 173. ♀.

Taxonomy: Linsley and MacSwain, 1955. Pan-Pacific Ent. 31: 172 (geogr. and floral records).
—Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 64, 65 (key). —Linsley,
MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 27 (key).

Biology: Linsley, MacSwain and Smith, 1955. Pan-Pacific Ent. 31: 182-183 (nest architecture,
parasites). —Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 68-69, tables
3-4, 7-11, 15 (nest architecture, floral relationships, male behavior, parasite). —Linsley,
MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 31, table 1 (floral
relationships).

Andrena flandersi Timberlake
South. Calif (Mojave Desert). Pollen: Collects pollen from the flowers of
Camisso7iia campestris, C. clavaeformis, C. contorta, C. kertiensis, but visits these and
other flowers for nectar including Chaenactis, Coreopsis bigelovii, C. califomica,
Cryptantha, Ericameria, Lasthenia chrysostoma, Layia glandulosa.
Andrena flandersi Timberlake, 1937. Pan-Pacific Ent. 12: 72. ♀.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 266, 278. 9 (key). —Linsley and
MacSwain, 1955. Pan-Pacific Ent. 31: 167 (geogr. and floral records). —Linsley, MacSwain
and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 64, 65 (key).

Biology: Linsley and MacSwain, 1956. Pan-Pacific Ent. 32: 118-120 (nest site, floral
relationships). —Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 72, tables
3, 7, 11-15 (floral relationships).

Andrena furva Linsley and MacSwain
South. Calif.; Mexico (Baja California). Pollen: Principally

gathers pollen from Camissonia campestris although it has been observed gathering
pollen from Clarkia cylindrica and occasionally with mixed pollen loads of Camissonia
and Clarkia.
Andrena {Onagrandrena) furva Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 117. ♀.

Taxonomy: Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 64, 73 (key,
geogr. records). —Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71:
27 (key).

Biology: MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 70: 46 (floral
relationships). —Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71:
32-33, fig. 15 (floral relationships).




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Andrena linsleyi Timberlake
South. Calif., southeast. Nev.; Mexico (Baja California). Parasite: Nomada
sp., Stylops sp. Pollen: Usually collects pollen from Oenothera deltoides, but occasionally
takes pollen from Camissonia clavaeformis including C. c. aurantiaca C. c. clavaeformis
and also occasionally from C. decorticans desertorum, Oenothera trichocalyx; visits
these and other flowers for nectar including Baileya, Dithyrea califomica, Encelia,
Geraea ca7iesce7is, Hyptis emoryi.
Andrena linsleyi Timberlake, 1937. Pan-Pacific Ent. 12: 71. ♀.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 266. 9 (key). -Linsley and
MacSwain, 1955. Pan-Pacific Ent. 31: 165, 166, 170-171 (key, geogr. and floral records).
—Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 6 (key).

Biology: Linsley, MacSwain and Smith, 1955. Pan-Pacific Ent. 31: 176 (floral relationships).
—Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 11-16, fig. 2, tables 1, 3,
7-12 (floral relationships, parasites). —Linsley, MacSwain and Raven, 1964. Calif. Univ.
Pubs. Ent. 3: 73 (floral relationships).

Andrena mojavensis Linsley and MacSwain
Calif. (Jawbone and Short Canyons, Kern County). Pollen:

Principal pollen source is Camissonia kemensis, but also takes pollen from C.

campestris, C. clavaeformis, C. deiitata johnstonii, C. d. parishii; visits these and other

flowers for nectar including Coreopsis bigelovii, Dithyrea califomica, Salix.
Andrena (Melandrena) mojavensis Linsley and MacSwain, 1955. Pan-Pacific Ent. 31: 171.

9, (J.

Taxonomy: Linsley and MacSwain, 1955. Pan-Pacific Ent. 31: 165 (key). —Linsley, MacSwain
and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 64, 65 (key).

Biology: Linsley, MacSwain and Smith, 1955. Pan-Pacific Ent. 31: 179-181 (nest architecture,
floral relationships). —Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33:
65-67, tables 5-10 (nest architecture, floral relationships, male behavior).

Andrena nevadae Linsley and MacSwain
Nev. (Eureka and White Pine Counties). Pollen: Unknown,
presumably collects pollen from the flowers of the subgenera Anogra or Pachylophis;
visits flowers of Camissonia clavaeformis and Stanleya pinnata for nectar.
Andrena (Onagrandrena) nevadae Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 125.
9.

Taxonomy: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 31 (key).

Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 39 (floral

relationships).

Andrena oenotherae Timberlake
Calif. (Coastal Santa Barbara Co. east to Colorado Desert); Mexico
(Baja California). Parasite: Lytta stygica (LeC.)?, Meloe sp.?, Stylops sp. Pollen:
Principally collects pollen from the flowers of Camissonia cheiranthifolia
cheiranthifolia on Santa Cruz Island, and C. c. siiffndicosa along the mainland coast
while more interior populations collect from C. bistorta; visits these and other flowers
for nectar including Cakile edentula, Calandrinia, Chaenactis, Chrysanthemum
coronarium, Coreopsis califomica, Croton califomicus, Cryptantha intermedia,
Eriogonum fasciculatum, Haplopappus cooperi, Hemizonia kelloggii, Lantana, Layia
platyglossa, Lupimis, Salix lasiolepis. Sisymbrium.

Andrena oenotherae Timberlake, 1937. Pan-Pacific Ent. 12: 69. ♀, ♂.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 266, 267. 9, 6 (key). -Linsley and
MacSwain, 1955. Pan-Pacific Ent. 31: 164, 166, 170 (key, geog. and floral records).
—Linsley and MacSwain, 1963. Pan-Pacific Ent. 39: 189-190. —Linsley, MacSwain, Raven
and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 27 (key).

Biology: Linsley, MacSwain and Smith, 1955. Pan-Pacific Ent. 31: 181-182 (nest architecture,
floral relationships, parasites). —Linsley and MacSwain, 1956. Pan-Pacific Ent. 32: 116-118,
1 fig. (nest architecture, floral relationships, parasite). —Linsley and MacSwain, 1963.
Pan-Pacific Ent. 39: 189-190 (floral relationships). —Linsley, MacSwain, Raven and Thorp,
1973. Calif. Univ. Pubs. Ent. 71: 33-34, fig. 14, table 9 (floral relationships).




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Andrena omninigra
***authority mismatch
clarkiae Linsley and MacSwain. Calif. (Monterey and San Luis Obispo Counties).
Pollen: Collects pollen from the flowers of Clarkia cylindrica, C. speciosa.
Andrena (Onagrandrena) omninigra clarkiae Linsley and MacSwain, 1961. Pan-Pacific
Ent. 37: 127. ♀, ♂.

Biology: MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 70: 15-16, fig. 1, tables
7-11 (floral relationships, mating behavior).

Andrena omninigra
***authority mismatch
omninigra Viereck. Calif, (west, slopes of Sierra Nevada Mts.). Pollen: Collects
pollen from the flowers of Clarkia including C. biloba, C. cylindrica, C. dudleyana, C.
purpurea quadrivuhiera, C. rhomboidea, C. speciosa polyantha, C. U7iguiculata, C.
xantiana; visits these and other flowers for nectar including Cryptantha flaccida, Gilia
capitata. Predator: Callinicus calcaneus (Loew).
Andrena (Aiidreiia) omninigra Viereck, 1917. Amer. Ent. Soc, Trans. 43: 385. ♀.
Ayidrena grundeli Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 274. ♀, ♂.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 265. 9 (key). —Linsley and
MacSwain, 1955. Pan-Pacific Ent. 31: 164, 165, 166 (key, geogr. records). —Linsley and
MacSwain, 1961. Pan-Pacific Ent. 37: 27 (type).

Biology: Linsley, 1972. Pan-Pacific Ent. 48: 94-96 (predator). —MacSwain, Raven and Thorp,
1973. Calif. Univ. Pubs. Ent. 70: 14, fig. 1, tables 1, 6-9, 13-18 (floral relationships).

Andrena oraria
***authority mismatch
actitis Linsley and MacSwain. Calif. (San Francisco). Pollen: Collects pollen from the
flowers of Camissonia cheiranthifolia cheiranthifolia.
Andrena (Onagrandrena) oraria actitis Linsley and MacSwain, 1963. Pan-Pacific Ent. 39:
196. ♀, ♂.

Biology: Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 35, fig. 15
(floral relationship).

Andrena oraria
***authority mismatch
oraria Linsley and MacSwain. Calif. (Marin and Sonoma Counties). Pollen: Collects
pollen from the flowers of Camissonia including C. cheira^ithifolia cheiranthifolia, C.
ovata.
Andrena {Onagrandrena) x>raria oraria Linsley and MacSwain, 1963. Pan-Pacific Ent. 39:
196. ♀, ♂.

Biology: Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 35, fig. 15
(floral relationships).

Andrena raveni Linsley and MacSwain
Oreg., Calif., Nev., Utah. Parasite: Nomada sp., Stylops sp.

Pollen: Collects pollen from the flowers of Camissonia clavaeformis clavaeforruis, C. c.
integrior, C. tanacetifolia, Oenothera deltoides piperi, but visits these and other flowers
for nectar including Agoseris, Madia, Stanleya pinnata, Taraxactim officinale.
Andrena (Onagrandrena) raveni Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 118. ♀.
S.

Taxonomy: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 31 (key).

Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 31-36, fig. 4, tables
14-19 (nest architecture, floral relationships, parasites).

Andrena rozeni Linsley and MacSwain
Ariz., south. Calif., Nev.; Mexico (Baja California), deserts.
Parasite: Nomada sp., Stylops sp. Pollen: Collects pollen from the flowers of
Camissonia including C. clavaeformis aurantiaca, C. c. clavaeformis, C. c. integrior, C.
c. peirsonii, C. decorticans, C. latifolia, C. tanacetifolia, C. trichocalyx and Oenothera
deltoides piperi; visits these and other flowers for nectar including Agoseris glauca,
Brassica, Cryptantha clevelandii, C. intermedia, Encelia farinosa, Geraea canescens.
Madia ramii, Mirabilis, Sisymbrium altissimum, Stanleya pinnata. Taraxacum
officinale.
Andrena (Melandrena) rozeni Linsley and MacSwain, 1955. Pan-Pacific Ent. 31: 168. ♀, ♂.

Taxonomy: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 5, 31 (key).
— Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 64, 65 (key).

Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 6-11, 36-37, figs. 1, 5,
tables 1-6, 17-18 (nest architecture, floral relationships, male behavior, parasites).




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— Linsley, MacSwain and Raven, 1964. Calif. Univ. Pubs. Ent. 33: 72-73, table 16 (floral

relationships).

Andrena rubrotincta Linsley
South. Calif., northwest. Ariz., south. Nev., southwest. Utah. Pollen:
Collects pollen regularly only from Camissonia brevipes pallidula and C. parryi,
although females have been taken at flowers of C. brevipes brevipes and C. clavaefonnis
au7-a7itiaca with pollen in their scopae.

Andrena rubrotincta Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 278. ♀.

Taxonomy: Linsley and MacSwain, 1955. Pan-Pacific Ent. 31: 165, 170 (key, geogr. records).
— Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 5, 6 (key).

Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 16-18, fig. 3, table 13
(floral relationships).

Andrena stagei Linsley and MacSwain
Wyo. (Sweetwater Co.). Pollen: Collects pollen from the flowers
o{ Oenothera trichocalyx.
Andreyia {Onagrandrena) stagei Linsley and MacSwain, 1962. Pan-Pacific Ent. 38: 52. ♀.

Taxonomy: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 31 (key).

Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 88 (floral
relationships).

Andrena thorpi Linsley and MacSwain
Nev. (Humboldt Co.). Parasite: Stylops sp. Pollen: Collects
pollen from the flowers of Oenothera deltoides piperi.
Andrena (Onagrandrena) thorpi Linsley and MacSwain, 1962. Pan-Pacific Ent. 38: 51. ♀.

Taxonomy: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 31 (key).

Biology: Linsley, MacSwain and Raven, 1963. Calif. Univ. Pubs. Ent. 33: 38-39, table 15
(floral relationships, parasite).

Andrena vanduzeei Linsley
Calif. Pollen: Possibly an oligolege of Gayophytum.

Andrena vanduzeei Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 280. ♀, ♂.

Taxonomy: Linsley and MacSwain, 1955. Pan-Pacific Ent. 31: 165, 166, 172 (key, geogr.
records).

Andrena vespertina Linsley and MacSwain
Calif. (Kern and Stanislaus Counties). Pollen: Collects pollen
only from the flowers of Camissonia boothii decorticans; visits flowers of Isomeris
arborea for nectar.
Andrena (Onagrandrena) vespertina Linsley and MacSwain, 1961. Pan-Pacific Ent. 37:
123. ♀, ♂.

Taxonomy: Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 27 (key).

Biology: Linsley, MacSwain, Raven and Thorp, 1973. Calif. Univ. Pubs. Ent. 71: 35-36, fig. 15,
table 10 (floral relationships).

Andrena yumorum Viereck
South. Calif.

Andrena (Andrena) yumonim Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 585. ♂.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 128-129 (type).

Genus ANDRENA Subgenus OPANDRENA Robertson

Opandrena Robertson, 1902. Amer. Ent. Soc, Trans. 28: 193.
Type-species: Andrena cressonii Robertson. Orig. desig.

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 210 (tax. characters). — LaBerge, 1964.
Nebr. Univ. State Mus., Bui. 4: 284, 289, 303 (tax. characters).

Andrena cressonii
***authority mismatch
cressonii Robertson. U. S. and southern Canada. Pollen: Apparently polylectic, visits
a wide variety of flowers including Amelanchier, Anemonella, Arabis, Aronia, Arimcus,
Brassica, Capsella, Ceanothus, Celastrus, Claytonia, Conius, Crataegus, Dentaria,
Fragaria, Heradeinn, Ilex, Oxalis, Pastinaca, Philadelphus, Polytaenia, Potentilla,
Prunus, Ptelea, Pyracantha, Radicula, RaiiuncHlus, Rhainnus, Rhus, Rosa, Rubus,
Salix, Saiiicula, Sisymbrium, Smilacina, Smilax, Stellaria, Taenidia, Taraxacum,
Thaspium, Tradescantia, Trifolium, Veronica, Viburnum, Zanthoxylum, Zizia.
Andrena Cressonii Robertson, 1891. Amer. Ent. Soc, Trans. 18: 56. ♀, ♂.




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Andrena kansensis Cockerell, 1899. Ent. News 10: 255. ♂.

Andrena bridweUi Cockerell, 1899. Ent. News 10: 255. ♂.

Ajidrena dubia Robertson, 1902. Canad. Ent. 34: 48. ♂.

Andrena {Trachandrena) trumani Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48:

12. ♀.
Andrena latisigna Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 22. ♂.
Andrena cressoni transformans Cockerell, 1934. Pan-Pacific Ent. 9: 157. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 193. 9, 6 (key). — Lanham, 1949.
Pan-Pacific Ent. 25: 147 (tax status). — Timberlake and Lanham, 1951. In Linsley, In
Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1063 (synonymy).
—Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 248-249, figs. 50, 52-54
(redescription).

Andrena cressonii
***authority mismatch
infasciata Lanham. Calif., Oreg., Wash. Pollen: Presumably gathers pollen from
Gilia, but visits other flowers for nectar including Brassica campestris, Ranunculus
califomicus, Pyracantha, Salix lasiolepis, S. nigra.
Andrena cressonii infasciata Lanham, 1949. Pan-Pacific Ent. 25: 147. ♀, ♂.

Biology: Linsley and MacSwain, 1959. Calif. Univ. Pubs. Ent. 16: 19 (floral relationships).

Andrena cressonii
***authority mismatch
ivanensis Mitchell. Va., N. C. Pollen: Unknown, but visits flowers of Brassica,
Padus.
Andrena (Opandrena) cressonii ivanensis Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui.
141: 249, fig. 52. ♂.

Andrena cressonii
***authority mismatch
tallahasseensis Mitchell. Fla., N. C. Pollen: Unknown, but visits flowers of Castanea
pjimila.
Andrena (Opandrena) cressonii tallahaseyisisi!) Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.

Bui. 141: 249, figs. 50, 53. ♂.
Andrena (Opandrena) cressonii tallahasseeyisis Mitchell, 1967. In Krombein and Burks, U.
S. Dept. Agr., Agr. Monog. 2, Second Suppl., p. 428. Emend.

Andrena prunifloris Cockerell
N. Mex. Pollen: Unknown, but visits flowers of Primus.
A7idrena prunifloris Cockerell, 1898. Canad. Ent. 30: 147. ♀.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 49. 9. (key).

Genus ANDRENA Subgenus PARANDRENA Robertson

Parandrena Robertson, 1897. Acad. Sci. St. Louis, Trans. 7: 337.

Type-species: Panurgus andrenoides Cresson. Desig. by Cockerell, 1897.

The species of this Nearctic subgenus are apparently oligoleges of Salix.

Revision: LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 307-346, figs. 5-9, 41-70 (N.
Amer. spp.).

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 202 (tax. characters). —LaBerge, 1964.

Nebr. Univ. State Mus., Bui. 4: 298 (tax. characters).

Andrena andrenoides (Cresson)
South. Canada (B. C, Sask., Man. and Ont.), south to Fla., Tex., N.
Mex. and Utah. Pollen: Oligelege o{ Salix including S. amygdaloides, S. cordata, S.
interior, S. nigra, but visits these and other flowers for nectar including Amelanchier
canadensis. Anemone patens, Antennaria plantaginifolia, Crataegus crus-galli,
Erigenia bulbosa, Heracleum maximum, Lomatium foeniculaceum, Prunus americana,
P. angustifolia, P. gracilis, Pyrus, Rubus, Sassafras variifolium. Sisymbrium, Stellaria
media, Viola.

Panurgiis andrenoides Cresson, 1878. Amer. Ent. Soc, Trans. 7: 62. ♂.

Panurgus rufocinctus Ashmead, 1890. Colo. Biol. Assoc, Bui. 1: 4. ♀.

Andrena andrenoides f. bicolor Robertson, 1898. Acad. Sci. St. Louis, Trans. 8: 47. Preocc.

Parandrena andrenoides var. clarigastra Viereck, 1908. Ent. News 19: 42. N. name.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 238-239, figs. 49-50, 52, 55,
table 6 (redescription).

Biology: Salt, 1927. Jour. Exp. Zool. 48: 250 (stylopization).




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Andrena arenicola LaBerge and Ribble
Kans., Okla., Tex. Pollen: Apparently an oligolege oi Saiix
including S. amygdaloides, S. longifolia, S. nigra, but visits other flowers including
Pyrus pyrus.
Andrena (Parandre7ia) arenicola LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98:
328, figs. 7, 53-54. ♀, ♂.

Biology: LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 330 (stylopization).

Andrena bucculenta LaBerge and Ribble
Calif. Pollen: Presumably an oligolege of Salix, but visits
other flowers including Eschscholzia californica, Layia platyglossa, Primus.
Andrena (Parandrena) bucculenta LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98:
334, figs. 6, 60-61. ♀, ♂.

Andrena concinnula (Cockerell)
Oreg., Calif., Ariz. (Chiricahua Mts.). Pollen: Oligolege oi Salix

including S. argophylla, S. cordata, S. exigua, S. gooddingii, S. hindsiaiia, S. laevigata,
S. lasiolepis, but visits other flowers presumably for nectar including Amsinckia
douglasiana, Capsella bursa-pastoris, Eschscholzia californica, Gilia, Sambucus,
Spiraea pninifolia, Tamarix gallica.

Parandrena concinnula Cockerell, 1898. Amer. Ent. Soc, Trans. 25: 189. ♂.

Andrena (Parandrena) mejidosa Viereck, 1916. Acad. Nat. Sci. Phila., Proc 68: 588. ♀.

Taxonomy: LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 331-334, figs. 8, 55-59
(redescription, synonymy).

Andrena gibberis Viereck
B. C. (Fairview), Oreg. (Irrigon), Calif. (Lassen and Mono Counties), Nev.
(Elko), Utah (Cedar City). Pollen: Presumably an oligolege oi Salix.
Andrena (Parandrena) gibberis Viereck, 1924. Canad. Ent. 56: 241. ♀, ♂.

Andrena nevadensis (Cresson)
B. C. to Calif., Nev. and Utah. Pollen: Presumably an oligolege oi Salix,
but visits other flowers including Acer, Ribes, Senecio, Tamarix.
Panurgus iievadensis Cresson, 1879. Amer. Ent. Soc, Trans. 7: 214. ♂.
Andrena (Parandrena) garretti Viereck, 1924. Canad. Ent. 53: 243. ♂.
Andrena (Parandrena) triangularis Viereck, 1924. Canad. Ent. 55: 243. ♀.

Taxonomy: Cockerell, 1898. Amer. Ent. Soc, Trans. 25: 188. 6. -LaBerge and Ribble, 1972.
Amer. Ent. Soc, Trans. 98: 341-344, figs. 7, 67-68 (redescription, synonymy).

Andrena nida Mitchell
Vt. to Ga., west to Mich., 111., Mo. and Miss. Pollen: Presumably an oligolege of
Salix including S. interior, but visits other flowers including Amelanchier, Crataegus,
Primus, Rubus, Viola.
A7idre7ia (Parandrena) nida Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 240, figs.
49,50,52,55. ♀, ♂.

Taxonomy: LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 321-324, fig. 7, 46-47
(redescription).

Andrena papagorum Viereck and Cockerell
Ariz., south. Calif., Utah. Pollen: Presumably an oligolege
of Salix, visits flowers of S. gooddingii, S. nigra.
Andrena papagorum Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 2. ♀.

Taxonomy: LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 338-341, figs. 9, 62-66
(redescription).

Andrena wellesleyana Robertson
Mass. and Conn., west to N. W. T., Alta., and Utah. Pollen: Unknown,
but visits flowers oi Primus virginiana, Salix.
Andrena wellesleyana Robertson, 1897. Acad. Sci. St. Louis, Trans. 7: 337. ♀, ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 241-242, figs. 50, 52, 55, table
6 (redescription). -LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 325-328, figs. 8,
42-52 (redescription).

Genus ANDRENA Subgenus PELICANDRENA LaBerge and Ribble

Andrena subg. Pelicandrena LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 346.
Type-species: Parandrena atypica Cockerell. Monotypic and orig. desig.

Revision: LaBerge and Ribble, 1972. Amer. Ent. Soc, Trans. 98: 346-351, figs. 71-76.




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Andrena atypica (Cockerell)
Oreg (Klamath Falls), Calif.; Mexico (Baja California). Pollen: Apparently
polylectic with some preference for flowers of Cryptantha, Salix, Rhus, Rhainnus and
Ceanothus in approximately that order; visitation records include Adenostoma
fasciculatum, Allimn haematochiton, Arctostaphylos bicolor, Baccharis, Brassica,
Ceanothus crassifolius, C. cuneatus, C. greggii, C. soredahcs, C. verrucosus, Cryptantha
intermedia, Emmenanthe penduliflora, Gilia capitata, G. multicaulis, Lomatium
dasycarpum, Marah macrocarpus, Oenothera, Plagiobothrys nothofulvus, Pnnius
ilicifolia, Quercus, Rhamnus califomica, R. crocea, Rhus diversiloha, R. integrifolitim,
R. ovata, R. trilobata, Rosa minutiflora, Salix argophylla. Salvia coiimibariae, S.
mellifera, Tamarix gallica.
Parandrena atypica Cockerell, 1941. San Diego Soc. Nat. Hist., Trans. 9: 347. ♀.

Taxonomy: Thorp, 1969. Calif. Univ. Pubs. Ent. 52: 123 (tax. characters and status).

Genus ANDRENA Subgenus PLASTANDRENA Hedickc

Andrena subg. Plastandrena Hedicke, 1933. Berlin Zool. Mus., Mitt. 19: 217.

Type-species: Melitta tibialis Kirby. Orig. desig.
Andrena subg. Schizandrena Hedicke, 1933. Berlin Zool. Mus., Mitt. 19: 218.

Type-species: Andrena aulica Morawitz. Orig. desig.

Revision: LaBerge, 1969. Amer. Ent. Soc, Trans. 95: 2-34, figs. 1-23, 3 tables (N. Amer. spp.).

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 201 (tax. characters, as Schizandrena).
— LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 297 (tax. characters, synonymy).

Andrena argemonis Cockerell
Nebr. and Colo, south to N. Mex. and Ariz.; Mexico (Chihuahua,

Coahuila, and Sonora south to Oaxaca). Pollen: Polylectic, visits flowers of Acacia,
Argemone mexicana, A. platyceras, Cleome serrulata, Medicago sativa, Melilotus alba,
Penstemon occidentalis, Persicaris pennsylvanicus, Sisymbrium irio.
Andrena argemonis Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 80. ♂.

Taxonomy: Cockerell, 1898. Ann. and Mag. Nat. Hist. (7) 2: 449. 9. —Cockerell, 1931. Amer.

Mus. Novitates 458: 16. 9, d (key). —Lanham, 1941. Ent. Soc. Amer., Ann. 34: 704, 707. ♀.

6. —LaBerge, 1969. Amer. Ent. Soc, Trans. 95: 12-15, figs. 24-28 (redescription).

Andrena crataegi Robertson
Transcont. in southern Canada (N. S. to B. C.) and northern U. S., south
to Ga., Ala., Ark., Tex., N. Mex., Utah, Nev. and north. Calif. Pollen: Polylectic with
some preference for the flowers of the family Rosaceae, visits flowers of Amelanchier,
Amorpha, Aruncus, Astragalus, Barbarea, Blephilia, Brassica, Caragana, Cardaria,
Carduus, Castanea, Ceanothus, Chamaebatia foliolosa. Chrysanthemum, Conium,
Coruus, Crataegus, Crepis, Cryptantha intermedia, Daucus, Deutzia, Eulophus,
Euphorbia, Fragaria, Frasera, Gilia achillaefolia, Hackelia, Heracleum lanatum,
Holodiscus, Hydrangea, Ilex, Iris, Ledum, Lomatiuin dasycarpum, Malvastrum,
Melilotus, Monarda, Narcissus, Opulaster, Pastinaca, Philadelphus, Polytaenia,
Potentilla, Primus, Ptelea, Purshia, Pyracantha, Pyrus, Ranunculus, Raphajius,
Rhamnus, Rhododendron, Rhus, Rorippa nasturtium-aqtiaticum, Rosa, Rubus, Salix,
Senecio, Solidago, Sorbaria, Spiraea, Symphoricarpos, Taenidia, Tamarix, Taraxacum,
Thaspium, Trifolium repens, Vaccinium, Viburnum, Zizia.

Andrena crataegi Robertson, 1893. Amer. Ent. Soc, Trans. 20: 273. ♀, ♂.

Andrena subcommoda Cockerell, 1902. Canad. Ent. 34: 45. ♀.

Andrena angusi Viereck, 1907. Ent. News 18: 284. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 190. 9, 6 (key). — Atwood, 1934.
Canad. Jour. Res. 10: 208, 210. 9, 6 (key). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141: 206-209, figs. 37-39, table 5 (redescription, synonymy). —LaBerge, 1967. Nebr.
Univ. State Mus., Bui. 7: 294 (subgeneric assignment, angusi). —LaBerge, 1969. Amer.
Ent. Soc, Trans. 95: 5-12, figs. 1, 24-28 (redescription).

Biology: Rau, 1922. Acad. Sci. St. Louis, Trans. 24: 33-34 (nest). —Clements and Long, 1923.
Carnegie Inst. Wash., Pub. 326: 249 (ecology). — Brittain, 1933. Canada Dept. Agr., Bui.
162: 94, figs, (floral relationships). —LaBerge, 1969. Amer. Ent. Soc, Trans. 95: 11, table 1
(floral relationships).




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Andrena fracta Casad and Cockerell
West. Tex. to south Calif.; Mexico (Distrito Federal, Puebla,

Zacatecas). Pollen: Polylectic, visits flowers of Acacia, Argemone, Baileya, Cryptantha
intermedia, Desconrainia sophia, Dithyrea, Encelia farinosa, Eschscholzia, Fallugia,
Geraea canescens, Lepidiuvi, Lesquerella, Medicago, Oenothera, Opuntia, Phacelia,
Physaria, Prosopis, Primus, Pyrus, Salix, Senecio.
Andrena fracta Casad and Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 84. ♂, ♀.

Taxonomy: LaBerge, 1969. Amer. Ent. Soc, Trans. 95: 30-34, figs. 14-18 (redescription).

Biology: LaBerge, 1969. Amer. Ent. Soc, Trans. 95: 34, table 3 (floral relationships).

Andrena mellea Cresson
Tex., N. Mex., Ariz. Pollen: Unknown, but visits flowers of Argemone,

Asclepias subverticillata, Baccharis glutinosa, Fallugia paradoxa, Melilotus alba, Salix.
Andrena mellea Cresson, 1868. Amer. Ent. Soc, Trans. 1: 384. ♀.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 48 (key). —Cockerell, 1931.

Amer. Mus. Novitates 458: 16. 9 (key). —LaBerge, 1969. Amer. Ent. Soc, Trans. 95: 16-18,

figs. 19-23 (redescription).

Andrena prunorum
***authority mismatch
prunorum Cockerell. Sask., west to B. C, south to west. Tex., N. Mex., Ariz, and
Calif.; Mexico (Baja California). Pollen: Polylectic, visits a wide variety of flowers
including Acer, Actinea, Adenostoma, Amsinckia douglasiana, Arabis, Arctostaphylos,
Argemone, Baccharis glutinosa, B. viminea, Barbarea orthoceras, Berberis, Brassica
campestris, B. incana, B. nigra, Calochortus venustiis, Calyptridium umbellatum,
Ceanothus integerrimus, Cercocarpus betuloides, Chaenactis glabriuscula, Chamaebatia
foliolosa, Chrysothamnus, Clarkia biloba, C. rhoniboidea. Clematis, Cleome serrulata,
Crataegus, Crepis, Cryptantha interynedia, Cymopteris, Encelia farinosa, Eriastrum
virgata, Erigeron, Eriodictyon trichocalyx laiiatum, Eriogonum fasciculatum, E.
nudum, Eriophylhim, Erysimum asperum, Eschscholzia califomica, Fallugia
paradoxa, Gilia capitata, G. davyi, Hackelia, Helianthus, Heracleum, Hyptis emoryi,
Lappula, Larrea tridentata, Lepidium flavuni, Lesquerella, Linaria, Lithospermum,
Lomatium, Malus, Melilotus, Mentzelia, Micrampelis, Moiiardella lanceolata,
Nemophila, Opuntia, Pastinaca, Penstemon, Petalostemon, Phacelia distaiis, P. minor,
Philadelphus lewisii califomica, Platystemon, Plumbago, Potentilla, Prosopis, Prunus,
Purshia, Ranunculus, Raphanus sativa, Rhamnus califomicus, Rhus, Ribes, Rorippa,
Rubus, Salix, Sambucus caerulea, Sanicula nevadensis, Sophia, Sphaeralcea ambigua,
Shepherdia, Sisymbrium irio, S. linifolia, Solidago, Spiraea, Symphoricarpos, Syriiiga,
Tamarix gallica. Taraxacum officinale, Thysanocarpus curvipes, Trichostema parishii,
Trifolium.

Andrena prunorum Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 81. ♀, ♂.

Andrena kincaidii Cockerell, 1897. Acad. Nat. Sci. Phila., Proc 49: 351. ♀.

Andrena pascoensis Cockerell, 1897. Entomologist 30: 305. ♀.

Andrena prunorum gillettei Cockerell, 1898. Ent. News 9: 172. ♂.

A7idrena venwni Viereck, 1904. Canad. Ent. 36: 190, 195. ♀, ♂.

Andrena arizonensis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 42. ♂.

Andrena prunorum var. mariformis Cockerell, 1916. Pomona Jour. Ent. Zool. 8: 46. ♀.

Andrena {Andrena) shasta Viereck, 1926. Calif. Acad. Sci., Proc. (4) 15: 402. ♀.

Andrena prunorum var. pauperatula Cockerell, 1938. Canad. Ent. 70: 6. ♀.

Taxonomy: Viereck, 1904. Canad. Ent. 36: 190, 195. 9, d (key, as kincaidii). —Cockerell,
1914. Ann. and Mag. Nat. Hist. (8) 14: 11. 9 (key). —Cockerell, 1931. Amer. Mus. Novitates
458: 16. 9,6 (key). — Lanham, 1941. Ent. Soc Amer., Ann. 34: 704, 707. 9, 6 (key).
— LaBerge, 1969. Amer. Ent. Soc, Trans. 95: 18-30, figs. 2-3, 9-13 (redescription,
synonymy, integumental color variation).

Biology: LaBerge, 1969. Amer. Ent. Soc, Trans. 95: 28-30, table 2 (floral relationships).

Andrena prunorum
***authority mismatch
sinaloa Viereck. Ariz.; Mexico (Baja California and Sinaloa). Pollen: Presumably
polylectic, visits flowers of Encelia farinosa, Larrea tridentata.
Andrena sinaloa Viereck, 1926. Calif. Acad. Sci., Proc. (4) 15: 403. ♀.

Taxonomy: LaBerge, 1969. Amer. Ent. Soc, Trans. 95: 30 (redescription).




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Genus ANDRENA Subgenus PTILANDRENA Robertson

Ptilandrena Robertson, 1902. Amer. Ent. Soc, Trans. 28: 192.
Type-species: Andrena erigeniae Robertson. Orig. desig.

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 215 (tax. characters). — LaBerge, 1964.
Nebr. Univ. State Mus., Bui. 4: 283, 290, 307-308 (tax. characters). —Davis and LaBerge,
1975. 111. Nat. Hist. Survey, Biol. Notes 95: 2 (included spp.).

Andrena distans Provancher
Canada; Minn, to New England, south to N. C. Pollen: Unknown, but
visits flowers of Geranium maculatum, Heracleum lanatum, Polemonium reptans.
Andrena distayis Provancher, 1888. Addit. Corr. Faune Ent. Canada, Hym., p. 307. ♀, ♂.
Andrena g. maculati Robertson, 1897. Acad. Sci. St. Louis, Trans. 7: 333. 9,6. Trinomial.
Andrena gerani-maculati Viereck, 1907. Ent. News 18: 282. ♀.

Taxonomy: Viereck, 1907. Ent. News 18: 282, 286. 9, 6 (key). —Mitchell, 1960. N. C. Agr.
Expt. Sta. Tech. Bui. 141: 154-155, figs. 27, 29-30, table 4 (redescription, synonymy).

Andrena erigeniae Robertson
Ont. to Ga., west to Minn., Mo. and Okla. Parasite: Leucophora obtusa

(Zett.), Stylops erigeniae Pierce. Pollen: Apparently collects pollen only from the flowers
of Claytonia virginica with which it is largely coterminous, but visits these and other
flowers for nectar including Barbarea vulgaris, Collinsia vema, Dicentra laciniata, D.
cucullaria, Erigenia bulbosa, Erythronium americanum, Hydrophyllum appendiculata,
Isopyrum bitematum, Prunus, Veronica hederaefolia.
Ayidrena erigeniae Robertson, 1891. Amer. Ent. Soc, Trans. 18: 52. ♀, ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 192. 9, 6 (key). —Mitchell, 1960. N.
C. Agr. Expt. Sta. Tech. Bui. 141: 155-156, figs. 27, 29-31, table 4 (redescription). —Davis
and LaBerge, 1975. 111. Nat. Hist. Survey, Biol. Notes 95: 13, figs. 13, 16-24 (immature
stages).

Biology: Davis and LaBerge, 1975. 111. Nat. Hist. Survey, Biol. Notes 95: 1-16, figs. 1-25, 1
table (nest, architecture, life history, provisioning, inquilinism, parasitism).

Andrena polemonii Robertson
111., Mich. Pollen: Unknown, but visits flowers of Geranium maculatum,
Polemonium reptans. Ranunculus septentrionalis.
Andrena polemonii Robertson, 1891. Amer. Ent. Soc, Trans. 18: 54. ♀, ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 190, 192. 9, 6 (key). —Robertson
1904. Canad. Ent. 36: 278. (tax. characters). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141: 157-158, figs. 27, 31, table 4 (redescription).

Genus ANDRENA Subgenus RHAPHANDRENA LaBerge

Andrena subg. Rhaphandrena LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 507.
Type-species: Andreyia prima Casad and Cockerell. Orig. desig.

Revision: LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 507-514, figs. 1, 3, 6, 59-68 (N. Amer.
spp.).

Andrena dapsilis LaBerge
Tex., Okla. Pollen: Unknown, but visits flowers of Descourainia pinnata
brachycarpa, Lesquerella.
Andrena (Rhaphandrena) dapsilis LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 511, figs. 6,
64-68. ♀, ♂.

Andrena prima Casad
West. Tex. to south. Calif, and north. Utah. Pollen: Apparently an oligolege of
the brassicaceous flowers, especially Lesquerella including L. gordonii, but visits other
flowers presumably for nectar including Alyssum, Brassica oleracea, Cryptantha
probably jamesii, Dithyrea wislizenii, Geraea canescens, Prunus.
Andrena prima Casad, 1896. Ann. and Mag. Nat. Hist. (6) 18: 78. ♀.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 48. 9 (key). — Linsley and
MacSwain, 1961. Pan-Pacific Ent. 37: 129 (subgeneric assignment).

Genus ANDRENA Subgenus SCAPHANDRENA Lanham

Andrena subg. Scaphaiidrena Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 200.

Type-species: Andrena montrosensis Viereck and Cockerell. Orig. desig.




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Andrena subg. Elandrena Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 203.

Type-species: Andrena atnplificata Cockerell. Orig. desig.
Andrena subg. Truncandrena Warncke, 1968. Est. Mus. Zool. Coimbra Univ., Mem. 307:
46.

Type-species: Andrena truncatilabris Morawitz. Orig. desig.

Revision: Ribble, 1974. Amer. Ent. Soc. Trans. 100: 101-189, 99 figs., 5 tables (N. Amer.
spp.).

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 296 (tax. characters). —Ribble,
1973. Ent. Soc. Amer., Ann. 66: 1281-1286 (tax. status of type species).

Andrena albiculta Viereck
Calif.

Andrena albiculta Viereck, 1917. Amer. Ent. Soc, Trans. 43: 366. ♀.

Taxonomy: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 173-174 (tax. status).

Andrena arabis Robertson
N. Y. to N. C, west to Mich, and 111. Pollen: Polylectic, often collected on

early spring flowering plants of the family Cruciferae; visitation records include Arabis
laevigata, Barbarea vulgaris, Brassica, Buxus, Capsella bursa-pastoris, Cardamine
bulbosa, Claytonia virginica, Dentaria diphylla, D. lanciniata, Erigenia bulbosa,
Erythronium americanum, Prunus, Salix microphylla. Spiraea, Stellaria media,
Taraxcum officinale, Tussilago farfara. Hybridizes with A. scurra Viereck and A.
capricomis Casad and Cockerell.

Andrena arabis Robertson, 1897. Acad. Sci. St. Louis, Trans. 7: 334. ♀.

Andrena (Pterandrena) brassicae Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 142,
figs. 24, 25, 28. ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 191. 9 (key). —Viereck, 1907. Ent.
News 18: 282. 9 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 251-252, figs.
54-56, table 7 (redescription). —Ribble, 1973. Ent. Soc. Amer., Ann. 66: 1281-1286
(hybridization). —Ribble, 1974. Amer. Ent. Soc, Trans. 100: 115-118, figs. 1, 14-18
(redescription, synonymy).

Biology: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 118, table 1 (floral relationships).

Andrena bruneri Viereck and Cockerell
Nebr., Wyo., Colo., Nev. Pollen: Unknown, but visits flowers of
Astragalus, Erysimum.
Andrena bruneri Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc 48: 15. ♀.
Andrena laramiensis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 22, 23. ♂.
Andrena viridibasis Cockerell, 1930. N. Y. Ent. Soc, Jour. 37: 444. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 9. — Timberlake, 1951. In Linsley, In

Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1059 (synonymy).

—Ribble, 1974. Amer. Ent. Soc, Trans. 100: 161-164, figs. 1, 80-84 (redescription,

synonymy).

Andrena capricornis Casad and Cockerell
Tex. to south. Calif. Parasite: Stylops sp. Pollen: Apparently
polylectic, often collected at the flowers of the family Cruciferae; visitation records
include Actiiiea, Alyssum maritimum, Brassica oleracea, Descourainia sophia,
Lesquerella gordonii, Physaria, Prunus, Salvia. Hybridizes with A. scurra Viereck and
A. arabis Robertson.

Andrena capricornis Casad and Cockerell, 1896. Canad. Ent. 28: 182. ♂.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 48. 6 (key). —Cockerell, 1899.

Ent. News 10: 255. 3 (key). —Ribble, 1973. Ent. Soc Amer., Ann. 66: 1281-1286

(hybridization). —Ribble, 1974. Amer. Ent. Soc, Trans. 100: 120-122, figs. 1, 19-23

(redescription).

Andrena chapmanae Viereck
Calif., Oreg., Idaho, Nev. and Utah. Pollen: Unknown, but visits flowers
of Ceanothus, Chaniaebatia foliolosa, Clarkia dudleyana, Cryptantha ambigua, C.
flaccida, C. intermedia, C. micrantha var. lepida, Descourainia sophia, Gilia capitata,
G. latiflora var. cana, Hackelia patens, Lupinus, Mentzelia, Nemophila maculata,
Phacelia linearis, Plagiobothrys canescens, P. nothofulvus, Prunus andersonii, Salix,
Senecio canus.

Andrena {Andrena) chapmanae Viereck, 1904. Canad. Ent. 36: 191. ♀.

Andrena yosemitensis Cockerell, 1924. Pan-Pacific Ent. 1: 61. ♀.




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Taxonomy: Ribble, 1974. Amer. Ent. Sec, Trans. 100: 166-170, figs. 1, 8, 90-94 (redescription).

Biology: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 168, table 5 (floral relationships).

Andrena cruciferarum Ribble
Calif. Pollen: Unknown, but visits flowers o{ Amsinckia, Brassica,
Capsella bursa-pastoris, Phacelia, Sisymbrium irio.
Andrena {Scaphandrena) cruciferarum Ribble, 1974. Amer. Ent. Soc, Trans. 100: 128, figs.
1-2,29-33. ♀, ♂.

Andrena gordoni Ribble
Northeast. Calif., Oreg., Wash., Wyo.

Andrena (Scaphandrena) gordoni Ribble, 1974. Amer. Ent. Soc, Trans. 100: 134, figs. 1-2,
39-43. ♀, ♂.

Andrena hicksi Cockerell
Colo., Wyo., Utah and Nev. (Ruby Mts.). Parasite: Stylops sp.
Andrena hicksi Cockerell, 1925. Ann. and Mag. Nat. Hist. (9) 16: 628. ♀.
Andrena unicula Cockerell, 1934. Pan-Pacific Ent. 9: 155. ♂.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc (4) 23: 265. 9 (key). — Lanham, 1941. Ent.
Soc. Amer., Ann. 34: 705, 707. 9, 6 (key). —Lanham, 1941. Ent. Soc, Amer., Ann. 34: 707.
6 (key, as unicula). —Ribble, 1974. Amer. Ent. Soc, Trans. 100: 164-166, figs. 1, 6, 85-89
(redescription, synonymy).

Biology: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 165 (stylopization).

Andrena kaibabensis Ribble
Ariz., Utah.

Andrena {Scaphandrena) kaibabensis Ribble, 1974. Amer. Ent. Soc, Trans. 100: 160, figs.
1,8. ♀.

Andrena lomatii Ribble
Calif. Pollen: Unknown, but visits flowers of Lomatium dasycarpum, Rhus
trilobata, Salvia sonomensis, Sanicula nevadensis.
Andrena (Scaphandrena) lomatii Ribble, 1974. Amer. Ent. Soc, Trans. 100: 131, figs. 1, 5,
34-38. ♀, ♂.

Andrena mackieae Cockerell
South. Oreg., Calif., Nev., Utah, Ariz.; Mexico (Baja California). Pollen:
Unknown, but visits flowers oi Aynsinckia, Arctostaphylos, Brassica, Ceanothus
cordulatus, C. crassifolius, C. cuneatus, C. integerrimus, C. verrucosus, Cryptantha,
Phacelia distayis, Plagiobothrys, Pninus virginiana, Rhamyms crocea, Rhus diversiloba,
Salix lasiolepis.
Andrena mackieae Cockerell, 1937. Amer. Mus. Novitates 899: 3. ♀.
Andrena (,i Scaphandrena) mackiae(\) Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 201.

Taxonomy: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 144-149, figs. 1, 6, 59-63 (redescription).

Biology: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 147, table 3 (floral relationships).

Andrena merriami Cockerell
B. C. to Calif., east to Mont., Nebr. Colo, and N. Mex. Pollen: Unknown,
but visits flowers of Acer, Astragalus, Cardaria draba, Claytonia, Daucus caroia,
Fraxinus, Gilia capitata, Hackelia patens, Lappula floribunda, Leptotaenia,
Lithophragma affinis, Lomatium ambiguum var. leptocarpuni, L. dissectum, L.
orientale, L. iritematum, Phacelia, Pninus demissa, P. virginiana, var. melanocarpa,
P. subcordata, Pyrus, Ranunculus, Salix, Taraxacum officinale, Thelypodium.

Andrena merriami Cockerell, 1901. Psyche 9: 284. ♀.

Andrena pullmani Viereck, 1904. Canad. Ent. 36: 191, 195. ♀, ♂.

Andrena amplificata Cockerell, 1910. Canad. Ent. 42: 368. ♀, ♂.

Andrena fulvinigra Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 42. ♀.

Taxonomy: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 150-155, figs. 1, 3, 64-69 (redescription,
synonymy).

Biology: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 153, table 4 (floral relationships).

Andrena mohavensis Ribble
South. Calif. (Mohave Desert). Pollen: Unknown, but visits flowers of
Lepidium flavum, Phacelia, Salix exigua.
Andrena {Scaphandrena) mohavensis Ribble, 1974. Amer. Ent. Soc, Trans. 100: 126, figs.
1,2,24-28. ♀, ♂.

Andrena plana Viereck
Oreg., Calif. Pollen: Apparently oligolectic on Trifolium including T.

involucratum, T. microcephalum , T. pratense, T. repens, T. tridentatum, T. variegatum,
but visits other flowers presumably for nectar including Brassica, Chamaebatia




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foliolosa, Cryptantha, Gilia, Lasthenia chrysostoma, Layia platygiossa var. breviseta,
Limnanthes douglasii, Nernophila, Orthocarpus faiicibarbatus, Plagiobothrys
nothofulvus, Ranunculus, Rhus trilobata, Sisyrinchium bellum, Vicia.
Andrena plana Viereck, 1904. Canad. Ent. 36: 193. ♀.

Taxonomy: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 141-144, figs. 1, 7, 54-58 (redescription).

Andrena primulifrons Casad
Tex. to Ariz, and south. Nev. (Hoover Dam). Pollen: Possibly an oligolege
of Lesquerella including L. densiflora, L. gordonii, L. ludoviciana, but visits other
flowers including Descourainia pinnata, Ditkyraea wislizenii, Larrea tridentata,
Lobularia, Machaeranthera tanacetifolia, Oreocarya, Phacelia crenulata, Physaria,
Prunus, Sisymbriuvi.
Andrena primulifrons Casad, 1896. Canad. Ent. 28: 183.
Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 48, 49. S, 9 (key). —Ribble,
1968. Nebr. Univ. State Mus., Bui. 8: 328-332, figs. 85-87, map 10, table 11 (redescription).
—Ribble, 1974. Amer. Ent. Soc, Trans. 100: 104, fig. 1 (subgeneric position).

Andrena santaclarae Ribble
Calif. Pollen: Unknown, but males visit flowers of Cryptantha, Lasthenia,
Nemophila, Prunus subcordata.
Andrena (Scapha7idrena) santaclarae Ribble, 1974. Amer. Ent. Soc, Trans. 100: 136, figs.
1,4,44-48. ♀, ♂.

Andrena scurra
***authority mismatch
Viereck. B. C. to Calif., east to Idaho and north. Utah. Pollen: Unknown, but visits

flowers of Achillea lanulosa, Baccharis, Brassica campestris, Capsella bursa-pastoris,
Cardaria draba, Ceanothus greggii, C. pahneri, Cosmos, Cotoneaster, Descourainia
Sophia, Lepidium, Lomatium dissectum, L. tritematum, Medicago sativa, Prunus
subcordata, Salix, Scrophularia califomica. Sisymbrium altissimum. Spiraea,
Taraxacum, Thlaspi arvense. Hybridizes with A. arabis Robertson and A. capricomis
Casad and Cockerell in the Great Basin area of the western United States and in the
Rocky Mountain area.
Andrena scurra Viereck, 1904. Canad. Ent. 36: 193, 195. ♀, ♂.

Taxonomy: Ribble, 1973. Ent. Soc. Amer., Ann. 66: 1281-1286 (hybridization). —Ribble, 1974.

Amer. Ent. Soc, Trans. 100: 110-115, figs. 1, 9-13 (redescription).

Andrena scurra
***authority mismatch
x arabis x capricomis Hybrids. Idaho, Wyo., Colo., Utah, Nev. and along the east slope
of the Rocky Mountains from north. N. Mex. to south. Alta; Calif. (Seven Oaks) and
Ariz. (Fredonia). Pollen: Apparently polylectic, exhibits a preference for the flowers of
the family Cruciferae, but visits a variety of flowers including Agoseris, Astragalus,
Brassica, Cardaria draba, Cleome semdata, Euphorbia, Lappula floribunda, Lepidium
virginicum var. medium, Lesquerella ovalifolia, Oxytropis, Philadelphus, Physaria
didymocarpa, Prunus, Ribes, Senecio canus. Sisymbrium altissimum, S. pinnatum,
Taraxacum officinale.

Andrena sieverti Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 436. ♀.

Andrena lappulae Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 437. ♂.

Andrena ellisiae Cockerell, 1914. Ann. and Mag. Nat. Hist. (8) 14: 9. ♀.

A7idrena montrosensis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc 48: 48. ♀.

Andrena speculifera Cockerell, 1930 (1929). N. Y. Ent. Soc, Jour. 37: 444. ♀.

Andre7ia physariae Cockerell, 1934. Pan-Pacific Ent. 9: 154. ♀.

Andrena sieverti var. opacicauda Cockerell, 1936. Amer. Mus. Novitates 831: 1. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 13. 9 (key, as sieverti). — Lanham,
1941. Ent. Soc Amer., Ann. 34: 706. 9 (key, as ellisiae). —Lanham, 1941. Ent. Soc. Amer.,
Ann. 34: 706. 9 (key, as physariae). —Ribble, 1973. Ent. Soc Amer., Ann. 66: 1281-1286
(hybridization). —Ribble, 1974. Amer. Ent. Soc, Trans. 100: 122-126 (synonymy).

Biology: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 124, table 2 (floral relationships,
stylopization).

Andrena shoshoni Ribble
Wyo. (Big Horn and Foxpark).

Andrena (S cap hand rena) shosho7ii Ribble, 1974. Amer. Ent. Soc, Trans. 100: 172, figs. 1,
8. ♀.




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Andrena sladeni Viereck
B. C. to Wyo. and Nev. Pollen: Unknown, but visits flowers of Taraxacum

officinale.
Andrena (Andrena) sladeni Viereck, 1924. Canad. Ent. 56: 239. ♀.

Taxonomy: Cockerell, 1932. Canad. Ent. 64: 286 (key). — Ribble, 1974. Amer. Ent. See, Trans.
100: 155-158, figs. 1, 4, 70-74 (redescription).

Andrena tildeni Ribble
Calif., Ariz. Pollen: Unknown, but visits flowers of Cryptantha intermedia,
Salix argophylla, S. exigiia.
Andrena (Scaphandrena) tildeni Ribble, 1974. Amer. Ent. Soc, Trans. 100: 139, figs. 1, 3,
49-53. ♂.

Andrena trapezoidea Viereck
Tex., Okla. Pollen: Unknown, but visits flowers of Descourainia pinnata,
Lesquerella derisiflora, L. gordonii, L. gracilis, L. recurvata, Rhus microphylla.
Andrena (Scrapter) trapezoidea Viereck, 1917. Amer. Ent. Soc, Trans. 43: 402. ♀.

Taxonomy: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 328-332, figs. 85-87, map 10, table 11
(redescription). —Ribble, 1974. Amer. Ent. Soc, Trans. 100: 104, fig. 1 (subgeneric
position).

Andrena vestali Cockerell
Colo., Nebr. Pollen: Unknown, but visits flowers of Viola.
Andrena vestali Cockerell, 1913. Ann. and Mag. Nat. Hist. (8) 11: 64. ♂.
Andrena vestali var. dolichocera Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48:
19. ♂.

Taxonomy: Lanham, 1941. Ent. Soc. Amer., Ann. 34: 707. 6 (key). —Ribble, 1974. Amer. Ent.
Soc, Trans. 100: 170-172, figs. 1, 8, 95-99 (redescription).

Andrena walleyi Cockerell
B. C. to Calif. (Alpine Co.), east to Idaho and Utah. Pollen: Unknown, but
visits flowers of Cardaria draba.
Andrena walleyi Cockerell, 1932. Canad. Ent. 64: 286. ♀.

Taxonomy: Ribble, 1974. Amer. Ent. Soc, Trans. 100: 158, figs. 1, 5, 75-79 (redescription).

Genus ANDRENA Subgenus SCOLIANDRENA Lanham

Andrena subg. Scoliandrena Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 223.

Type-species: Andrena osmioides Cockerell. Monotypic and orig. desig.

The species of this subgenus chiefly visit the flowers of Cryptantha upon which the females
are apparently solely dependent for pollen.

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 283, 291, 313 (tax. characters).

Andrena cryptanthae Timberlake
South. Calif. Pollen: Oligolege of Cryptantha including C. barbigera,
C. intermedia, but visits other flowers for nectar.
Andrena (Scoliandrena) cryptanthae Timberlake, 1951. U. S. Natl. Mus. Proc. 101: 401. ♀.
6.

Andrena osmioides
***authority mismatch
benitonis Lanham. North. Calif. Pollen: Oligolege of Cryptantha.

Andrena osmioides benitonis Lanham, 1949. Pan-Pacific Ent. 25: 148. 9,

Andrena osmioides
***authority mismatch
osmioides Cockerell. South. Calif. Pollen: Oligolege of Cryptantha including C.

intermedia, but visits other flowers for nectar including Plagiobothrys califomica, P.
nothofulvus.
Andrena osmioides Cockerell, 1916. Pomona Jour. Ent. Zool. 8: 45. ♂.

Genus ANDRENA Subgenus SCRAPTEROPSIS Viereck

Atidrena subg. Scrapteropsis Viereck, 1922. Boston Soc. Nat. Hist., Occas. Papers 5: 42.

Type-species: Andrena (Scrapteropsis) fenningeri Viereck. Monotypic.
Andrena subg. Mimandrena Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 217.

Type-species: Andrena imitatrix Cresson. Orig. desig.

Revision: LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 441-504, figs. 1-53, 1 table (N. Amer.
spp.).

Taxonomy: Warneke, 1968. Coimbra Univ. Est. Mus. Zool, Mem. 307: 68-69 (synonymy).
—LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 235, 237-240 (phylogeny, tax. status).




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Andrena alamonis Viereck
West. Tex., N. Mex., Ariz. Pollen: Unknown, but visits flowers of
Lesquerella.
A7idrena (Scrapter) alamonis Viereck, 1917. Acad. Nat. Sci. Phila., Proc. 68: 599. ♀.

Andrena alleghaniensis Viereck
Ont. and Maine to S. C. west to Sask. and Colo. Pollen: Apparently
polylectic, visits flowers of Amelanchier, Barbarea, Brasska, Chrysanthemum
leucanthemum, Conius, Crataegus, Euphorbia cyparissias, Fragaria vesca, F.
virgiyiiana, Ledum, Melilotus alba, Prunus, Ranunculus, Rubus idaeus, Salix,
Viburnum opuius.
Andrena (Trachandreva) alleghaniensis Viereck, 1907. Ent. News 18: 280, 286. ♀.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 177 (redescription). — Knerer
and Atwood, 1963. Canad. Ent. 95: 583, figs. 1, 3. 6. —Knerer and Atwood, 1964. Ent. See.
Ontario, Proc. 94: 47-48. -LaBerge, 1971. Ent. Soc. Amer., Trans. 97: 474-479, figs. 1, 5,
27-31 (redescription).

Andrena angusticrus LaBerge
Calif. Pollen: Unknown, but visits flowers of Lasthenia chrysostoma,
Platystemon califomicus.
Andrena (Scrapteropsis) angusticrus LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 499, figs.
1,6. ♀.

Andrena aquila LaBerge
Calif. (Fresno County). Pollen: Unknown, but visits flowers of Platystemon
califomicus.
Andrena (Scrapteropsis) aquila LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 502, figs. 1, 6.
9.

Andrena atlantica Mitchell
N. J. to Fla., west to Miss. Pollen: Apparently polylectic, visits flowers of
Crataegus, Diospyros virginiana, Fragaria, Ilex glabra, Ligustrum, Linaria, Lyssa
sylvatica, Polycodimn, Pyracantha, Rubus, Salix nigra, Senecio, Spiraea,
Toxicodendron.
Andrena {Trachandrena) atlantica Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
178, figs. 35-37. ♀, ♂.

Taxonomy: LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 479-481, figs. 1, 5 (redescription).

Andrena biareola LaBerge
Calif. Pollen: Unknown, but visits flowers of Cryptantha, Meconella,
Platystemon califomicus.
Andrena {Scrapteropsis) biareola LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 494, figs. 1, 3.
9,6.

Andrena buccata LaBerge
Calif. Pollen: Unknown, but visits flowers of Blennosperma nana,
Lasthenia, Platystemon.
Andrena (Scrapteropsis) buccata LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 497, figs. 1, 6,
49-53. ♀, ♂.

Andrena daeckei Viereck
N. J., Pa.

Andrena (Trachandrena) daeckei Viereck, 1907. Ent. News 18: 280, 285, 286. ♀, ♂.

Taxonomy: LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 486-489, figs. 1, 4, 37-41 (redescription,
distribution).

Andrena fenningeri Viereck
Mass. to Ga., west to III, Tenn. and Tex. Pollen: Polylectic, with a strong
preference for Salix including S. humilis, S. sericea, S. tristis and a secondary
preference for Prunus including P. americanus, P. persica, but visits other flowers
including Acer rubrum, Alsijie media, Amelanchier, Aronia, Pyrus mains.
A7idre7ia (Scrapteropsis) fenningeri Viereck, 1922. Boston Soc. Nat. Hist., Occas. Papers 5:

42, figs. 9,6.
Andrena (Trachandrena) indianensis Cockerell, 1929. Ent. Soc. Amer., Ann. 22: 757. ♀.
Andrena (mimandrena) vema Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 206,
figs. 37, 38. ♂.
Taxonomy: LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 465-470, figs. 1, 6, 17-21 (redescription,
synonymy).

Andrena flaminea LaBerge
Tex. (Beeville and Edinburg).

Andrena (Scrapteropsis) flaminea LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 472, fig. 1.
9.




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ilicis Mitchell. N. Y. to north. Fla., west to Kans. and Tex. Pollen: Unknown, but visits flowers
o{ Batodendron, Castanea pumila, Celastrus, Crataegus, Diospyros virginiana. Ilex,
Pninus, Rhiis glabra, Salix.
Andre7ia (Trachandreiia) ilicis Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 186,
fig. 35. ♀.

Taxonomy: LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 459-463, figs. 1, 3, 12-16

(redescription).

Andrena imitatrix Cresson
Que. to Fla. west to Alta., Mont., Utah and Ariz. Pollen: Polylectic, although
exhibits a strong preference for the flowers of the Rosaceae and Salicaceae; visitation
records include Acer negundo, Alsine media, Amelanchier arborea, A. canadensis,
Amorpha fruticosa, Angelica atropurpiirea, Antennaria dioicimt. Astragalus, Barbarea
vulgaris. Benzoin aestivale. Cassia fasciculata, Ceanothus ovatus, Claytonia, Conius,
Crataegus mollis, C. occidentalis, Dentaria laciniata, Descourainia interynedia,
Diospyros virginiana, Erytkronium, Euphorbia, Exochoroa racemosa, Forsythia,
Grossularia, Heracleum, Ilex, Laurocerasus caroliniana, Lepargyraea canadensis,
Lomatium daucifolium, L. foeniculaceum, Lonicera morrowi, Medicago sativa,
Pastinaca sativa, Paeonia, Phacelia, Physocarpus opulifolius, Populus, Prunus
americana, P. gracile, P. pennsylvanica, P. serotina, P. virginiana, Ptelea, Pyracantha,
Pyrus mains, Rhamnus utilis, Rhus aromatica, R. canadensis, Ribes gracile, Rosa,
Rubus, Salix amygdaloides, S. babylonica, S. bebbiana, S. discolor, S. longifolia, S.
nigra, Senecio, Spiraea anaicus, S. thunbergi, S. vanhouttei, Tamarix gallica.
Taraxacum officinale, Thlaspi arvense, Vaccinium, Viburnum deniatum, V.
prunifolium, Zygadenia, Zizia.

Andrena imitatrix Cresson, 1872. Amer. Ent. Soc, Trans. 4: 258. ♀.

Andrena claytoniae Robertson, 1891. Amer. Ent. Soc, Trans. 18: 59. ♀, ♂.

Trachandrena albofoveata Graenicher, 1903. Canad. Ent. 35: 166. ♀.

Andrena johnsoniana Cockerell, 1906. Ann. and Mag. Nat. Hist. (7) 17: 224. ♀.

Andrena {Trachandrena) crataegiphila Viereck and Cockerell, 1914. U. S. Natl. Mus.,
Proc48:7. ♂.

Andrena (Trachandrena) titusi Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc 48: 12.
9.

Andrena (Scrapter) imitatrix var. profunda Viereck, 1917. Amer. Ent. Soc, Trans. 43: 398.
9.

Andrena (Trachandrena) tardula Cockerell, 1930. N. Y. Ent. Soc, Jour. 37: 447. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 190. 9, 6 (key). —Cockerell, 1929.
Ent. Soc. Amer., Ann. 22: 755 (tax. characters). —Cockerell, 1931. Canad. Ent. 63: 22. 9 (as
johnsoniana). —Cockerell, 1931. Amer. Mus. Novitates 458: 11. 9 {zs johnsoniana).
— Lanham and Timberlake, 1951. In Muesebeck, Krombein and Townes, U. S. Dept. Agr.,
Agr. Monog. 2: 1068 (synonymy). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
205-206, fig. 37-39, table 5 (redescription, synonymy). —Yager and Rozen, 1966. Amer.
Mus. Novitates 2265: 6 (pupa). —LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 451-459, figs.
1-2, 7-11 (redescription, synonymy).

Biology: Rau, 1922. Acad. Sci. St. Louis, Trans. 24: 34 (nesting habits). —Clements and Long,
1923. Carnegie Inst. Wash., Pub. 336: 249 (ecology). —LaBerge, 1971. Amer. Ent. Soc,
Trans. 97: 457-459, table 1 (floral relationships).

Andrena kalmiae Atwood
N. S., Que. and Ont. south to Mass. and Conn. Pollen: Apparently an

oligolege of Kalmia including K. angustifolia, but visits other flowers presumably for
nectar including Ledum, Lyonia ligustrina, Vaccinium.
Andrena kalmiae Atwood, 1934. Canad. Jour. Res. 10: 210, fig. 9,6.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 186, fig. 35, table 5
(redescription). —LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 489-491, figs. 1, 4
(redescription, distribution).

Andrena morrisonella Viereck
Ont. to Fla., west to 111. and Ala. Pollen: Unknown, but visits flowers of
Amelanchier, Crataegus, Gypsophila, Ilex, Mains, Populus, Prunus, Ranunculus acris,
Rubus, Salix.
Andrena (Scrapter) morrisonella Viereck, 1917. Amer. Ent. Soc, Trans. 43: 399. ♀.




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Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 192, fig. 35, table 5

(redescription). — LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 463-465, fig. 1 (redescription).

Andrena phenax Cockerell
South. Calif.

Andrena phenax Cockerell, 1898. Amer. Ent. Soc. Trans. 25: 188. ♀.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 129 (tax. status). —Thorp, 1969.
Calif. Univ. Pubs. Ent. 52: 121-123 (tax. status). —LaBerge, 1971. Amer. Ent. Soc, Trans.
97: 503-504, fig. 1 (redescription).

Andrena rubi Mitchell
N. C. to Ga., west to Tex., Okla., Kans. and Nebr. Pollen: Unknown, but visits

flowers of Aronia, Crataegus, Descourainia intermedia, D. pinnata, Prunus americana,
P. gracilis, Ritbiis, Salix nigra, Spiraea.
Andrena {Trachandrena) rubi Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 198, fig.
35. ♀.

Andrena stipator LaBerge
Calif. Pollen: Unknown, but visits flowers of Cryptantha, Meconella,
Platystemon califomicus.
Andrena (Scrapteropsis) stipator LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 491, figs. 1, 3,
44-48. ♀, ♂.

Andrena unicostata LaBerge
Tex., Okla. Pollen: Unknown, but visits flowers of Salix.

Andrena (Scrapteropsis) unicostata LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 481, figs. 1,
5,32-36. ♀, ♂.

Genus ANDRENA Subgenus SIMANDRENA Perez

Simandrena Perez, 1890. Soc. Linn. Bordeaux, Actes. 44: 174.

Type-species: Andre^ia propinqua Schenck. Desig. by Hedicke, 1933.
Andrena subg. Platandrena Viereck, 1924. Canad. Ent. 56: 21.

Type-species: Andrena vestita Provancher. Monotypic and orig. desig. (,=Andrena

Andrena nasonii Robertson
)
Andrena subg. Stenandrena Timberlake, 1949. In Lanham, Calif. Univ. Pubs. Ent. 8: 213.

Type-species: Andrena pallidifovea (Viereck). Orig. desig.

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 285, 286, 293, 304 (tax. characters,

synonymy).

Andrena angustitarsata Viereck
Wash, to Calif. Pollen: Apparently polylectic, visits a wide variety of
flowers including Achillea lanulosa, A. millefolium, Arenaria douglasii, Ceanothus
cuneatus, C. greggii, C. integerrimus, C. leucodermis, C. orcuttii, C. palmeri, C.
soredatus, Chamaebatia foliolosa, Claytonia spatulata, Eriodictyon californicum,
Eriophyllum confertiflorum, Gilia capitata, Grossularia califomica, Horkelia bolanderi,
Lasthenia chrysostoma, Linanthus aureus, Lomatium utriculatum, Paeonia califomica,
Plagiobothrys nothofulvus, Potentilla glandidosa, Prunus ilicifolia. Ranunculus
califomicus, Rhamnus califomicus, R. crocea, Rhus diversiloba, Salix laevigata, S.
lasiayidra, S. lasiolepis, Sanicula bipinnatifida, S. nevadensis, S. tuberosa, Saxifraga
integrifolia, Swertia parryi, Thysanocarpus curvipes.

Andrena angustitarsata Viereck, 1904. Canad. Ent. 36: 189, 196. ♀.

Pterandrena nudiscopa Viereck, 1904. Canad. Ent. 36: 227, 228. ♀.

Andrena essigi Timberlake
South. Calif. Pollen: Unknown, but visits flowers of Cryptantha intermedia,
Lasthenia aristata, L. gracilis, Layia elegans, Rhamnus crocea, Rhus trilobata,
Sisymbrium irio.

Andrena {Stenandrena) essigi Timberlake, 1951. U. S. Natl. Mus., Proc 101: 384. ♀, ♂.

Andrena friesei Viereck
Calif. Ecology: Unknown, but visits flowers oi Amsinckia douglasiana,
Brassica, Calandrinia menziesii, Eschscholzia califomica, Lasthenia, Raphanus
sativus.

Andrena {Andrena) friesei Viereck, 1917. Acad. Nat. Sci. Phila., Proc. 68: 558. ♀.

Andrena huardi Viereck
Calif. Pollen: Possibly an oligolege of Salix including S. argophylla, S.

gooddingii, S. hindsiana, S. laevigata, S. lasiolepis, S. nigra, but visits other flowers
possibly for nectar only including Alyssum maritimum, Sambucus.

Andrena {Andrena) angustitarsata huardi Viereck, 1917. Amer. Ent. Soc, Trans. 43: 368.
$.




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Andrena hypoleuca Cockerell
South. Calif. (Catalina Island).

Andreiia hypoleuca Cockerell, 1939. Pomona Jour. Ent. Zool. 31: 25. ♀.

Andrena nasonii Robertson
Eastern Canada and Minn., south to Ga. and Tex., Colo. Pollen: Apparently
polylectic, visits a wide variety of flowers including Acer, Aynelanchier, Apocynum,
Arabia, Capsella, Chaerophyllum, Claytonia, Crataegus, Forsythia, Fragaria, Hex,
Ligustrum, Polemonium, Polytaenia, Potentilla, Primus, Pyracantha, Ranunculus,
Rhamnus, Rubus, Senecio, Sisymbrium, Spiraea, Taenidia, Toxicodeyidron, Viburnum,
Viola, Zizia.

Andrena vestita Provancher, 1888. Addit. Corr. Faune Ent. Canada, Hym., p. 310. ♂.
Preocc.

Andrena nasonii Robertson, 1895. Amer. Ent. Soc, Trans. 22: 120. ♀.

Andrena harifordensis Cockerell, 1902. Ann. and Mag. Nat. Hist. (7) 9: 103. ♀.

Andrena {Andrena) nasonii var. fulvodorsata Viereck, 1917. Amer. Ent. Soc, Trans. 43:
385. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 190, 192. 9, 6 (key). —Viereck,
1907. Ent. News 18: 283, 285. 9, 6 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui.
141: 244-245, figs. 51-52, 54 (redescription).

Andrena opacella Timberlake
Calif. (Mt. Hamilton Range). Pollen: Unknown, but visits flowers of
Salix.
Ayidrena (Siynandreyia) opacella Timberlake, 1951. U. S. Natl. Mus., Proc 101: 406. ♀.

Andrena opacibasis Cockerell
Calif.

Andreyia {Platandrena) opacibasis Cockerell, 1936. Pan-Pacific Ent. 12: 146. ♀.

Andrena opacissima Cockerell
Idaho, Wash, to Utah and Calif. Pollen: Unknown, but visits flowers of
Ceanothus cordulatus, Cryptantha intermedia, Lomatium, Salix lasiolepis.
Andrena opacissima Cockerell, 1918. Ann. and Mag. Nat. Hist. (9) 1: 165. ♀.

Taxonomy: Cockerell, 1933. Pan-Pacific Ent. 9: 158. 9. —Cockerell, 1936. Ann. and Mag. Nat.
Hist. (10) 18: 632.

Andrena opaciventris Cockerell
Calif. Pollen: Unknown, but visits flowers of Ceanothus crassifolins, C.
integerrimus, C. vestitus, Cryptantha intermedia, Lomatium dasycarpum, Pi-unus,
Ranunculus occidentalis, Rhamnus crocea, Salix lasiolepis, Sanicula tuberosa, S.
bipinnatifida.
Andrena opaciventris Cockerell, 1916. Pomona Jour. Ent. Zool. 8: 47. ♀.

Andrena orthocarpi Cockerell
Calif. Pollen: Unknown, but visits flowers of Achillea millefolium,

Calycadenia multiglandulosa, Ceanothus cuneatus, C. soredatus, Limnanthes douglasii,
Lomatium utriculatum, Orthocarpus densiflorus, 0. lithospermoides. Ranunculus
califomicus, Salix lasiolepis, Phoradendron, Thysanocarpus curvipes.
Andrena {Platandrena) orthocarpi Cockerell, 1936. Pan-Pacific Ent. 12: 147. ♀.

Taxonomy: Lanham, 1947. Pan-Pacific Ent. 23: 72. ♂.

Andrena pallidifovea (Viereck)
B. C. to Calif., Utah. Pollen: Collects pollen from the flowers of
Compositae including Chaenactis, Ericameria cooperi, Eriophyllum, Helianthus
gracilentus, Layia, but visits other flowers for nectar including Gilia capitata,
Lasthenia chrysostoma.
Pterandrena pallidifovea Viereck, 1904. Canad. Ent. 36: 195, 227, 228. ♂, ♀.
Andrena plumifera Cockerell, 1916. Canad. Ent. 48: 393. ♀.

Taxonomy: Timberlake, 1951. In Linsley, In Muesebeck, Krombein and Townes, U. S. Dept.
Agr., Agr. Monog. 2: 1075 (synonymy). —Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 383.
6.

Andrena pensilis Timberlake
Calif. Pollen: Unknown, but visits flowers of Eriodictyon califomicum,
Eschscholzia califomica, Lasthenia chrysostoma, Limnanthes douglasii, Layia
platyglossa, Orthocarpus densiflorus, 0. lithospermoides, Ranunculus califomicus,
Sisyrinchium bellum.
Andrena (Platandrena) pensilis Timberlake, 1938. Pan-Pacific Ent. 14: 27. ♀, ♂.

Andrena runcinatae Cockerell
B. C. and Alta. to Colo., N. Dak.

Andrena runcinatae Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 434. ♀, ♂.




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Andrena uvulariae Mitchell
Md. (Beltsville). Pollen: Unknown, but visits flowers of Uvularia
sessilifolia.
Andrena (^Simandrena) uvulariae Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
245, fig. 54. ♀.

Andrena vexabilis Timberlake
North. Calif. Pollen: Unknown, but visits flowers of Achillea lanulosa,
Agoseris heterophylla, Chaviaebatia foliolosa. Coreopsis maritima, Cryptantha
micraniha, Encelia californica, Eriophyllum confertiflorum, Lasthenia chrysostoma, L.
gracilis, Layia platyglossa, Helianthella californica, Horkelia fusca, Senecio
califomicus, Viguiera, Wyethia angustifoiia.

Andrena (Stenandreyia) vexabilis Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 385. ♀, ♂.

Andrena wheeleri Graenicher
N. S. and Maine to Alta., south to Colo., Minn., Wis. and N. C. Pollen:
Unknown, but visits flowers of Centaurea, Chrysanthemum leucanthemum, Camus,
Rhododendron, Rubus, Sedum acre.

Andrena wheeleri Graenicher, 1904. Ent. News 15: 65. ♀, ♂.

Andrena wheeleri pallidior Cockerell, 1938. Canad. Ent. 70: 7. ♀.

Taxonomy: Atwood, 1934. Canad. Jour. Res. 10: 208, 209. 9, 6 (key). —Mitchell, 1960. N. C.
Agr. Expt. Sta. Tech. Bui. 141: 246-247, figs. 51-52 (redescription, synonymy).

Genus ANDRENA Subgenus TAENIANDRENA Hedicke

Andrena subg. Taeniandrena Hedicke, 1933. Berlin Zool. Mus., Mitt. 19: 219.
Type-species: Melitta ovatula Kirby. Orig. desig.

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 286, 292, 306-307 (tax. characters).

Andrena wilkella (Kirby)
Holarctic; N. S. to Wis., south to Va. and Ohio. Presumably introduced.

Pollen: Apparently polylectic, visits flowers of Achillea, Amorpha, Brassica, Cirsium,
Crataegus, Daucus, Epilobium, Fragaria, Leontodon, Malus, Philadelphus, Pinus,
Primus, Raphanus, Rhododendron, Rhodura, Rubus, Solidago, Stellaria, Taraxacum,
Trifolium, Vaccinium.
Melitta wilkella Kirby, 1802. Monog. Apum Angliae, v. 2, p. 145. ♀, ♂.
Andrena winkleyi Viereck, 1907. Ent. News 18: 283, 285. ♀, ♂.

Taxonomy: Malloch, 1918. Biol. Sec. Wash., Proc. 31: 61. —Atwood, 1934. Canad. Jour. Res.
10: 207, 210. 9, 6 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 250-251,
figs. 52, 54-55 (redescription).

Biology: Brittain, 1933. Canada Dept. Agr., Bui. 162: 94 (floral relationships). —Atwood, 1933.
Canad. Jour. Res. 9: 456 (floral relationships). — BrittAin and Newton, 1933. Canad. Jour.
Res. 9: 341 (flower records). —Brittain and Newton, 1934. Canad. Jour. Res. 10: 261
(flower records).

Genus ANDRENA Subgenus THYSANDRENA Lanham

Andrena subg. Thysandrena Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 213.
Type-species: Andrena Candida Smith. Orig. desig.

Taxonomy: LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 287, 293, 294, 306 (tax. characters).

Andrena argentiscopa Viereck
Calif.

Andrena (Andrena) argentiscopa Viereck, 1917. Amer. Ent. Soc, Trans. 43: 368. ♀.

Andrena beckeri Cockerell
Colo.

Andrena beckeri Cockerell, 1921. Ann. and Mag. Nat. Hist. (9) 7: 211. ♀.

Taxonomy: Cockerell, 1932. Pan-Pacific Ent. 8: 175. —Lanham, 1941. Ent. Soc. Amer., Ann.
34: 706. ♀.

Andrena bisalicis Viereck
N. Dak. to New England states, south to Ga. Parasite: Nomada sp. Pollen:

Unknown, but visits flowers of Amelanchier, Aronia, Brassica, Crataegus, Ilex, Prunus,
Salix.
Andrena salicis Robertson, 1891. Amer. Ent. Soc, Trans. 18: 53. ♀, ♂. Preocc.
Ajidrena bisalicis Viereck, 1908. Ent. News 19: 42. N. name.
Andrena (Andrena) adelae Viereck, 1922. Boston Soc. Nat. Hist., Occas. Papers 5: 37. ♀.




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Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 191, 192. 9, 6 (key). —Mitchell,
1958. In Krombein, U. S. Dept. Agr., Agr. Monog 2, First Suppl. p. 212 (synonymy).
—Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 219-220, figs. 42-44, table 6
(redescription). —Yager and Rozen, 1966. Amer. Mus. Novitates 2265: 5, figs. 1-2 (pupa).

Biology: Rozen, 1966. Amer. Mus. Novitates 2244: 26 (parasite).

Andrena brevipalpis Cockerell
Colo.

Andrena brevipalpis Cockerell, 1930. Ann. and Mag. Nat. Hist. (10) 5: 109. ♀.

Taxonomy: Cockerell, 1932. Pan-Pacific Ent. 8: 175. 9. —Cockerell, 1937. Amer. Mus.
Novitates 899: 4. ♀.

Andrena campanulae
***authority mismatch
Viereck and Cockerell. Colo., Nebr.

Andrena campanulae Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 43: 38, 43. ♂, ♀.

Taxonomy: Cockerell, 1932. Pan-Pacific Ent. 8: 177. ♀, ♂.
Candida Smith. B. C. to south. Calif., Colo, and Utah; Mexico (Baja California). Pollen:

Polylectic, visits a wide variety of flowers during both the first and second broods
including Acer grandidentatum, A. platynoides, Adenostoma fasciculatum, Alyssiim
maritimmn, Amelanchier, Arctostaphylos glauca, Baccharis glutinosa, B. viminea,
Brassica alba, B. adpressa, B. arvensis, B. campestris, B. nigra, Calandrinia menziesii,
Ceanothus eordulatus, C. crassifolius, C. cuneatus, C. integerrimus, C. orcuttii, C.

. palmeri, C. verrucosus, Chamaebatia foliolosa, Conium maculatum, Cryptantha
flaccida, C. lepida, Cymoptenis longipes, Daucus carota, Eriogonum fasciculatum, E.f.
polifolii, Eriophyllum tridactyllum, Gnaphalium bicolor, Isomeris arborea, Lomaiium
dasycarpum, L. grayi, Ligusticum vulgare, Lupinus paynei, Mains pumila, Medicago
sativa, Melilotus alba, Nemophila menziesii, Phacelia affinis, P. distayis, P. minor, P.
ramosissima, Plagiobotkrys tenelius. Primus f rem ontii, P. ilicifolia, P. lyonii, P.
subcordata. Ranunculus califomicus, Rhamnus califomica, R. crocea, R. ilicifolia,
Salix argophylla, S. discolor, S. exigua, S. hindsiana, S. laevigata, S. lasiolepis, S. nigra.
Salvia mellifera, Sambucus caerulea, Sisymbrium irio, S. officinale.

Andrena Candida Smith, 1879. Descr. New Species Hym. Brit. Mus., p. 56. ♀.

^Andrena (Andrena) Candida tranioserica Viereck, 1917. Acad. Nat. Sci. Phila., Proc. 68:
533. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 155 (type). —Cockerell, 1906. Psyche
13: 10 (type). — Lanham, 1941. Ent. Soc. Amer., Ann. 34: 706, 708. 9, 3 (key). — Youssef
and Bohart, 1968. Kans. Ent. Soc, Jour. 41: 450-455, figs. 2-7, 11-22 (immature stages).

Biology: Linsley, 1937. Brooklyn Ent. Soc, Bui. 32: 127 (floral relationships, spring and early
summer broods). —Youssef and Bohart, 1968. Kans. Ent. Soc, Jour. 41: 442-455, 22 figs,
(nest architecture, life history, floral relationships).

Andrena ceanothina Cockerell
Calif Pollen: Unknown, but visits flowers of Arctostaphylos, Ceanothus
crassifolius, Ribes indecorum.
Andrena ceanothina Cockerell, 1936. Pan-Pacific Ent. 12: 138. ♀.

Andrena cercocarpi Cockerell
Calif. Pollen: Unknown, but visits flowers of Cercocarpus.
Andrena (Platandrena) cercocarpi Cockerell, 1936. Pan-Pacific Ent. 12: 140. ♀.

Taxonomy: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 332 (subgeneric position).

Andrena chlorinella Viereck
Oreg.

Andrena chlorinella Viereck, 1904. Canad. Ent. 35: 189. ♀.
Andrena xanthostigma Viereck, 1904. Canad. Ent. 35: 193. ♀.

Taxonomy: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 333 (subgeneric position).

Andrena claremonti Viereck
Calif.

Andrena claremonti Viereck, 1926. Pomona Jour. Ent. Zool. 18: 2. ♂.

Andrena coloradina Viereck and Cockerell
Colo.

Andrena coloradina Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 53, 56. ♀.

Taxonomy: Cockerell, 1933. Pan-Pacific Ent. 9: 154. 9 (key).

Andrena fulvihirta Viereck and Cockerell
B. C. to Colo, and Calif

Andrena fulvihirta Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 54. ♀.




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Andrena washingtoni vianitouensis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc.
48: 31. ♂.

Taxonomy: Timberlake, 1951. In Linsley, In Muesebeck, Krombein and Townes, U. S. Dept.
Agr., Agr. Monog. 2: 1066 (synonymy).

Andrena geranii Robertson
New England states and Va., west to Minn, and Colo. Pollen: Unknown,
but visits flowers of Arabis, Blephiiia, Geranium, Hydrophylhim, Osmorrhiza,
Polemonium.
Andrena geranii Robertson, 1891. Amer. Ent. Soc, Trans. 18: 54. ♀, ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 190, 192. 9, i (key). —Mitchell,
1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 221-222, figs. 44-45, table 6 (redescription).

Andrena haroldi Timberlake
Calif. Pollen: Unknown, but visits flowers of Lasthenia.

Andrena (Micrandrena) haroldi Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 398. ♀, ♂.

Taxonomy: Ribble, 1968. Nebr. Univ. State Mus., Bui. 8: 333 (subgeneric position).

Andrena knuthiana Cockerell
Oreg., Calif. Pollen: Unknown, but visits flowers of Ceanothus

cordulatus, Cryptantha lepida, Rhamnus califoniica, Salix, Sanibucus velutina.
Andrena knHthia7ia Cockerell, 1906. Ztschr. System. Hym. Dipt. 1: 80. ♀, ♂.

Taxonomy: Cockerell, 1901. Ent. News 12: 74. ♀.

Andrena lata Viereck
N. S. to Minn., south to Ga. Pollen: Unknown, but visits flowers of Brassica,
Hydrangea, Potentilla, Primus, Pyrus, Rubus.
Andrena (Andre^ia) lata Viereck, 1922. Boston Soc. Nat. Hist., Occas. Papers 5: 39. ♀.
Andrena (Andrena) vulgaris Viereck, 1922. Boston Soc. Nat. Hist., Occas. Papers 5: 40. ♂.

Taxonomy: Atwood, 1934. Canad. Jour. Res. 10 (2): 208, 210. 9, 6 (key). —Mitchell, 1960. N.
C. Agr. Expt. Sta. Tech. Bui. 141: 222-224, figs. 44-45, table 6 (redescription, synonymy).

Andrena medionitens Cockerell
B. C. and Alta. to Calif, and Colo. Pollen: Unknown, but visits flowers
of Cymopterus terebinthus, Eriogonum latifoiium var. midum, E. subscaposum.
Erysimum asperuyyi, Gilia tenuiflora, Hackelia jessicae, Holodiscus discolor, Horkelia
bemardina, H.fusca, Oxytropis, Potentilla glandiilosa, P. wheeleri, Sisymbrium
incisum, Spraguea umbellata, Streptanthus tortuosus.
Andrena medionitens Cockerell, 1902. Ann. and Mag. Nat. Hist. (7) 9: 101. ♀.

Taxonomy: Viereck, 1904. Canad. Ent. 36: 192, 194. 9,6. —Viereck and Cockerell, 1914. U.
S. Natl. Mus., Proc. 48: 48. 9 (variation).

Biology: Clements and Long, 1923. Carnegie Inst. Wash., Pub. 336: 249 (ecology, as
madronitens).

Andrena phaceliae Mitchell
Md., Va., 111. Pollen: Unknown, but visits flowers of Alsine pubera,
Phacelia dubia, P. purshii.
Andrena (Thysandrena) phaceliae Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 224,
figs. 44, 45. ♀, ♂.

Andrena phocata Cockerell
Colo.

Andrena phocata Cockerell, 1910. Canad. Ent. 42: 369. ♀.

Andrena subcandida Viereck
B. C. to Calif.

Andrena subcandida Viereck, 1904. Canad. Ent. 36: 193. ♀.

Andrena subdistans Viereck
Wash., Oreg.

Andrena subdistans Viereck, 1904. Canad. Ent. 36: 193. ♀.

Andrena subtrita Cockerell
Nev.

Andrena subtrita Cockerell, 1910. Ann. and Mag. Nat. Hist. (8) 5: 263. ♀.

Andrena taeniata Viereck
Calif.

Andretia (Atidrena) taeniata Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 538. ♀.

Andrena trizonata (Ashmead)
Colo.

Cilissa trizonata Ashmead, 1890. Colo. Biol. Assoc, Bui. 1: 6. ♀.

Andrena vierecki Cockerell
Utah, Colo., Calif. Parasite: Myopa rubida (Bigot).

Andrena vierecki Cockerell, 1904. Ann. and Mag. Nat. Hist. (7) 14: 26. ♀.

Andrena boharti Linsley, 1939. Pan-Pacific Ent. 15: 159. ♀, ♂.




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Taxonomy: Lanham, 1941. Ent. Soc. Amer., Ann. 34: 706, 707. 9, 6 (key). — Timberlake,
1951. In Linsley, In Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2:
1085 (synonymy).

Biology: MacSwain and Bohart, 1947. Pan-Pacific Ent. 23: 30 (parasite).

Andrena w-scripta Viereck
B. C. to Calif. Pollen: Unknown, but visits flowers of Acacia greggii,
Adenostoma fascicidatum, Arctostaphylos glandulosa, A. mariposa, A. patula, A.
pungens, Ceanothus cordulatus, C. cmieatus, C. greggii, C. soredatus, Chamaebatia
foliolosa, Claytonia spathulata, Cryptantha intermedia, C. lepida, Daucus carota,
Eriogonum fascicidatum, Eriophyllum confertiflorum, Fritallaria, Heteromeles
arbutifolia, Holodiscus discolor, Horkelia, Lasthenia chrysostoma, Lomatiiim, Lotus
scoparius, Pliacelia dista^is, Prunus ilicifolia, Ranunculus califomicus, Rhamnus
califomicus, Salix laevigata.
Andrena w-scripta Viereck, 1904. Canad. Ent. 36: 193, 194. ♀, ♂.

Genus ANDRENA Subgenus TRACHANDRENA Robertson

Trachandrena Robertson, 1902. Amer. Ent. Soc, Trans. 28: 189.
Type-species: Andrena rugosa Robertson. Orig. desig.

All the species of this subgenus, except the Eurasian Andrena lagopus Latreille, are centered
in America north of Mexico and are not known to occur south of the United States.

Revision: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 235-371, 69 figs., 2 tables, 15 maps (N.
Amer. spp.).

Taxonomy: Lanham, 1949. Calif. Univ. Pubs. Ent. 8: 216 (tax. characters). —LaBerge, 1964.

Nebr. Univ. State Mus., Bui. 4: 282, 292, 308 (tax. characters). — Warncke, 1968. Coimbra

Univ. Est. Mus. Zool., Mem. 307: 68-69 (tax. status).

Andrena amphibola (Viereck)
B. C. south to Calif., east to Alta., Mont. Wyo., Nebr. and Colo. Pollen:
Apparently polylectic, visits flowers of Astragalus, Daucus pusilla, Drymocallis,
Eriogonum flavum, Geum triflorum, Holodiscus discolor. Iris, Lappula floribunda,
Lonicera, Lupinus, Medicago sativa, Penstemon, Phacelia, Potentilla, Salix, Senecio,
Symphoricarpos, Taraxacum officinale, Thermopsis divaricata.

Trachandrena amphibola Viereck, 1904. Canad. Ent. 35: 158, 159. 9, tJ.

Trachandrena crassihirta Viereck, 1904. Canad. Ent. 35: 158. ♀.

Trachandrena perdensa Viereck, 1904. Canad. Ent. 35: 158. ♀.

Trachandrena hadra Viereck, 1904. Canad. Ent. 35: 158. ♀.

Trachandrena limarea Viereck, 1904. Canad. Ent. 36: 158, 159. ♀, ♂.

Andrena {Trachandrena) eriogoni Cockerell, 1927. Ent. Soc Amer., Ann. 20: 397. ♀.

Andrena {Trachandrena) seneciophila Cockerell, 1928. Psyche 35: 62. ♀.

Taxonomy: Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 10. 9 (as perdensa).
—Lanham, 1914. Ent. Soc. Amer. Ann. 34: 704. 9 (key, as seneciophila). —LaBerge, 1973.
Amer. Ent. Soc, Trans. 99: 284-292, figs. 1, 7, 34-35 (redescription, synonymy, variation in
color of vestiture).

Andrena ceanothi Viereck
N. S. and Que., south to Ga., west to Minn, and Colo. Pollen: Polylectic, visits
flowers of Anielanchier, Amorpha fndicosa, Barbarea vulgaris, Ceanothus americamis,
Cerasus serotina, Chamaedaphne, C hrysanthemum leucanthemum, Comus
candidissimus, Crataegus, Fragaria, Ledum-, Prunus, Ptelea, Rhus, Rubus argidus, R.
villosus, Salix, Taraxacum officinale, Vaccinium angustifolium. Viburnum,
Waldsteinia.

Andrena {Trachandrena) ceanothi Viereck, 1917. Amer. Ent. Soc, Trans. 43: 404. 9 {6
misdet.).

Andrena {Trachandrena) compacta Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
181. ♂.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 17. 9 (key). — Atwood, 1934. Canad.
Jour. Res. 10: 212. 9,6. —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 179-180,
figs. 35-36, table 5 (redescription, synonymy). —LaBerge, 1973. Amer. Ent. Soc, Trans. 99:
346-349, figs. 1, 12, 58, 59 (redescription, synonymy).




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Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 349 (floral relationships).

Andrena cleodora
***authority mismatch
cleodora (Viereck). B. C. to north. Calif., east to Mont., Wyo. and Colo., ?Ariz. (Santa
Catalina Mts.). Pollen: Oligolege of Ceanothus including C. cuneatus, C. integerrimus, C.
laevigatus, but visits other flowers presumably for nectar including Arctostapkylos,
Aster, Chamaebatia foUolosa, Conium maciilatum, Eriogonum nudmn, Lupinus,
Phacelia, Prunus, Rhamnus, Ribes, Salix, Sorbus.
Trachandrena cleodora Viereck, 1904. Canad. Ent. 35: 158. ♀.
Andrena {Trachandreyia) lutzi Cockerell, 1931. Amer. Mus. Novitates 458: 12. ♀.

Taxonomy: Cockerell, 1932. Pan-Pacific Ent. 8: 173. 6. —LaBerge, 1973. Amer. Ent. Soc,
Trans. 99: 323-326, figs. 1, 12, 48-49 (redescription).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 327 (floral relationships).

Andrena cleodora
***authority mismatch
melanodora Cockerell. South. Calif. Pollen: Oligolege of Ceanothus including C.

cordulatus, C. integerrimus, C. palmeri, but visits other flowers presumably for nectar
including Arctostapkylos patula, Barbarea orthoceras, Cryptantha lepida, Eriodictyon
califomicum, Prunus emarginata, P. ilicifolia, Pyrus mains, Rhannius califomica.
Andrena (Trachandrena) melanodora Cockerell, 1932. Pan-Pacific Ent. 8: 173. ♀.

Taxonomy: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 326-327, figs. 48-49 (redescription).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 327 (floral relationships).

Andrena cupreotincta Cockerell
B. C, Alta. and Sask., south to Calif., Nev., Utah and N. Mex. Pollen:
Apparently polylectic, visits flowers of Abies concolor, Arctostaphylos nevadensis,
Ceanothus integerrirmis, Eriogonum, Hackelia, Lesquerella, Penstemon, Prunus

Andrena virginiana
***authority mismatch
demissa, Pyrus malus, Rhamnus califomica, R. rubra, Ribes inerme,
Rubus, Salix, Taraxacum officinale. •

Andrena cupreotincta Cockerell, 1901. Canad. Ent. 33: 153. ♀.

Trachandrena ochreopleura Viereck, 1904. Canad. Ent. 35: 158. ♀.

Andrena {Trachandrena) swenki Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc 48: 3.
9.

Andrena (Trachandrena) jockorum Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48:
3. ♂.

Trachandrena abjuncta Cockerell, 1929. N. Y. Ent. Soc, Jour. 37: 445. ♀.

Taxonomy: Viereck, 1904. Canad. Ent. 36: 158. 9 (key). —Cockerell, 1937. Ent. News 49: 257.
9 (as abjuncta). —LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 315-321, figs. 1, 11, 46-47
(redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 321 (floral relationships).

Andrena cyanophila Cockerell
B. C. and Alta., south to Calif., Nev. and Ariz. Pollen: Apparently
polylectic, visits flowers oi Argentina anserina, Ceanothus fendleri. Euphorbia,
Geraniiiyn fremontii, Oreocarya nana, Pinus ponderosa, Polemonium, Potentilla
arguta, P. glandulosa, Prunus melanocarpa. Ranunculus Montensis, Ribes
longiflorum, Senecio perplexis, Taxaxacum officinale.
Andrena cyanophila Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 431. ♀.
Andrena tacitula Cockerell, 1910. Ann. and Mag. Nat. Hist. (8) 5: 262. ♂.

Taxonomy: Cockerell, 1916. Canad. Ent. 48: 253. — Lanham, 1941. Ent. Soc. Amer., Ann. 34:
704, 707. 9, 6 (key). -LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 353-357, figs. 1, 10, 64-65
(redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 357 (flora! relationships).

Andrena forbesii Robertson
N. S. to B. C, south to Ga., west to Wash., Oreg. and Calif. Pollen:

Polylectic, apparently prefers pollens of Rosaceae and Salicaceae over other sources;
visitation records include Acer negundo, Alliaria officinalis, Amelanchier arborea, A.
canadensis, Aronia melanocarpa. Astragalus, Barbarea vulgaris, Berberis vulgaris,
Brauneria purpurea, Callirhoe digitata, Cardamine bulbosa, Ceanothus ovatus, Cercis
canadensis, Claytonia caroliniana, C. virginica, Conius florida, C. mas, Crataegus,
Dentaria laciniata, Erythronium americanum, Euphorbia comniutata, Forsythia
suspe7isa, Fragaria virginiana, Heracleum lanatum, Hieracium, Ilex, Lappula




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heterosperma, Lepargyrea canadensis, Lomatium daucifolium, L. foenimlaceum,
Lonicera, Meiilofus alba, M. officinalis, Paeonia, Pastinaca sativa, Physocarpus
opulifolius, Populus deltoides, Prunus americanus, P. demissa, P. melanocarpa, P.
serotina, P. virginiana, Ptelea, Pyracantha, Pyrus leonis, P. mains. Ranunculus
abortivus, Rhamnus lanceolatus, Rhus aromatica, R. canadensis, R. toxicodendroyi,
Ribes gracile, Rosa, Rubus argenteus, Salix discolor, S. nigra, Sedum tematum,
Shepherdia argentea. Spiraea aru7icus, S. latifolia, Symphoricarpos, Tamarix gallica.
Taraxacum officinale, Thlaspi arvetise, Vaccinium, Viburnum opulus, V. prunifolium,
Zizia a urea.

Andrena forbesii Robertson, 1891. Amer. Ent. Soc, Trans. 18: 59. ♀.

Trachandrena indotata Viereck, 1904. Canad. Ent. 36: 158, 159. ♀, ♂.

Andre7ia (Trachandrena) lincolni Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48:
13. ♀.

Andrena (Trachandrena) rodecki Cockerell, 1929. Ent. Soc. Amer., Ann. 22: 755. ♀.

Andrena (Trachandrena) pyracanthae Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
194, fig. 35. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 189, 190. 9,6. —Cockerell, 1931.
Amer. Mus. Novitates 458: 16. 9. (as rodecki). —Cockerell, 1931. Amer. Mus. Novitates
458: 17 (key, as lincolni). — Atwood, 1934. Canad. Jour. Res. 10: 208, 209, figs. 9, 6 (key).
— Lanham, 1941. Ent. Soc. Amer., Ann. 34: 704, 707. 9, 6 (key). — Lanham, 1941. Ent. Soc.
Amer., Ann. 34: 704. 9 (key, as lincolni). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141: 183-184, figs. 35-36, table 5 (redescription). — LaBerge, 1973. Amer. Ent. Soc,
Trans. 99: 261-267, figs. 1, 4, 24-25 (redescription, synonymy, geogr. variation).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 266-267, table 1 (floral relationships).

Andrena fuscicauda (Viereck)
B. C. to Calif, and west. Nev. and Ariz (Yuma). Pollen: Apparently

polylectic, visits flowers of Arctostaphylos patula, Berberis, Ceanothus cordulatus, C.

integerrimus, C. leucodermis, C. palmeri, C. soredatus, Chamaebatia foliolosa,

Cryptantha intermedia, Eriodictyon califomicum, Eriogonum, Gilia, Horkelia pan-yi,

Malus, Nemophila menziesii, Pinus ponderosa, Potentilla, Prunus demissa, P.

eynarginata, P. ilicifolia, P. subcordata, Quercus dumosa, Rhamnus califomica, R.

crocea, Salix laevigata, Tai-axacum denselionis.
Trachandrena fuscicauda Viereck, 1904. Canad. Ent. 35: 159. ♀.
Andrena (Trachandrena) califomica Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 373. ♀.

6.
Andrena (Trachandrena) califomica wickhami Timberlake, 1951. U. S. Natl. Mus., Proc.

101: 375. ♀.

Taxonomy: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 357-361, figs. 1, 14, 66-67
(redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 361 (floral relationships).

Andrena heraclei Robertson
N. Y. to N. C, west to Kans. Pollen: Unknown, but visits flowers of
Camassia esculenta, Celastrus scayidens, Cercis canadeyisis, Crataegus, Coiiius,
Heracleum, Pastinaca sativa, Prunus, Ptelea, Salix, Taenidia, Washingtonia
longistylus.
Andrena heraclei Robertson, 1897. Acad. Sci. St. Louis, Trans. 7: 336. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 189. 9. —Mitchell, 1960. N. C. Agr.

Expt. Sta. Tech. Bui. 141: 184, fig. 35, table 5 (redescription). -LaBerge, 1973. Amer. Ent.

Soc, Trans. 99: 343-346, figs. 1, 3, 56-57 (redescription).

Andrena hippotes Robertson
N. S. to B. C, south to Ga., west to Wash., Oreg. and north. Calif. Pollen:
Polylectic, visits a wide variety of flowers including Acer, Aesculus glabra sargentii,
Amelanchier arborea, Amorpha canescens, Angelica atropurpurea, Barbarea vulgaris,
Berberis, Caltha palustyis, Camassia esculenta, Ceanothus ovatus, Celastrus, Cercis
canadeiisis, Chamaedaphne, Clayfonia, Comus ynas, Crataegus ynollis, Daucus carota,
Dentaria laciniata, Descourainia intermedia. Echinacea angustifolia, E. purpurea,
Encelia, Euphorbia, Fragaria, Fraxinus, Hieracium, Ligustruyn vulgare, Lomatium
tritematum, Lonicera fragrantissima, Melilotus alba, M. officinalis, Nyssa sylvatica,




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Paeonia, Pastinaca sativa, Phacelia, Physocarpus opulifolius, Potentilla anserina,
Primus americana, P. demissa, P. serotina, P. virginiana, Ptelea, Pyracantha
yunannensis, Pyrus maliis, Ranunculus abortivus, Rhamnus, Rhus glabra, Ribes
aureum, Rosa Carolina, Rubus, Salix nigra, Shepherdia argentea. Spiraea discolor, S.
sorbifolia, Sphaeralcea coccinea, Symphoricarpos, Taraxacum officinale, Thlaspi
arvense, Trollius laxa, Vaccinium, Viburnum dentatum, V. opulus, Zizia aurea.

Andrena hippotes Robertson, 1895. Amer. Ent. Soc, Trans. 22: 120. ♀, ♂.

Andrena (Trachandrena) perforatella Cockerell, 1906. Ann. and Mag. Nat. Hist. (7) 17:
306. ♀.

Andrena (Trachandrena) arenakensis Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
177, fig. 35. ♀.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 184-185, figs. 35, 37, table 5
(redescription, synonymy). — LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 292-299, figs. 1, 8,
36-37 (redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 298, table 2 (floral relationships).

Andrena mariae Robertson
N. S. to N. W. T. and B. C, south to Ga., west to N. Mex., Utah, Idaho and

Wash. Pollen: Oligolege of Salix including S. babylonica, S. bebbiana, S. nigra, S. tristis,

but visits other flowers including Amelanchier, Astragahcs, Barbarea vulgaris,

Cogswellia, Comandra, Crataegus mollis, Erigenia, Erigeron philadelphicus,

Erythronium, Fragaria, Hackelia floribunda, Kalmia, Lomatium daucifolium, L.

foeniculaceum, Pastinaca sativa, Prunus americanus, Pulsatilla, Pyrus malus, Ribes

gracile, Rubus, Sisymbrium, Spiraea, Taraxacum officinale.
Andrena mariae Robertson, 1891. Amer. Ent. Soc, Trans. 18: 58. ♀, ♂.
Andrena sphecodina Cockerell and Casad, 1896. Ann. and Mag. Nat. Hist. (6) 18: 78. ♀, ♂.
Andrena mariae var. concolor Robertson, 1898. Acad. Sci. St. Louis, Trans. 8: 46. ♀, ♂.
Andrena (Trachandrena) submariae Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc

48:4. ♀.
Andrena (Trachandrena) martialis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48:

5. ♂.
Andrena (Trachandrena) sphecodiniformis Viereck and Cockerell, 1914. U. S. Natl. Mus.,

Proc. 48: 6. ♂.
Andreyia (Trachandrena) profundiformis Viereck and Cockerell, 1914. U. S. Natl. Mus.,

Proc. 48: 8. ♂.
Andrena (Trachandrena) stricklandi Cockerell, 1936. Canad. Ent. 68: 275. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 189, 190. 9,6. —Viereck, 1907. Ent.
News 18: 284, 286. 9, 6 (key). -Cockerell, 1929. Ent. Soc. Amer., Ann. 22: 754 (as
martialis). —Cockerell, 1929. Ent. Soc Amer., Ann. 22: 754 (as sphecodina). — Atwood,
1934. Canad. Jour. Res. 10: 208, 210. 9, 6 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 187-188, figs. 35, 38, table 5 (redescription). —LaBerge, 1973. Amer. Ent.
Soc, Trans. 99: 299-305, figs. 1, 9, 39-41 (redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 305 (floral relationships).

Andrena miranda Smith
N. S. to N. J. and Pa., west to Alaska, south to N. Mex., Ariz., Nev. and Calif.
(Alta and Carrville). Parasite: Stylops sp. Pollen: Polylectic, visits a wide variety of
flowers including Achillea millefolia, Angelica, Aralia latifolia, Argemone, Astragalus,
Barbarea vulgaris, Blepharipappus, Brassica campestris, Ceanothus, Cirsium
undulatum, Cornus, Crataegus, Drymocallis arguta, Euvhorbia, Geranium richardsoni,
Heracleum lanatum, Lappula heierosperma, Lotus corniculatus, Malvastrum, Melilotus
alba, M. officinalis, Pastinaca sativa, Potentilla fruticosa, P. recta, Prunus virginiaria
demissa. Ranunculus septentrionalis, Ribes, Rosa arkansana, R. Carolina, R.
multiflora, Rubus adoratus, Rudbeckia hirta, Sanicula marilandica, Spiraea latifolia,
S. sorbifolia, Symphoricarpos occidentalis. Taraxacum officinale, Vaccinium,,
Viburnum opulus, Waldsteinia fragarioides, Zizia aptera, Z. aurea.

Andrena miranda ^mith, 1879. Desc. New Species Hym. Brit. Mus., p. 54. ♀, ♂.

Andrena grandior Cockerell, 1897. Entomologist 30: 307. ♀.

Andre7ia multiplicata Cockerell, 1902. Canad. Ent. 34: 46. ♀.

Andrena multiplicatiformis Viereck, 1907. Ent. News 18: 284. ♀.




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Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 153. 9, 6 (types). — Cockerell, 1906.
Psyche 13: 34 (type). -Atwood, 1934. Canad. Jour. Res. 10: 208, 210. 9, 6 (key).
— Atwood, 1934. Canad. Jour. Res. 10 (2): 208, 210 (as grandior). — Timberlake, 1951. hi
Linsley, hi Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1072
(synonymy). —Mitchell, 1958. In Krombein, U. S. Dept. Agr., Agr. Monog. 2, First Suppl.
p. 215 (synonymy). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 190-191, figs. 35,
38, table 5 (redescription, synonymy). — LaBerge, 1973. Amer. Ent. Sec, Trans. 99:
267-273, figs. 1, 5, 26-27 (redescription).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 272-273 (floral relationships,

stylopization).

Andrena nuda Robertson
Maine to Ga., west to Tex., N. Mex., Utah and Wyo. Pollen: Polylectic,
although with a preference for flowers of the Rosaceae: visitation records include
Aruncus, Benzoin aestivale, Cercocarpus, Claytonia, Conium, Crataegus mollis,
Erigenia, Euphorbia, Heracleum, Ilex, Lomatium dissectum, Melilotus officinalis,
Myosotis, Pastinaca saliva, Peiisiemon, Prunus americana, P. serotina, P. virginica,
Ptelea, Pyracantha, Pynis leonis, P. malus, P. pyrus, Rhamnus lanceolata, Rhus,
Rubus argutus, Salix, Sanicula, Sassafras, Staphylea, Symplocus, Taenidia, Viburnum
prunifoiium, Zizia.

Andrena nuda Robertson, 1891. Amer. Ent. Soc, Trans. 18: 57. ♀.

Andrena (Trachandrena) davisiana Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc.
48:6. ♂.

Andrena {Trachandrena) pseudobscura Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui.
141: 193, fig. 36. ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 189. 9 (key). — Lanham, 1941. Ent.
Soc. Amer., Ann. 34: 704. 9 (key). —Lanham, 1941. Ent. Soc. Amer., Ann. 34: 707. S (key,
as davisiana). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 192-193, fig. 35
(redescription). -LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 333-338, figs. 1, 14, 52-53
(redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 337-338 (floral relationships).

Andrena quintiliformis Viereck
B. C. to Calif., east to Idaho, Nev. and Utah. Pollen: Apparently
polylectic, visits flowers of Achillea millefolia. Allium, Arctostaphylos, Ceanothus
cuneatus, C. laevigatus, Chamaebatia foliolosa, Cirsium, Eriogonum, Lappula,
Lomatium tritematum, Phacelia humilis. Ranunculus, Salix, Solidago califomica.
Andrena (Trachandrena) quintiliformis Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 606.

9.
Andrena (Trachandrena) coactifera Viereck, 1926. Calif. Acad. Sci., Proc. (4) 15: 399. ♀.

Taxonomy: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 278-281, figs. 1, 6, 30-31 (redescription,

synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 281 (floral relationships).

Andrena quintilis Robertson
Wis. and 111., west to N. Dak., S. Dak., Nebr., Colo, and Kans. Pollen:

Polylectic, although exhibits a decided preference for the pollen and nectar of Amorpha
including A. canescens, A. fruticosa, but visits other flowers including Apocynum
cannabinum. Chrysanthemum leucanthemum, Crataegus, Drymocallis arguta.
Echinacea angustifolia, Erigeron ramosus, Pastinaca sativa, Ptelea, Pycnanthemum
flexuosum, P. linifolium, Salix, Symphoricarpos, Zizia aurea.
Andrena quintilis Robertson, 1898. Acad. Sci. St. Louis, Trans. 8: 46. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 189. 9 (key). —Mitchell, 1960. N. C.
Agr. Expt. Sta. Tech. Bui. 14: 195-196, fig. 35, table 5 (redescription). —LaBerge, 1973.
Amer. Ent. Soc, Trans. 99: 281-284, figs. 1, 6, 32-33 (redescription).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 284 (floral relationships).

Andrena rehni Viereck
Que. and Ont., south to G., west to Wis. Pollen: Apparently polylectic, visits
flowers of Aruncus dioecium, Castanea pumila, Ceanothus, Chrysanthemum
leucanthemum, Cicuta maniculata, Hydrangea, Ilex opaca, Quercus prinoides, Salix.
Andrena rehni Viereck, 1907. Ent. News 18: 284. ♀.




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Andrena (Trachandrena) votula Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 204.

9.

Taxonomy: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 349-853, figs. 1, 15, 60-63

(redescription, synonymy).

Andrena rugosa Robertson
Que. and New England, south to Ga., west to Minn, and Nebr. Pollen:
Polylectic, visits flowers of Acer, Amelanchier arborea, A. canadensis, Angelica,
Astragalus, Barbarea vulgaris, Brassica arvensis, Claytonia virginica, Comus mas,
Crataegus punctata, Dentaria laciniata, Dirca, Erigenia bulbosa, Fragaria, Geranium,
Hepatica, Heracleum, Isopyrum, Macrocalyx nyctelea, Melilotus officinalis, Paeonia,
Pastinaca sativa, Physocarpus opulifolius, Prunus americana, P. virginiana, Ptelea,
Pyracantha, Pyrus malus, Rhus, Ribes, Rosa multiflora, Rubus argutus, R.
oecidentalis, Salix, Sanicula, Spiraea aru7icus, Symphoricarpos, Taraxacum officinale.
Viburnum, Zanihoxyluyn, Zizia aurea.

Andrena rugosa Robertson, 1891. Amer. Ent. Soc, Trans. 18: 58. 9, 3 ■

Andrena paenerugosa Viereck, 1907. Ent. News 18: 285. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 189, 190. 9, S (key). — Atwood,
1934. Canad. Jour. Res. 10: 208, 210. 9, 6 (key). —Mitchell, 1960. N. C. Expt. Sta. Tech.
Bui. 141: 198-199, figs. 35-36, table 5 (redescription, synonymy). —LaBerge, 1973. Amer.
Ent. Soc, Trans. 99: 327-332, figs. 1, 13, 50-51 (redescription).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 332 (floral relationships).

Andrena salicifloris Cockerell
B. C. and Alta., south to Calif., Nev., Utah and Colo. Pollen: Polylectic,
but with some predilection for the flowers of Salix; visitation records include Acacia,
Acer, Amelanchier, Arctostaphylos, Berberis, Brassica, Ceanothus, Cerastium
oreophyllum, Chamaebatia foliolosa, Cicuta, Cistus, Epilobium angustifolium,
Eriogonum, Geranium, Grossularia, Hackelia patens, Hieracium lanatum, Lasthenia,
Lepidium, Lomatium dissectum, L. tritematum, Pastinaca sativa, Prunus demissa, P.
ilicifolia, P. virginiana, Pulsatilla patens, Pyracantha yunannensis, Pyrus malus.
Ranunculus califomicus, Ribes lacustre, Rubtis yiutkana, R. ursinus, R. vitifolius.
Taraxacum densileonis, T. officinalis.

Andrena salicifloris Cockerell, 1897. Acad. Nat. Sci. Phila., Proc. 49: 351. ♀.

Trachandrena auricauda Viereck, 1904. Canad. Ent. 35: 159, 161. ♀.

Andrena (Trachandrena) tacitula var. grossulariae Viereck and Cockerell, 1914. U. S.
Natl. Mus., Proc. 48: 9. ♂.

Andrena (Trachandrena) nortoni Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 602. ♀, ♂.

Andrena (Trachandrena) veris Cockerell, 1934. Pan-Pacific Ent. 9: 156. ♀, ♂.

Taxonomy: Viereck, 1904. Canad. Ent. 36: 158, 159. 9, 6 (variation). — Lanham, 1941. Ent.
Soc. Amer., Ann. 34: 704, 707. 9, 6 (key, as veris). —LaBerge, 1973. Amer. Ent. Soc,
Trans. 99: 255-261, figs. 1, 3, 22-23 (redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 260 (floral relationships).

Andrena semipunctata Cockerell
Wash, to south. Calif., east to southeast. Utah and southwest. N. Mex.
Pollen: Oligolege of Salix including S. exigua, S. hindsiana, S. laevigata, S. lasiolepis, S.
nigra, but visits other flowers presumably for nectar including Baccharis, Ceanothus
soredatus, Prunus, Pyrus malus, Rhus trilobata, Sambucus, Senecio salignifolia,
Sisymbrium irio, Tamarix gallica. Taraxacum officinale.
Andrena semipunctata Cockerell, 1902. Ann. and Mag. Nat. Hist. (7) 9: 102. ♀.

Taxonomy: Viereck, 1904. Canad. Ent. 36: 159. 9, 6 (key). —LaBerge, 1973. Amer. Ent. Soc,
Trans. 99: 311-315, figs. 1, 9, 44-45 (redescription).

Biology: Linsley, 1937. Brooklyn Ent. Soc, Bui. 32: 127 (floral relationships, adult behavior).

Andrena sigmundi Cockerell
N. B. south to Ga., west to B. C, south to Calif., Utah and N. Mex. Pollen:
Oligolege of Salix including S. bebbiana, S. discolor, but visits other flowers including
Acer saccharinum, Amelanchier canadensis, Barbarea vulgaris, Ceanothus fendleri,
Crataegus, Fragaria virginiana, Fraxinus, Kalmia latifolia, Lepargyrea argentea, L.
canadensis, Melilotus alba, Populus, Potentilla, Prunus demissa, P. virginiana.




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Pulsatilla ludoviciana, Spiraea, Taraxacum officinale, Vaccinium angustifolium,

Waldsteinia fragarioides.
Andrena sigmundi Cockerell, 1902. Canad. Ent. 34: 45. ♀.
Andrena radiatula Cockerell, 1902. Canad. Ent. 34: 46. ♀.
Andrena weedi Viereck, 1907. Ent. News 18: 284, 285. ♀, ♂.
Andrena (Trachandrena) pnmicola Cockerell, 1913. Ann. and Mag. Nat. Hist. (8) 12: 375.

9.
A7idre7ia (Trachandrena) moscovensis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc.

48: 13. ♀.
Andrena (Trachandrena) corrugata Cockerell, 1931. Amer. Mus. Novitates 458: 9. ♀.

Taxonomy: Cockerell, 1931. Amer. Mus. Novitates 458: 17 (key, as moscovenis). —Mitchell,
1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 200-202, figs. 36, table 5 (redescription,
synonymy). — LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 249-255, figs. 1, 2, 16-21
(redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 255 (floral relationships).

Andrena spiraeana Robertson
Ont. and Maine to Ga., west to Minn., Nebr. and Kans. Pollen: Polylectic,
visits flowers of Alliaria officinalis, Angelica, Anmcus, Azalea, Barbarea vulgare,
Castanea pumila, Ceanothus americanus, Cornus, Crataegus, Cryptotaenia, Erigeron
philadelphicus. Euphorbia commutata, Heracleum, Hydrangea, Melilotus alba,
Pastinaca sativa, Physocarpus opulifolius, Ptelea, Rhus glabra, R. typhine, Rosa,
Spiraea aruncus, S. latifolia, Sorbaria sorbifolia, Vibtirnum nudum, Zizia.

Andrena spiraeana Robertson, 1895. Amer. Ent. Soc, Trans. 22: 120. ♀, ♂.

Andrena (Trachandrena) montensis Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
191, fig. 35. ♀.

Andrena (Trachandrena) unica Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 203,
fig. 36. ♂.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 189, 190. 9, 6 (key). —Viereck,
1907. Ent. News 23: 107 (key). — Lanham, 1941. Ent. Soc Amer., Ann. 34: 704, 707. 9, S
(key). - Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 202-203, figs. 35, 38, table 5
(redescription). —LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 338-342, figs. 1, 11, 53-54
(redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 342 (floral relationships).

Andrena striatifrons Cockerell
B. C. and Alta., south to Ariz, and Calif. Pollen: Oligolege of Salix, but
visits other flowers including Arctostaphylos, Eriogonum, Hackelia floribunda,
Lomatium, Primus, Pyrus malus, Ranuncuhis, Rubus parviflorus, Symphoricarpos,
Taraxacum officinale, Vaccinium.

Andrena striatifrons Cockerell, 1897. Entomologist 30: 308. ♀.

Trachandrena pemuda Viereck, 1904. Canad. Ent. 35: 159. ♀.

Andrena trachandrenoides Viereck, 1904. Canad. Ent. 36: 190, 195. ♀, ♂.

Andrena (Trachandrena) marioides Viereck, 1917. Acad. Nat. Sci. Phila., Proc. 68: 601. ♀.

Andrena (Trachandrena) politissima Cockerell, 1918. Ann. and Mag. Nat. Hist. (9) 166. ♀.

Andrena dolichotricha Cockerell, 1924. Ent. News 35: 348. ♂.

Andrena (Trachandrena) brevibasis Cockerell, 1931. Amer. Mus. Novitates 458: 8. ♀.

Andrena (Trachandrena) postnitens Cockerell, 1931. Amer. Mus. Novitates 458: 15. ♀.

Taxonomy: Viereck, 1904. Canad. Ent. 36: 159. 9 (key). —LaBerge, 1973. Amer. Ent. Soc,
Trans. 99: 305-311, figs. 1, 10, 42-43 (redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 311 (floral relationships).

Andrena virginiana Mitchell
P. E. I., Que, and Ont, south to Ga., west to Minn., 111. and Ark. Pollen:
Apparently polylectic, visits flowers of Ceanothus americanus, Cerasus serotina,
Daucus carota. Ilex verticillata, Kalmia, Melilotus alba, Pyc7ianthemum linifolium.
Spiraea latifolia, Solidago.
Trachandrena obscura Robertson, 1902. Amer. Ent. Soc, Trans. 35: 189. ♀. Preocc.
Andrena (Trachandrena) virginiana Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:

203. ♀.
Andrena (Biareolina) obscurida Warncke, 1970. Bayer. Ent., Nachr. 19: 30. N. name.




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Taxonomy: Viereck, 1907. Ent. News 18: 284. 9 (key, as obscura). — Cockerell, 1929. Ent.
Soc. Amer., Ann. 22: 757. 9 (as obscura). — LaBerge, 1973. Amer. Ent. Soc, Trans. 99:
273-277, figs. 1, 6, 28-29 (redescription, synonymy).

Biology: LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 277 (floral relationships).

Andrena winnemuccana LaBerge
Nev., Oreg. Pollen: Unknown, but visits flowers of Thelypodium
laciniatum.
Andrena (Trachandrena) winnemuccana LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 366,
fig. 1. ♀.

Andrena zionensis LaBerge
South. Utah. Pollen: Unknown, but visits flowers of Amelanchier, Salix.
Andrena {Trachandrena) zionensis LaBerge, 1973. Amer. Ent. Soc, Trans. 99: 361, figs. 1,
68-69. ♀, ♂.

Genus ANDRENA Subgenus TYLANDRENA LaBerge

Andrena subg. Tylandrena LaBerge, 1964. Nebr. Univ. State Mus., Bui. 4: 312.
Type-species: Cilissa erythrogaster Ashmead. Orig. desig.

Revision: LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 543-605, 75 figs., 5 maps

(N. Amer. spp.).
coraeina LaBerge and Bouseman. South. Calif. (Kramer Hills).

Andrena (Tylandrena) coraeina LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans. 96:
591, figs. 1,61-65. ♀, ♂.

Andrena erythrogaster (Ashmead)
Que. to B. C, south to N. C, Ga., Ark., Okla., N. Mex. and Utah.
Pollen: Apparently an oligolege of Salix including S. amygdaloides, S. bebbiana, S.
cordata, S. humilis, S. interior, S. longifolia, S. nigra, S. sericea, but visits other flowers
presumably for nectar including Amelanchier canadensis. Anemone canadensis,
Antennaria plantaginifolia, Aronia, Claytonia virginica, Crataegus coccinea, C.
crus-galli, Descourainia intermedia, Heracleum lanatum, Lomatium foeniculacenm.
Primus americana, P. serotina, P. virginiana, Pyrus, Radicula obtusa, Rhamnus
lanceolata, Rhus aromatica, R. canadensis, R. trilobata, Ribes, Rubus, Shepherdia
argentea. Spiraea, Taraxacum officinale. Viburnum prunifolium, Zizia aurea.

Cilissa erythrogaster Ashmead, 1890. Colo. Biol. Assoc, Bui. 1: 6. ♂.

Andrena perezi Robertson, 1891. Amer. Ent. Soc, Trans. 18: 51. ♀, ♂.

Andrena rhodura Cockerell, 1898. Ent. News 9: 171. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 190, 192. 9, i (key). — Lanham,
1941. Ent. Soc. Amer., Ann. 34: 704, 707. 9, 6 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 114-115, figs. 18-19 21, table 3 (redescription). —LaBerge and Bouseman,
1970. Amer. Ent. Soc, Trans. 96: 550-556, figs. 1-2, 6-10 (redescription).

Biology: Rau, 1935. Ent. News 45: 35 (nesting habits, floral relationships).

Andrena fuscipennis LaBerge and Bouseman
Tex. (Kerrville). Pollen: Unknown, but visits flowers of
Prunus.
Andrena (Tylandrena) fuscipennis LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans.
96: 564, figs. 1, 21-25. ♀, ♂.

Andrena hallii Dunning
Nebr. and Kans., west to Wash, and Nev. Pollen: Apparently an oHgolege of
cruciferous flowers, especially Stanleya pinnata, but also visits flowers of Brassica
nigra, Malvastrum coccineum, Melilotus, Opuntia, Penstemon, Primus, Romneya,
Sisymbrium linifolium.
Andre7ia Hallii Dunning, 1898. Canad. Ent. 30: 268. ♀.
A7idrena heterura Cockerell, 1930. N. Y. Ent. Soc, Jour. 37: 445. ♀.

Taxonomy: Viereck, 1904. Canad. Ent. 36: 191. 9 (key). —Cockerell, 1931. Amer. Mus.

Novitates 458: 10. 9. —LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 588-591,
figs. 1, 4, 56-60 (redescription, synonymy).

Andrena hurdi Lanham
Calif. Pollen: Unknown, but visits flowers of Brassica, Layia. A. waldmerei
LaBerge and Bouseman is possibly a subspecies or synonym of this species which is
perhaps the most elegant N. Amer. species of the genus Andrena.
Andrena hurdi Lanham, 1949. Pan-Pacific Ent. 25: 33. ♀, ♂.




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Taxonomy: LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 583-584, figs. 1, 46-50
(redescription).
jcssicae Cockerell. N. Mex., Ariz. Pollen: Unknown, but visits flowers of Lesquerella gordonii.
Andrena jessicae Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 79. ♂.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 48 (key). —LaBerge and
Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 571-574, figs. 1, 3, 31-35 (redescription).

Andrena layiae Timberlake
Calif.; Mexico (Baja California). Pollen: Unknown, but visits flowers of
Brassica, Lasthenia chrysostoma, L. minor, Layia glandulosa, Senecio califomicus.
Andrena {Ptilandrend) layiae Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 381. ♀.

Taxonomy: LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 594-597, figs. 1, 3,
66-70 (redescription).

Andrena mesillae Cockerell
Tex. to Ariz. Pollen: Apparently an oligolege of Lesquerella including L.
gordonii, L. gracilis, but visits other flowers including Descourainia sophia, Phacelia,
Physaria, Senecio saiigyius.
Andrena mesillae Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 90. ♂.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 49. 6. (key). —LaBerge and

Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 584-588, figs. 1, 4, 51-55 (redescription).

Andrena perplexa Smith
Occupies a disjunct range; in the east occurs from Ont. and Conn., south to
Ga., west to Minn. Iowa, Okla. and Tex.; in the west occurs from Colo, and Mont., west
to B. C, Wash., Oreg. and north. Calif. (Shasta County). Pollen: Apparently polylectic,
visits flowers of Acer, Amelanchier, Aronia, Aruncus, Barbarea, Brassica, Crataegus,
Crepis nincinata, Euonymus japonica, Fragaria virginiana. Geranium, Heracleum,
Lomatium dissectum. Mains, Melilotus, Prunus americana, P. demissa, P.
pennsylvanica, P. virginiana, Ptelea, Pyracantha, Pyriis, Rhamnus, Rhododendron,
Rubus deliciosus, Salix, Senecio, Spiraea, Stellaria pubera, Stenophragma thaliana,
Taraxacum officinale, Viburyium.

Andrena perplexa Smith, 1853. Cat. Hym. Brit. Mus. v. 1, p. 148. ♀.

Andrena belfragei Cresson, 1872. Amer. Ent. Soc, Trans. 4: 256. ♀.

Andrena brunniventris Cresson, 1872. Amer. Ent. Soc, Trans. 4: 258. ♂.

Andrena vibumella Graenicher, 1903. Canad. Ent. 35: 165. ♀, ♂.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 150 (type). — Viereck, 1904. Canad.
Ent. 36: 189, 194. 9, S (key, as vibumella). —Cockerell, 1906. Psyche 13: 10 (type).
—Viereck, 1907. Ent. News 18: 365 (as vibumella). — Lanham, 1941. Ent. Soc. Amer.,
Ann. 34: 705, 708. 9, cJ (as vibumella). — Michener, 1947. Amer. Midland Nat. 38: 445. ♀.
6 (as vibumella). —Lanham, 1949. Ent. News 60: 67 (as belfragei). —Mitchell, 1958. In
Krombein, U. S. Dept. Agr., Agr. Monog 2, Suppl. 1: 215 (synonymy). —Mitchell, 1960. N.
C. Agr. Expt. Sta. Tech. Bui. 141: 116-117, figs. 18-19, 21, table 3 (redescription).
—Stephen, 1966. Kans. Ent. Soc, Jour. 39: 51-53, fig. 1 (larva, pupa, as vibumella).
—LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 559-564, figs. 1, 3, 16-20
(redescription, synonymy).

Biology: Parker and Boving, 1924. U. S. Natl. Mus., Proc. 64: 1 (nesting habits). —Stephen,
1966. Kans. Ent. Soc, Jour. 39: 42-51, 2 figs, (nest architecture, life history, as vibumella).
subaustraiis Cockerell. B. C. to Calif., east to Idaho, Utah and east. N. Mex. Pollen: Oligolege
of Salix including S. gooddingii, S. hindsiana, S. laevigata, S. lasiolepis, S. nigra, but
visits other flowers for nectar including Baccharis viminea, Capsella bursa-pastoris,
Descourainia sophia, Hackelia floribunda, Malus, Prunus americanus, Pyrus, Ribes
aureum, R. inenne, Senecio saligjius. Sisymbrium irio, Tamarix, Taraxacum officinale.
Predator: Philanthus solivagus Say.
A7idrena subaustraiis Cockerell, 1898. Canad. Ent. 30: 146. ♀, ♂.
Andrena viridinitens Cockerell, 1936. Pan-Pacific Ent. 12: 152. ♀.

Taxonomy: LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 556-559, figs. 1, 2,
11-15 (redescription, synonymy).




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Andrena sublayiae LaBerge and Bouseman
Cent. Calif. Pollen: Unknown, but visits flowers of
Hemizonia, Layia.

Andrena (Tylandrena) sublayiae LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans.
96:597, figs. 1,3,71-75. ♀, ♂.

Andrena subtilis Smith
B. C. to Calif., east to Mont., Wyo. and Colo. Pollen: Polylectic, visits flowers of
Acer macrophylla, Baccharis viminea, Brassica campestris, Calandrinia caulescens,
Caragana, Conium maculatum, Crepis, Gilia, Grossularia, Hackelia floribunda,
Lasthenia, Limnanthes douglasii var. douglasii, Lomatium dissectum, Primus deynissa,
P. subcordata, Purshia tridentaia, Pyracantha, Pyrus, Rayiu7icnlus califomicus, Rosa,
Salix lasiolepis, Senecio, Spiraea, Tamarix gallica, Taraxacum.

Andrena subtilis Smith, 1879. Descr. New Species Hym. Brit. Mus., p. 55. ♀.

Andrena seminigra Viereck, 1904. Canad. Ent. 36: 190. ♀.

Andrena semicyanea Cockerell, 1924. Pan-Pacific Ent. 1: 58. ♀.

Taxonomy: Morice and Cockerell, 1901. Canad. Ent. 33: 155 (type). —Cockerell, 1906. Psyche
13: 6 (type). —LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 574-580, figs. 1, 5,
36-40 (redescription, synonymy).
Biology: Linsley and MacSwain, 1959. Calif. Univ. Pubs. Ent. 16: 19 (floral relationships).
waldraerei LaBerge and Bouseman. South. Calif. Pollen: Unknown, but visits flowers of

Allium haematochiton, Encelia califomica, Lomatium dasycarpum, Rhus trilobata.
Possibly a subspecies or synonym of ^4. hurdi Lanham.
Andrena (Tylandrena) waldmerei LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans.
96:580, figs. 1,41-45. ♀, ♂.

Andrena wilmattae Cockerell
Pa. to N. C, west to Wyo. and Colo. Pollen: Unknown, but visits flowers
of Amorpha canescens, Crataegus, Crepis runcinata, Erigeron, Melilotus alba, M.
officinale, Primus angustifolia. Spiraea vanhouteii, Pastinaca sativa. Viburnum
lentago.
Andrena Wilmattae Cockerell, 1906. Ann. and Mag. Nat. Hist. (7) 17: 224. ♀.
A7idre7ia subaustralifonnis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc 48: 28. ♂.
Andrena beutenmuelleri Viereck, 1916. Amer. Mus. Nat. Hist., Bui. 35: 729. ♀.
Andrena (B ythandrena) acra Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 111, figs.
18, 19,21. ♂.
Taxonomy: LaBerge and Bouseman, 1970. Amer. Ent. Soc, Trans. 96: 567-571, figs. 1, 4,
26-30 (redescription, synonymy).

Genus ANDRENA Subgenus XIPHANDRENA LaBerge

Andrena subg. Xiphandrena LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 504.

Type-species: Andrena (Trachandrena) mendica Mitchell. Monotypic and orig.
desig.
Revision: LaBerge, 1971. Amer. Ent. Soc, Trans. 97: 504-507, figs. 1, 4, 54-58.

Andrena mendica Mitchell
111., Ohio, Tenn., Va., N. C, Ga. Pollen: Unknown, but visits flowers of
Arimcus arimcus, Ceanothus including C. americanus.
Andrena (Trachandrena) mendica Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
189, figs. 35-37, 40. ♀, ♂.

Genus ANDRENA Subgenus UNASSIGNED

Although some of these species have been assigned and even reassigned to some of the sub-
genera in Andrena, their correct assignments must await further studies of the genus.

Andrena agricolarum Viereck and Cockerell
Colo.

Andrena agricolarum Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 29. ♂.

Andrena albuginosa (Viereck)
Oreg.

Pterandrena albuginosa Viereck, 1904. Canad. Ent. 36: 227, 228. ♀.

Andrena angustella
***authority mismatch
angustella Cockerell. Calif.

Andrena angustella Cockerell, 1936. Pan-Pacific Ent. 12: 136. ♂.




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Andrena angustella
***authority mismatch
quercina Cockerell. Calif. (Live Oak Canyon, Redlands). Pollen: Unknown, but
visits flowers of Ceanothus, Salix.

Andrena angustella quercina Cockerell, 1939. Ann. and Mag. Nat. Hist. (11) 3: 183. ♂.

Andrena angustifovea Viereck
B. C. to Oreg.

Andre7ia angustifovea Viereck, 1904. Canad. Ent. 36: 194. ♂.

Andrena atala Viereck
N. Mex.

Andrena atala Viereck, 1903. Amer. Ent. Soc, Trans. 23: 54. ♀.

Taxonomy: Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 430. 9,8.

Andrena azygos Viereck
N. Mex.

Andrena (Andrena) azygos Viereck, 1917. Acad. Nat. Sci. Phila., Proc. 68: 550. ♂.
banks! Malloch. N. J. to Fla., west to Tex. Pollen: Unknown, but visits flowers of Prunus
august if 1 ia.
Andrena bayiksi Malloch, 1917. Brooklyn Ent. Soc, Bui. 12: 89. ♂, ♀.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 254-255, figs. 51, 54, 56-57,

table 7 (redescription).

Andrena caeruleonitens Viereck
Cahf.

Andrena caeruleonitens Viereck, 1926. Pomona Jour. Ent. Zool. 18: 1. ♂.

Andrena casadae Cockerell
N. Mex., Colo.

Andrena casadae Cockerell, 1896 Ann. and Mag. Nat. Hist. (6) 18: 83. ♂.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 48. 6 (key).

Andrena cerebrata Mitchell
N. C, Ark. Pollen: Unknown, but visits flowers o{ Malus, Philadelphns,
Rapha7ius sativa, Spiraea.
A7idrena cerebrata Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 255, figs. 51, 56, 57.
6.

Taxonomy: Chandler, 1966. Pan-Pacific Ent. 42: 89-90. ♀.

Andrena cragini Cockerell
Kans.

A7idrena cragini Cockerell, 1899. Ent. News 10: 254. ♂.

Andrena dallasiana Cockerell
Tex.

Andrena dallasiana Cockerell, 1910. Ann. and Mag. Nat. Hist. (8) 5: 262.

Andrena didelta Viereck
N. Mex.

Andrena delta Viereck, 1903. Amer. Ent. Soc, Trans. 29: 55. ♀. Preocc.

A7idre7ia didelta Viereck, 1908. Ent. News 19: 42. N. name.

Andrena dimorpha Mitchell
N. J., N. C, Fla.

Andrena (Pterandrena) dimorpha Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 143,
figs. 24-26,28. ♀, ♂.

Taxonomy: LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 294 (subgeneric status).

Andrena dolomellea Lanham
Tex. Pollen: Unknown, but visits flowers of Crataegus.

Andrena dolomellea Lanham, 1949. Ent. News 60: 65. ♀, ♂.

Andrena dreisbachi Mitchell
Mich.

A7idre7ia (Gonandrena) dreisbachi Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 230.
9.

Taxonomy: Knerer and Atwood, 1963. Canad. Ent. 95: 586, figs. 6, 8. ♂.

Andrena elongatula Viereck
Calif.

Andrena (Andrena) elongatula Viereck, 1917. Amer. Ent. Soc, Trans. 43: 378. ♂.
enceiiarum Cockerell. Ariz. Pollen: Unknown, but visits flowers of Encelia farinosa.

Andrena enceiiarum Cockerell, 1937. Amer. Mus. Novitates 948: 13. ♂.

Andrena enocki (Cockerell)
Calif.

Paraiidrena enocki Cockerell, 1898. Amer. Ent. Soc, Trans. 25: 189. tJ.

Andrena hirsutula Cockerell
N. W. T. (Hay River).

Andrena hirsutula Cockerell, 1936. Canad. Ent. 68: 282. ♀.

Andrena inclinata Viereck
Calif.

Andrena (Andrena) inclinata Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 559. ♀.




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interrogationis Viereck and Cockerell. Colo.

Andrena interrogationis Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 35. ♂.

Andrena jennei Viereck
Wash.

Andrena (Andrena) jennei Viereck, 1917. Acad. Nat. Sci. Phila., Proc. 68: 561. ♂.

Andrena lauracea Robertson
111. Pollen: Unknown, but visits flowers of Sassafras variifolium.

Andrena lauracea Robertson, 1897. Acad. Sci. St. Louis, Trans. 7: 331. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 194. 9. —Mitchell, 1960. N. C. Agr.

Expt. Sta. Tech. Bui. 141: 148 (redescription).

Andrena littlefieldi Viereck
Colo.

Andrena (Andrena) littlefieldi Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 563. ♂.

Andrena mellitarsis Viereck
Nev.

Andrena (Andrena) mellitarsis Viereck, 1917. Amer. Ent. Soc, Trans. 43: 384. ♂.

Andrena microsoma Viereck
Pacific Northwest.

Andrena microsoma Viereck, 1904. Canad. Ent. 36: 194. ♂.

Andrena monilicornis Cockerell
N. Mex.

Aridrena monilicornis Cockerell, 1896. Canad. Ent. 28: 181. ♂.

Taxonomy: Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 48. 6 (key).

Andrena moquiorum Viereck and Cockerell
Ariz.

Andrena moquiorum Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 21. ♂.

Andrena mustelicolor Viereck
Idaho, Wash, to Calif.

Andrena mustelicolor Viereck, 1904. Canad. Ent. 35: 189, 195. ♀, ♂.

Andrena navajorum Viereck and Cockerell
N. Mex.

Ayidrena navajorum Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 29. ♂.

Andrena neurona Viereck
B. C, Wash.

Andrena neurona Viereck, 1904. Canad. Ent. 36: 191. ♀.

Andrena nigerrima Casad
N. Mex.

Andrena nigerrima Casad, 1896. Ann. and Mag. Nat. Hist. (6) 18: 83. ♀.

Taxonomy: Linsley, 1938. Calif. Acad. Sci., Proc. (4) 23: 265, 272. 9 (key).

Andrena nothoscordi Robertson
111. (Carlinville). Pollen: Unknown, but visits flowers of Notkoscordum
bivalve.
Andrena nothoscordi Robertson, 1897. Acad. Sci. St. Louis, Trans. 7: 331. ♀.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 191. 9 (key). —Mitchell, 1960. N. C.
Agr. Expt. Sta. Tech. Bui. 141: 165 (redescription of type). — Ribble, 1968. Nebr. Univ.
State Mus., Bui. 8: 333 (subgeneric position).

Andrena nubilifascia Viereck
Calif.

Andrena (Ptilandrena) nubilifascia Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 597. ♀.

Andrena obscuripostica Viereck
Nev.

Andrena obscuripostica Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 568. ♀.

Andrena occidentalis Cockerell
N. Mex.

Andrena platyparia occidentalis Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 87. ♂.

Andrena oniscicolor (Viereck)
Oreg.

Pterandrena oniscicolor Viereck, 1904. Canad. Ent. 36: 227, 228. ♀.

Andrena paduocorum Viereck and Cockerell
Colo.

Andrena paduocorum Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 38. ♂.

Andrena pediculihirta Viereck
Calif.

Andrena (Ptilandrena) pediculihirta Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 598. ♀.

Andrena perindotata Viereck
Wash.

Andrena indotata Viereck, 1904. Canad. Ent. 36: 190. ♀. Preocc.
Andrena perindotata Viereck, 1908. Ent. News 19: 42. N. name.

Andrena pineti Cockerell
Wyo.

Andrena nigerrima pineti Cockerell, 1931. Amer. Mus. Novitates 458: 13. ♀.




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Andrena plumiscopa Timberlake
South. Calif. Pollen: Unknown, but visits flowers of Cryptantha
intermedia, Platystemon califomicus.
Andrena {Pterandrena) plumiscopa Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 376. ♀.
6.
Taxonomy: LaBerge, 1967. Nebr. Univ. State Mus., Bui. 7: 294 (subgeneric position).
polygon! Viereck and Cockerell. Colo.

Andrena polygoni Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 18. ♂.
potentillarutn Viereck. Alta. to Colo. Pollen: Unknown, but visits flowers of Potentilla.

Andrena (Andrena) potentillarum, Viereck, 1924. Canad. Ent. 56: 79. ♀.

Andrena pulverea Viereck
Calif.

Andrena (Andrena) pulverea Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 569. ♀.

Andrena purpurina Viereck and Cockerell
Colo.

Andrena purpurina Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 16. ♀.

Andrena robertsonii Dalla Torre
Colo, and Minn, to N. S., south to Ark. and Ga. Pollen: Unknown, but
visits flowers of Apocynum, Aronia, Aruncus, Brassica, Ceanothus, Cryptotaenia,
Pastinaca, Ptelea, Pyracantha, Rhus, Rubus, Taenidia, Viburnum. Predator:
Philanthus albopilosiis Cress.
Andrena serotina Robertson, 1893. Amer. Ent. Soc, Trans. 20: 148. 9 (cJ misdet.). Preocc.
Anthrena robertsonii Dalla Torre, 1896. Cat. Hym., v. 10, p. 149. N. name.

Taxonomy: Robertson, 1902. Amer. Ent. Soc, Trans. 28: 193. 9, 6 (key). — Atwood, 1934.
Canad. Jour. Res. 10: 208, 209. 9, 6 (key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141: 237-238, figs. 46-47, 49, table 6 (redescription).

Biology: Evans, 1975. Ent. Soc. Amer., Ann. 68: 891 (predator).

Andrena rufojugata Cockerell
Wyo.

Andrena rufojugata Cockerell, 1931. Amer. Mus. Novitates 458: 17. ♀.

Andrena sancta Viereck
Calif.

Andrena (Andrena) sancta Viereck, 1916. Acad. Nat. Sci. Phila., Proc. 68: 571. ♀.

Andrena scotoptera Cockerell
Tex.

Andrena scotoptera Cockerell, 1934. Amer. Mus. Novitates 732: 1. ♀.

Andrena stenosoma Viereck
Nev.

Andrena (Andrena) stenosoma Viereck, 1925. Amer. Ent. Soc, Trans. 51: 135. tJ.

Andrena stictigastra Viereck
Calif.

Andrena (Andrena) stictigastra Viereck, 1917. Acad. Nat. Sci. Phila., Proc. 68: 579. ♀.

Taxonomy: Linsley and MacSwain, 1961. Pan-Pacific Ent. 37: 129 (subgeneric position).

Andrena subarctica Cockerell
Alta. (Hay River).

Andre7ia subarctica Cockerell, 1936. Canad. Ent. 68: 282. ♀.

Andrena tetonorum
***authority mismatch
Viereck and Cockerell. Nebr.

Andrena tetonorum Viereck and Cockerell, 1914. U. S. Natl. Mus., Proc. 48: 24. ♂.

Andrena trivialis Viereck
Calif.

Andrena (Andrena) trivialis Viereck, 1917. Amer. Ent. Soc, Trans. 43: 388. ♀.

Andrena variantia Linsley
B. C.

Andrena (Andrena) varia Viereck, 1924. Canad. Ent. 56: 81. ♀. Preocc.

Andreyia variantia Linsley, 1951. U. S. Dept. Agr., Agr. Monog. 2: 1084. N. name.

Andrena vicinoides Viereck
B. C, Wash.

Andrena vicinoides Viereck, 1904. Canad. Ent. 36: 191. ♀.

Nomina Nuda in Andrena Fabricius

Andrena alienoides (Viereck) Smith, 1910. Ins. New Jersey, p. 690.
Andrena delawarearum (Viereck). Smith, 1910. Ins. New Jersey, p. 690.
Andrena sparsipilosa Pierce, 1908. U. S. Natl. Mus., Bui. 66: 108.
Andrena tuberculata (Ashmead) Smith, 1910. Ins. New Jersey, p. 690.




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Genus MEGANDRENA Cockerell

Genus MEGANDRENA Subgenus MEGANDRENA Cockerell

Andrena subg. Megandrena Cockerell, 1927. Pan-Pacific Ent. 4: 42.

Type-species: Andrena {Megandrena) eyiceiiae Cockerell. Monotypic.

Megandrena enceliae (Cockerell)
Ariz., south. Calif., deserts. Pollen: Oligolectic on flowers of Larrea

tridentata, but visits other flowers for nectar including Argemone platyceros, Aster,
Cirsium, Encelia calif ortiica, E.farinosa, Eschscholzia darwinensis, Geraea canescens,
Ferrocactus ancanthodes, Opimtia basilaris, 0. echinocarpa.
Andrena (Megandrena) enceliae Cockerell, 1927. Pan-Pacific Ent. 4: 43. ♀.

Biology: Kurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 23-24, table 14 (floral
relationships, sleep).

Genus MEGANDRENA Subgenus ERYTHRANDRENA Zavortink

Megandrena subg. Erythrandrena Zavortink, 1972. Ent. Soc. Wash., Proc. 74: 61.

Type-species: Megandrena (Erythrandrena) meiitzeliae Zavortink. Monotypic and
orig. desig.

Megandrena mentzeliae Zavortink
Nev. (Spring Mts., northwest of Las Vegas). Pollen: Oligolectic on
flowers of Mentzelia tricuspis.
Megandrena (Erythrandrena) mentzeliae Zavortink, 1972. Ent. Soc. Wash., Proc. 74: 64,
figs. 1-3. V, d.

Biology: Zavortink, 1972. Ent. Soc. Wash., Proc. 74: 69-75, figs. 2-3 (floral relationships,
mating, sleep).

Genus ANCYLANDRENA Cockerell

Andrena subg. Ancylandrena Cockerell, 1930. Pan-Pacific Ent. 7: 5.

Type-species: Andrena (Ancylandrena) heterodoxa Cockerell. Monotypic.

Revision: Zavortink, 1974. CaHf. Acad. Sci., Occas. Papers, 109: 1-36, 6 figs. (N. Amer. spp.).

Ancylandrena atoposoma (Cockerell)
South. Calif.; Mexico (Baja California and Sonora). Pollen: Apparently
collects pollen from the flowers of the leguminous genera Lotus and Lupinus including
Lotus glaber, L. scoparius, Lupinus formosus, L. hallii, L. paynei, but visits these and
other flowers for nectar including Calochortus splendens, Ceanothus, Convolvulus,
Cryptantha intermedia, Eriodictyon trichocalyx, Eriogonum fasciculatum, Gilia
virgata, Penstemon, Phacelia ramosissima, Sphaeralcea ambigua.
Andrena (Ancylandrena) heterodoxa Cockerell, 1930. Pan-Pacific Ent. 7: 6. ♂, ♀. Preocc.
Andrena atoposoma Cockerell, 1934. Pan-Pacific Ent. 10: 82. N. name.

Ancylandrena koebelei (Timberlake)
Southeast. Calif, and south. Nev. Pollen: Possibly ohgolectic on flowers
of Dalea including D.fremontii, but visits other flowers presumably for nectar including
Echinocactus, Oenothera refracta, Stanleya pinnata.
Megandrena (Ancylandrena) koebelei Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 413.
6.

Biology: Zavortink, 1974. Calif. Acad. Sci., Occas. Papers, 109: 28 (matinal behavior).

Ancylandrena larreae Timberlake
N. Mex. (Lordsburg), west to south. Calif, and south. Nev., deserts.

Parasite: Hexepeolus rhodogyne Linsley and Michener? Pollen: Oligolectic on flowers of
Larrea tridentata, but visits these flowers and others for nectar including Dalea greggii,
Prosopis jnliflora.
Megandrena (Ancylandrena) larreae Timberlake, 1951. U. S. Natl. Mus., Proc. 101: 411. ♀.
6.

Biology: Zavortink, 1974. Calif. Acad. Sci., Occas. Papers 109: 23 (floral relationships).
—Kurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 22-23, fig. 4, tables 2, 9, 11, 12-14
(floral relationships, parasite).

Ancylandrena timberlakei Zavortink
Ariz, to south. CaHf. and south. Nev. Pollen: Apparently polylectic,
visits flowers of Dalea fremontii, Eriodictyon trichocalyx, Hyptis emoryi, Larrea




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tridentata, Lepidium fremontii, Malacotkrix, Mentzelia involucrata, M. tricuspis,
Phacelia, Sphaeralcea ambigua, Verbesina.
Ancylandreyia timberlakei Zavortink, 1974. Calif. Acad. Sci., Occas. Papers 109: 16, figs. 1,
4. ♂, ♀.
Biology: Zavortink, 1974. Calif. Acad. Sci., Occas. Papers 109: 19 (floral relationships).

Subfamily PANURGINAE

Taxonomy: Rozen, 1966. Amer. Mus. Novitates 2259: 1-22, 54 figs, (larvae). —Yager and
Rozen, 1966. Amer. Mus. Novitates 2265: 6-13, figs. 7-18 (pupae). —Rozen, 1970. Amer.
Mus. Novitates 2416: 1-16, 19 figs, (immature stages).

Biology: Rozen, 1967. Amer. Mus. Novitates 2297: 1-44, 18 figs, (review of N. Amer. spp.).

Genus PROTANDRENA Cockercil

Protaiidrena Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 91.

Type-species: Andrena mauriila Cockerell. Desig. by Sandhouse, 1943.
Austrandrena Timberlake, 1906. Psyche 13: 37.

Type-species: Andrena modesta Smith. Monotypic and orig. desig.
Proioandrena Cresson, 1928. Amer. Ent. Soc, Mem. 5: 58. Emend.

Revision: Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 133-227, 93 figs. (N. Amer. spp.).

Taxonomy: Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 92 (tax. characters). —Cockerell,
1896. Canad. Ent. 28: 184 (tax. characters). —Timberlake, 1955. Bol. Lab. Zool. Gen. Agr.
33: 398-409 (key to females, as Psaenythia). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141: 286-287, figs. 58, 67-68, table 8 (east. U. S. spp.). —Timberlake, 1973. Calif. Univ.
Pubs. Ent. 72: 1 (tax. status).

Protandrena amplipennis Timberlake
Ariz. (10 mi. S.W. of Patagonia). Pollen: Unknown, but visits flowers
of Croton.
Protandrena amplipennis Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 146. ♀.

Protandrena bancrofti Dunning
Ariz., N. Mex., Tex., Kans., Mo., Okla., Nebr., Colo., Wyo., Iowa, N. Dak.,
Ind., Ill, Wis. Pollen: Unknown, but visits flowers oi Amorpha, Asclepias, Astragalus,
Baccharis, Eysenhardtia, Giiardiola, Helianthus, Heterotheca, Lepidium, Medicago,
Melilotus, Monarda, Petalostemon, Psoralea, Solanum, Symphoricarpos.
Protandrena Bancrofti Dunning, 1897. Canad. Ent. 29: 264. ♀.

Taxonomy: Cockerell, 1921. Amer. Mus. Novitates 24: 13. 9. —Timberlake, 1976. Amer. Ent.
Soc, Trans. 102: 150, figs. 4, 5, 6 (geogr. and floral records).

Biology: Chandler, 1962. Kans. Ent. Soc, Jour. 35: 314 (nest).

Protandrena bicolor (Timberlake)
Tex. to Ariz.; Mexico (Coahuila, Durango). Pollen: Collects pollen from
flowers of the composite genus Bahia, but also visits flowers of Verbesina.
Psaenythia bicolor Timberlake, 1955. Bol. Lab. Zool. Gen. Agr. 33: 401. ♀.

Taxonomy: Rozen, 1970. Amer. Mus. Novitates 2416: 9-13, figs. 9-17 (larva, pupa).
—Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 150-152, figs. 7, 8, 9. ♂.

Biology: Rozen, 1967. Amer. Mus. Novitates 2297: 17-19, fig. 6 (nesting activity, nest
architecture, floral relationships). —Rozen, 1970. Amer. Mus. Novitates 2416: 6-7, fig. 2
(nest architecture, nesting habits).

Protandrena bishoppi Crawford
Tex.

Protandrena bishoppi Crawford, 1916. Ent. Soc. Wash., Proc. 19: 128. ♀.

Taxonomy: Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 152-153 (notes on type).

Protandrena cockerelli Dunning
N. J., Ind., 111., Iowa, Mich., Nebr., Mo., Kans., Colo., N. Mex., Tex. Pollen:
Unknown, but visits flowers o{ Acerates, Amphiachyris, Asclepias, Cassia, Euphorbia
corollata, Gaillardia, Melilotus, Monarda, Petalostemon, Solanum, Teucrium.
Protandrena Cockerelli Dunning, 1897. Canad. Ent. 29: 47. ♀.




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Taxonomy: Cockerell, 1899. Ent. News 10: 3. 6. — Timberlake, 1955. Bol. Lab. Zool. Gen.
Agr. 33: 400 (tax. status). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 287 (tax.
characters and status). —Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 156-157 (tax.
characters and status).

Protandrena cognata Timberlake
Ariz. (Cochise County).

Proiandrena cognata Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 157. ♀.

Protandrena duplonotata (Timberlake)
Ariz. (Pinal Co.).; Mexico (Sonora). Pollen: Unknown, but visits
flowers of Hymenothrix.

Psaenythia duplonotata Timberlake, 1955. Bol. Lab. Zool. Gen. Agr. 33: 407. ♀.

Taxonomy: Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 159 (geogr. and floral record).

Protandrena euphorbiae (Timberlake)
Ariz. (Hope and Picacho Pass); Mexico (Sonora). Pollen: Unknown,
but visits flowers of Euphorbia, including E. capitellata.
Psaenythia euphorbiae Timberlake, 1955. Bol. Lab. Zool. Gen. Agr. 33: 406. ♀.

Taxonomy: Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 161-162, figs. 16, 17, 18. ♂.

Protandrena heteromorpha (Cockerell)
N. Mex., Ariz. Pollen: Unknown, but visits flowers of Aplopappus
gracilis, Gutierrezia microcephala, Heterotheca subaxillaris, Verbesi7ia encelioides.
Andrena heteromorpha Cockerell, 1896. Canad. Ent. 28: 180. ♀, ♂.

Taxonomy: Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 168-171, figs. 25, 26, 27
(redescription).

Protandrena hurdi Timberlake
N. Mex. (Rodeo and 25 miles east of Steins), Ariz. (Chiricahua and

Huachuca Mts.). Pollen: Unknown, but visits flowers o{ Asclepias subverticillata.
Protandrena hurdi Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 171, figs. 28, 29, 30. ♂.

Protandrena kansensis Timberlake
Kans. (Douglas and Greenwood Counties). Pollen: Unknown, but visits
flowers of Am phiachyris dracunculoides.
Protandrena kansensis Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 175, figs. 34, 35, 36.
9, (J.

Protandrena leucopus Timberlake
Ariz. (Cochise and Pima Counties).

Protandrena leucopus Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 178, figs. 40, 41, 42.
6.
mauruia (Cockerell). Tex.

Andrena mauruia Cockerell, 1896. Canad. Ent. 28: 179. ♀.

Protandrena mexicanorum (Cockerell)
N. Dak., Nebr. Colo, and Tex., west to Ariz.; Mexico (Chihuahua,
Coahuila, Durango and Sonora). Pollen: Unknown, but visits flowers of Asclepias,
Kallstroemia grandiflora, Solanum. Protandrena durangoensis Timberlake is possibly
a subspecies of this.
Andrena mexicanorum Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 85. 9
Andrena asclepiadis Cockerell, 1896. Ann. and Mag. Nat. Hist. (6) 18: 86. ♂.

Taxonomy: Timberlake, 1955. Bol. Lab. Zool. Gen. Agr. 33: 400 (synonymy, tax. status, flower
records). —Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 188-189, figs. 55, 56, 57 (tax.
characters and status).

Protandrena pectidis (Timberlake)
Ariz., N. Mex.; Mexico (Sonora). Pollen: Unknown, but visits flowers of
Pedis papposa, Viguiera annua.
Psaenythia pectidis Timberlake, 1955. Bol. Lab. Zool. Gen. Agr. 33: 405. ♀.

Taxonomy: Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 191-192, figs. 58, 59, 60. ♂.

Protandrena sorocula Timberlake
N. Mex. (7 miles southeast of Rodeo).

Protandrena sorocula Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 205. ♀.

Protandrena sphaeralceae Timberlake
Ariz. (Cochise County). Pollen: Unknown, but visits flowers of
Sphaeralcea.
Protandrena sphaeralceae Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 207. ♀.

Protandrena swenki Crawford
Nebr.

Protandrena swenki Crawford, 1913. U. S. Natl. Mus., Proc 45: 241. ♀.

Taxonomy: Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 191-192, figs. 58, 59, 60. ♂.




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Protandrena texana Timberlake
Tex. (Archer, Gillespie and Throckmorton Counties). Pollen: Unknown, but
visits flowers of Teucrium laciniatum.
Protandrena texana Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 211, figs. 76, 77, 78. ♂.

Protandrena tidestromiae Timberlake
N. Mex., Ariz. Pollen: Collects pollen from flowers of Tidestromia
including T. lanuginosa, but visits other flowers presumably for nectar including
Verbesi7ia.
Protandrena tidestromiae Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 212, figs. 79, 80,
81. ♀, ♂.

Protandrena trifoliata (Cockerell)
N. Mex., Ariz. Pollen: Unknown, but visits flowers of Baccharis
glutinosa, Euphorbia capitellata, Solanum, Trifolium, Verbesina.
Andrena trifoliata Cockerell, 1896. Canad. Ent. 28: 179. ♀.

Taxonomy: Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 215-217, figs. 82, 83, 84. ♂.

Protandrena verbesinae (Timberlake)
West Tex. to Ariz.; Mexico (Coahuila and Durango). Pollen:

Unknown, but visits flowers of Asclepias, Baileya, Heterotheca, Pedis, Verbesina
encelioides.
Psaenytkia verbesinae Timberlake, 1955. Bol. Lab. Zool. Gen. Agr. 33: 402. ♀, ♂.

Taxonomy: Timberlake, 1976. Amer. Ent. Soc, Trans. 102: 222-224, figs. 88, 89, 90 (geogr. and
floral records).

Genus PANURGINUS Nylander

Panurginus Nylander, 1848. Notiser Sallskapet Flora Fauna Fenn. 1: 228.

Type-species: Panurginus niger Nylander. Monotypic.
Greeleyella Cockerell, 1904. Entomologist 37: 235.

Type-species: Greeleyella beardsleyi Cockerell. Monotypic and orig. desig.
Birkmania Viereck, 1909. Ent. Soc. Wash. Proc. 11: 50.

Type-species: Panurginus polytrichus Cockerell. Monotypic and orig. desig.
(=Birkmania andrenoides Viereck).

Revision: Crawford, 1926. Ent. Soc. Wash., Proc. 28: 207-214. — Michener, 1935. Canad. Ent.
67: 275-278 (species with blackfaced males).

Taxonomy: Rozen, 1966. Amer. Mus. Novitates 2259: 5, 10-12, figs. 17-28 (larvae).

Panurginus armaticeps Cockerell
South. Calif. Pollen: Unknown, but visits flowers of Ceanothus

integerrimus, Cryptantka intermedia, Gilia integrifolia, Phacelia davidsonii, P. distans,
Pote7itilla glandulosa, Prunus demissa, Rhamnus crocea, Salix laevigata.
Panurginus armaticeps Cockerell, 1916. Ann. and Mag. Nat. Hist. (8) 17: 280. "♀"=♂.

Taxonomy: Michener, 1937. Ann. and Mag. Nat. Hist. (10) 19: 322. ♂, ♀.

Panurginus atramontensis Crawford
Md. to Ga., Miss. Pollen: Unknown, but visits flowers of Polycodium.
Panurginus atramontensis Crawford, 1926. Ent. Soc. Wash., Proc. 28: 210. cJ, 9.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 259-260, figs. 59-61, table 8
(redescription).

Panurginus atriceps (Cresson)
Idaho, Wash., Oreg., Calif. Pollen: Collects pollen from flowers of Dowingia
bella or D. cuspidata, but also visits flowers of Ceanothus dentatus, C. integei-rinnis,
Cryptantha intermedia, Lasthenia chrysostoma, Montia perfoliata, Nemophila
maculata, N. pulchella, Ranuncuhis, Rhamnus crocea, Rhus diversiloba, Tamarix
gallica. Predator: Philanthus pulcher Dalla Torre.
Calliopsis atriceps Cresson, 1878. Amer. Ent. Soc, Trans. 7: 67. ♂.

Taxonomy: Cockerell, 1898. Amer. Ent. Soc, Trans. 25: 194. 9. —Cockerell, 1925. Calif Acad.
Sci., Proc (4) 14: 198. i, 9. —Rust, 1976. Pan-Pacific Ent. 52: 165, figs. 3, 10-16 (larva).

Biology: Rust, 1976. Pan-Pacific Ent. 52: 161-166, figs. 2-9 (nest site, nest architecture, cell
provisions, adult and immature habits).

Panurginus beardsleyi (Cockerell)
N. Dak., Nebr., Colo. Pollen: Unknown, but visits flowers of
Malvastrum.
Greeleyella beardsleyi Cockerell, 1904. Entomologist 37: 236. ♀.




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Panurginus maivastri Swenk and Cockerell, 1907. Ent. News 18: 179. ♂, ♀.

Taxonomy: Cockerell, 1906. Canad. Ent. 38: 164. ♂.

Panurginus bilobatus Michener
South. Calif. Pollen: Unknown, but visits flowers of Astragalus,

Crypta7itha intermedia, Gilia integrifolia, Lastbenia gracilis, Phacelia distans, P.
hispida, Rha^nnus crocea, R. ilicifolia. Salvia coiumbariae, Salix laevigata. Sisymbrium
irio.
Panurginus bilobatus Michener, 1937. Ann. and Mag. Nat. Hist. (10) 19: 322. ♂, ♀.

Panurginus ceanothi Michener
South. Calif. Pollen: Unknown, but visits flowers of Ceanotkus.

Pannrginns ceanothi Michener, 1935. Pan-Pacific Ent. 11: 180. ♂.

Panurginus cressoniellus Cockerell
Colo., N. Mex. Predator: Philanthus pulcher Dalla Torre.
Calliopsis clypeata Cresson, 1878. Amer. Ent. Soc, Trans. 7: 67. ♂. Preocc.
Panurginus cressoniellus Cockerell, 1898. Canad. Ent. 30: 29. N. name.
Panurginus cressoniellus calockorti Cockerell, 1906. Ann. and Mag. Nat. Hist. (7) 17: 222.

9.
/Panurginus verus Cockerell, 1901. Psyche 9: 163. ♀.

Taxonomy: Cockerell, 1897. Acad. Nat. Sci. Phila., Proc. 49: 350. 6,9. — Viereck, 1903. Amer.
Ent. Soc, Trans. 29: 49. 6. —Cockerell, 1912. Ent. News 23: 445 (synonymy).

Panurginus emarginatus Michener
Calif.

Panurginus emargiiiatus Michener, 1935. Pan-Pacific Ent. 11: 179. ♂, ♀.

Panurginus gabrielis Michener
South. Calif.; Mexico (Baja California). Pollen: Unknown, but visits flowers
of Arctostaphylos glauca, Ceanothus crassifolius, C. greggii, C. leucodennis, C.
vei~rucosus, Rhainnus californica, R. crocea.
Panurginus gabrielis Michener, 1935. Canad. Ent. 67: 276. ♂.

Taxonomy: Michener, 1937. Ann. and Mag. Nat. Hist. (10) 19: 321 (tax. characters).

Panurginus gracilis Michener
Calif.; Mexico (Baja California). Pollen: Unknown, but visits flowers of
Arctostaphylos pungens, Baccharis, Ceanothus crassifolius, C. verrucosus, Lasthenia
chrysostoma, Rhainnus crocea, Salix lasiolepis, Salvia mellifera.
Panurginus gracilis Michener, 1935. Canad. Ent. 67: 276. ♂, ♀.

Panurginus ineptus Cockerell
Colo., Wash., Oreg., Utah.

Panurginus ineptus Cockerell, 1922. Amer. Mus. Novitates 36: 8. ♀.
Panurginus bakeri Crawford, 1926. Ent. Soc. Wash., Proc. 28: 213. ♂. Preocc.
Panurginus rohweri Crawford, 1932. Ent. Soc. Wash., Proc. 34: 78. N. name.

Taxonomy: Timberlake, 1971. Pan-Pacific Ent. 47: 148 (tax. status, synonymy).

Panurginus maritimus Michener
Calif.

Panurginus maritimus Michener, 1935. Pan-Pacific Ent. 11: 178. ♂.

Taxonomy: Michener, 1935. Canad. Ent. 67: 277. ♀.

Panurginus melanocephalus (Cockerell)
Calif. Parasite: Zodion cinereiventre Van Duzee, Z. nigrifrons
Krober. Pollen: Collects pollen of Ranunculus califomicus, but visits these and other
flowers for nectar.
Panurgus melanocephalus Cockerell, 1926. Pan-Pacific Ent. 3: 80. ♂, ♀.
Panurginus morrisoni Crawford, 1926. Ent. Soc. Wash., Proc. 28: 209. ♂.

Taxonomy: Michener, 1953. Kans. Univ. Sci. Bui. 35: 1036, figs. 86-88 (larva). — Rozen, 1966.
Amer. Mus. Novitates 2259: 12, figs. 21-23 (larva).

Biology: MacSwain and Bohart, 1947. Pan-Pacific Ent. 23: 30 (parasite). — Linsley and
MacSwain, 1959. Calif. Univ. Pubs. Ent. 16: 20-21, 28 pi. 6 (nest, life history, mating, sex
ratio, floral relationships, parasite).

Panurginus nigrellus Crawford
Calif. Pollen: Unknown, but visits flowers of Arctostaphylos, Baccharis
glutinosa, Cryptantha intermedia, Lasthenia chrysostoma, Nemophila pulchella.
Ranunculus califomicus, Salix lasiolepis, Tamarix gallica.
Panurginus nigrellus Crawford, 1926. Ent. Soc. Wash., Proc. 28: 210. ♂.

Taxonomy: Michener, 1935. Pan-Pacific Ent. 11: 180. ♀.

Panurginus nigrihirtus Michener
Calif.

Panurginus nigrihirtus Michener, 1935. Pan-Pacific Ent. 11: 179. ♂, ♀.




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Panurginus occidentalis (Crawford)
Calif. Pollen: Collects pollen from flowers oi Lunnanthes douglasii.
Greeleyella occidentalis Crawford, 1916. Insecutor Inscitiae Menstruus 4: 105. ♂, ♀.

Biology: Rust, 1976. Pan-Pacific Ent. 52: 159, 165-166, fig. 1 (mating behavior, pollen and
nectar source).

Panurginus polytrichus Cockerell
Miss., La., Tex.

Panurginus polytrichus Cockerell, 1909. Ann. and Mag. Nat. Hist. (8) 4: 28. ♂.
Birkmania andrenoides Viereck, 1909. Ent. Soc. Wash., Proc. 11: 50. ♀.
Greeleyella resinata Cockerell, 1910. Ann. and Mag. Nat. Hist. (8) 5: 361. ♀.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 260-262, figs. 59-61, table 8
(redescription).

Panurginus potentillae (Crawford)
N. J., Md., Va., N. C. Pollen: Collects pollen from the flowers of
Potentilla, but visits these and other flowers for nectar including Fragaria,
Ranunculus.
Greeleyella potentillae Crawford, 1916. Insecutor Inscitiae Menstruus 4: 104. ♂, ♀.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 262-263, figs. 59-61, table 8
(redescription). — Rozen, 1966. Amer. Mus. Novitates 2259: 10-12, figs. 17-20 (larva).
— Rozen, 1967. Amer. Mus. Novitates 2297: 28-29, figs. 12-13, table 2 (egg, larva).

Biology: Rozen, 1967. Amer. Mus. Novitates 2297: 27-30, tables 1-2 (nest architecture, life
history, floral relationship).

Genus PSEUDOPANURGUS Cockerell

Pseudopanurgus Cockerell, 1897. Canad. Ent. 29: 290.

Type-species: Panurgus aetliiops Cresson. Orig. desig.
Protandrenopsis Crawford, 1903. Canad. Ent. 35: 337.

Type-species: Panurgus aethiops Cresson. Monotypic and orig. desig.
(=Protandrenopsis fuscipennis Crawford).

Revision: Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 1-58, 4 plates, 56 figs. (N. Amer. spp.).

Taxonomy: Cockerell, 1898. Amer. Ent. Soc, Trans. 25: 195-198. —Cockerell, 1922. Amer.
Mus. Novitates 36: 9. -Robertson, 1922. Psyche 29: 164 (Illinois spp.). —Mitchell, 1960. N.
C. Agr. Expt. Sta. Tech. Bui. 141: 266-270, figs. 62, 65-66, table 8 (eastern U. S. spp.).
—Rozen, 1966. Amer. Mus. Novitates 2259: 5, 7-9, figs. 8-16 (larva). —Timberlake, 1967.
Amer. Mus. Novitates 2298: 1-10 (Ariz. spp.). —Timberlake, 1975. Calif. Univ. Pubs. Ent.
77: 1, 2 (key, tax. status).

Biology: Rozen, 1967. Amer. Mus. Novitates 2297: 19-27, figs. 1-3, 6-8, tables 1-2 (nest
architecture, life history, floral relationships, parasites).

Pseudopanurgus aethiops (Cresson)
Nebr., Kans., Colo., Tex., N. Mex., Ariz. Pollen: Collects pollen from the
flowers of Compositae including Baileya, Encelia, Grindelia, Helianthus, Heterotheca,
Melampodium, Prionopsis, Verbesina encelioides, but visits these and other flowers for
nectar including Asclepias.
Panurgus aethiops Cresson, 1872. Amer. Ent. Soc, Trans. 4: 259. ♀, ♂.
Protandrenopsis fuscipennis Crawford, 1903. Canad. Ent. 35: 337. ♀.

Taxonomy: Michener, 1951. In Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr.
Monog. 2: 1098 (synonymy). —Rozen, 1966. Amer. Mus. Novitates 2259: 9 figs. 9-10 (larva).
—Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 8-10 figs. 1-2 (redescription).

Biology: Hicks, 1931. Canad. Ent. 63: 172 (nest). —Rozen, 1967. Amer. Mus. Novitates 2297:
19-27, figs. 1-2, 7, tables 1-2 (nest architecture, life history, floral relationship).

Taxonomy: Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 10 (pollen sources).

Pseudopanurgus bradleyi Timberlake
Tex. (Blanco County).

Pseudopanurgus bradleyi Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 10, figs. 3-4. ♂.

Pseudopanurgus cazieri
***authority mismatch
cazieri Timberlake. Ariz. (Cochise County). Another subspecies, P. c. coahuilensis
Timberlake, occurs in Mexico.
Pseudopanurgus cazieri cazieri Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 13, figs. 7-8.
9,6.




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Pseudopanurgus dicksoni Timberlake
N. Mex., Ariz.; Mexico (Baja California and Sonora). Pollen: Unknown,
but visits flowers of Aplopappus, Argemone intermedia, Crotalaria pumila, Milleria
quinquefolia, Pedis papposa.
Pseudopanurgus dicksoni Timberlake, 1967. Amer. Mus. Novitates 2298: 8. ♀, ♂.

Taxonomy: Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 19, figs. 15-16 (tax. characters,
flower record).

Pseudopanurgus fraterculus
***authority mismatch
fraterculus (Cockerell). West. Tex., N. Mex.; Mexico (Aguascalientes, Coahuila,
Durango and San Luis Potosi). Pollen: Unknown, but visits flowers of Gutierrezia
lucida, Tidestromia.
Calliopsis fraterculus Cockerell, 1896. Canad. Ent. 28: 159. ♂, ♀.

Taxonomy: Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 21-23 (redescription, tax. status).

Pseudopanurgus fraterculus
***authority mismatch
timberlakei Cockerell. Colo., N. Mex., Ariz., Calif. (San Diego County); Mexico
(Baja California). Parasite: Holcopasites arizonicus (Linsley)?, H. insoletus (Linsley)?
Pollen: Apparently an oligolege of the Compositae, including Aplopappus gracilis,
Baileya multiradiata, B. pleniradiata, Gutierrezia califomica, G. microcepliala,
Heliopsis, Heterotheca subaxillaris, Hymenothrix wislizeni. Pedis papposa, but visits
these and other flowers for nectar including Allionia, Eriogomim.
Pseudopanurgus timberlakei Cockerell, 1931. Ent. Soc. Wash., Proc. 33: 201.

Taxonomy: Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 23-24, figs. 2, 19-20 (redescription,
floral and geogr. records).

Biology: Rozen, 1965. N. Y. Ent. Soc, Jour. 73: 88 (parasite). — Rozen, 1967. Amer. Mus.
Novitates 2297: 19-27, figs. 1-2, tables 1-2 (nest architecture, life history, pollen source,
parasite). — Hurd and Linsley, 1972. Smithsn. Contrib. Zool. 114: 39 (parasites).

Pseudopanurgus pectidellus Cockerell
N. Mex., Ariz.; Mexico (Chihuahua). Pollen: Apparently an oligolege of
the Compositae, visits flowers of Baileya, Pedis papposa, Verbesina.
Pseudopanurgus pectidellus Cockerell, 1904. Ann. and Mag. Nat. Hist. (7) 14: 26.

Taxonomy: Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 32-34, figs. 33-34 (redescription,
geogr. and floral records).

Pseudopanurgus pectiphilus (Cockerell)
N. Mex., Ariz. Pollen: Apparently an oligolege of the Compositae,
visits flowers of Baileya multiradiata, Heterotheca subaxillaris. Pedis papposa,
Tidestromia lanuginosa.
Panurginus pectiphilus Cockerell, 1913. Ann. and Mag. Nat. Hist. (8) 11: 193. ♀.

Taxonomy: Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 34-35, figs. 35-36 (redescription,

geogr. and floral records).

Pseudopanurgus perarmatus Timberlake
South. Ariz.; Mexico (Chihuahua and Sonora). Pollen: Apparently an

oligolege of the Compositae, visits flowers of Aplopappus gracilis, Heterotheca

subaxillaris, Verbesina encelioides, Zexmenia podocephala.

Pseudopanurgus perarmatus Timberlake, 1967. Amer. Mus. Novitates 2298: 5. ♀, ♂.

Taxonomy: Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 35, figs. 37-38 (tax. characters).

Pseudopanurgus perpunctatus Timberlake
N. Mex., Ariz. Pollen: Presumably an oligolege of Compositae, visits
flowers of Heterotheca, Pedis papposa.
Pseudopanurgus perpunctatus Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 35, figs. 39-40.
9,i.

Pseudopanurgus rugosus (Robertson)
Md. to Ga., west to 111., Nebr., Kans., Tex. Pollen: Apparently an oligolege
of the Compositae, especially the Helianthae, visits flowers of Bidens aristosa,
Helianthus annuus, H. divaricatus, H. mollis, H. petiolaris, H. subaxillaris, Heliopsis
helianthoides, Pontederia cordata, Rudbeckia laciniata, R. triloba, Silphium
perfoliatum, S. speciosuyyi, but visits flowers of Melilotus officinalis for nectar.
Calliopsis rugosus Robertson, 1895. Amer. Ent. Soc, Trans. 22: 121. ♀, ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 269-270, figs. 62, 65-66
(redescription). -Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 38, figs. 41-42 (tax.
characters, geogr. and floral records).




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Pseudopanurgus texanus Timberlake
Tex.; Mexico (Tamaulipas).

Pseudopanurgus texanus Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 40, figs. 45-46. ♀.
i.

Pseudopanurgus verticalis Timberlake
N. Mex., Ariz., south. Calif. (Riverside County); Mexico (Sonora).
Parasite: Hokopasites insoletus (Linsley). Pollen: Apparently an oligolege of the
Compositae; visits flowers of Bahia absinthifolia. Pedis papposa, Verbesina encelioides,
but also visits the flowers of Kallstroemia grandiflora presumably for nectar.
Pseudopanurgus verticalis Timberlake, 1967. Amer. Mus. Novitates 2298: 2. ♀, ♂.

Taxonomy: Rozen, 1966. Amer. Mus. Novitates 2259: 9, figs. 11-16 (larva, as Pseudopanurgus
sp. B). —Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 50, figs. 55-56 (tax. characters).

Biology: Rozen, 1967. Amer. Mus. Novitates 2297: 27 (parasite of Pseudopanurgus sp. B).
— Hurd and Linsley, 1972. Smithsn. Contrib. Zool. 114: 39 (parasite).

Genus ANTHEMURGUS Robertson

Anthemurgus Robertson, 1902. Canad. Ent. 34: 321.

Type-species: Anthemurgus passiflorae Robertson. Monotypic and orig. desig.

Taxonomy: Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 1 (tax. status).

Anthemurgus passiflorae Robertson
111. to N. C. Pollen: Unknown, but visits flowers of Passiflora.
Anthemurgus passiflorae Robertson, 1902. Canad. Ent. 34: 321. ♀, ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 265-266, figs. 62, 65-66
(redescription).

Genus HETEROSARUS Robertson

Heterosarus Robertson, 1918. Ent. News 29: 91.

Type-species: Caliiopsis parvus Robertson. Monotypic and orig. desig.

Revision: Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 1-56, 8 plates, 87 figs. (N. Amer. spp.).

Taxonomy: Timberlake, 1964. Amer. Mus. Novitates 2185: 2-5 (key to spp. in Amer. north of
Mexico). —Timberlake, 1973. Calif. Univ. Pubs. Ent. 72: 1, 2 (tax. status).

Heterosarus arizonicus (Timberlake)
Ariz., N. Mex. Pollen: Unknown, but visits flowers of Eriogonum.
Pseudopanurgus arizonicus Timberlake, 1964. Amer. Mus. Novitates 2185: 23. ♂, ♀.

Taxonomy: Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 14, figs. 9-10 (tax. characters).

Heterosarus bakeri (Cockerell)
Colo., N. Mex., Ariz.; Mexico (Chihuahua, Michoacan, Morelos and Puebla).
Pollen: Unknown, but visits flowers of Calochortus gunnisonii. Euphorbia, Frasera
speciosa, Helenium hoopesii, Heterotheca subaxillaris, Linum lewisii, Lopezia, Monarda
austromontana, Oxalis, Phacelia leucophylla, Potentilla filipes, P.fruticosa,
Pseudocymopterus montanus, Sedum steiiopetalum, Solidago.
Caliiopsis bakeri Cockerell, 1896. Ent. News 7: 221. ♂.

Taxonomy: Viereck, 1903. Amer. Ent. Soc, Trans. 29: 50. —Cockerell, 1910. Psyche 17: 245.
6, 9. —Timberlake, 1964. Amer. Mus. Novitates 2185: 7-8 (geogr. and floral records).
—Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 15-16, figs. 13-14 (tax. characters, geogr.
and floral records).

Heterosarus dawsoni (Timberlake)
Tex., N. Mex., Colo. Pollen: Unknown, but visits flowers of an
unspecified crucifer.
Pseudopanurgus dawsoni Timberlake, 1964. Amer. Mus. Novitates 2185: 10. ♀, ♂.

Taxonomy: Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 20, figs. 23-24 (tax. characters).

Heterosarus euphorbiae Timberlake
Ariz. (Pima County); Mexico (Chiapas, Michoacan, Morelos, Nayarit
and Puebla).
Heterosarus euphorbiae Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 24, figs. 29-30. ♂, ♀.

Heterosarus flavotinctus (Cockerell)
N. Mex., Colo., Ariz.; Mexico (Zacatecas). Pollen: Unknown, but visits
flowers of Bigelovia, Sphaeralcea.
Panurginus pauper \&r. flavotinctus Cockerell, 1898. Denison. Univ. Sci. Labs., Bui. 11: 51.

3.




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Taxonomy: Cockerell, 1898. N. Mex. Univ., Bui. 1: 51. 6. — Cockerell, 1899. Entomologist 32:
129. 9. — Timberlake, 1964. Amer. Mus. Novitates 2185: 20-23 (redescription).
— Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 26, figs. 33-34 (tax. characters and status).

Heterosarus illinoiensis (Cresson)
Que. to Fla., west to III, Kans., Okla. and Tex. Pollen: Unknown, but
visits flowers of Amphiachyris dracunculoides, Bellis integrifolia, Chrysanthemum
leucayithemum, Descourainia, Erigeron, Gaillardia, Heterotheca subaxillaris, Lepidinm.
Calliopsis illinoiensis Cresson, 1878. Amer. Ent. Soc, Trans. 7: 66. ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 274, figs. 64-66, table 8
(redescription). —Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 30-31, figs. 43-44 (tax.
status, geogr. and floral records).

Heterosarus nanulus (Timberlake)
N. Mex., Ariz.; Mexico (Baja California, Chihuahua, Durango and

Sonora). Parasite: Holcopasites tegularis Hurd and Linsley. Pollen: Unknown, but visits
flowers of Euphorbia, Physalis, Wislizenia refracta.
Pseudopanurgus nanulus Timberlake, 1964. Amer. Mus. Novitates 2185: 5. ♀, ♂.

Taxonomy: Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 37, figs. 55-56 (tax. characters).

Biology: Hurd and Linsley, 1972. Smithsn. Contrib. Zool. 114: 26, 39 (parasite).

Heterosarus neomexicanus (Cockerell)
N. Mex., Ariz.; Mexico (Distrito Federal, Durango, Guanajuato,

Hidalgo, Mexico, Michoacan, Morelos, Puebla and Tlaxcala). Pollen: Unknown, but visits
flowers of Cacalia decomposita, Ceanothus, Erigeron, Geranium, Helenium hoopesii,
Monarda austromontana, Penstemon, Solidago.
Panurginus neomexicanus Cockerell, 1898. Ann. and Mag. Nat. Hist. (7) 2: 451. ♂.
Panurginus nigrinus Viereck, 1903. Amer. Ent. Soc, Trans. 29: 50. ♀.

Taxonomy: Timberlake, 1964. Amer. Mus. Novitates 2185: 8-10 (redescription, geogr. and
floral records). —Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 37-38, figs. 57-58 (tax.
characters).

Heterosarus opacicollis (Timberlake)
Ariz. Pollen: Unknown, but visits flowers of Eriogonum.
Pseudopanurgus opacicollis Timberlake, 1964. Amer. Mus. Novitates 2185: 15. ♀.

Taxonomy: Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 38 (tax. characters).

Heterosarus opaculus (Cockerell)
Ariz. (Mud Springs, Santa Catalina Mts.).

Panurginus opaculus Cockerell, 1922. Amer. Mus. Novitates 36: 3. ♀.

Taxonomy: Timberlake, 1964. Amer. Mus. Novitates 2185: 18-20 (redescription).
—Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 39, fig. 87 (tax. characters).

Heterosarus parvus (Robertson)
Pa., N. C, Ga., Miss., Mo., 111., Wis., N. Dak., Alta., Colo., N. Mex. Pollen:
Unknown, but visits flowers of Aster, Eulophus, Geranium, Gerardia tenuifolia,
Gilleyiia, Monarda, Solidago, Thaspium.
Calliopsis parvus Robertson, 1892. Amer. Nat. 26: 273. ♀, ♂.
Pseudopanurgus gerardiae Crawford, 1932. Ent. Soc. Wash., Proc. 34: 77. ♂, ♀.
Pseudopanurgus stevensi Crawford, 1932. Ent. Soc. Wash., Proc. 34: 78. ♂, ♀.
Panurginus borealis Cockerell, 1937. Canad. Ent. 69: 33. ♀.

Taxonomy: Crawford, 1932. Ent. Soc. Wash., Proc. 34: 77 (tax. characters). —Mitchell, 1960.
N. C. Agr. Expt. Sta. Tech. Bui. 141: 277-278, figs. 64-66, table 8 (synonymy, redescription).
—Timberlake, 1964. Amer. Mus. Novitates 2185: 25 (synonymy, as stevensi).
—Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 40, figs. 63-64 (synonymy, tax. characters).

Heterosarus pauper (Cresson)
New England to Fla., west to 111. and Mo. Pollen: Collects pollen especially
from the flowers of Ceanothus americanus, but also visits flowers of Rubus presumably
for nectar.
Calliopsis pauper Cresson, 1878. Amer. Ent. Soc, Trans. 7: 66. ♀, ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 278-279, figs. 64-66, table 8
(redescription). —Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 40, figs. 65-66 (tax.
characters).

Biology: Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 40 (pollen source).




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Heterosarus pernitens (Cockerell)
Ariz., N. Mex.

Panurginus pernitens Cockerell, 1922. Amer. Mus. Novitates 36: 4. ♀.

Taxonomy: Timberlake, 1964. Amer. Mus. Novitates 2185: 14-15 (redescription).
— Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 40-41 (tax. characters).

Heterosarus subglaber Timberlake
Tex. (Del Rio).

Heterosarus subglaber Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 45, figs. 71-72. ♂.

Heterosarus townsendi (Cockerell)
Tex., N. Mex., Ariz.

Calliopsis townsendi Cockerell, 1897. Amer. Ent. Soc, Trans. 24: 152. ♂.

Taxonomy: Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 48-49, figs. 77-78. cJ, 9
(redescription).

Heterosarus virginicus (Cockerell)
Md. to Ga., W. Va. and Ala. Pollen: Unknown, but visits flowers of
Ceanothus, Houstonia purpurea.
Panurginus virginicus Cockerell, 1907. Entomologist 40: 137. ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 281-282, figs. 64-66, table 8
(redescription). —Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 52, figs. 83-84 (tax.
characters).

Genus PTEROSARUS Timberlake

Pterosarus Timberlake, 1967. Amer. Mus. Novitates 2298: 10.

Type-species: Calliopsis rudbeckiae Robertson. Orig. desig.

Taxonomy: Rozen, 1966. Amer. Mus. Novitates 2259: 5-8, figs. 1-8 (larvae, as

Pseiidopanurgus). —Timberlake, 1967. Amer. Mus. Novitates 2298: 19-22 (Key to spp.).
—Timberlake, 1975. Calif. Univ. Pubs. Ent. 77: 1, 2 (key, tax. status).

Pterosarus aestivalis (Provancher)
Que.

Panurgus aestivalis Provancher, 1882. Nat. Canad. 13: 205. ♀, ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 271-272 (tax. characters).

Pterosarus albitarsis (Cresson)
Conn, to Ga., west to III, Colo., N. Mex. and Tex., ?Calif. Pollen:

Unknown, but visits flowers of Aster ericoides villosus, Brauneria pallida, B. purpurea.
Coreopsis palmata, Helianthus divaricatus, H. mollis, Lepackys pinnata, Rudbeckia
hirta, R. laciniata, R. submentosa, Verbesina helianthoides.
Panurgus albitarsis Cresson, 1872. Amer. Ent. Soc, Trans. 4: 260. ♀, ♂.
Panurgus picipes Cresson, 1872. Amer. Ent. Soc, Trans. 4: 261. ♂.

Taxonomy: Cockerell, 1900. Canad. Ent. 32: 364. 6. —Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 266-267, figs. 62, 65-66, table 8 (synonymy, redescription). —Timberlake,
1967. Amer. Mus. Novitates 2298: 11 (tax. position).

Pterosarus altissimus (Cockerell)
Colo.

Panurginus altissimus Cockerell, 1922. Amer. Mus. Novitates 36: 3. ♂, ♀.

Pterosarus andrenoides (Smith)
N. B. to N. C, west to Minn, and III, ?N. Mex. Pollen: Unknown, but
visits flowers o{ Aster, Rudbeckia triloba, Soiidago. Predator: Philanthus lepidus
Cress.
Scrapter Andrenoides Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 121. ♀.
Calliopsis asteris Robertson, 1895. Amer. Ent. Soc, Trans. 22: 121. ♀, ♂.

Taxonomy: Cockerell, 1904. Canad. Ent. 36: 303. 9. —Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 272-273, figs. 64-66, table 8 (synonymy, redescription).

Pterosarus atricornis (Cresson)
Colo., N. Mex.

Calliopsis atricornis Cresson, 1878. Amer. Ent. Soc, Trans. 7: 67. ♂.

Panurginus atricornis (!) Viereck, 1903. Amer. Ent. Soc, Trans. 29: 50 (tax. characters).

Taxonomy: Timberlake, 1967. Amer. Mus. Novitates 2298: 11 (tax. position).

Pterosarus aurifodinae (Michener)
South. Calif., mountains.

Pseudopanurgus aurifodinae Michener, 1937. Ann. and Mag. Nat. Hist. (10) 19: 320. ♂.

Pterosarus barberi (Cockerell)
N. Mex.

Panurgiiius barberi Cockerell, 1899. Entomologist 32: 129. ♂.




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Pterosarus boylei (Cockerell)
Colo., N. Mex., Ariz. Parasite: Holcopasites insoletus (Linsley)? Pollen:
Collects pollen from the flowers of Verbesma encelioides .
Calliopsis boylei Cockerell, 1896. Canad. Ent. 28: 161. ♂.

Taxonomy: Cockerell, 1897. Acad. Nat. Sci., Phila., Proc. 49: 350. —Crawford, 1912. Canad.
Ent. 44: 368 (tax. characters). — Rozen, 1966. Amer. Mus. Novitates 2259: 6-7, figs. 1-7
(larva).

Biology: Rozen, 1967. Amer. Mus. Novitates 2297: 19-27, figs. 1-2, tables 1-2 (nest
architecture, life history, parasite).

Pterosarus californicus (Cresson)
Calif.; Mexico (Baja California). Pollen: Unknown, but visits flowers of
Chaenactis glabriuscula, Encelia califomica, Gilia muUicaidis, Lasthenia californica,
L. gracilis, Layia elegans, L. platyglossa, Ranunculus californicus.
Calliopsis californicus Cresson, 1878. Amer. Ent. Soc, Trans. 7: 67. ♂.

Pterosarus citrinifrons (Viereck)
N. Mex.

Panurginus citrinifrons Viereck, 1903. Amer. Ent. Soc, Trans. 29: 51. ♂.

Pterosarus citripes (Ashmead)
Colo.

Calliopsis citripes Ashmead, 1890. Colo. Biol. Assoc, Bui. 1: 5. ♂.

Pterosarus compositarum (Robertson)
Md. to Ga., west to 111. Pollen: Unknown, but visits flowers of
Aster, Bidens aristosa, Boltonia asteroides, Polygonum scandens, Rudbeckia triloba,
Solidago yiemoralis, S. ulmifolia.
Calliopsis compositarum Robertson, 1893. Amer. Ent. Soc, Trans. 20: 274. ♀.

Taxonomy: Robertson, 1895. Amer. Ent. Soc, Trans. 22: 121. 6. —Mitchell, 1960. N. C. Agr.
Expt. Sta. Tech. Bui. 141: 273-274, figs. 64-66 (redescription).

Pterosarus didirupa (Cockerell)
Idaho, Colo., N. Mex.

Panurginus didirupa Cockerell, 1908. Entomologist 41: 293. ♂, ♀.

Pterosarus expallidus (Swenk and Cockerell)
Nebr.

Panurginus expallidus Swenk and Cockerell, 1907. Ent. News 18: 181. ♂.

Pterosarus helianthi (Mitchell)
Ind. (Tippecanoe Co.). Pollen: Unknown, but visits flowers of Helianthus
annuus.
Pseudopanurgus (Pseudopanurgus) helianthi Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141:267. ♀.

Pterosarus horizontalis (Swenk and Cockerell)
Nebr

Panurginus horizontalis Swenk and Cockerell, 1907. Ent. News 18: 183. ♂.

Pterosarus illustris (Timberlake)
Ariz. (Chiricahua Mts.). Pollen: Unknown, but visits flowers of
Verbesina encelioides, Viguiera.
Pseudopanurgus illustris Timberlake, 1967. Amer. Mus. Novitates 2298: 18. ♀.

Pterosarus innuptus (Cockerell)
Alta., N. Dak., S. Dak., Nebr., Colo., N. Mex., Ariz. Pollen: Unknown, but
visits flowers of Helianthus.
Calliopsis innuptus Cockerell, 1896. Ent. News 7: 222. "♀"=♂.
Panurginus albitarsis \ar.fortior Cockerell, 1899. Entomologist 32: 129. ♂.
Panurginus innuptus var. absonus Cockerell, 1912. Ent. News 23: 446. ♂.

Taxonomy: Cockerell, 1898. Amer. Ent. Soc, Trans. 25: 194. ♂, ♀.

Pterosarus irregularis (Cockerell)
Colo.

■Panurginus irregularis Cockerell, 1922. Amer. Mus. Novitates 36: 2. ♂.

Pterosarus labrosiformis
***authority mismatch
distractus (Cockerell). Nebr.

Panurginus labrosiformis distractus Cockerell, 1912. Ent. News 23: 447. ♂.
labrosifromis labrosiformis (Robertson). N. C, Ga., 111. Pollen: Unknown, but visits flowers of
Actinomeris alternifolia, Bidens aristosa. Coreopsis tripteris, Helianthus divaricatus,
H. tuberosus, Heliopsis helianthoides, Rudbeckia laci7iiata, R. triloba, Silphium
perfoliatum, Solidago canadensis.
Panurginus labrosiformis Robertson, 1898. Acad. Sci. St. Louis, Trans. 8: 49. ♀, ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 274-275, figs. 64-66, table 8
(redescription).




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Pterosarus labrosus (Robertson)
N. Y. to N. C, west to Minn. Pollen: Unknown, but visits flowers of
Helianthtis divaricatus, H. tuberosus, Heliopsis, Rudbeckia lanceolata, R. triloba.
Calliopsis labrostis Robertson, 1895. Amer. Ent. Soc, Trans. 22: 122. ♀, ♂.

Taxonomy: Robertson, 1898. Acad. Sci. St. Louis, Trans. 8: 48, 9 , 6 . —Mitchell, 1960. N. C.
Agr. Expt. Sta. Tech. Bui. 141: 268-269, figs. 62, 65-66, table 8 (redescription).

Pterosarus leucopterus (Cockerell)
Colo.

Panurginus leucopterus Cockerell, 1923. Ent. News 34: 49. ♂.

Pterosarus lutzae (Cockerell)
Colo.

Panurginus lutzae Cockerell, 1922. Amer. Mus. Novitates 36: 7. ♀.

Pterosarus nebrascensis
***authority mismatch
muesebecki (Michener). N. C. to Fla., Miss. Pollen: Unknown, but visits flowers
of Aster, Chrysopsis, Haplopappus.
P seudopanurgus nebrascensis timberlakei Michener, 1947. Amer. Midland Nat. 38: 446. ♂.

9. Preocc.
P seudopanurgus (Heterosarus) nebrascensis muesebecki Michener, 1951. In Muesebeck,
Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1100. N. name.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 276-277, fig. 65, table 8
(redescription, geogr. range).

Pterosarus nebrascensis
***authority mismatch
nebrascensis (Crawford). Maine to N. J., west to Colo, and Alta. Pollen:
Unknown, but visits flowers of Aster, Solidago.
Panurginus nebrascensis Crawford, 1903. Canad. Ent. 35: 335. ♂, ♀.

Taxonomy: Crawford, 1912. Canad. Ent. 44: 368 (tax. characters). —Mitchell, 1960. N. C. Agr.
Expt. Sta. Tech. Bui. 141: 275-276, figs. 64-66, table 8 (redescription).

Pterosarus nubis (Cockerell)
N. Mex.

Pamirgimis nubis Cockerell, 1913. Ann. and Mag. Nat. Hist. (8) 11: 193. ♀.

Pterosarus occidus (Timberlake)
Ariz. Parasite: Holcopasites arizonicus (Linsley)? Pollen: Aplopappus
gracilis, Helianthus, Heterotheca subaxillaris, Verbesina encelioides.
P seudopanurgus (Pterosarus) occidus Timberlake, 1967. Amer. Mus. Novitates 2298: 14.
9,6.

Taxonomy: Rozen, 1966. Amer. Mus. Novitates 2259: 7-8, fig. 8 (larva, as species A).

Biology: Rozen, 1965. N. Y. Ent. Soc, Jour. 73: 88 (parasite). — Hurd and Linsley, 1972.
Smithsn. Contrib. Zool. 114: 39 (parasite).

Pterosarus ornatipes (Cresson)
Nebr., Colo., Tex.

Calliopsis ornatipes Cresson, 1872. Amer. Ent. Soc, Trans. 4: 260. ♂.

Taxonomy: Crawford, 1912. Canad. Ent. 44: 368. (tax. characters).

Pterosarus pecki (Cockerell)
Alta.

Panurginus pecki Cockerell, 1937. Canad. Ent. 69: 114. ♂.

Pterosarus perlaevis (Cockerell)
Colo., N. Mex., Ariz. Parasite: Holcopasites insoletus (Linsley)?

Calliopsis perlaevis Cockerell, 1896. Canad. Ent. 28: 160. ♀.

Taxonomy: Cockerell, 1922. Amer. Mus. Novitates 36: 5.

Biology: Hurd and Linsley, 1972. Smithsn. Contrib. Zool. 114: 39 (parasite).

Pterosarus piercei
***authority mismatch
albertensis (Cockerell). Alta.

Panurginus piercei albertensis Cockerell, 1937. Canad. Ent. 69: 33. ♀.

Pterosarus piercei
***authority mismatch
piercei (Crawford). N. Dak., Nebr., Colo., N. Mex.

Pamirginus piercei Crawford, 1903. Canad. Ent. 35: 335. ♂, ♀.

Biology: Pierce, 1904. Nebr. Univ., Studies 4: 185. —Hicks, 1936. Canad. Ent. 68: 47 (nesting

habits).
porterae (Cockerell). N. Mex., Colo., Ariz.

Panurginus Porterae Cockerell, 1900. Canad. Ent. 32: 364. ♂, ♀.

Pterosarus pulchricornis (Cockerell)
Colo.

Panurgimis pulchricornis Cockerell, 1922. Amer. Mus. Novitates 36: 8. ♀.

Pterosarus renimaculatus (Cockerell)
N. Dak., Wyo., Nebr., Colo., N. Mex., Tex.

Calliopsis renimaculatus Cockerell, 1896. Ent. News 7: 222. ♀.




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Taxonomy: Swenk and Cockerell, 1907. Ent. News 18: 180. 9 (only). -Crawford, 1915. U. S.
Natl. Mus., Proc. 94: 579. 9, cJ. —Stevens, 1919. Canad. Ent. 51: 208. ♂.

Pterosarus rudbeckiae (Robertson)
Wis., 111. Pollen: Unknown, but visits flowers of Aster, Bidens

aristosa, Helianthus divaricatns, Rudbeckia hirta, R. laciniata, R. siihmentosa, R.
triloba.
Calliopsis rudbeckiae Robertson, 1895. Amer. Ent. Soc, Trans. 22: 122. ♀, ♂.

Biology: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 279-280, figs. 64-66, table 8
(redescription).

Pterosarus simulans (Swenk and Cockerell)
N. Dak., Nebr.

Pannrginus simulans Swenk and Cockerell, 1907. Ent. News 18: 182. ♂, ♀.

Pterosarus solidaginis (Robertson)
New England west to 111., south to Miss. Pollen: Unknown, but visits
flowers oi Bidens aristosa, Boltonia asteroides. Coreopsis tripteris, Helianthus
grosse-serratus, H. tuberosus, Rudbeckia submeyitosa, Solidago canadensis.
Calliopsis solidaginis Robertson, 1893. Amer. Ent. Soc, Trans. 20: 274. ♀.

Taxonomy: Robertson, 1898. Acad. Sci. St. Louis, Trans. 8: 50. 6. —Mitchell, 1960. N. C. Agr.
Expt. Sta. Tech. Bui. 141: 280-281, figs. 64-66, table 8 (redescription).

Pterosarus stathamae (Timberlake)
Ariz. (Southwestern Research Station).

Pseudopanurgus {Pterosarus) stathamae Timberlake, 1967. Amer. Mus. Novitates 2298:
16. ♀.

Pterosarus stigmalis (Swenk and Cockerell)
Nebr.

Pannrginus stigmalis Swenk and Cockerell, 1907. Ent. News 18: 180. ♂, ♀.

Pterosarus vicinus (Timberlake)
Ariz. (Cochise County). Pollen: Unknown, but visits flowers of
Aplopappus gracilis.
Pseudopanurgus (Pterosarus) vicinus Timberlake, 1967. Amer. Mus. Novitates 2298: 11.
9, 6.

Pterosarus ximenesiae (Cockerell)
N. Mex.

Pannrginus ximenesiae Cockerell, 1913. Ann. and Mag. Nat. Hist. (8) 11: 195. ♀.

Genus METAPSAENYTHIA Timberlake

Metapsaenythia Timberlake, 1969. Ent. News 80: 89.

Type-species: Calliopsis abdoininalis Cresson. Orig. desig.

Taxonomy: Timberlake, 1969. Ent. News 80: 89-92 (N. Amer. spp.). —Timberlake, 1973. Calif.
Univ. Pubs. Ent. 72: 1 (tax. relationships).

Metapsaenythia abdominalis
***authority mismatch
abdominalis (Cresson). Tex., Kans. Pollen: Collects pollen from flowers of

Monarda, but visits other flowers for nectar including Aster tanacetifolius, Brazoria
truncata, Dalea aurea, Gaillardia, Helianthus annuus.
Calliopsis abdominalis Cresson, 1878. Amer. Ent. Soc, Trans. 7: 68. ♀, ♂.
Camptopoeum semirufum Cockerell, 1937. Amer. Mus. Novitates 899: 1. ♀.

Biology: Timberlake, 1969. Ent. News 80: 90 (floral relationships).

Metapsaenythia abdominalis
***authority mismatch
tricolor (Cockerell). N. J. and Pa. to Ga. Pollen: Apparently an oligolege of
Monarda including M. punctata.
Calliopsis tricolor Cockerell, 1897. Amer. Ent. Soc, Trans. 24: 151. ♀.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 270-271, figs. 62-63, 66, table
8 (redescription). —Timberlake, 1969. Ent. News 80: 90-91 (tax. status).

Genus NOMADOPSIS Ashmead

Revision: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 1-202, 218 figs., 17 maps (monograph,
includes information on biology, systematics and ecology).

Taxonomy: Cockerell, 1898. Amer. Ent. Soc, Trans. 25: 195-198 (tax. status and characters).
—Timberlake, 1952. Ent. Soc. Amer., Ann. 45: 104-107 (tax. status). —Rozen, 1959. Ent.
Soc Wash., Proc. 61: 255-259 (geogr. and floral records). —Rozen, 1966. Amer. Mus.
Novitates 2259: 5, 16 (larvae). —Yager and Rozen, 1966. Amer. Mus. Novitates 2265: 6-8,
figs. 3-10 (pupae).




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Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 4-54 (life history, nest architecture, adult
behavior, comparative ecology, floral relationships, parasites, predators). —Rozen, 1963.
Amer. Mus. Novitates 2142: 1-17, 20 figs., 2 tables (nest architecture, life history, floral
relationships). —Rozen, 1967. Amer. Mus. Novitates 2297: 1-44, 18 figs. 2 tables (review).

Genus NOMADOPSIS Subgenus NOMADOPSIS Ashmead

Nomadopsis Ashmead, 1898. Psyche 8: 285.

Type-species: Peridita zo7ialis Cresson. Monotypic and orig. desig. Lapsus for
Calliopsis zojialis Cresson.
Spinoliella subg. Claremontiella Cockerell, 1933. Pan-Pacific Ent. 9: 25.

Type-species: Calliopsis zonalis Cresson. Monotypic and orig. desig. (=Spinoliella
euxantha Cockerell).

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 60-63 (tax. status).

Nomadopsis cincta
***authority mismatch
cincta (Cresson). Calif, (east side of Sierra Nevada, Tulare and Trinity Counties),
Nevada. Pollen: Apparently an oligolege of Calochortus including C. liiteus.
Calliopsis cindus Cresson, 1879. Amer. Ent. Soc, Trans. 7: 201. ♀, ♂.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 83, figs. 94-98, map 4 (redescription, tax.
status).

Nomadopsis cincta
***authority mismatch
hurdi Rozen. Calif. (Sacramento). Pollen: Collects pollen from the flowers of

Calochortus luteus.
No7Hadopsis cincta hurdi Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 85. ♂, ♀.

Nomadopsis comptula (Cockerell)
Calif. (Tulare, Inyo, and San Bernardino Counties). Pollen: Collects

pollen from the flowers Potentilla, including P. bolanderi var. bernardina, P. b. var.

parryi, P. gracilis, but visits these and other flowers including Calochortus palnieri var.

paludicola for nectar.
Spinoliella comptula Cockerell, 1916. Ann. and Mag. Nat. Hist. (8) 17: 284. ♀.
Spinoliella edivardsii media Michener, 1937. Ann. and Mag. Nat. Hist. (10) 19: 325. ♀, ♂.
Spinoliella edwardsii bernardina Michener, 1937. Ann. and Mag. Nat. Hist. (10) 19: 326. ♀.

6.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 75-77, figs. 79-83, map 3 (redescription,
synonymy).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 7, 31, 75 (nest site, mating, floral
relationships).

Nomadopsis edwardsii (Cresson)
Calif., Oreg., Wash., Idaho. Pollen: Collects pollen from flowers of

Calochortus, Chamaebatia foliolosa, Potentilla including P. bolanderi var. parryi, P.

congesta, P.fusca, P. glandulosa, P. tridentata, but visits these and other flowers for

nectar including Clarkia rhomboidea, Eriogonum marifolium, Fragaria califomica. Iris

hartwegii, Lathyrus sulphureus, Spraguea umbellata.
Calliopsis Edwardsii Cresson, 1878. Amer. Ent. Soc, Trans. 7: 64. ♀, ♂.
Calliopsis lateralis Cresson, 1878. Amer. Ent. Soc, Trans. 7: 65. ♀.
Spinoliella triangulifera Cockerell, 1925. Calif. Acad. Sci., Proc (4) 14: 195. ♀.

Taxonomy: Michener, 1937. Ann. and Mag. Nat. Hist. (10) 19: 325, S (as triangulifera).
—Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 77-81, figs. 4-5, 28-29, 64, 84-88, map 3
(redescription, synonymy, larva).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 6, 28-31, 81 (nest site, mating behavior,
floral relationships).

Nomadopsis linsleyi Rozen
Calif.; Mexico (Baja California). Pollen: Collects pollen from the flowers of

Eriodictyon, including E. a7igustifoliut)i, E. crassifoliuni, E. parryi, E. trichocalyx, but
visits these and other flowers for nectar including Arctostaphylos, Ceanothus.
Nomadopsis linsleyi Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 85, figs. 6, 7, 30, 31, 53,
99-103, map 5. cJ, 9.

Nomadopsis obscurella (Cresson)
Calif., Oreg., Ariz. (Tucson), ?Wash.; Mexico (Baja California). Pollen:

Apparently oligolectic on Eschscholzia, especially E. califomica, but visits other flowers




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including Agoseris glauca, Caleyidula, Ceanotkus, Chaenactis glabmiscula, Coreopsis
lanceolata, Cryptantha intermedia, Lasthenia, Layia platyglossa, L. p. breviseta,
Malacothrix califomica, Nemopliila menziesii, Phacelia davidsonii, P. distans, P.
tanacetifolia, Sida malvaeflora, Sphaeralcea.
Calliopis obscurellus Cresson, 1879. Amer. Ent. Soc, Trans. 7: 201. ♀, ♂.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 91-93, figs. 104-108, map 7 (redescription).

Biology: Timberlake, 1952. Ent. Soc. Amer., Ann. 45: 107 (mating). —Rozen, 1958. Calif.
Univ. Pubs. Ent. 15: 32, 93 (mating, floral relationships).

Nomadopsis puellae (Cockerell)
Tex., N. Mex., Ariz., Nev., Calif., Idaho.; Mexico (Baja California). Parasite:
Oreopasites vanduzeei Cockerell. Pollen: Evidently collects pollen chiefly from the
flowers of Malacothrix including M. califomica, M. exigua, M. glabrata, but visits other
flowers including Anisocoma acaulis, Aster abatvs. Astragalus, Baileya pleniradiata,
Calycoseris wrightii, Chaenactis fremontii, Encelia farinosa, Haplopappus cooperi, H.
linearifolius, Rafinesquia neomexicana.
Spinoliella puellae Cockerell, 1933. Pan-Pacific Ent. 9: 25. ♀, ♂.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 89-91, figs. 89-93, map 6 (redescription).
— Rozen, 1959. Ent. Soc. Wash., Proc. 61: 255 (geogr. records). —Yager and Rozen, 1966.
Amer. Mus. Novitates 2265: 8 (pupa).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 8, 31, 54, 89 (nest site, mating behavior,
parasites, floral relationships).

Nomadopsis timberlakei Rozen
Calif. (Los Angeles County), desert. Pollen: Possibly an oligolege of
Monardella exilis.
Notnadopsis timberlakei Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 73, map 2. cJ, 9.

Nomadopsis zonalis
***authority mismatch
sierrae Rozen. Calif. (Sierra Nevada Mts.). Parasite: Oreopasites sp. Pollen: Collects

pollen from the flowers of Monardella lanceolata, but visits these and other flowers for
nectar and or pollen including Solidago califomica.
Nomadopsis zonalis sierrae Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 71-73, map 2. ♂, ♀.

Taxonomy: Rozen, 1963. Amer. Mus. Novitates 2142: 3-5, figs. 1-2, table 1 (immature stages).
—Yager and Rozen, 1966. Amer. Mus. Novitates 2265: 7-8, figs. 7-8 (pupa).

Biology: Rozen, 1963. Amer. Mus. Novitates 2142: 2-6 (nest site, nest architecture, floral
relationships, parasite).

Nomadopsis zonalis
***authority mismatch
zonalis (Cresson). Calif, (central and south coasts). Pollen: Collects pollen from the
flowers of Monardella douglasii, M. lanceolata.
Calliopsis zonalis Cresson, 1879. Amer. Ent. Soc, Trans. 7: 201. ♂.
Spinoliella euxantha Cockerell, 1916. Ann. and Mag. Nat. Hist. (8) 17: 283. ♀.

Taxonomy: Timberlake, 1951. In Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr.

Monog. 2: 1102 (synonymy). —Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 69-71, figs. 66, 74-78,

map 2 (redescription).
Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 70-71 (floral relationships).

Genus NOMADOPSIS Subgenus MACRONOMADOPSIS Rozen

Nomadopsis subg. Macronomadopsis Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 93.
Type-species: Nomadopsis micheneri Rozen. Grig, desig.

Nomadopsis anthidia
***authority mismatch
anthidia (Fowler). Oreg., Calif, (west slope of Sierra Nevada, northern Coast Ranges,
Sacramento Valley, part of San Joaquin Valley). Parasite: Euphytomima nomiivora
James, Oreopasites vanduzeii melanantha Linsley. Pollen: Collects pollen from the
flowers of Trifolium including T. involucratum, T. melananthum, T. microcephalum, T.
monanthum var. parvum, T. repens, T. tridentatum, T. variegatum, but visits these and
other flowers for nectar including Eriodictyon trichocalyx, Ltipinus bicolor. Predator:
Solenopsis sp.
Calliopsis anthidius Fowler, 1899. Psyche 8: 407. 6 ( 9 misdet.).




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Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 99-102, figs. 8, 9, 56, 63, 119-123, map 9
(tax. status, redescription, larva). —Rozen, 1959. Ent. Soc. Wash., Proc. 61: 258 (gender of
name). —Yager and Rozen, 1966. Amer. Mus. Novitates 2265: 8, figs. 9-10 (pupa).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 8-9, 32-34, 53, 101 (nest site, nest
architecture, mating behavior, floral relationships, parasite, predator, larva).
— Moradeshaghi and Bohart, 1968. Kans. Ent. Soc, Jour. 41: 456-473, 15 figs., 3 tables
(parasite).

Nomadopsis anthidia
***authority mismatch
lutea Rozen. Calif, (east of Sierra and southern Calif.), Oreg., Idaho, Utah. Pollen:
Collects pollen from the flowers of Trifoliuni including T. involucratum, T. repens.
Nomadopsis anthidius lutea Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 102, figs. 2, 3, 10, 11,
34, 35, 57, 124-128, map 9. cj, 9.

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 9, 34, 101 (nest site, mating behavior, floral
relationships).

Nomadopsis barri Rozen
Idaho, Oreg. Pollen: Unknown, but visits flowers of Melilotus.

Nomadopsis (Macronomadopsis) barri Rozen, 1959. Ent. Soc. Wash., Proc. 61: 256, figs. 1,
4. ♂, ♀.
niiorum Rozen. Calif. (Mono Co.). Pollen: Probably collects pollen from the flowers of
Trifolium.
Nomadopsis {Macronomadopsis) filiorum Rozen, 1963. Amer. Mus. Novitates 2142: 6, figs.
3-6. ♂, ♀.

Biology: Rozen, 1963. Amer. Mus. Novitates 2142: 3-6, tables 1-2 (nest site, nest architecture,
floral relationships).

Nomadopsis micheneri Rozen
Calif. (Sierra Nevada and Trinity Mt.). Pollen: Collects pollen from the

flowers of Trifolium including T. gracilentiim, T. monanthus var. pay~vum, T. repeals.
Nomadopsis micheneri Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 95, figs. 12, 13, 32, 33, 55,
67, 109-113, map 8. ♂, ♀.

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 9-12, 34-36, 43, 53, 98 (nest site, nest
architecture, mating behavior, floral relationships).

Nomadopsis zebrata
***authority mismatch
bobbae Rozen. Ariz. (Navajo and Coconino Counties).

Nomadopsis zebrata bobbae Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 107, map 8. ♂, ♀.

Nomadopsis zebrata
***authority mismatch
zebrata (Cresson). S. Dak., Wyo., Colo., N. Mex. Pollen: Collects pollen from the
flowers of Astragalus.
Calliopsis zebratus Cresson, 1878. Amer. Ent. Soc, Trans. 7: 64. ♀.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 103-107, figs. 114-118, map 8 (tax. status,
redescription).

Biology: Cockerell, 1915. Ent. News 26: 366 (nest site).— Hicks, 1936. Canad. Ent. 68: 47
(nest).— Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 35 (nest site).

Genus NOMADOPSIS Subgenus MICRONOMADOPSIS Rozen

Nomadopsis subg. Micronomadopsis Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 107.
Type-species: Nomadopsis fracta Rozen. Orig. desig.

Nomadopsis australior (Cockerell)
S. Dak., Nebr., Colo., Utah, N. Mex., Tex., Ariz.; Mexico (Chihuahua).

Pollen: Collects pollen from the flowers of Lepidium alyssoides, L. montanum, but may
also obtain pollen from the flowers of Dithyrea wislizeni.
Calliopsis australior Cockerell, 1897. Acad. Nat. Sci. Phila., Proc. 49: 349. ♀ (♂ misdet.).

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 149-152, figs. 71, 182-185, map 15. d, 9
(redescription). —Rozen, 1959. Ent. Soc. Wash., Proc. 61: 259 (geogr. rec).

Biology: Custer, 1927. Psyche 34: 199 (nesting activities). —Custer, 1929. Canad. Ent. 61: 49
(nest). —Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 8, 152 (nest site, floral relationships).




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Nomadopsis barbata Timberlake
South. Calif., north into Central Valley. Parasite: Oreopasites sp. Pollen:
Collects pollen from the flowers of Phacelia including P. distans, P. tanacetifolia, but
visits these and other flowers for nectar including Cryptantha, Linanthus dianthiflonis.
Nomadopsis barbata Timberlake, 1952. Ent. Sec. Amer., Ann. 45: 107. ♂, ♀.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 114-117, figs. 18-19, 42-43, 59, 133-136,
map 11 (redescription, larva).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 8, 36, 40, 117, table 2 (nest site, nesting
activity, mating behavior, floral relationships, parasite).

Nomadopsis beamerorum Rozen
Tex. (Reeves and Pecos Counties). Pollen: Unknown, but holotype male
was visiting flowers of Prosopis.
Nomadopsis (Micronomadopsis) beamerorum Rozen, 1963. Amer. Mus. Novitates 2142: 13,
figs. 12-15. ♂, ♀.

Nomadopsis callosa Timberlake
Tex. (El Paso Co.); Mexico (Baja California).

Nomadopsis callosa Timberlake, 1952. Ent. Soc. Amer., Ann. 45: 110. ♂.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 140-142, figs. 174-177, map 8
(redescription). —Rozen, 1959. Ent. Soc. Wash., Proc. 61: 259 (type).

Nomadopsis cazieri Rozen
Calif. (San Jacinto Mts. and San Diego County).

Nomadopsis cazieri Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 135, figs. 165-169, map 14. ♂.
9.

Nomadopsis foleyi Timberlake
South. Calif., deserts. Pollen: Collects pollen from the flowers of Larrea

iridentata, but visits these and other flowers for nectar including Heliotropium, Nama
hispida.
Nomadopsis foleyi Timberlake, 1952. Ent. Soc. Amer., Ann. 45: 112. ♀, ♂.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 119-121, figs. 141-144, map 10
(redescription).

Biology: Rozen, 1958. Calif Univ. Pubs. Ent. 15: 121 (floral relationships).

Nomadopsis fracta Rozen
Calif, (widely distributed at lower altitudes). Pollen: Collects pollen from the

flowers of Eriodictyon angustifolium, E. califoniicnm, E. crassifolium, E. trichocalyx,
but visits these and presumably other flowers for nectar including Ceanothus
integerrivius, Chaenactis, Chamaebatia foliolosa, Cryptantha intermedia, Erigeron
divergens, Eriogonum fascicidattim, Gilia exilis, M imulus fremontii, Nama parryi,
Phacelia brachyloba, P. douglasii.
Nomadopsis fracta Rozen, 1952. Kans. Ent. Soc, Jour. 25: 144, 4 figs. 6,9.

Taxonomy: Michener, 1953. Kans. Univ. Sci. Bui. 35: 1036, figs. 92-98 (larva). -Rozen, 1958.
Calif Univ. Pubs. Ent. 15: 111-113, figs. 36-37, 68-69, 129-132, map 10 (redescription, larva).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 36, 39, 48-50, 55, 60, 113 (mating behavior,

sleep, flower relationships, voltinism).

Nomadopsis helianthi (Swenk and Cockerell)
Nebr. and N. Mex., west to Calif; Mexico (Baja California).
Parasite: Oreopasites euphorbiae Ckll. Pollen: Collects pollen from the flowers of
Euphorbia albomarginata, E. polycarpa, but visits these and other flowers for nectar
including Cryptantha intermedia, Eremocarpus setigerus, Eriogonum fasciculatum, E.
gracile, Gutierrezia sarothrae, Helianthus, Heliotropium curassavicum, Hemizonia
paniculata.

Spinoliella helianthi Swenk and Cockerell, 1907. Ent. News 18: 178. ♀.

Spinoliella euphorbiae Cockerell, 1925. Pan-Pacific Ent. 1: 179. ♀.

Taxonomy: Michener, 1953. Kans. Univ. Sci. Bui. 35: 1036, figs. 99, 106-108 (larva, as
euphorbiae). —Rozen, 1958. Calif Univ. Pubs. Ent. 15: 133, map 13 (type). —Rozen, 1958.
Calif Univ. Pubs. Ent. 15: 129-133, figs. 16-17, 40-41, 61, 65, 157-160, map 13 (as
euphorbiae, redescription, larva). —Rozen, 1963. N. Y. Ent. Soc, Jour. 7: 142-143
(synonymy).




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Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 11, 16-17, 37, 40, 43, 48-49, 50, 55, 59, 133
(adult activity, defecation, floral relationships, habitat, hibernation, mating behavior, sleep,
parasitism, voltinism).

Nomadopsis hesperia
***authority mismatch
equina (Cockerell). Calif. (Great Valley), Nev. (Virginia City). Parasite: Oreopasites
vanduzeei Ckll. Pollen: Collects pollen from the flowers of Heliotropium curassavicum,
but visits these and other flowers for nectar including Erodium cicutarium, Hemizonia
pungens, Medicago sativa.
Spinoliella equina Cockerell, 1925. Calif. Acad. Sci., Proc. (4) 14: 196. ♀, ♂.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 148-149 (redescription). —Yager and
Rozen, 1966. Amer. Mus. Novitates 2265: 6-7, figs. 5-6 (pupa).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 49, 59, 147 (flower relationships, parasite).

Nomadopsis hesperia
***authority mismatch
hesperia (Swenk and Cockerell). South. Calif. Pollen: Collects pollen from the flowers
of Heliotropium curassavicum oculatum.
Spinoliella hesperia Swenk and Cockerell, 1907. Ent. News 18: 186. ♀, ♂.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 147-148, figs. 20-21, 44-46, 70, 178-181,
map 15 (redescription).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 49, 59, 147 (floral relationships).

Nomadopsis larreae Timberlake
South. Calif. (Colorado and Mojave Deserts). Ecology: Nest sites are
estabhshed in sand dune areas. Pollen: Collects pollen from the flowers of La rrea
tridentata.
Nomadopsis larreae Timberlake, 1952. Ent. Soc. Amer., Ann. 45: 111. ♀.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 126-128, figs. 153-156, map 12
(redescription).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 11, 37, 41, 43, 49, 55, 128 (nest site, nest
architecture, nesting activities, mating behavior, floral relationships).

Nomadopsis macswaini Rozen
Ariz. Pollen: Evidently collects pollen from the flowers of Wislize^iia
refracta.
Nomadopsis macswaini Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 152, figs. 195-198, map 16.
6, 9.

Nomadopsis meliloti (Cockerell)
Tex., N. Mex., Ariz. (Willcox). Pollen: Pollen-laden females have been
collected at the flowers of Dalea scoparia, but also visits flowers of Melilotus.
Calliopsis meliloti Cockerell, 1896. Canad. Ent. 28: 158. ♀.

Taxonomy: Cockerell, 1933. Pan-Pacific Ent. 9: 158. S, 9. —Rozen, 1958. Calif. Univ. Pubs.
Ent. 15: 138-140, figs. 170-173, map 14 (redescription). —Rozen, 1959. Ent. Soc. Wash.,
Proc. 61: 259 (geogr. record).

Nomadopsis mellipes Timberlake
South. Calif. Pollen: Apparently collects pollen from the flowers of Lotus
including L. glaber, L. hamatus, L. salsuginosus, L. scoparitis, L. strigosus, but visits
these and apparently other flowers for nectar including Camissonia dentata, Encelia.
Nomadopsis mellipes Timberlake, 1952. Ent. Soc. Amer., Ann. 45: 109. ♂, ♀.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 125-126, figs. 149-152, map 12
(redescription).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 126 (floral relationships).

Nomadopsis nigromaculata Timberlake
South. Calif., Ariz., deserts. Pollen: Apparently collects pollen from
the flowers of Euphorbia polycarpa including E. p. var. hirtella, but visits other flowers
including Eriogonum trichopes, Pedis papposa possibly for nectar.
Nomadopsis nigromaculata Timberlake, 1952. Ent. Soc. Amer., Ann. 45: 115. ♀, ♂.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 133-135, figs. 161-164, map 14
(redescription).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 37-38, 50, 52, 135 (mating behavior, adult
behavior, floral relationships, voltinism).




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Nomadopsis personata (Cockerell)
Wash., Oreg., Idaho, Wyo. Pollen: Pollen laden females have been
collected at the flowers of Cleome lutea, C. semdata.
Calliopsis personafus Cockerell, 1897. Acad. Nat. Sci. Phila., Proc. 49: 349. ♀.

Ta.xonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 165-167, figs. 186-189, map 15
(redescription). —Yager and Rozen, 1966. Amer. Mus. Novitates 2265: 7 (pupa).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 167 (floral relationships).

Nomadopsis phaceliae Timberlake
Calif, (chiefly east side of Sierra Nevada and Mojave Desert). Pollen:
Apparently collects pollen from the flowers of Phacelia fremontii, but visits these and
other flowers for nectar including Euphorbia albomarginata, Hesperochiron
califomicum, Nama demissum.
Nomadopsis phaceliae Timberlake, 1952. Ent. Soc. Amer., Ann. 45: 116. ♂.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 117-119, figs. 137-140, map 11
(redescription). —Rozen, 1959. Ent. Soc. Wash., Proc. 61: 258-259 (geogr. and floral
records).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 119 (floral relationships).

Nomadopsis scitula
***authority mismatch
lawae (Michener). Calif. (Inyo, Lassen and Mono Counties), Nev. (Fallon). Pollen: A

female laden with pollen has been taken at the flowers of Thelypodium brachycarpum,
others at flowers of Cleome and a male at the flowers of Cleomella oocarpa.
Spinoliella lawae Michener, 1937. Ann. and Mag. Nat. Hist. (10) 19: 324. ♀.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 156-157, 159-160, map 16 (redescription).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 160 (floral relationships).

Nomadopsis scitula
***authority mismatch
scitula (Cresson). Wyo., Idaho, Utah, Colo., N. Mex.; variants occur in B-C, Oreg.,
northeast. Calif., northeast. Nev., Colo, and Wyo. Parasite: Oreopasites scitiili Ckll.
Pollen: Females with pollen-laden hind legs have been collected at the flowers of
Cleoyne semdata, Sisymbrium altissiynum.
Calliopsis scitidiis Cresson, 1878. Amer. Ent. Soc, Trans. 7: 64. ♀.
Calliopsis pictipes Cresson, 1878. Amer. Ent. Soc, Trans. 7: 65. ♂.

Taxonomy: Cockerell, 1897. Acad. Nat. Sci. Phila., Proc. 49: 348 (tax. characters).
—Timberlake, 1951. hi Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr. Monog.
2: 1102 (synonymy). —Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 21, 154-159, figs. 72, 199-203,
map 16 (redescription, tax status, larva). —Yager and Rozen, 1966. Amer. Mus. Novitates
2265: 7 (pupa).

Biology: Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 440 (parasite). —Rozen, 1958. Calif.

Univ. Pubs. Ent. 15: 12, 40, 47, 159 (nest site, nesting activities, floral relationships.)

Nomadopsis scutellaris
***authority mismatch
scutellaris (Fowler). Wyo., Idaho, Utah, Nev., Wash., Oreg., Calif., Ariz.; Mexico
(northern Baja California). Parasite: Euphytomima nomiivora James, Myrmosula
rutilans (Fowler), Oreopasites sp. Pollen: Apparently polylectic, pollen-laden females
have been collected from the flowers oi Heliotr opium curassavicum,
Mesembryanthemum, Sesuvium sessile, Wislizenia refracta, alsike clover, ladino clover,
carrots, but also visits other flowers including Alyssum maritimum, Brassica incana,
Chrysanthemum, Cressa truxillense, Gutierrezia sarothrae, Melilotus indica, Phacelia
fremontii, Raphamis sativus, Sesuvium verrucosum, Solidago, Tamarix. Another
subspecies, A^. scidellaris peninsularis (Cockerell), occurs in southern Baja California.

Calliopsis scutellaris Fowler, 1899. Psyche 8: 406. ♂.

Calliopsis visaliensis Fowler, 1899. Psyche 8: 406. ♀.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 20, 23, 24, 25, 27, 30, 160-165, figs. 24-25,
47-48, 62, 73, 190-194, map 17 (redescription, larva). —Yager and Rozen, 1966. Amer. Mus.
Novitates 2265: 7 (pupa).

Biology: Snelling, 1954. Pan-Pacific Ent. 30: 124 (parasite). —Rozen, 1958. Calif. Univ. Pubs.
Ent. 15: 12, 15, 38, 40, 48, 53-54, 164 (nest, site, nest architecture, life history, mating
behavior, parasite). — Moradeshaghi and Bohart, 1968. Kans. Ent. Soc, Jour. 41: 456-473,
15 figs., 3 tables (parasite).




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Nomadopsis snellingi Rozen
Calif. (Squaw Valley). Pollen: Unknown, but visits flowers of Salvia.

Nomadopsis {Micronomadopsis) snellingi Rozen, 1963. Amer. Mus. Novitates 2142: 9, figs.
7-11. ♂, ♀.

Nomadopsis trifolii Timberlake
Calif. Pollen: Collects pollen from the flowers of Mimulus fremontii,

Trifolium mela7iantku7n, T. tridentatum, T. variegatum, but also visits other flowers
including Cryptantha, E riodictyon, Salix.
Nomadopsis trifolii Timberlake, 1952. Ent. Soc. Amer., Ann. 45: 113. ♀, ♂.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 14, 19, 23, 26, 30, 121-123, figs. 14-15,
38-39, 60, 145-148, map 12 (redescription, egg, larva).

Biology: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 10, 15, 17, 35, 40, 48-50, 123 (nest site, nest
architecture, life history, mating behavior, hibernation, sleep, voltinism, floral
relationships).

Genus NOMADOPSIS Subgenus UNASSIGNED

Nomadopsis boharti Rozen
Calif. (Eldorado, Napa and Tuolumne Counties).

Nomadopsis boharti Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 171, figs. 204-208, map 7. ♂.
9.

Taxonomy: Yager and Rozen, 1966. Amer. Mus. Novitates 2265: 8 (pupa).
interrupta (Provancher). Ont. (Toronto).

Calliopsis interrupta Provancher, 1888. Addit. Corr. Faune Ent. Canada, Hym., p. 320. ♀.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 174. (tax. status).

Nomadopsis quadrilineata
***authority mismatch
provancher). Ont. (Toronto).

Calliopsis Jt-lineata Provancher, 1888. Addit. Corr. Faune Ent. Canada, Hym., p. 319. ♀.
Calliopsis quadrilineata Dalla Torre, 1896. Cat. Hym., v. 10, p. 174. Emend.

Taxonomy: Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 174 (tax. status).

Nomadopsis smithi Rozen
Calif. (San Benito County). Pollen: Unknown, but visits flowers of Cryptantha.

Nomadopsis smithi Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 169, figs. 214-218, map 7. ♂, ♀.

Nomadopsis solitaria Rozen
Calif. (Lassen County).

Nomadopsis solitaria Rozen, 1963. Amer. Mus. Novitates 2142: 15, figs. 16-20. ♂.

Nomadopsis xenus Rozen
Wash., Calif. (Lake Tahoe). Pollen: Pollen-laden females have been collected
from the flowers of Phacelia.
Nomadopsis xenus Rozen, 1958. Calif. Univ. Pubs. Ent. 15: 167, figs. 209-213. ♂, ♀.

Genus CALLIOPSIS Smith

Revision: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 753-936, 156 figs., 8 maps, 9 tables (N. and
Cent. Amer. spp., includes biological information).

Taxonomy: Cockerell, 1898. Amer. Ent. Soc, Trans. 25: 195-198 (U. S. spp.). —Mitchell, 1960.
N. C. Agr. Expt. Sta. Tech. Bui. 141: 287-293, figs. 58, 69-71, table 8 (eastern U. S. spp.).
—Rozen, 1966. Amer. Mus. Novitates 2259: 5, 13-16, figs. 29-40 (larvae). —Yager and
Rozen, 1966. Amer. Mus. Novitates 2265: 8-10, figs. 11-12 (pupae).

Biology: Rozen, 1967. Amer. Mus. Novitates 2297: 1-44, 18 figs., 2 tables (review).
Genus CALLIOPSIS Subgenus CALLIOPSIS Smith

Calliopsis Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 128.

Type-species: Calliopsis andreniformis Smith. Desig. by Ashmead, 1899.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 782-816, figs. 5, 6, 8-45, maps 1-3
(redescription, included spp.).

Calliopsis andreniformis Smith
Southeast. Canada including the Maritime Provinces, south to Fla., west
to Mont., S. Dak., Colo., Utah, Okla. and east Tex. Parasite: Holcopasites calliopsidis
(Linsley), H. illinoie^isis (Robertson), Sphecodes brachycephalus Mitchell?, Sphecodes
sp., Parabombylius coquilletti (Will.)?, Trochometridium tribulatum Cross, Villa




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sinuosa (Wied.)- Pollen: Polylectic, known sources include Aster ericoides villosus,
Convolvulus, Desmodium marilandicum, D. paniculatum, Erigeron canadensis,
Gerardia temiifolia, Hedyotis purpurea, Lippia lanceolata, Lycopus sinuatus, Lythnim
alaturn, Malva neglecta, M. rotu7idifolia, Melilotus alba, M. officinalis, Oxalis stricta,
Polygala sanguinea, Polygonum buxifomie, Psoralea onybrichis, Pycnanthemuyn
pilosum, Trifoliuni pratense, T. procumbens, T. repens. Verbena bracteata, V. hastata,
V. urticifolia, Verbesina helianthoides, but visits these and other flowers for nectar
including Achillea, Ailanthus altissirna, Arnmania coccinea, Amorpha canescens,
Anaphalis margaritacea, Asclepias, Bidens aristosa, Boltonia asteroides, Brassica,
Castanea pumila, Ceanothus americanus. Chrysanthemum leucanthemum, Chrysopsis,
Cleome, Coreopsis palmata, C. tripteris, Cryptotaenia canadensis, Cucumis, Dianthera
americana, Epilobium, Eryngiiim yuccifolium, Geum album, Gillenia stipulacea,
Hedeoma pulegioides, Hedyotis nigricans, Helenium, Hypericum perforatum, Lindemia
dubia riparia, Lespedeza capitata, L. procumbens, L. repens, L. reticulata, Ligustrum,
Medicago sativa, Nepeta cataria, Oenothera laciniata, Oxalis dillenii, Penstemon,
Petalostemon candidum. Polygonum convolvulus, P. pennsylvanicum, Portulaca,
Potentilla monspeliensis, P. recta. Prunella vulgaris. Primus, Psoralea tenuiflora, P.
tenuiflora floribunda, Pycnajithemum flexuosum, P. virginianum, P. lanceolatum,
Raphanus sativus, Rhus glabra, Rosa, Rubus, Rudbeckia triloba, Serinea oppositifolia,
Sisymbriinn repandum, Solanum carolinense, Solidago canadensis, Spiranthes gracilis,
Stachys palustris, Stellaria, Strophostyles pauciflora, Stylosaiithes biflora,
Symphoricarpos, Tamarix, Trifolium hybridum. Verbena bracteosa, Veronica spicata,
Vicia. Predator: Philanthus gibbosus (Fabr.), P- politus Say.

Calliopsis andreniformis Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 128. ♀.

Calliopsis flavipes Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 128. ♂.

Calliopsis lepidus Cresson, 1878. Amer. Ent. Soc, Trans. 7: 68. ♀.

Panurgus vemalis Provancher, 1882. Nat. Canad. 13: 204. ♀, ♂.

Taxonomy: Michener, 1953. Kans. Univ. Sci. Bui. 35: 1038, figs. 100-105 (larva). —Mitchell,
1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 288-289, figs. 69-71, table 8 (redescription,
synonymy). — Rozen, 1966. Amer. Mus. Novitates 2259: 16, fig. 40 (larva). —Yager and
Rozen, 1966 Amer. Mus. Novitates 2265: 8-10, figs. 11-12 (pupa). — Shinn, 1967. Kans. Univ.
Sci. Bui. 46: 786-791, figs. 5-6, 8-13, map 1 (redescription).

Biology: Graenicher, 1905. Wis. Nat. Hist. Soc, Bui. 3: 159 (nest). — Swenk, 1907. Ent. News
18: 297 (nest, parasite). — Reinhard, 1924. Ann. Rept. Smithsn. Inst. 2738: 371-373
(predator). —Robertson, 1926. Psyche 33: 118 (nest, parasite). — Hendrickson, 1930. Iowa
State Coll. Jour. Sci. 4: 163 (nest). —Pearson, 1933. Ecol. Monog. 3: 387 409-411 (floral
relationships). — Ainslie, 1937. Canad. Ent. 69: 97-100 (nest, floral relationships, parasite).
— Lovell and Lovell, 1939. Rhodora 41: 185 (ecology). — Crandall and Tate, 1947. Amer.
Soc. Agronomy, Jour. 39: 161-163 (nest site, floral relationships). —Mitchell, 1956. Elisha
Mitchell Sci. Soc. 72: 207 (parasite). —Michener and Rettenmeyer, 1956. Kans. Univ. Sci.
Bui. 37: 645 (age of nest site). —Montgomery, 1957. Ind. Acad. Sci., Proc. 66: 129 (ecology).
— Byers, 1962. Kans. Ent. Soc, Jour. 35: 320 (ecology). —Rozen, 1966. Amer. Mus.
Novitates 2244: 33 (parasite). —Shinn, 1967. Kans. Univ. Sci. Bui. 46: 895-935 figs. 144-156,
tables 4-9 (nest architecture, life history, immature stages, odor production, sleep, floral
relationships, parasites). —Rozen, 1967. Amer. Mus. Novitates 2297: 32-33, fig. 9, table 1
(nest architecture, life history, parasite). — Hurd and Linsley, 1972. Smithsn. Contrib. Zool.
114: 39, table 1 (parasites).

Calliopsis empelia Shinn
Ariz. (Douglas and Mt. Lemmon).

Calliopsis (Calliopsis) empelia Shinn, 1967. Kans. Univ. Sci. Bui. 46: 811, figs. 38-41, map
2. ♀, ♂.

Calliopsis helenae Shinn
Tex. (10 mi. s. Alice); Mexico (Piedras Negras, Coahuila).

Calliopsis (Calliopsis) helenae Shinn, 1967. Kans. Univ. Sci. Bui. 46: 814, map 1. ♀.

Calliopsis hondurasica Cockerell
La. (Robson), Tex.; Mexico to Panama. Pollen: Unknown, but visits
flowers of Cassia, Kallstroemia hirsutissima, Lippia, Nama tmdulattmi. Phyla
strigosa, Teucrium, Trifolium repens. Verbena.
Calliopsis hoiidurasica Cockerell, 1949. U. S. Natl. Mus., Proc 98: 437. ♀.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 800-804, figs. 30-33, map 2 (redescription).




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Calliopsis rhodophila Cockerell
N. Mex., Ariz. (Chine Valley), south. Calif. (Riverside, San Bernardino
and San Diego Counties); Mexico (Durango and Zacatecas). Pollen: Unknown, but visits
flowers of Solidago, Sphaeralcea.
Calliopsis andreniformis rhodophilus Cockerell, 1897. Acad. Nat. Sci. Phila., Proc. 49: 350.

(J, 9.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 796-799, figs. 22-25, map 1 (redescription).

squatnifera Timberlake. Ariz. Pollen: Unknown, but visits flowers of Baccharis glutinosa,
Euphorbia, Medicago sativa.
Calliopsis squamifera Timberlake, 1947. Pan-Pacific Ent. 23: 28. ♂.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 804-807, figs. 42-45, map 3 (redescription).

Calliopsis teucrii Cockerell
N. Mex., Ariz., Colo.; Mexico (Distrito Federal, Durango, Guerrero, Mexico,
Michoacan, Morelos, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Tlaxcala
and Zacatecas). Pollen: Unknown, but visits flowers of Asclepias, Convolvulus incanus,
Heterotheca chrysopsidis, Melilotus officinalis, Oxalis, Potentilla, Stylosanthes,
Taraxacum, Teucrium laciniatum.
Calliopsis teucrii Cockerell, 1899. hi Cockerell and Porter, Ann. and Mag. Nat. Hist. (7) 4:
412. ♀.

Genus CALLIOPSIS Subgenus PERISSANDER Michener

Calliopsis subg. Perissander Michener, 1942. N. Y. Ent. Soc, Jour. 50: 275.

Type-species: Calliopsis (Perissander) anomoptera Michener. Monotypic and orig.
desig.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 816-834, figs. 7, 46-57, maps 3-4
(redescription, included spp.).

Calliopsis anomoptera Michener
East. N. Mex., west to south. Calif.; Mexico (Baja California and

Sonora). Pollen: Polylectic, collects pollen and nectar from a variety of flowers including
Cladothrix lanuginosa, Eriogonum, Euphorbia albomargi^iata, E. capitellata, E.
hirtella, E. pleniradiata, E. polycarpa hirtella, E. polycarpa typica, Lepidium thurberi,
Tidestromia.
Calliopsis (Perissander) anomoptera Michener, 1942. N. Y. Ent. Soc, Jour. 50: 275. ♂, ♀.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 818-823, figs. 7, 46-49, map 4 (redescription).

Biology: Krombein, 1961. Ent. News 72: 82-83 (floral relationships with Euphorbia
albo7nargi7iata). —Shinn, 1967 Kans. Univ. Sci. Bui. 46: 823 (floral relationships).

Calliopsis fulgida Shinn
N. Mex. (4.8 mi. N. of Rodeo), Ariz. (5 mi. E. Portal). Pollen: Unknown, but
visits flowers of Euphorbia, Tidestro^uia lanuginosa.
Calliopsis (.Perissander) fulgida Shinn, 1967. Kans. Univ. Sci. Bui. 46: 831, map 3 9.

Calliopsis gilva Shinn
Ariz. (Douglas and Pima Counties), N. Mex. (Hidalgo County). Pollen: Unknown,
but visits flowers of Euphorbia, Verbesiiia, Tidestromia la7iugi7iosa.
Calliopsis (Perissajider) gilva Shinn, 1967. Kans. Univ. Sci. Bui. 46: 829, figs. 54-57, map 4.
9, 6.

Calliopsis limbus Shinn
Ariz. (Santa Catalina Mts.), N. Mex. (Granite Pass). Pollen: Unknown, but visits
flowers of Euphorbia.
Calliopsis (Perissander) limbus Shinn, 1967. Kans. Univ. Sci. Bui. 46: 827, map 4. ♀, ♂.

Calliopsis rogeri Shinn
Ariz. (Cochise County). Pollen: Unknown, but visits flowers of Euphorbia
albomarginata, Lepidium thurberi.
Calliopsis (Perissander) rogeri Shinn, 1967. Kans. Univ. Sci. Bui. 46: 823, figs. 50-53, map

4. ♀, ♂.

Genus CALLIOPSIS Subgenus CALLIOPSIMA Shinn

Calliopsis subg. Calliopsima Shinn, 1967. Kans. Univ. Sci. Bui. 46: 834.
Type-species: Calliopsis rozeni Shinn. Orig. desig.




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Calliopsis bernardinensis Michener
South. Calif. (Riverside and San Bernardino Counties). Pollen:
Apparently an oligolege of autumnal flowering Compositae including Gutierrezia
sawthrae, Heterotheca grand if lora, Senecio ionophyllus.
Calliopsis bernardinensis Michener, 1937. Ann. and Mag. Nat. Hist. (10) 19: 323. ♂, ♀.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 852-855, figs. 77-82, map 6 (redescription,
floral records).

Calliopsis chlorops Cockerell
N. Mex., Colo., Wyo., Idaho, Utah, Nev., Ariz. Pollen: Apparently an
oligolege of the Compositae; visits flowers of Chrysopsis, Grindelia squarrosa,
Haplopappus gracilis, Medicago sativa, Ratibida, Verbesina, Viguiera annua.
Calliopsis chlorops Cockerell, 1899. In Cockerell and Porter, Ann. and Mag. Nat. Hist. (7)
4:413. ♂.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 860-862, figs. 95-98, map 6 (redescription,
floral records).

Calliopsis coloradensis Cresson
Alta. to south. Utah and south. N. Mex., east to Mississippi River,

thence through east. Tex. and Gulf States to east slope of the Appalachian Mts. in N. C.
Parasite: Holcopasites arizonicus (Linsley). Pollen: Apparently an oligolege of the
Astereae and Heliantheae (Compositae); collects pollen from the flowers of Bidens
aristosa, Boltonia asteroides. Coreopsis tripteris, Rudbeckia triloba, Solidago
canadensis, but visits other flowers some of which may also serve as pollen sources
including Anthemis cotula, Aplopappus pluriflorus, Aster dumosus, A. praetus, Bidens
laevis, Chrysopsis, Chrysothamnus nauseosus, Eriocarpum gracile, Grindelia perennis,
G. squarrosa, Haplopappus, Helenium nudiflorum, Heterotheca subaxillaris, Macrotera,
Silphiuni, Solidago rigida, S. serotina.

Calliopsis coloradensis Cresson, 1878. Amer. Ent. Soc, Trans. 7: 63. ♀, ♂.

Calliopsis coloratipes fedorensis Cockerell, 1909. Ann. and Mag. Nat. Hist. (8) 4: 28. ♀.

Taxonomy: Cockerell, 1908. Canad. Ent. 40: 147. 6. — Timberlake, 1951. In Muesebeck,
Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1103 (synonymy). —Mitchell, 1960.
N. C. Agr. Expt. Sta. Tech. Bui. 141: 289-290, figs. 69-71 (redescription, floral records).
—Shinn, 1967. Kans. Univ. Sci. Bui. 46: 841-846, figs. 63-66, map 5, tables 2, 3
(redescription).

Biology: Hicks, 1926. Colo. Univ., Studies 15: 223 (nest). —Robertson, 1922. Psyche 29: 169
(floral relationships). —Shinn, 1967. Kans. Univ. Sci. Bui. 46: 846 (floral relationships).
— Hurd and Linsley, 1972. Smithsn. Contrib. Zool. 114: 39, table 1 (parasite).

Calliopsis coloratipes Cockerell
N. Mex., Ariz., Utah, Nev. Pollen: Unknown, but visits flowers of

Asclepias subverticillata, Bigelovia hartwegii, B. wrightii, Chrysothamnus nauseosus
consimilis, Heterotheca.
Calliopsis flavifrons race coloratipes Cockerell, 1898. Denison Univ. Sci. Labs., Bui. 11: 52.
6.

Taxonomy: Cockerell, 1898. Univ. N. Mex., Bui. 1: 52. 6. —Cockerell. 1899. Entomologist 33:
64. 9. -Shinn, 1967. Kans. Univ. Sci. Bui. 46: 862-865, figs. 99-102, map 5 (redescription,
floral records).

Calliopsis crypta Shinn
Ariz. (Chiricahua Mts.); Mexico (Chihuahua). Parasite: Holcopasites stevensi

(Cwfd.). Pollen: Collects pollen from Heterotheca subaxillaris, but also visits the flowers
of Cirsium, Helianthus.
Calliopsis crypta Shinn, 1965. Amer. Mus. Novitates 2211: 15, figs. 9-12. ♀, ♂.

Taxonomy: Rozen, 1966. Amer. Mus. Novitates 2259: 14, figs. 29-33 (larva). —Shinn, 1967.
Kans. Univ. Sci. Bui. 46: 858-860, figs. 87-90, map 6 (redescription).

Biology: Rozen, 1967. Amer. Mus. Novitates 2297: 30-31, fig. 10, tables 1-2 (nest architecture,
life history, floral relationship, parasite). —Hurd and Linsley, 1972. Smithsn. Contrib. Zool.
114: 39, table 1 (parasite).

Calliopsis deserticola Shinn
South. Calif., deserts. Pollen: Unknown, but visits flowers of Encelia
farinosa.
Calliopsis (Calliopsima) deserticola Shinn, 1967. Kans. Univ. Sci. Bui. 46: 865, figs.
103-106, map 5. ♀, ♂.




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Calliopsis pectidis Shinn
west. N. Mex., Ariz., south. Calif.; Mexico (Baja California). Parasite:

Holcopasites arizonicus (Linsley)? Pollen: Unknown, but visits flowers of Aplopappus,
Baccharis, Baileya pleniradiata, Helianthus, Heterotheca subaxillaris, Hymenothrix
wislizeni, Melilotus alba, Pedis angustifolia, P. papposa, Tidestromia lanuginosa,
Verbesina encelioides.
Calliopsis pectidis Shinn, 1965. Amer. Mus. Novitates 2211: 10, figs. 5-8. ♀, ♂.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 846-850, figs. 67-71, map 6 (redescription).

Biology: Shinn, 1965. Amer. Mus. Novitates 2211: 15 (parasite). —Shinn, 1967. Kans. Univ.
Sci. Bui. 46: 850, 920 (floral records, parasite). —Kurd and Linsley, 1972. Smithsn. Contrib.
Zool. 114: 39, table 1 (parasite).

Calliopsis pugionis Cockerell
South. Calif, (cismontane). Pollen: Apparently an oligolege of Compositae,
visits flowers of Chaenactis artemisiaefolia, C. glabriuscula. Coreopsis (cultivated), C.
lanceolata, Encelia farinosa, Hemizonia wrightii, Oenothera veitchiana.
Calliopsis pugionis Cockerell, 1925. Calif. Acad. Sci., Proc. (4) 14: 197. ♀.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 868-870, figs. 107-110, map 5 (redescription).

Calliopsis rozeni Shinn
Tex., N. Mex., Ariz.; Mexico (Chihuahua and Coahuila). Parasite: Holcopasites
stevensi (Cwfd.)? Pollen: Collects pollen from Heterotheca subaxillaris primarily, but
visits other flowers including Baccharis, Baileya pleniradiata, Chamaesaracha
conioides, Eriogonum gracile, Gaillardia, Helianthus, Hymenoxys odorata, Melilotus
alba, Parkinsonia, Pedis papposa, Psilostrophe cooperi, Sphaeralcea emoryi, Verbesina
exauria.
Calliopsis rozeni Shinn, 1965. Amer. Mus. Novitates 2211: 2, figs. 1-4. ♀, ♂.

Taxonomy: Rozen, 1966. Amer. Mus. Novitates 2259: 14-16, figs. 34-39 (larva, as rozeni?).
—Shinn, 1967. Kans. Univ. Sci. Bui. 46: 838-841, figs. 58-62, map 5 (redescription).

Biology: Rozen, 1965. N. Y. Ent. Soc, Jour. 73: 88 (parasite). —Rozen, 1967. Amer. Mus.
Novitates 2297: 31-32, tables 1-2 (nest architecture, life history, floral relationships,
parasite). — Hurd and Linsley, 1972. Smithsn. Contrib. Zool. 114: 39, table 1 (parasite).

Calliopsis timberlakei Shinn
West. Tex., N. Mex., Ariz., Utah. Pollen: Apparently an oligolege of

Compositae, visits flowers of Bahia absinthifolia, Bigelovia, Chrysothamnus nauseosus,
Gutierrezia longifolia, G. microcephala, G. sarothrae, Haplopappus spinulosus, H.
wrightii.
Calliopsis (Calliopsima) timberlakei Shinn, 1967. Kans. Univ. Sci. Bui. 46: 850, figs. 72-76,
map 7. ♀, ♂.

Calliopsis unca Shinn
N. Mex. (3 mi. W. Bingham). Pollen: Unknown, but visits flowers of Baileya
pleniradiata.
Calliopsis (Calliopsima) unca Shinn, 1967. Kans. Univ. Sci. Bui. 46: 855, figs. 83-86, map 7.
6.

Genus CALLIOPSIS Subgenus VERBENAPIS Cockerell and Atkins

Calliopsis subg. Verbenapis Cockerell and Atkins, 1902. Ann. and Mag. Nat. Hist. (7) 10:
44.

Type-species: Calliopsis verbenae Cockerell and Porter. Monotypic.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 876-891, figs. 123-143, map 8 (redescription,
included spp.).

Calliopsis hirsutifrons Cockerell
West. Tex. (El Paso), N. Mex.; Mexico (Chihuahua, Guanajuato, Hidalgo
and Mexico).
Calliopsis hirsutifrons Cockerell, 1896. Canad. Ent. 28: 158. ♂.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 887-889, figs. 134-138. 9, 6 (redescription).

Calliopsis micheneri Shinn
Tex. (Carrizo Springs). Pollen: Unknown, but visits flowers of Verbena
cloveri.
Calliopsis (Verbenapis) micheneri Shinn, 1967. Kans. Univ. Sci. Bui. 46: 889, figs. 139-143,

map 8. ♀, ♂.




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Calliopsis nebraskensis Crawford
N. J., Wis. west to N. Dak., S. Dak., Nebr., Colo., south to 111., Ark.
and Kans. Parasite: Holcopasites heliopsis (Robt.), Sphecodes sp.? Pollen: Apparently
polylectic since mixed pollen loads from flowers of Leguminosae and Verbenaceae have
been observed; visitation records include flowers of Ambrosia, Asclepias, Medicago
sativa. Verbena hastata, V. stricta, V. urticaefolia, Vernonia.
Calliopsis verbenae var. Nebraskensis Crawford, 1902. Canad. Ent. 34: 240. ♀, ♂.

Taxonomy: Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 291-292, figs. 69-71, table 8
(redescription). — Shinn, 1967. Kans. Univ. Sci. Bui. 46: 882-887, figs. 129-133, map 8
(redescription).

Biology: Robertson, 1914. Ent. News 25: 72 (floral relationships, as verbenae). — Rau and
Rau, 1916. Jour. Anim. Behavior 6: 368 (nesting habits, mating). —Robertson, 1922. Psyche
29: 171 (floral relationships, as verbenae). —Rau, 1922. Acad. Sci. St. Louis, Trans. 24 (7):
33 (nesting habits, mating behavior, parasite). —Shinn, 1967. Kans. Univ. Sci. Bui. 46:
886-887 (summary of bionomics including floral relationships). — Hurd and Linsley, 1972.
Smithsn. Contrib. Zool. 114: 39, table 1 (parasite).

Calliopsis verbenae Cockerell and Porter
West. Tex. to southeast. Ariz.; Mexico (Durango). Pollen:

Unknown, but visits flowers of C hamaesaracha conioides, Sphaeralcea lobata. Verbena

bipinnatifida, V. macdougalli.
Calliopsis verbenae Cockerell and Porter, 1899. Ann. and Mag. Nat. Hist. (7) 4: 412. ♀, ♂.

Taxonomy: Shinn, 1967. Kans. Univ. Sci. Bui. 46: 878-881, figs. 123-128, map 8 (tax. status,
redescription, flower records).

NoMEN Nudum in Calliopsis Smith

Calliopsis bridwelli Bridwell, 1899. Kans. Acad. Sci., Trans. 16: 210.

Unplaced Taxon in Calliopsis Smith

Calliopsis flavifrons Smith
Va., Fla.

Calliopsis flavifrons Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 129. ♂.

Taxonomy: Cockerell, 1905. Amer. Ent. Soc, Trans. 31: 321. 9. —Mitchell, 1960. N. C. Agr.
Expt. Sta. Tech. Bui. 141: 293 (tax. status). —Shinn, 1967. Kans. Univ. Sci. Bui. 46: 892
(tax. status).

Genus HYPOMACROTERA Cockerell and Porter

Hypomacrotera Cockerell and Porter, 1899. Ann. and Mag. Nat. Hist. (7) 4: 418.
Type-species: Hypomacrotera callops Cockerell and Porter. Orig. desig.

Biology: Rozen, 1970. Amer. Mus. Novitates 2416: 1-16, 19 figs., table 1 (nest architecture,
life history, mating habits, immature stages, nest associates).

Hypomacrotera callops
***authority mismatch
callops Cockerell and Porter. Colo., N. Mex., Ariz. Parasite: Holcopasites illinoiensis
minimus (Linsley)? Pollen: Collects pollen from the flowers of Sphaeralcea.
Hypomacrotera callops Cockerell and Porter, 1899. Ann. and Mag. Nat. Hist. (7) 4: 419. ♂.
9.

Taxonomy: Cockerell, 1937. Amer. Mus. Novitates 948: 3. —Rozen, 1970. Amer. Mus.
Novitates 2416: 7-9, 15-16, figs. 3-8, 18-19 (larva, pupa).

Biology: Rozen, 1970. Amer. Mus. Novitates 2416: 2-5, table, 1 (nest architecture, nesting
habits, pollen source). —Hurd and Linsley, 1972. Smithsn. Contrib. Zool. 114: 36, table 1
(parasite).

Hypomacrotera callops
***authority mismatch
persimilis Cockerell. Ariz., south. Calif.; Mexico (Baja California and Sonora). Parasite:
Holcopasites illinoiensis yninimus (Linsley). Pollen: Collects pollen from the flowers of
Physalis, but visits other flowers for nectar including Melilotus alba, Sida hederacea.
Hypomacrotera callops persimilis Cockerell, 1899. Entomologist 33: 64. ♀, ♂.

Biology: Hurd and Linsley, 1972. Smithsn. Contrib. Zool. 114: 36, table 1 (parasite).

Hypomacrotera subalpinus
***authority mismatch
andradensis Cockerell. N. Mex. to south. Calif.; Mexico (Baja California and ^

Sonora). Pollen: Collects pollen from the flowers of Sphaeralcea including S. ambigua, S.




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angustifolia, S. emoryi, S. orcutti, but visits other flowers for nectar including Baileya

multiradiata, B. pleniradiata.
Hypomacrotera andradensis Cockerell, 1937. Amer. Mus. Novitates 948: 3. ♀, ♂.

Hypomacrotera subalpinus
***authority mismatch
subalpinus (Cockerell). N. Mex., Ariz.; Mexico (Sonora). Parasite: Oreopasites sp.

Pollen: Collects pollen from the flowers of Spkaeralcea including S. ambigua, S. orcutti.
Calliopsis subalpinus Cockerell, 1894. Ent. News 5: 235. ♂.
Calliopsis seminifus Cockerell, 1896. Ent. Monthly Mag. 32: 219. ♀.

Biology: Rozen, 1970. Amer. Mus. Novitates 2416: 2-5, table 1 (nest architecture, nesting
habits, pollen source, parasite).

Genus XENOPANURGUS Michener

Xenopanurgus Michener, 1952. Kans. Ent. Soc, Jour. 25: 24.

Type-species: Xenopanurgus readioi Michener. Orig. desig.

Taxonomy: Shinn, 1964. Ent. News 75: 73-78 (included sp.).

Xenopanurgus readioi Michener
Ariz. (Huachuca Mts.); Mexico (Toluca).

Xenopanurgus readioi Michener, 1952. Kans. Ent. Soc, Jour. 25: 25, 5 figs. S.

Taxonomy: Shinn, 1964. Ent. News 75: 74-78. 9, cj.

Genus PERDITA Smith

This is a very large genus of pollen-collecting bees that has estabhshed through its component
species a predominantly oligolectic relationship with the flora of North America. Even though
the genus occurs transcontinentally in southern Canada, the United States and Mexico to as far
south as Central America (Guatemala), it is centered in the more arid areas of northern Mexico
and adjacent southwestern United States. The nesting habits of only a few species have been
studied and these, like other species of the family, make their nests in the ground, although one
species, Perdita (Cockerellula) opuntiae Cockerell, is known to excavate its nests in sandstone.
Several females of at least some species occupy a single nest, but there is no evidence of caste
development.

In recognition of utilitarian advantages, the subgenera are arranged alphabetically in this
catalog.

Revision: Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 25-107. —Cockerell, 1922. Amer.
Mus. Novitates 33: 14 (partial key). — Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 345-432,
172 figs, (all subgenera except Hexaperdita Timberlake, Perditella Cockerell, Alloperdita
Viereck, and typical Perdita Smith). —Timberlake, 1956. Calif. Univ. Pubs. Ent. 11:
247-350, figs. 173-367 (treats subgenera Alloperdita Viereck, Perditella Cockerell,
Hexaperdita Timberlake, and Pygoperdita Timberlake). —Timberlake, 1958. Calif. Univ.
Pubs. Ent. 14: 303-410, figs. 368-521 (treats zonalis and halictoides groups of subgenus
Perdita). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 1-156, figs., 522-732 (treats

Perdita octomaculata
***authority mismatch
group of subgenus Perdita). — Timberlake, 1962. Calif. Univ. Pubs. Ent. 28:
1-124, figs. 733-904, 2 text figs, (treats ventralis group of subgenus Perdita, and suppl. to
Parts I-IV). —Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 125-388, figs. 905-1190, 77 text
figs, (treats sphaeralceae group of subgenus Perdita). — Timbelake, 1968. Calif. Univ.
Pubs. Ent. 49: 1-196, figs. 1191-1360, 31 text figs, (suppl. to preceding parts, bibliography,
index to Parts I-VII, corrigenda). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 1-63, figs.
1361-1417 (suppl. to Parts I-VII, treats subgenera Cockerellia, Epimacrotera,
Hexaperdita, Pentaperdita, Perdita, Procockerellia, Pygoperdita, Xeromacrotera).

Taxonomy: Cockerell, 1922. Amer. Mus. Novitates 33: 1-15 (west. U. S. spp., species
formation). —Timberlake, 1952. Ent. Soc. Wash., Proc. 54: 199-204 (eastern U. S. spp.).
—Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 961-985 (treats spp. of the subgenera
Cockerellia, Cockerellula, Macroteropsis, Xerophasma). —Mitchell, 1960. N. C. Agr. Expt.
Sta. Tech. Bui. 141: 295-330, figs. 58, 72-79, table 9 (eastern U. S. spp.). —Rozen, 1966.
Amer. Mus. Novitates 2259: 5, 16-21, figs. 41-54 (larvae). —Yager and Rozen, 1966. Amer.
Mus. Novitates 2265: 12 (pupae).




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Biology: Linsley, 1958. Hilgardia 27: 562, tables 4, 5 (pollen sources of some subgenera).
— Rozen, 1967. Amer. Mus. Novitates 2297: 1-44, 18 figs., 2 tables (review). — Hurd and
Linsley, 1975. Smithsn. Contrib. Zool. 193: 24-26, figs. 5-7 {Larrea visiting spp.).

Genus PERDITA Subgenus ALLOMACROTERA Timberlake

Perdita subg. Allomacrotera Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 131.

Type-species: Perdita stephanomeriae Timberlake. Monotypic and orig. desig.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 5-6 (key to spp. of subgenera
Allomacrotera, Procockerellia).

Perdita stephanomeriae Timberlake
Calif. (Colorado and Mojave Deserts). Pollen: Possibly an

oligolege of Stephanomeria including S. pauciflora, a male has been collected at the
flowers of Dalea spinosa.
Perdita stephanomeriae Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 404. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 60: 131-132, figs. 658, 659, 731. ♂.

Genus PERDITA Subgenus ALLOPERDITA Viereck

Perdita subg. Alloperdita Viereck, 1917. Amer. Mus. Nat. Hist., Bui. 37: 241.
Type-species: Perdita novae-angliae Viereck. Monotypic.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 348 (key). —Timberlake, 1956. Calif.
Univ. Pubs. Ent. 11: 271-275 (redescription, key to spp.). —Timberlake, 1958. Calif. Univ.
Pubs. Ent. 14: 393 (modification of key). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17:
134. —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 30.

Perdita bradleyi Viereck
N. J. to Fla. Pollen: Unknown, but visits flowers of Batodendroyi, Crataegus,
Hydrocotyle umbellatiim, Ilex, Nyssa sylvatica, Pyracantha, Sarracenia flava.
Perdita bradleyi Viereck, 1907. Ent. News 18: 393. ♂.

Taxonomy: Timberlake, 1928. Amer. Mus. Novitates 321: 7 (as obscurata). —Timberlake,
1956. Calif. Univ. Pubs. Ent. 11: 274, figs. 211, 212, 306 (as obscurata). —Timberlake, 1958.
Calif. Univ. Pubs. Ent. 14: 394 (as obscurata). —Timberlake, 1960. Calif. Univ. Pubs. Ent.
17: 134 (type). —Mitchell, 1964. N. C. Agr. Expt. Sta. Tech. Bui. 141: 310-311, figs. 72-73,
table 9 (redescription).

Perdita floridensis Timberlake
N. C. to Fla. Pollen: Unknown, but visits flowers of Batodendroyi, Ilex
glabra, Leucothoe.
Perdita floridensis Timberlake, 1928. Amer. Mus. Novitates 321: 1. ♂, ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 273, 274 (key, geogr. records).
—Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 393 (key). -Mitchell, 1960. N. C. Agr.
Expt. Sta. Tech. Bui. 141: 311, figs. 72-74, 79, table 9 (redescription).

Perdita gertschi Timberlake
Tex. (Brewster County); Mexico (Coahuila). Pollen: Unknown, but visits
flowers of Prosopis juliflora.
Perdita (Alloperdita) gertschi Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 393, figs. 468,
469,519. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 30-31. ♀.

Perdita mitchelli Timberlake
N. C, Fla., Ala., Miss. Pollen: Unknown, but visits flowers of Ceanothus,
Cyrilla racemiflora, Oxydendrum.
Perdita mitchelli Timberlake, 1947. Ent. Soc. Wash., Proc. 49: 81. ♀, ♂.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 273, 274, figs. 213-214, 307 (key,
geogr. records). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 311-312, figs. 73-74,
table 9 (redescription).

Perdita novaeangliae Viereck
Mass. to Fla. Pollen: Unknown, but visits flowers of Batodendron,
huckleberry, Lyonia ligustrina.
Perdita novae-angliae Viereck, 1907. Ent. News 18: 394. ♀, ♂.
Perdita novaeangliae Viereck, 1917. Amer. Mus. Nat. Hist., Bui. 37: 241. Emend.




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Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 273, 274, 275, figs. 219, 220, 310 (key,
genitalia). -Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 393 (key). -Mitchell, 1960. N. C.
Agr. Expt. Sta. Tech. Bui. 141: 312-313, figs. 72-73, table 9 (redescription).

Perdita obscurata Cresson
N. C. to Tex. Parasite: Neolarra cockerelli (Cwfd.)? Pollen: Unknown, but
visits flowers of Batodendron, Diospyros virginiana, Stillmgia sylvatica.
Perdita obscurata Cresson, 1878. Amer. Ent. Soc, Trans. 7: 70. ♀, ♂.
Perdita Carolina Timberlake, 1952. Ent. Soc. Wash., Proc. 54: 199. ♀, ♂.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 273, 274, figs. 215, 216, 308 (key,
genitalia, geogr. and floral records, as Carolina). —Timberlake, 1960. Calif. Univ. Pubs.
Ent. 17: 134 (synonymy). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 313-314,
figs. 72-74, 79, table 9 (synonymy, redescription).

Perdita townesi Timberlake
N. C. to Fla. Pollen: Unknown, but visits flowers of Batodendron,
Ceanothus niicrophyllus, Diospyros virginiana. Ilex, Nyssa sylvatica.
Perdita townesi Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 31, figs. 1209-1210, 1308. ♂.

9.

Genus PERDITA Subgenus COCKERELLIA Ashmead

Cockerellia Ashmead, 1898. Psyche 8: 284.

Type-species: Perdita albipennis Cresson. Monotypic and orig. desig.
(=Perdita hyalina Cresson).
Philoxanthus Ashmead, 1898. Psyche 8: 285.

Type-species: Perdita beata Cockerell. Monotypic and orig. desig.

The species of this subgenus are found almost exclusively at flowers of Compositae, mostly of
the genera Baileya, Coreopsis, Gaillardia, Geraea, Helianthus, Ratibida, Rudbeckia and Ver-
besina, but also occur at the flowers oi Erigeron, Heterotheca, Pedis and Prionopsis.

Taxonomy: Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 971-974 (key to females of included
spp.). —Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 347, 385-402 (key to included spp.).
—Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 326. —Timberlake, 1958. Calif. Univ. Pubs.
Ent. 14: 382-383. -Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 19-22 (modification of
key). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 2-5 (key to spp. allied to hilaris).

Perdita albihirta
***authority mismatch
albihirta Timberlake. South. Calif. (Colorado Desert). Pollen: Apparently an

oligolege of Geraea canescens, but visits the flowers of Encelia farinosa and Melilotus

presumably for nectar.
Perdita albihirta albihirta Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 395, fig. 79. ♀, ♂.

Perdita albihirta
***authority mismatch
geraeae Timberlake. Calif. (Colorado and Mojave Deserts), Nev. (Statehne). Pollen:

Apparently an oligolege of Geraea canescens, but visits the flowers of Acamptopappus

sphaerocephaius var. hirtellus and Larrea tridentata presumably for nectar.
Perdita albihirta geraeae Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 396. ♀, ♂.

Perdita albipennis
***authority mismatch
albipennis Cresson. N. Dak. south to Tex., west to Wyo., Colo, and Ariz. Pollen:

Unknown, but visits flowers of Helianthus annuus, H. petiolaris, Hymenopappus,

Nolina, Prionopsis ciliata. Predator: Philanthus ventilabris Fabr.
PerditaC.) albipennis Cresson, 1868. Amer. Ent. Soc, Trans. 1: 386. ♀.
Perdita hyalina Cresson, 1878. Amer. Ent. Soc, Trans. 7: 68. ♂.
Perdita albipennis var. helianthi Cockerell, 1896. Acad. Nat. Sci. Phila., Proc 48: 104. ♂.

9.
Perdita albipennis var. helianthi mut. pasonis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc.

48: 105. ♂.
Perdita lacteipennis Swenk and Cockerell, 1907. Ent. News 18: 51. ♀, ♂.

Taxonomy: Cockerell, 1899. Psyche 8: 419. 6 (as hyalina). —Cockerell, 1899. Psyche 8: 419.
6 (as pasonis). —Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 977-978 (geogr. and floral
records, as lacteipennis lacteipennis). —Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 388,
390, 391, 398, 400 (geogr. and floral records, in part, as albipennis, lacteipennis
lacteipennis, pasonis and hyalina). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 130-131
(types, synonymy, status discussed). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 20
(synonymy, status discussed).




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Biology: Alcorn and Gamboa, 1975. Ariz. Acad. Sci. 10: 163 (predator).

Perdita albipennis
***authority mismatch
canadensis Crawford. Alta., Idaho, N. Dak., Utah (Uinta and Wasatch Counties).
Perdita canadensis Crawford, 1912. Canad. Ent. 44: 360. ♀.

Taxonomy: Stevens, 1919. Canad. Ent. 51: 205. 6. — Timberlake, 1954. Calif. Univ. Pubs. Ent.
9: 388, 390, 398-399 (key, tax. status, as lacteipennis canadensis). —Timberlake, 1960.
Calif. Univ. Pubs. Ent. 17: 130-131 (status discussed).

Perdita albipennis
***authority mismatch
heliophila Cockerell. Utah, Colo., N. Mex.; Mexico (Chihuahua). Pollen: Unknown,
but visits flowers oi Heliantlius including H. annuiis.
Perdita heliophila Cockerell, 1916. Ann. and Mag. Nat. Hist. (8) 17: 281. ♀.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 390, 397, figs. 82, 83, 158 (tax.
characters and tax. status). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 20 (tax.
characters). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 2 (tax. characters).

Perdita albipennis
***authority mismatch
pallidipennis Graenicher. Ind. and Wis., west to east. Kans. and Nebr. Pollen:
Unknown, but visits flowers of Helianthus annuus.
Perdita pallidipennis Graenicher, 1910. Canad. Ent. 42: 101. ♀, ♂.

Taxonomy: Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 978 (tax. characters and status, as

Perdita lacteipennis
***authority mismatch
pallidipennis). —Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 388, 390, 399
(key, geogr. records). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 131 (status
discussed). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 302-303, table 9
(redescription).

Perdita beata
***authority mismatch
beata Cockerell. Tex. (Davis Mts. and El Paso), N. Mex. (Las Cruces and Mesilla Park).
Pollen: Unknown, but visits flowers of Verbesina encelioides. Another subspecies,
Perdita beata signata Timberlake, occurs in Mexico (Chihuahua, Coahuila and Durango).
Perdita beata Cockerell, 1895. Psyche 1 (sup.): 10. ♀.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 386, 391 (key, geogr. and floral
record). -Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 130 (tax. status).

Perdita bequaerti
***authority mismatch
bequaerti Viereck. N. J. to Fla., west to Minn, and Miss. Pollen: Unknown, but visits
flowers of Bideyis, Helianthus divaricatus, Physalis lanceolata, P. virginiana.
Perdita {Cockerellia) bequaerti Viereck, 1917. Amer. Mus. Nat. Hist., Bui. 37: 241. ♀, ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 397, figs. 80, 157 (tax. characters).
—Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 300, figs. 72-74, 79, table 9
(redescription).

Perdita bequaerti
***authority mismatch
indianensis Cockerell. Ind. (Gary and Lafayette), Okla. (South McAlester), Tenn.
(Knoxville).
Perdita pallidipennis indianeiisis Cockerell, 1922. Amer. Mus. Novitates 33: 2. Perdita (Cockerellia) wickhami Cockerell, 1922. Amer. Mus. Novitates 33: 13. ♀.

Taxonomy: Timberlake, 1952. Ent. Soc. Wash., Proc. 54: 204 (tax. status). —Timberlake, 1953.
Kans. Univ. Sci. Bui. 35: 976 (tax. characters, synonymy). —Timberlake, 1954. Calif. Univ.
Pubs. Ent. 9: 388, 390, 397 (key, tax. status, geogr. range). —Mitchell, 1960. N. C. Agr.
Expt. Sta. Tech. Bui. 141: 302, fig. 72 (status discussed).

Perdita coreopsidis
***authority mismatch
collaris Cockerell. N. Mex.; Mexico (Chihuahua).

Perdita verbesinae collaris Cockerell, 1916. Ann. and Mag. Nat. Hist. (8) 17: 281. ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 389, 390, 401 (key, redescription).

Perdita coreopsidis
***authority mismatch
coreopsidis Cockerell. Kans., Tex.; Mexico (Coahuila). Pollen: Unknown, but visits
flowers of Gaillardia which it apparently prefers, but also has been t^aken at the
flowers of Coreopsis and Monarda punctata coryi.
Perdita coreopsidis Cockerell, 1906. Entomologist 39: 126. ♀, ♂.

Taxonomy: Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 983-984 (geogr. and floral records).
—Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 389, 390, 401 (key, geogr. and floral records).
—Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 4 (key).




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Perdita coreopsidis
***authority mismatch
kansensis Timberlake. Kans. (Grove and Ford Counties), Tex. (Van Horn), N.
Mex. (Roosevelt County), Utah (Iron County). Pollen: Unknown, but visits flowers of
Gaillardia.
Perdita coreopsidis kansensis Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 984. ♀, ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 389, 390, 401 (key, geogr. range).
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 20 (geogr. records).

Perdita coreopsidis
***authority mismatch
obscurior Timberlake. Gulf Coast of Tex. (Galveston to Brownsville, but a male

has been taken inland at Lytle, Atacosa County). Pollen: Unknown, but visits flowers of
Gaillardia.
Perdita coreopsidis obscurior Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 984. ♀, ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 389, 390, 401 (key, geogr. records).

Perdita fracticincta Timberlake
Tex. (Sarita). Pollen: Unknown, but visits flowers of Coreopsis.
Perdita fracticincta Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 976. female.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 388, 397 (key).
hiiaris Timberlake. Utah (Dixie State Park), N. Mex. (San Juan County). Pollen: Unknown, but
visits flowers of ChrysothaninMS naitseosus.
Perdita hiiaris Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 21, figs. 1207, 1208, 1307. ♂.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 2-3. ♀.

Perdita imbellis Timberlake
Ariz. (Coconino County).

Perdita imbellis Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 21, figs. 1205, 1206, 1306. ♂.

Perdita incana Timberlake
Tex. (Culberson County), N. Mex. (Quay County).

Perdita iyicaiia Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 383. ♀, ♂.

Perdita lepachidis
***authority mismatch
lepachidis Cockerell. Kans., Tex., Colo., N. Mex. Pollen: Unknown, but visits
flowers of Ratibida tagetes.
Perdita lepachidis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 106. ♂.

Taxonomy: Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 982-983. 9. —Timberlake, 1954. Calif.
Univ. Pubs. Ent. 9: 389, 391, 401 (key). -Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 4
(key).

Perdita lepachidis
***authority mismatch
levifrons Timberlake. Tex. (Brownsville).

Perdita lepachidis levifrons Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 983. ♀, ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 389, 391, 401 (key).

Perdita lingualis Cockerell
Nebr., Kans., Tex., N. Mex., Colo., Utah; Mexico (Tamaulipas). Pollen:

Apparently an oligolege oi Helianthus including H. a)inuus, H. petiolaris, but may also
visit other large flowered Compositae.
Perdita albipennis var. lingualis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 105. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 130, 131 (tax. status). — Michener,
1963. Kans. Ent. Soc, Jour. 36: 117-118, figs. 2-6 (prepupa). — Rozen, 1966. Amer. Mus.
Novitates 2259: 21, figs. 52-54 (larva). —Yager and Rozen, 1966. Amer. Mus. Novitates
2265: 12, figs. 17-18 (pupa). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 4 (key).

Biology: Michener, 1963. Kans. Ent. Soc, Jour. 36: 114-118, 6 figs, (nest architecture, colonial
behavior, floral relationships).

Perdita luculenta Timberlake
South. Calif. (Imperial County).

Perdita luculenta Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 19. ♀.

Perdita perpulchra
***authority mismatch
flavidior Timberlake. Kans., (Barton and Rice Counties), Tex. (Somervel County).
Pollen: Unknown, but visits flowers of Aphanostephus skirrhobasis, Heterotheca
subaxillaris, Rudbeckia bicolor.
Perdita perpulchra flavidior Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 975. ♀.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 395. —Timberlake, 1956. Calif. Univ.
Pubs. Ent. 11: 327 (geogr. and floral records).

Perdita perpulchra
***authority mismatch
perpulchra Cockerell. Tex. (El Paso), N. Mex. (Mesilla Valley), Ariz. (Holbrook and
Tucson); Mexico (Aguascalientes and Durango). Pollen: Unknown, but visits flowers of
Asclepias, Baileya, Haplopappus heterophylla, Verbesina encelioides.
Perdita perpulchra Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 99. ♀.




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Perdita baileyae Cockerell, 1909. Canad. Ent. 41: 151. ♂.

Taxonomy: Cockerell, 1897. Ann. Mag. and Nat. Hist. (6) 19: 398. 6. — Timberlake, 1953.
Kans. Univ. Sci. Bui. 35: 975 (tax. characters). —Timberlake, 1954. Calif. Univ. Pubs. Ent.
9: 387, 389, 395 (key, tax. characters). -Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 327
(synonymy, tax. status).

Perdita perpulchra
***authority mismatch
punctissima Timberlake. Tex. (Bexar, Wood, Robertson, Lamar Counties). Pollen:
Unknown, but visits flowers of Heterothpca subaxillaris.
Perdita perpulchra ptmctissima Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 327. ♀, ♂.

Perdita pulchella Timberlake
Tex. (Carrizo Springs).

Perdita pulchella Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 393. ♀.

Perdita purpurascens Timberlake
Tex. Pollen: Unknown, but visits flowers of Aster tanacetifolius,
Brazoria truncata, Gaillardia, Monarda.
Perdita purpurascens Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 328. ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 382-383. 9 (tax. characters, geogr.
and floral records).

Perdita scopata Timberlake
Gulf Coast of Tex. (Galveston to Brownsville). Pollen: Unknown, but
visits flowers of Coreopsis, Helianthus annuus.
Perdita scopata Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 980. ♀, ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 401, fig. 88 (tax. characters).

Perdita shinnersi Timberlake
Tex. Pollen: Unknown, but visits flowers o{ Brazoria truncata.
Coreopsis basalis, Gaillardia, Hymenopappus artemisaefolius, Monarda.
Perdita shi^inersi Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 326. ♀, ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 383 (geogr. and floral records).

Perdita tricincta Timberlake
South. Tex. (Cameron, Hidalgo and Starr Counties). Pollen: Unknown,
but visits flowers of Helianthus anriuus. This may prove to be a race of Perdita

Perdita albipennis
***authority mismatch
Cress.
Perdita tricincta Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 978. ♀, ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 388, 390, 400, fig. 86 (key, tax.
status).

Perdita utahensis Cockerell
N. Mex. (Rodeo), Ariz. (Tucson), southwest. Utah, south. Calif. (Colorado
Desert). Pollen: Unknown, but visits flowers of Pectis papposa, Tidestromia, Verbesina
encelioides.
Perdita utahensis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 106. ♀.

Taxonomy: Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 974. 6. —Timberlake, 1954. Calif.

Univ. Pubs. Ent. 9: 387, 391, 394, figs. 76, 155 (tax. characters, geogr. and floral records).

—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 20 (geogr. and floral records).

Perdita verbesinae Cockerell
Tex. (Barstow), N. Mex. (Deming and Las Cruces), Ariz. (Douglas,

Portal, Tucson and Santa Rita Mts.); Mexico (Chihuahua). Pollen: Usually visits flowers
of Verbesiiia encelioides, but has been recorded from the flowers o{ Helianthus.

Perdita verbesinae Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 102. ♀, ♂.

Perdita verbesinae mut. intermedia Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 103. ♀.

Perdita verbesinae mut. nigrior Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 103. ♀.

Perdita verbesinae mut. cyanella Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 104. ♂.

Perdita verbesinae mut. maculata Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 104. ♂.

Taxonomy: Cockerell, 1899. Psyche 8: 323 (color variation). —Timberlake, 1954. Calif. Univ.
Pubs. Ent. 9: 388, 391, 397, fig. 81 (key, tax. characters, geogr. and floral records).

Genus PERDITA Subgenus COCKERELLULA Strand

Perdita subg. Lutziella Cockerell, 1922. Amer. Mus. Novitates 47: 1. Preocc.

Type-species: Perdita (Lutziella) opuntiae Cockerell. Monotypic and orig. desig.
Perdita subg. Cockerellula Strand, 1932. Folia Zool. Hydrobiol. Riga 4: 196. N. name.




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Taxonomy: Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 963-968 (tax. characters, key to
included spp.). —Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 346, 354-356 (key to included
spp., tax. characters). -Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 122-123 (key to males
of included spp.). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 2 (modified key to males).

Perdita bidenticauda Timberlake
Tex. (Big Bend Park). Pollen: Unknown, but visits flowers of
Opiintia.
Perdita bidenticauda Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 965. ♂, ♀.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 356, figs. 13, 14, 122 (tax. characters,
key). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 122 (key). —Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 2 (key).

Perdita knulli Timberlake
Tex. (Davis Mts.). Pollen: Unknown, but the female allotype bears dark
brown pollen, with spherical and moderately fine grains.
Perdita knulli Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 123, figs. 652, 653, 728. ♂, ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 2. 6 (key).

Perdita laticauda Timberlake
Tex. (Dryden); Mexico (Coahuila). Pollen: Unknown, but visits flowers
oi Cha7uaesaraclia conioides, Gaillardia, Gilia acerosa.
Perdita laticauda Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 967. ♂, ♀.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 356, figs. 15, 16, 123 (tax. characters,
key). -Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 122. 6 (key).

Perdita lobata Timberlake
Tex. (Hidalgo and Starr Counties). Pollen: Unknown, but visits flowers of
Opuntia.
Perdita lobata Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 967. ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 354, 356, figs. 17, 18, 124 (tax.
characters, key). —Timberlake, 1960. CaHf. Univ. Pubs. Ent. 17: 122. 6, 9 (tax. characters,
key).

Perdita opuntiae Cockerell
Colo., Wyo., S. Dak. (Pine Ridge). Ecology: Nests in soft sandstone rocks
of the Laraniie formation. Pollen: Collects pollen from the flowers of Opuntia.
Perdita (Lutziella) opiintiae Cockerell, 1922. Amer. Mus. Novitates 47: 2. ♂, ♀.

Taxonomy: Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 964 (key). —Timberlake, 1954. Calif.
Univ. Pubs. Ent. 9: 354, 355, figs. 9, 10, 120 (tax. characters, key). —Timberlake, 1960.
Calif. Univ. Pubs. Ent. 17: 122. 6 (key^

Biology: Custer, 1928. Psyche 35: 67-84 (nesting habits). —Custer, 1929. Canad. Ent. 61: 49
(nesting habits). —Custer, 1929. Psyche 36: 293 (nesting habits).

Genus PERDITA Subgenus EPIMACROTERA Timberlake

Perdita subg. Epimacrotera Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 377.
Type-species: Perdita ainsliei Crawford. Orig. desig.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 378-379 (key to included spp.).
—Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 380 (modified key to included spp.).
—Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 129 (tax. characters, relationship to
Glossoperdita). —Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 92-93 (revised key to
included spp.). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 14-15 (key to males).

Perdita ainsliei Crawford
Iowa, N. Mex. (Bernalillo County).

Perdita ainsliei Crawford, 1932. Ent. Soc. Wash., Proc. 34: 74. ♂, ♀.

Taxonomy: Timberlake, 1954. CaHf. Univ. Pubs. Ent. 9: 378, 379, figs. 55, 56, 144 (key, tax.
characters). —Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 380 (key, geogr. record).
—Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 93 (key). —Timberlake, 1968. CaHf. Univ.
Pubs. Ent. 49: 14 (key).

Perdita biguttata Timberlake
Ariz. (Cochise County). Pollen: Unknown, but visits flowers of
Eriogonum abertianum var. neomexicanuiu, Euphorbia.
Perdita biguttata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 95. ♀.
Perdita parvula Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 15, figs. 1202, 1304. ♂.

Taxonomy: Timberlake, 1968. CaHf. Univ. Pubs. Ent. 49: 15 (synonymy in footnote).




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Perdita binotata Timberlake
Ariz.; Mexico (Baja California). Pollen: Unknown, but visits flowers of
Physalis.
Perdita binotata Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 385, figs. 73, 74, 153. ♀, ♂.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 93 (key). —Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 14 (key).
crassula Timberlake. N. Mex. (Las Cruces), Ariz. (Willcox). Pollen: Unknown, but visits
flowers of Euphorbia.
Perdita crassula Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 381, figs. 462, 463, 516. ♂.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 93-94. 9 (key). —Timberlake, 1968.
Calif. Univ. Pubs. Ent. 49: 14 (key).

Perdita diversa Timberlake
Tex. (El Paso); Mexico (Coahuila, Nuevo Leon). Pollen: Unknown, but
visits flowers of Coldenia greggii.
Perdita diversa Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 384, figs. 69, 70, 151. ♂, ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 15, 19 (key).

Perdita euphorbiae Timberlake
Ariz. (Picacho Pass); Mexico (Sonera). Pollen: Unknown, but visits
flowers of Euphorbia, Kallstroemia.
Perdita euphorbiae Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 380, figs. 59, 60, 146. ♀, ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 130 (geogr. and floral records).
—Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 93 (key). —Timberlake, 1968. Calif. Univ.
Pubs. Ent. 49: 15, 17 (key, geogr. record).

Perdita namatophila Timberlake
Calif. (Riverside County). Pollen: Unknown, but visits flowers of
Nama demissum.
Perdita namatophila Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 379, figs. 57, 58, 145. ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 380 (key). —Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 15 (key).

Perdita nanula Timberlake
Ariz. (Douglas). Pollen: Unknown, but visits flowers of Euphorbia
albomarginata.
Perdita nanula Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 129. ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 93 (key). —Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 15, 16. 6 (key).

Perdita nigrocaerulea Timberlake
CaUf. (Oro Grande Wash). Pollen: Unknown, but visits flowers of
Salazaria mexicana.
Perdita nigrocaendea Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 382, figs. 65, 66, 149. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 15 (key).

Perdita peculiaris Timberlake
Tex. (Terrell County). Pollen: Unknown, but visits flowers of
C hamaesaracha conioides.
Perdita pectdiaris Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 970. ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 384, figs. 71, 72, 152 (tax. characters).
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 15 (key).

Perdita polycarpae Timberlake
South. Calif.; Mexico (Baja California). Pollen: Possibly an oligolege of
Euphorbia including E. polycarpa, but also visits flowers of Pedis papposa.
Perdita polycarpae Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 381, figs. 61, 62, 147. ♂, ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 14 (key).

Genus PERDITA Subgenus GLOSSOPERDITA Cockerell

Glossoperdita Cockerell, 1916. Pomona Jour. Ent. Zool. 8: 43.

Type-species: Glossoperdita pelargoides Cockerell. Monotypic.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 347, 392-394 (redescription, key to
included spp.). —Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 379 (key to included spp.).

Perdita blaisdelli Timberlake
Calif. (Calaveras County).

Perdita blaisdelli Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 373. ♀.




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Perdita giliae Timberlake
Ariz. Pollen: Unknown, but visits flowers of Gilia, possibly G. aggregata.
Perdita giliae Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 374, figs. 51, 52, 141. ♀, ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 379 (key).

Perdita hurdi Timberlake
South. Calif., Ariz.; Mexico (Sonora). Ecology: Cuts unopened flowers of
Proboscidea in order to obtain pollen. Pollen: Collects pollen from the flowers of
Proboscidea althaeafolia, P. arenaria.
Perdita hurdi Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 324, figs. 283, 284, 335. 9,
Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 379 (key). —Timberlake, 1960. Calif.
Univ. Pubs. Ent. 17: 127-128 (tax. characters, geogr. and floral records).

Biology: Hurd and Linsley, 1963. Kans. Ent. Soc, Jour. 36: 248-252, 5 figs, (corolla cutting
habits, floral relationships).

Perdita navarretiae
***authority mismatch
angusticeps Timberlake. Calif. (Tuolumne County).

Perdita navarretiae angusticeps Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 128. ♂, ♀.

Perdita navarretiae
***authority mismatch
navarretiae Timberlake. Calif. (Mariposa County). Pollen: Unknown, but visits
flowers of Navarretia viscidula,
Perdita navarretiae Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 379, figs. 460, 461, 515. ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 128 (tax. status).

Perdita navarretiae
***authority mismatch
powelli Timberlake. Calif. (Nevada County).

Perdita navarretiae powelli Timberlake, 1962. Calif. Univ. Pubs. Ent. 91. ♂, ♀.

Perdita pelargoides (Cockerell)
South. Calif, (cismontane). Pollen: Unknown, but visits flowers of
Hugelia virgata, Navarretia atractyloides.
Glossoperdita pelargoides Cockerell, 1916. Pomona Jour. Ent. Zool. 8: 43. ♀.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 373, figs. 49, 50, 140. ♂.
—Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 379 (key).

Genus PERDITA Subgenus HESPEROPERDITA Timberlake

Perdita subg. Hesperoperdita Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 374.
Type-species: Perdita nificauda Cockerell. Orig. desig.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 375 (key to included spp.).

Perdita pyrifera Cockerell
Calif. (Pleyto, Monterey County).

Perdita pyrifera Cockerell, 1925. Calif. Acad. Sci., Proc. (4) 14: 192. ♀.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 376, fig. 142. ♂.

Perdita trisignata
***authority mismatch
ornata Timberlake. North. Calif. Pollen: Possibly an oligolege of Lotus, visits
flowers of L. glaber, L. nevadensis, L. strigosus var. hirtellus.
Perdita ruficauda ornata Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 377. ♀, ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 127 (tax. status).

Perdita trisignata
***authority mismatch
trisignata Cockerell. South. Calif.; Mexico (Baja California). Pollen: Possibly an

oligolege of Lotus including L. americamis, L. argophyllus, L. davidsonii, L. scoparius,
L. strigosus, but also visits other flowers including Cryptantha, Heteromeles.
Perdita trisignata Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 76. ♀.
Perdita ruficauda Cockerell, 1916. Pomona Jour. Ent. Zool. 8: 44. ♀.

Taxonomy: Cockerell, 1922. U. S. Natl. Mus., Proc. 60: 17. 9 , cJ (as nificauda).
—Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 375, 376 (key, type). —Timberlake, 1954.
Calif. Univ. Pubs. Ent. 9: 375, 376, figs. 53, 54, 143 (key, tax. characters, geogr. and floral
records, as ruficauda). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 127 (synonymy).

Genus PERDITA Subgenus HETEROPERDITA Timberlake

Perdita subg. Heteroperdita Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 365.
Type-species: Perdita rhodogastra Timberlake. Orig. desig.

The species of this subgenus apparently collect pollen exclusively from flowers of the
boraginaceous genera Coldenia and Heliotropium.




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Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 347, 366 (key to included spp.).

— Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 126 (modified key to included spp.).

—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 10-11 (key to females of included spp.).

Perdita arenaria Timberlake
Ariz. (Yuma), Nev. (Washoe County), south. Calif. (Colorado Desert).

Pollen: Apparently an oligolege of Coldenia including C. nuttallii, C. palmeri.

Perdita arenaria Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 368, figs. 39, 40, 135. ♂, ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 126 (key). -Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 11, 13 (key, geogr. and floral records).

Perdita bellula Timberlake
South. Calif. (Colorado Desert). Pollen: Apparently an oligolege of
Coldenia including C. palmeri, C. plicata.
Perdita bellula Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 369, figs. 41, 42, 136. ♂, ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 11 (key).

Perdita coldeniae Timberlake
South. Calif. (Colorado Desert). Pollen: Apparently an oligolege of
Coldenia including C. palmeri, but visits flowers of Pedis papposa presumably for

Perdita coldeniae Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 368, figs. 37, 38, 134. ♂, ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 11 (key).

Perdita frontalis Timberlake
South. Calif. (Colorado Desert). Pollen: Apparently an oligolege of

Coldenia including C. plicata, but visits flowers of Eriogonum deserticola presumably
for nectar.
Perdita frontalis Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 12. ♀.

Perdita maculosa Timberlake
South. Calif. (Colorado Desert). Pollen: Apparently an oligolege of
Coldenia including C. plicata.
Perdita macidosa Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 378, fig. 514. ♀, ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 126 (key). -Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 11 (key).

Perdita optiva Timberlake
South. Calif. (Colorado Desert). Pollen: Apparently an oligolege of Coldenia
including C. palmeri.
Perdita optiva Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 371, figs. 47, 48, 139. 6 (9
misdet.).
Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 13. 6 (tax. status, as sexfasciata).

Perdita rhodogastra Timberlake
South. Calif. (Colorado Desert). Pollen: Apparently collects pollen

from the flowers of Coldenia palmeri and Heliotropium oculatum, but visits these and
other flowers for nectar including Pedis papposa, Phichea sericea.
Perdita rhodogastra Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 367, figs. 35, 36, 133. ♀.
6.
Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 11 (key).

Perdita scutellaris Timberlake
Southern Calif. (Colorado Desert). Pollen: Apparently an oligolege of
Coldenia including C. plicata, but visits the flowers of Eriogonum deserticola
presumably for nectar.
Perdita scutellaris Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 89, figs. 829, 830, 893, text
fig. B. 9,cJ.
Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 11 (key).

Perdita sexfasciata Timberlake
Ariz. (Yuma County), south. Calif. (Riverside and San Bernardino
Counties). Pollen: Apparently an oligolege of Coldenia including C. palmeri, but visits
these and other flowers for nectar including Cryptantha, Nama.
Perdita sexfasciata Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 370, figs. 43, 44, 137. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 11, 13. 9 (key).

Perdita trifasciata Timberlake
Tex. (Dryden), N. Mex. (White Sands); Mexico (Coahuila).
Perdita trifasciata Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 969. ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 366, 370, figs. 45, 46, 138. ♀.
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 11, 14 (key).




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Perdita vesca Timberlake
Nevada (Wadsworth). Pollen: Possibly an oligolege of Coldenia, visits
flowers of C. nuttallii.
Perdita vesca Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 11, figs. 1199, 1200, 1303. ♀, ♂.

Perdita wasbaueri Timberlake
South. Calif. (Colorado Desert). Pollen: Apparently an oligolege of
C oldcttid j)lic(xtci.
Perdita wasbaueri Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 126, figs. 656, 657, 730. ♀.
6.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 11 (key).

Genus PERDITA Subgenus HEXAPERDITA Timberlake

Perdita subg. Hexaperdita Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 416.
Type-species: Perdita ignota Cockerell. Orig. desig.

These bees are exclusively oligoleges of the family Compositae.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 348 (key). —Timberlake, 1956. Calif.
Univ. Pubs. Ent. 11: 247-252 (key to included spp.).

Perdita agasta Timberlake
Tex. (Hidalgo Co.).

Perdita agasta Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 389. ♀.

Perdita alexi Timberlake
Tex. (College Station), Kans. (Barber County). Pollen: Unknown, but visits
flowers of Helianthus petiolaris, Heterotheca subaxillaris.
Perdita alexi Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 26. ♀.

Perdita asteris Cockerell
West. Tex., N. Mex., Ariz. Pollen: Unknown, but visits flowers of

Aplopappus, Aster canescens, Chrysopsis, Heterotheca subaxillaris, Nolina microcarpa,
Verbesina encelioides.

Perdita asteris Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 93. ♀.

Perdita vespertilio Cockerell, 1896. Acad. Nat. Sci. PhDa., Proc. 48: 95. ♂.

Perdita crassiceps Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 97. ♂.

Perdita vagans Cockerell, 1896. Acad. Nat. Sci. PhDa., Proc. 48: 101. ♂.

Perdita mellina Cockerell, 1900. Entomologist 33: 62. ♂.

Taxonomy: Cockerell, 1906. Entomologist 39: 148. 9 (as vespertilio). —Timberlake, 1956.
Calif. Univ. Pubs. Ent. 11: 249, 258 (key, tax. characters, geogr. and floral records).
—Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 389-390 (tax. characters, geogr. and floral
records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 27 (synonymy, geogr. and floral
records).

Perdita bebbiae Timberlake
Calif. (San Diego County). Pollen: Unknown, but visits flowers of Bebbia
juncea.
Perdita bebbiae Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 255, figs. 179, 180, 292. ♀, ♂.

Perdita bishoppi
***authority mismatch
bishoppi Cockerell. N. C. to Fla., west to Tex. Pollen: Unknown, but visits flowers of
Heterotheca latifolia, H. subaxillaris, Isopappus divaricatus.
Perdita bishoppi Cockerell, 1906. Entomologist 39: 148. ♀, ♂.

Taxonomy: Cockerell, 1922. Amer. Mus. Novitates 33: 7. —Timberlake, 1956. Calif. Univ.
Pubs. Ent. 11: 249, 250, 261, figs. 193, 194, 299 (key, tax. characters, geogr. and floral
records). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 303-304, figs. 72-74, 79,
table 9 (redescription).

Perdita bishoppi
***authority mismatch
planorum Timberlake. Kans., Tex. Pollen: Unknown, but visits flowers of
Heterotheca latifolia, H. subaxillaris, Isopappus divaricatus.
Perdita bishoppi planorum Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 261. ♀, ♂.

Perdita blatchleyi Timberlake
Fla. (Dunedin). Pollen: Unknown, but visits flowers of Chrysopsis.
Perdita blatchleyi Timberlake, 1952. Ent. Soc. Wash., Proc. 54: 201. ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 248, 258 (key). —Mitchell, 1960. N.
C. Agr. Expt. Sta. Tech. Bui. 141: 304, fig. 72, table 9 (redescription).

Perdita boltoniae
***authority mismatch
boltoniae (Robertson). 111., La., Miss. Pollen: Unknown, but visits flowers of
Boltonia.
Perditella boltoniae Robertson, 1902. Canad. Ent. 34: 321. ♀, ♂.




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Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 249, 252, 265 (key, tax. characters,
geogr. records). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 304-305, fig. 73,
table 9 (redescription).

Perdita boltoniae
***authority mismatch
chrysopsina Timberlake. N. J. to Fla. Pollen: Unknown, but visits flowers of
Chrysopsis mariana, Coreopsis, Erigeron, Gerardia, Haplopappus.
Perdita chrysopsina Timberlake, 1928. Hawaii. Ent. Soc, Proc. 7: 155. ♂, ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 250, 252, 265, figs. 199, 200, 302 (key,
tax. characters). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 305-306, figs. 72, 74,
table 9 (redescription).

Perdita callicerata
***authority mismatch
Cockerel). Tex. to south. Calif.; Mexico (Chihuahua, Durango). Pollen: Unknown,

most frequently has been collected at the flowers of Baileya multiradiata, but also

visits flowers oi Baileya pleniradiata, Baccharis, Dithyrea wislizenii, Haplopappus.
Perdita callicerata Cockerell, 1896. N. Y. Ent. Soc, Jour. 4: 206. Nomen nudum, validated

by Cockerell, 1897. Amer. Ent. Soc, Trans. 24: 153. male.
Perdita callicerata var. chrysoceras Cockerell, 1897. N. Mex. Agr. Expt. Sta., Bui. 24: 43.

6.
Perdita callicerata var. lencura Cockerell, 1909. Canad. Ent. 41: 152. ♀.
Perdita lutzi Cockerell, 1922. Amer. Mus. Novitates 33: 7. ♀.

Taxonomy: Cockerell, 1897. Acad. Nat. Sci. Phila., Proc. 49: 354. 9. -Timberlake, 1956. Calif.
Univ. Pubs. Ent. 11: 247, 248, 250, 255, figs. 181, 182, 293 (key, tax. characters, geogr. and
floral records).

Perdita cambarella
***authority mismatch
cambarella Cockerell. Tex. Pollen: Unknown, but visits flowers of Heterotheca
siibaxillaris, Isopappus divaricatus.
Perdita cambarella Cockerell, 1906. Entomologist 39: 150. ♀, ♂.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 248, 251, 256-257 (key, tax.
characters, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 29
(color variation, geogr. records).

Perdita cambarella
***authority mismatch
platyura Cockerell. Kans., Tex., N. Mex. Pollen: Unknown, but visits flowers of
Heterotheca latifolia, Isopappus divaricatus.
Perdita platyura Cockerell, 1922. Ann. and Mag. Nat. Hist. (9) 10: 547. ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 248, 251, 257, figs. 183, 184, 294. S
(key, tax. characters, geogr. and floral records).

Perdita cara Timberlake
Tex. (Reeves County). Pollen: Unknown, but visits flowers of Prosopis.
Perdita cara Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 388, figs. 466, 467, 518. ♂.

Perdita compacta Timberlake
Ariz. (Graham County). Pollen: Unknown, but visits flowers of Pedis
papposa.
Perdita compacta Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 9. ♀.

Perdita fedorensis Cockerell
Tex. (Lee County).

Perdita fedorensis Cockerell, 1916. Ann. and Mag. Nat. Hist. (8) 17: 280. ♀.

Taxonomy: Timberlake, 1956. Calif. -Univ. Pubs. Ent. 11: 249, 259 (key).

Perdita foveata
***authority mismatch
brachycephala Timberlake. Tex. Pollen: Unknown, but visits flowers of
Aphanostephus skirrhobasis, Aster tanacetifolius. Coreopsis basalis.
Perdita foveata brachycephala Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 266. ♀, ♂.

Perdita foveata
***authority mismatch
foveata Timberlake. Fla. (Dade and Pasco Counties), Tex. (Montgomery County), Ark.
(Sevier and Scott Counties), Kans. (Reno County). Pollen: Unknown, but visits flowers
of Coreopsis tinctoria, Hymenopappus.
Perdita foveata foveata Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 265, figs. 201, 202,
303. ♀, ♂.
Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 392 (geogr. and floral records).
—Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 307. figs. 72-74, 79, table 9
(redescription).

Perdita foveata
***authority mismatch
persimilis Timberlake. Tex. (Comanche County), N. Mex. (Chaves County).
Perdita foveata persimilis Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 267. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 29 (geogr. record).




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Perdita georgica Timberlake
N. C. to Fla., west to Miss. Pollen: Unknown, but visits flowers of Aster,
Clirysopsis mariana, C. microcephala, Haplopappiis, Heterotlieca, Isopappiis.
Perdita georgica Timberlake, 1928. Hawaii. Ent. Soc, Proc. 7: 158. ♂, ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 249, 251, 257, figs. 185, 186, 295 (key,
tax. characters, geogr. and floral records). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141: 307-308, figs. 72-74, table 9 (redescription).

Perdita graenicheri Timberlake
Fla. (Miami). Pollen: Unknown, but visits flowers of Clirysopsis
tracyi.
Perdita graenicheri Timberlake, 1947. Ent. Soc. Wash., Proc. 49: 82. ♀, ♂.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 249, 252, 264, figs. 197, 198, 301
(key). -Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 308-309, figs. 72, 73, table 9
(redescription).

Perdita heterothecae
***authority mismatch
heterothecae Cockerell. Ariz., Utah (San Juan County), south. Calif. (Colorado
Desert), Mexico (Baja California). Pollen: Unknown, but visits flowers of Aster
tephrodes, Grindelia, Gutierrezia califomica, G. sarothrae, Heliotropium curassavicum,
Heterotheca grandiflora, H. siibaxillaris, Tamarix.
Perdita heterothecae Cockerell, 1900. Entomologist 33: 62. ♀, ♂.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 248, 251, 260-261, figs. 191, 192, 298
(key, tax. characters, geogr. and floral records). —Timberlake, 1971. Calif. Univ. Pubs. Ent.
66: 10 (geogr. and floral records).

Perdita heterothecae
***authority mismatch
trizonata Timberlake. Ariz. (Willcox). Pollen: Unknown, but visits flowers of
Aster intricatus, Euphorbia.
Perdita heterothecae trizonata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 96. ♀, ♂.
ignota basalis Timberlake. Tex. (Hidalgo County); Mexico (Tamaulipas).

Perdita ignota basalis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 391. ♀, ♂.
ignota crawfordi Cockerell. Colo., Nebr., Kans., Okla., Mo. Pollen: Unknown, but visits flowers
of Chrysopsis, Grindelia, Heterotheca subaxillaris, Isopappus divaricatus, Prionopsis
ciliata, Sideranthus nibiginosus.
Perdita crawfordi Cockerell, 1901. Entomologist 34: 190. ♂, ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 262-263, figs. 195, 196, 300 (tax.
characters, geogr. and floral records).
ignota ignota Cockerell. Tex., Colo., Ariz.; Mexico (Chihuahua and Zacatecas). Pollen:

Unknown, but visits flowers of Baccharis, Chrysopsis nitteri, Erigeron, Heterotheca,
Sideranthus, Solidago.
Perdita ignota Cockerell, 1896. Ent. Monthly Mag. 32: 220. ♀.

Taxonomy: Cockerell, 1907. Colo. Univ., Studies 4: 247. 9, cJ. —Cockerell, 1922. Amer. Mus.
Novitates 33: 7 (tax. characters). —Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 250, 252,
262 (key, tax. characters, geogr. and floral records). —Timberlake, 1971. Calif. Univ. Pubs.
Ent. 66: 10 (geogr. record).
ignota isopappi Timberlake. East. Tex., La. Pollen: Unknown, but visits flowers of

Aphanostephus skirrhobasis. Aster tanacetifolius, Chrysanthemum, Chrysopsis

nuttallii, Grindelia, Helenium tenuifolium, Heterotheca latifolia, H. pilosa, H.

subaxillaris, Isopappus divaricatus, Prionopsis ciliata, Xanthisma texanum var.

dmmmondii.
Perdita isopappi Timberlake, 1928. Hawaii. Ent. Soc, Proc. 7: 152. ♂, ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 249, 252, 263-264 (key, tax.
characters, geogr. and floral records).

Perdita nubila Timberlake
Fla., Ark., Kans. Pollen: Unknown, but visits flowers of Erigeron ramosus.
Perdita nubila Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 392. ♀, ♂.

Taxonomy: Timberlkake, 1968. Calif Univ. Pubs. Ent. 49: 29 (geogr. records).

Perdita pratti Cockerell
Tex., Okla., Kans. Pollen: Unknown, but visits flowers of Helenium,
Helianthus cucumerifolius, H. petiolaris, Heterotheca latifolia, H. subaxillaris.
Perdita pratti Cockerell, 1906. Entomologist 39: 125. ♀.




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Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 248, 250, 252-253, figs. 173, 174, 289.
6 (key, tax. characters, geogr. and floral records).

Perdita xanthisma
***authority mismatch
Cockerel!. Wyo., Nebr., Kans., Tex., N. Mex., Ariz. Pollen: Unknown, but visits

flowers of Aplopapptis divaricatus, A. stibruginosus, Croton monanthogynus, Grindelia,
Gutierrezia, Helianthus annuus, Heterotkeca pilosa, H. subaxillaris, Isopappus
divaricatus, Prionopsis ciliata, Sideranthus, Xanthisma texanum.
Perdita xanthismae Cockerell, 1905. Ent. News 16: 331. ♀.
Perdita xanthismae sideranthi Cockerell, 1906. Entomologist 39: 178. ♀.
Perdita xanthisma Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 29. Emend.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 247, 249, 251, 258-259, figs. 187, 188,
296. 9 , 6 (key, tax. characters, synonymy, geogr. and floral records). —Timberlake, 1958.
Calif. Univ. Pubs. Ent. 14: 390 (geogr. and floral records). —Timberlake, 1968. Calif. Univ.
Pubs. Ent. 49: 29-30 (geogr. and floral records).

Genus PERDITA Subgenus MACROTERA Smith

Macrotera Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 130.
Type-species: Macrotera bicolor Smith. Monotypic.

The species of this subgenus are apparently oligoleges of Opuntia.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 346, 352-353 (tax. characters, key to
included spp.). —Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 375-378 (key to included
spp.).

Perdita crassa Timberlake
Tex. (Sutton and Gillespie Counties). Pollen: Unknown, but possibly an
oligolege of Opuntia.
Perdita crassa Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 376, figs. 456, 457, 512. ♀, ♂.

Perdita texana
***authority mismatch
ablusa Timberlake. N. Mex., Tex. Pollen: Collects pollen from flowers of Opuntia
phaeacantha var. major.
Perdita texana abhisa Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 376. ♀, ♂.

Biology: Barrows, Chabot, Michener and Snyder, 1976. Kans. Ent. Soc, Jour. 49: 275-279, 4
figs, (mating behavior, foraging, floral relationship, variation in head width, as texana).

Perdita texana
***authority mismatch
texana (Cresson). Tex. (Bastrop, Gonzales, Lee and possibly Travis Counties). Pollen:
Unknown, but visits flowers of Opuntia.
Macrotera texana Cresson, 1878. Amer. Ent. Soc, Trans. 7: 70. ♀.
Macrotera megacephala Cresson, 1878. Amer. Ent. Soc, Trans. 7: 71. ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 353, figs. 5, 6, 118 (tax. characters).
—Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 375, 376 (tax. characters, status,
synonymy).

Genus PERDITA Subgenus MACROTERELLA Timberlake

Perdita subg. Macroterella Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 360.
Type-species: Perdita mortuaria Timberlake. Orig. desig.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 346, 361 (key to included spp.).
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 7-8 (key to females of included spp.; key to
males of carinata, mortuaria, opacella).

Perdita carinata Timberlake
Calif. (Colorado Desert). Pollen: Unknown, but visits flowers of
Eriogonum, Eschscholzia minutiflora, Ferrocactus acanthodes.
Perdita carinata Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 9, figs. 1197, 1198, 1302. ♀.
6.
mellca Timberlake. Ariz., south. Calif. (Colorado Desert); Mexico (Baja California and Sonora).
Pollen: Apparently an oligolege of Euphorbia including E. polycarpa, E. p. var. hxHella.
Perdita mellea Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 364, figs. 33, 34, 132. ♂, ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 125 (geogr. and floral records).
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 7 (key).




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Perdita mortuaria Timberlake
South. Calif., Ariz.; deserts. Pollen: Unknown, but visits flowers of
Echinocereus engelmanni, Eriogomim inflatiim, Eschscholzia ealifomica, E.
minutiflora var. darwinensis, Eucnide sirens, Larrea tride7itata, Prosopis juliflora.
Perdita mortuaria Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 362, figs. 27, 28, 129. ♂, ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 324 (tax. characters, geogr. record,
variation). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 126 (geogr. and floral record,
variation). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 7, 8, 10 (key, geogr. and floral
records).
nigrrella Timberlake. South. Calif.; deserts. PoUen: Unknown, but visits flowers of Phacelia
including P. cremilata.
Perdita nigrella Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 362, figs. 29, 30, 130. cj, 9.

Taxonomy: Timberlake, 1968 Calif. Univ. Pubs. Ent. 49: 7 (key).

Perdita opacella Timberlake
Ariz. (Grand Canyon), Utah (Moab). Pollen: Unknown, but visits flowers
of Cleome lutea, Stanley a.
Perdita opacella Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 324. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 7, 8, figs. 3, 1195, 1196, 1301, 1353.
S.

Perdita solitaria Cockerell
N. Mex. (Organ Mts.), Ariz. (Santa Catalina Mts.).

Perdita solitaria Cockerell, 1896. N. Y. Ent. Soc, Jour. 4: 206. Nomen nudum.
Perdita solitaria Cockerell, 1897. Amer. Ent. Soc, Trans. 24: 152. ♀.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 361, 365 (key, geogr. records).
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 7 (key).

Perdita tristella Timberlake
South. Calif. (Riverside County). Pollen: Unknown, but visits flowers of
Calochortus splendens, Eriogonum fasciciilatum, Hugelia virgata, Sphaeralcea.
Perdita tristella Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 363, figs. 31, 32, 131. ♂, ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 7 (key).

Genus PERDITA Subgenus MACROTEROPSIS Ashmead

Macroteropsis Ashmead, 1899. Amer. Ent. Soc, Trans. 26: 85.

Type-species: Perdita latior Cockerell. Monotypic and orig. desig.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 346, 356-357 (redescription, key to
included spp.). —Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 87 (key to males of included
spp.).

Perdita arcuata
***authority mismatch
arcuata Fox. Calif. (San Benito County); Mexico (Baja California). Pollen: Apparently
an oligolege of Sphaeralcea.
Perdita arcuata Fox, 1893. Calif. Acad. Sci., Proc (2) 4: 18. ♂.

Taxonomy: Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 54 (tax. characters).
—Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 357, 358 (key, tax. characters, tax. status).
—Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 87 (key). —Timberlake, 1968. Calif. Univ.
Pubs. Ent. 49: 4 (geogr. and floral record).

Perdita arcuata
***authority mismatch
dinognatha Cockerell. Ariz. (Tempe), Nev. (Charleston Mts.), south. Calif.; deserts.
Pollen: Apparently collects pollen only from the flowers of Sphaeralcea ambigua, but
visits other flowers for nectar including Encelia frutescens, Eriogonum fasciculatum
var. polifolium.
Perdita dinognatha Cockerell, 1922. U. S. Natl. Mus., Proc 60 (18): 19. ♂.

Taxonomy: Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 968 (tax. status, geogr. and floral
records). —Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 357, 359, figs. 21, 22, 126. 9 (key,
geogr. and floral records).

Perdita echinocacti Timberlake
Ariz.; Mexico (Sinaloa and Sonora). Pollen: Unknown, but visits
flowers q{ Antigonon leptopus, Jacquemontia, Kallstroemia grandiflora.
Perdita echinocacti Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 360, figs. 25, 26, 128. tJ, 9.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 125 (geogr. and floral records).
—Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 87 (key).




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Perdita haplura Cockerell
Tex. (Terrell and Val Verde Counties), N. Mex. (Lincoln County). Pollen:
Unknown, but visits flowers of Sphaeralcea angustifolia var. cuspidata.
Perdita haplura Cockerell, 1922. U. S. Natl. Mus., Proc. 60 (18): 19. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 124 (type). -Timberlake, 1962.
Calif. Univ. Pubs. Ent. 28: 87-89, text fig. A. 6. -Timberlake, 1968. Calif. Univ. Pubs. Ent.
49: 4 (geogr. and floral records).

Perdita latior Cockerell
N. Mex., Ariz., Nev. (Charleston Mts.). Pollen: Apparently an oligolege of
Sphaeralcea including S. angustifolia, but also visits flowers of Sida hederacea
presumably for nectar.
Perdita latior Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 53. cJ, 9.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 357, 359, figs. 23, 24, 127.
—Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 87 (key).

Perdita magniceps Timberlake
N. Mex. (Luna County). Pollen: Unknown, but visits flowers of
Sphaeralcea.
Perdita magniceps Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 124, flgs. 654, 655, 729. ♂.

Perdita portalis Timberlake
N. Mex. (Deming), Ariz. (Cochise and Coconino Counties); Mexico
(Zacatecas). Pollen: Apparently an oligolege of Sphaeralcea.
Perdita poHalis Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 357, figs. 19, 20, 125. ♂, ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 323. 6 (variation). —Timberlake,
1960. Calif. Univ. Pubs. Ent. 17: 125 (geogr. and floral record). —Timberlake, 1962. Calif.
Univ. Pubs. Ent. 28: 87. 6 (key). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 4 (geogr.
record).

Perdita robertsi Timberlake
Tex. (Val Verde County). Pollen: Possibly collects pollen from the flowers
of Opuntia.
Perdita robertsi Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 6, fig. 2. ♀.

Genus PERDITA Subgenus PENTAPERDITA Cockerell and Porter

Perdita subg. Pentaperdita Cockerell and Porter, 1899. Ann. and Mag. Nat. Hist. (7) 4: 415.
Type-species: Perdita albovittata Cockerell. Monotypic.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 347, 404-406 (redescription, key to
included spp.). —Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 329 (modified key to
included spp.). —Timberlake, 1958. Calif. Uniy. Pubs. Ent. 384-385 (key to spp. allied to
melanochlora). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 22-24 (key to females of
included spp.).

Perdita albovittata Cockerell
N. Mex., Ariz., south. Calif. (Colorado and Mojave Deserts); Mexico
(Chihuahua, Coahuila, Durango). Pollen: Unknown, but visits flowers of Aplopappus
spinulosus, Baileya multiradiata, B. pleniradiata, Cevallia sinuata, Coldenia greggii,
Eriogonum injlatum, Gutierrezia califomica, G. liicida, Haplopappus acradenius, H.
gracilis, Pectis papposa, Physalis, Stephanomeria, Verbesina encelioides.
Perdita albovittata Cockerell, 1895. Acad. Nat. Sci. PhUa., Proc. 47: 15. ♀.
Perdita laticeps Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 98. ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 405, 406, 411, figs. 105, 106, 168 (key,
geogr. and floral records, possible synonymy). —Timberlake, 1968. Calif. Univ. Pubs. Ent.
49: 23, 24 (synonymy, geogr. and floral record).

Perdita amoena Timberlake
Ariz. (Cochise and Santa Cruz Counties), Utah (Washington County);
Mexico (Sonora).
Perdita amoena Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 330, figs. 287, 288, 336. ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 385, 386. 9 (key). -Timberlake,
1960. Calif. Univ. Pubs. Ent. 17: 132 (geogr. record). —Timberlake, 1968. Calif. Univ. Pubs.
Ent. 49: 23, 24 (key, geographic records).

Perdita annexa Timberlake
N. Mex. (Pecos and Santa Fe). Pollen: Unknown, but visits flowers of
Hymenoxys richardsoni.
Perdita annexa Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 133. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 23, 24. 9 (key).




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Perdita bradleyana Timberlake
Tex. (Eagle Pass and El Paso); Mexico (Zacatecas).

Perdita bradleyana Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 409, figs. 99, 100, 165. ♂.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 329. 9. —Timberlake, 1958. Calif.

Univ. Pubs. Ent. 14: 385 (key). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 23, 25 (key,

geogr. record). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 7 (geogr. record).

Perdita chrysophila
***authority mismatch
chrysophila Cockerell. Tex. (Culberson County). N. Mex. (Hollywood, Organ and
Pecos), Ariz. (Safford); Mexico (Hidalgo, San Luis Potosi and Zacatecas). Pollen:
Unknown, but visits flowers of Flourensia, Hymenoxys, Picradenia florihunda,
Verbesina evcelioides, Zaluzania globosa. Another subspecies, Perdita chrysophila
quadricincta Timberlake, occurs in Mexico (Nuevo Leon).

Perdita chrysophila Cockerell, 1896. N. Y. Ent. Soc, Jour. 4: 206. ♂.

Perdita nigrifacies Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 406 9.

Taxonomy: Cockerell, 1904. Entomologist 37: 6. 6 . —Timberlake, 1954. Calif. Univ. Pubs.
Ent. 9: 406, figs. 93, 94, 162 (key, type). —Timberlake, 1958. Calif. Univ. Pubs. Ent. 14:
386-387 (tax. characters, as nigrifacies). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17:
132-133, figs. 660, 661, 732. i (redescription). —Timberlake, 1962. Calif. Univ. Pubs. Ent.
28: 95-96 (tax. characters, as nigrifacies). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49:
23, 25 (key, synonymy, geogr. and floral records).

Perdita coahuilensis Timberlake
Tex. (El Paso); Mexico (Coahuila).

Perdita coahuilensis Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 409, figs. 101, 102, 166. ♀.


Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 329, 330 (key, geogr. record).
—Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 385 (key). —Timberlake, 1968. Calif. Univ.
Pubs. Ent. 49: 24 (key).

Perdita idahoensis Timberlake
Idaho (Cassia County), Calif. (Inyo County). Pollen: Unknown, but
visits flowers oi Chrysothaninus, Gutierrezia microcephala.
Perdita idahoensis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 385, figs. 464, 465, 517. ♂.
9.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 23 (key). —Timberlake, 1971. Calif.
Univ. Pubs. Ent. 66: 9 (geogr. and floral record).

Perdita mandibularis Timberlake
South. Calif. (Colorado and Mojave Deserts). Pollen: Unknown, but
visits flowers of Bebbia juncea, Chaenactis carphoclinia, Dalea, Geraea canescens.
Perdita mandibularis Timberlake, 1954. calif. Univ. Pubs. Ent. 9: 411, figs. 107, 108, 169.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 23 (key).

Perdita melanochlora Cockerell
Ariz. (Santa Rosa Valley and Tucson), south. Calif. (Colorado and

Mojave Deserts), Nev. (Arden and Charleston Mts.). Pollen: Unknown, but visits flowers
of Baileya multiradiata, Bebbia juncea, Chrysothamnus, Encelia farinosa, Sphaeralcea,
Viguiera deltoides, V. nevadensis.
Perdita melanochlora Cockerell, 1922. Amer. Mus. Novitates 33: 7. ♀.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 405, 406, 408, figs. 97, 98, 164. ♀, ♂.
—Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 329 (key). —Timberlake, 1958. Calif. Univ.
Pubs. Ent. 14: 385 (key). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 23 (key).
—Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 9 (geogr. records).

Perdita nigroviridis Timberlake
Tex. Pollen: Unknown, but visits flowers of Gail lardia pulchella,
Helenium, Helianthus annuus.
Perdita nigroviridis Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 407, figs. 95, 96, 163. ♀, ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 386 (geogr. and floral records).
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 23, 26 (key, geogr. and floral record).

Genus PERDITA Subgenus PERDITA Smith

Perdita Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 128.

Type-species: Perdita halictoides Smith. Monotypic.
Neoperdita Ashmead, 1899. Amer. Ent. Soc, Trans. 26: 85.




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Type-species: Perdita zebrata Cresson. Monotypic and orig. desig.
Perdita subg. Geoperdita Cockerell and Porter, 1899. Ann. and Mag. Nat. Hist. (7) 4: 415.

Type-species: Perdita chamaesarachae Cockerell. Monotypic.
Perdita subg. Tetraperdita Cockerell and Porter, 1899. Ann. and Mag. Nat. Hist. (7) 4: 415.

Type-species: Perdita sexviaculata Cockerell. Monotypic and orig. desig.
Zaperdita Robertson, 1918. Ent. News 29: 91.

Type-species: Perdita jnaura Cockerell. Monotypic and orig. desig.

This is the largest subgenus of Perdita. Six groups of species are currently recognized and,
although some of these groups have been divided into subgroups in the existing classification, no
attempt has been made to reflect these subsidiary units in this catalog. The six recognized spe-
cies groups are arranged in accordance with the current classification of this subgenus.

Revision: Timberlake, 1958. Cahf. Univ. Pubs. Ent. 14: 303-365 (zonalis group).
— Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 365-374 (halictoides group). —Timberlake,
1960. Calif. Univ. Pubs. Ent. 17: 1-121 (.octomaculata group). —Timberlake, 1960. Calif.
Univ. Pubs. Ent. 17: 135 {halictoides group). —Timberlake, 1962. Calif. Univ. Pubs. Ent.
28: 1-86 (veiitralis group). —Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 1, 97-101 (zonalis
group). —Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 1, 101-102 (halictoides group).
—Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 1, 102-106 (octomaculata group).
—Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 106-107 {valida group). —Timberlake, 1964.
Calif. Univ. Pubs. Ent. 28: 1, 125-362 (sphaeralceae group). —Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 40-57 (zonalis group). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49:
57-62 (halictoides group). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 62-95
(octomaculata group). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 95-112 (ventralis
group). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 112-161 (sphaeralceae group).
—Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 11-56 (suppl. studies on the species of the
subgenus Perdita).

Species Group Zonalis

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 303-315 (tax. characters, key to
included spp.). —Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 1, 97-101 (tax. characters,
modified key to included spp.). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 40-43 (suppl.
key to included spp.). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 11-13 (suppl. key to
included spp.).

Perdita accepta Timberlake
Oreg. (Crook County).

Perdita accepta Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 340. ♀.

Perdita adjuncta Timberlake
Nev. (Austin), Calif. (Modoc County), Oreg. (Lake County), Idaho
(Bingham, Cassia and Elmore Counties). Pollen: Unknown, but visits flowers of
Chrysotham7ius.
Perdita adjuncta Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 327, figs. 390, 391, 479. ♀.
6.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 97 (key).

Perdita aemula
***authority mismatch
aemula Timberlake. Oreg., Utah. Pollen: Unknown, but visits flowers of
Chrysothaynnus, Solidago.
Perdita aemula Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 330, figs. 396, 397, 482. ♂, ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 98 (key). —Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 42 (key).

Perdita aemula
***authority mismatch
quadrifasciata Timberlake. Calif. (Mono County). Pollen: Unknown, but visits flowers
of Chrysothavinus.
Perdita aemula quadrifasciata Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 43. ♀, ♂.

Perdita affecta Timberlake
Ariz. (Tuba City). Pollen: Unknown, but visits flowers of Artemisia
filifolia.
Perdita affecta Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 14. ♀.

Perdita albopicta Timberlake
Utah (Millard County). Pollen: Unknown, but visits flowers of
Chrysothamnus.
Perdita albopicta Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 341. ♀.




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Perdita ampla Timberlake
N. Mex. (McKinley and San Juan Counties). Pollen: Unknown, but visits
flowers of Ckrysofhamniis nauseosus, Senecio.
Perdita ampla Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 14. ♀.

Perdita baccharidis Cockerell
Ariz. (Maricopa and Pima Counties), Calif. (Riverside and Imperial
Counties). Pollen: Apparently an oligolege of Baccharis including B. emoryi, B.
sarothroides.
Perdita baccharidis Cockerell, 1900. Entomologist 33: 61. ♀.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 309, 313, 352-353, figs. 428, 429, 498.
9, 6 (redescription). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 44 (geogr. and floral
records).

Perdita basinicola Timberlake
Nev. (Churchill County), Calif. (Inyo County).

Perdita basinicola Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 44, figs. 1213, 1214, 1310.
cJ, 9. \

Perdita chrysothamni Timberlake
Calif. (Mojave Desert). Pollen: Apparently an oligolege of
Chrysothamnus nauseosus.
Perdita chrysothamni Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 319, figs. 374, 375, 473.
9,c5.

Perdita ciliata Timberlake
North. Calif., Oreg., Idaho. Pollen: Unknown, but visits flowers of Aster,
Chrysothamnus nauseosus, Eriogonum, Gutierrezia, Haplopappus, Heterotheca
grandiflora, Lessingia glandulifera, Melilotus alba, Senecio douglasii, Solidago
occidentalis. Predator: Philanthus pacificus arizonae Dunning.
Perdita interserta ciliata Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 334, figs. 404, 405,
486. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 43, 45 (key, tax. status, geogr.
records, variation).

Perdita confinis Timberlake
Calif. (Mono County).

Perdita confinis Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 15. ♀.

Perdita crassihirta Timberlake
Wash. (Chelan County).

Perdita crassihirta Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 46. ♀.

Perdita craterognatha Timberlake
Calif. (Mono County).

Perdita craterognatha Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 46, figs. 1215, 1216,
1311. ♀, ♂.

Perdita depressa Timberlake
Utah (Monument Valley), Ariz. (Havasu Canyon). Pollen: Unknown, but
visits flowers of Cleome jonesii.
Perdita depressa Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 48, figs. 7, 1217, 1218, 1312.
9, 6.

Perdita dicksoni Timberlake
Calif. (Inyo County). Pollen: Unknown, but visits flowers of
Chrysothamnus albidus.
Perdita dicksoni Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 328, figs. 392, 393, 480. ♀, ♂.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 97 (key).

Perdita dubia
***authority mismatch
dubia Cockerell. Colo. (Garfield and Pitkin Counties), Ariz. (Yavapai County). Pollen:
Unknown, but visits flowers of Gutierrezia microcephala, G. sarothrae.
Perdita dubia Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 75. ♂.
Perdita fratema Timberlake, 1929. N. Y. Ent. Soc, Jour. 39: 123. ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 306, 312, 329, figs. 394, 395, 481. ♀.
6 (redescription, key, as fratema). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 43, 50
(key, tax. status). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 16-17 (tax. status,
synonymy).

Perdita dubia
***authority mismatch
parilis Timberlake. Ariz. (Coconino and Mohave Counties), Utah, Nev. (Clark County),
Idaho (Cassia and Franklin Counties), Wyo. (Franklin County). Pollen: Unknown, but
visits flowers oi Aplopappus, Chrysothamnus, Eriogonum, Gutierrezia.
Perdita parilis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 342, figs. 414, 415, 491. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 42, 43, 52 (key, geogr. and floral
records). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 17-18 (tax. status)




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Perdita ericameriae Timberlake
South. Calif. (Los Angeles and Riverside Counties). Pollen: Unknown,
but visits flowers of Aplopappus palmeri var. pachylepis, Gutierrezia califoniica,
Lepidospartum squamatum.
Perdita ericameriae Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 317, figs. 372, 373, 472.
9, 6.

Perdita festiva Timberlake
Utah (Emery County).

Perdita festiva Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 321. ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 98 (key).

Perdita foleyi Timberlake
Calif. (Kern and Tulare Counties). Pollen: Unknown, but visits flowers of
Chrysotharnnus nauseosus.
Perdita foleyi Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 347, figs. 418, 419, 493. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 50 (geogr. and floral record).

Perdita haigi Timberlake
Nev. (Churchill County).

Perdita haigi Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 100, figs. 833, 834, 895. ♀, ♂.

Perdita impunctifrons Timberlake
Calif. (Inyo and San Bernardino Counties).

Perdita impunctifrons Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 355. ♀.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 18 (geogr. record).

Perdita interserta Cockerell
South. Calif, (cismontane and Mojave Desert). Pollen: Unknown, but
visits flowers of Aplopappiis palmeri var. pachylepis, A. veneta var. venionioides,
Baccharis emoryi, Chrysotharnnus nauseosus, Eschscholzia californica, Lepidospartum
squamatum, Polygonum lopathifolium, Rhus diversiloba, Satureja occidentalis,
Solidago occidentalis.
Perdita interserta Cockerell, 1922. U. S. Natl. Mus., Proc. 60 (18): 20. ♀.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 307, 311, 312, 333-334, figs. 402, 403,
485. 6 , 9 (key, tax. characters, tax. status, geogr. and floral records). —Timberlake, 1968.
Calif. Univ. Pubs. Ent. 49: 45 (tax. status).

Perdita irregularis Timberlake
N. Mex. (McKinley County). Pollen: Unknown, but visits flowers of
Chrysothamnus nauseosus.
Perdita irregularis Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 18, figs. 1367, 1368, 1402.
9, 6.

Perdita isocomae Timberlake
Calif. (Riverside). Pollen: Unknown, but visits flowers of Aplopappus
venetus var. venionioides, Baccharis emoryi, Polygonum lopanthifolium , Solidago
occidentalis.
Perdita isocomae Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 335, figs. 408, 409, 488. ♂.
9.

Perdita lepidosparti
***authority mismatch
lepidosparti Timberlake. Calif. (Mojave Desert and Lassen County), Nev.

(Washoe County), Idaho (Bingham, Cassia, Fremont and Owyhee Counties). Pollen:
Unknown, but visits flowers of Chrysotharnnus, Eriogonum heermannii, Gutierrezia
lucida, Lepidospartum squamatum, Tetradymia comosa.
Perdita lepidosparti lepidosparti Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 325, figs.
386,387,478. ♂, ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 98 (key). -Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 50 (geogr. and floral records).

Perdita lepidosparti
***authority mismatch
novella Timberlake. South. Calif. (San Bernardino and San Gabriel Mts.).

Perdita lepidosparti novella Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 326, figs. 388,
389. ♀, ♂.
Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 98 (key).

Perdita lompocensis Timberlake
Calif. (Lompoc).

Perdita lompocensis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 337, figs. 410, 411, 489.

6, 9.

Perdita melanderi Timberlake
Calif. (Morro Bay).

Perdita melanderi Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 50, fig. 8. ♀.

Perdita munda Timberlake
Utah, Ariz. (Apache County).

Perdita munda Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 340. ♀.




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Perdita nigrocincta Timberlake
Calif. (Mono County).

Perdita nigrocincta Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 351. ♀.

Perdita obispoensis Timberlake
Calif. (San Luis Obispo County).

Perdita obispoensis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 335, figs. 406, 407, 487. ♂.

Perdita oregonensis
***authority mismatch
expleta Timberlake. Calif. (Sierra Nevada Mts.). Pollen: Unknown, but visits
flowers of Clirysothamyius.

Perdita oregoyiensis expleta Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 99. ♀.

Perdita oregonensis
***authority mismatch
oregonensis Timberlake. Wash, and Idaho, south to Calif, and Nev. Pollen:

Unknown, but visits flowers of Chrysotliamnns viscidiflorus, Gntierrezia.

Perdita oregonensis Timberlake, 1929. Pan-Pacific Ent. 6: 52. ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 306, 311, 320-321, figs. 378, 379, 475.
9, S (key, redescription, geogr. and floral records). —Timberlake, 1962. Calif. Univ. Pubs.
Ent. 28: 98 (key, tax. status). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 41, 52 (geogr.
and floral records).

Perdita pallidiventris Timberlake
Calif. (Fresno County). Pollen: Unknown, but visits flowers of
Eriogonion fasciculatum.
Perdita pallidiventris Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 99. ♀.

Perdita placida Timberlake
Calif. (Inyo County).

Perdita placida Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 339. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 42 (key).

Perdita polita Timberlake
Calif. (Kings, Tulare and Stanislaus Counties). Pollen: Unknown, but visits
flowers of He » I izonia pungens.
Perdita polita Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 332, figs. 400, 401, 484. ♀, ♂.

Perdita primula Timberlake
Calif. (Indio). Pollen: Unknown, but visits flowers of Melilotus alba.

Perdita primula Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 347, figs. 420, 421, 494. ♂.

Perdita proxima Timberlake
Calif. (Mohave Desert). Pollen: Unknown, but visits flowers of

Chrysotliavinus nauseosiis, C. n. consiniilis, Cleomella obtusifolia, Haplopappus
acradeniiis, Solidago occidentalis.
Perdita proxima Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 322, figs. 380, 381, 476. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 52 (geogr. and floral records).

Perdita punctifrons Timberlake
Calif. (Los Angeles and San Bernardino Counties). Pollen: Unknown,
but visits flowers of Lessingia gennanoruui var. glandulifera.
Perdita punctifrons Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 350, figs. 426, 427, 497. ♀.
6.

Perdita repens Timberlake
Calif. (Mt. Hamilton). Pollen: Unknown, but visits flowers of Haplopappus
arborescens.
Perdita repens Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 19, figs. 1369, 1370, 1403. ♂.

Perdita rivalis Timberlake
Calif. (San Bernardino, San Gabriel and Sierra Nevada Mts.), Nev.

(Daggett Pass). Pollen: Unknown, but visits flowers of Aster adscendens, A. canescens,
Erigeron divergens, E. foliolosus var. stenophyllus, Gayopkytum heterozygos, G.
nuttallii.
Perdita rivalis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 349, figs. 424, 425, 496. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 43, 52 (key, geogr. and floral
records).

Perdita scotti Timberlake
Calif. (Inyo, Los Angeles and Kern Counties). Pollen: Unknown, but visits
flowers of Chrysothamnus nauseosus.
Perdita scotti Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 338, figs. 412, 413, 490. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 53 (geogr. and floral records).

Perdita similis
***authority mismatch
pascoensis Timberlake. Wash. (Pasco).

Perdita similis pascoensis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 324, figs. 384, 385.
6, 9.

Perdita similis
***authority mismatch
similis Timberlake. Oreg., Idaho, and Wyo., south to N. Mex., Ariz, and south. Calif.
(Inyo, Riverside and Imperial Counties). Pollen: Unknown, but visits flowers of




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Chrysothamnus nauseosus, C. viscidiflorus, Cleome semilata, Erigeron, Gutierrezia
sarothrae, Haplopappus heterophylhis, Prosopis juliflora.

Perdita similis similis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 323 fies 382 383 477
9,6. 8 , , •

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 22 (collection site). —Timberlake,
1968. Calif. Univ. Pubs. Ent. 49: 41, 53 (key, geogr. records, floral preferences).
ocia Timberlake. Calif. (Inyo County).

Perdita socia Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 42, 43, 53-54, plate figs. 1219
1220, 1313. 9,6. . -F s .

itottleri Cockerel]. Sask., south to N. Mex. west to Idaho, Nev. and Calif. (Mono County).

Pollen: Unknown, but visits flowers of Aster, Chrysothamnus graveolens, C. g. var.

glabrata, C. 7iatiseosiis, Gutierrezia sarothrae, Solidago missouriensis.
Perdita stottleri Cockerell, 1896. N. Y. Ent. Soc, Jour. 4: 205. ♀.
Perdita stottleri Jlavida Swenk and Cockerell, 1907. Ent. News 18: 58. ♀, ♂.

Taxonomy: Cockerell, 1904. Entomologist 37: 6. 6. —Timberlake, 1958. Calif. Univ. Pubs.
Ent. 14: 304, 310, 315 (tax. characters, key, geogr. and floral records). —Timberlake, 1958.
Calif. Univ. Pubs. Ent. 14: 304, 306, 310, 315-316, figs. 368, 369, 470 (tax. characters, key,
geogr. and floral records, as stottleri Jlavida). —Timberlake, 1962. Calif. Univ. Pubs. Ent.
28: 98 (key, as stottleri Jlavida). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 54
(synonymy, geogr. and floral records).

Perdita subvestita Timberlake
N. Mex. (Citron County), Ariz. (Coconino County), Utah (Kane County).

Pollen: Unknown, but visits flowers of Aplopappus gracilis, Chrysothamnus

viscidijlonis, Gutierrezia microcephala.
Perdita subvestita Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 55, figs. 1221, 1222, 1314.

9,6.

Perdita swezeyi Timberlake
Calif., Nev. (Ormsby County). Pollen: Unknown, but visits flowers
Chrysothamnus, Erigeron ramosus.
Perdita swezeyi Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 348, figs. 422, 423, 495. ♀, ♂.

Perdita taeniata Timberlake
N. Mex. (McKinley and San Juan Counties). Pollen: Unknown, but visits
flowers of Chrysothamnus 7iauseos2is, Senecio longi.
Perdita taeniata Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 20, figs. 1371, 1372, 1404. ♀.
6.

Perdita toschiae Timberlake
Nev. (Douglas County).

Perdita toschiae Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 56. ♀.

Perdita townsendi Cockerell
N. Mex. (Otero and Rio Arriba Counties), Ariz. (Apache County). Pollen:
Unknown, but visits flowers of Aster, Chrysothamnus, Haplopappus heterophylhis.
Perdita townsendi Cockerell, 1896. N. Y. Ent. Soc, Jour. 4: 204. ♀, ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 305, 310, 320, figs. 376, 377, 474 (tax.
characters, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 56
(geogr. and floral records).

Perdita vestita Timberlake
Utah (Grand and Kane Counties), Ariz. (Coconino County). Pollen:
Unknown, but visits flowers of Chrysothamnus, Gutierrezia.
Perdita vestita Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 316, figs. 370, 371, 471. ♀, ♂.

Taxonomy: Timberiake, 1968. Calif. Univ. Pubs. Ent. 49: 41, 42, 57 (key, geogr. and floral
records).

Perdita zonalis
***authority mismatch
aequalis Timberlake. Oreg., northern Calif., ?Nev., ?Mont. Pollen: Unknown, but visits
flowers of Chrysothamnus including C. nauseosus speciosus, Solidago.
Perdita zonalis aequalis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 344. ♀, ♂.

Perdita zonalis
***authority mismatch
bernardina Timberlake. Calif. (Bear Valley and Santa Ana Canyon in San Bernardino
Mts.). Pollen: Unknown, but visits flowers of Aster canescens, Chrysothamnus
nauseosus, C. viridulus, C. viscidiflorus, Eriogonum molestum var. davidsonii, E.
wrightii var. subscaposum, Gutierrezia califomica, G. lucida, Solidago califomica.
Perdita zonalis bernardina Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 345, figs. 416, 417,
492. ♀, ♂.




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Perdita zonalis
***authority mismatch
monticola Timberlake. Calif. (Sierra, Shasta, Nevada, Plumas and Placer Counties).
Pollen: Unknown, but visits flowers of Chrysothamnus, Solidago.
Perdita zonalis monticola Timberlake, 1958 Calif. Univ. Pubs. Ent. 14: 345. ♀, ♂.

Perdita zonalis
***authority mismatch
pallescens Timberlake. Calif. Pollen: Unknown, but visits flowers of Gutierrezia
iiicida, Haplopappus acradenius.
Perdita zonalis pallescens Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 344. ♀, ♂.

Perdita zonalis
***authority mismatch
zonalis Cresson. Calif. (Riverside and Kern Counties). Pollen: Unknown, but visits
flowers of Eriogonum, Gutierrezia califomica, Lepidospartum squamatum.
Perdita zonalis Cresson, 1879. Amer. Ent. Soc. Trans. 7: 202. ♀.

Taxonomy: Timberlake, 1929. Pan-Pacific Ent. 6: 49. 9, cJ. —Timberlake, 1958. Calif. Univ.
Pubs. Ent. 14: 308, 313, 343-344 (key, redescription, geogr. and floral records).
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 41, 43 (key).

Species Group Halictoides

The species of this group collect pollen from flowers of the solanaceous genera Physalis and
Chamaesaracha, but visit these and other flowers for nectar.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 365-367 (tax. characters, key to
included spp.). —Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 1 (key). —Timberlake, 1968.
Calif. Univ. Pubs. Ent. 49: 57-59 (key to included spp.).

Perdita beameri Timberlake
Tex. Pollen: Collects pollen from the flowers of Chamaesaracha including
C. conioides, but visits other flowers for nectar including Astragalus, Gilia acerosa,
Physalis lobata.
Perdita beameri Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 276, figs. 56, 1041, 1042,
1164. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 57, 59 (key, geogr. and floral
records).

Perdita chamaesarachae Cockerell
N. Mex. (Albuquerque, High Rolls, Las Vegas, Roswell and Santa
Fe), Ariz. (Cochise and Yavapai Counties); Mexico (Durango). Pollen: Apparently
collects pollen from the flowers of Chamaesaracha coronopus, but visits other flowers
including Physalis.
Perdita chamaesarachae Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 65. ♂, ♀.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 367, 371, figs. 450, 451, 509 (key,
redescription, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 58,
59 (key).

Perdita fidissima Timberlake
Tex. (Kenedy County). Pollen: Unknown, but visits flowers of

Parkinsonia.
Perdita fidissima Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 59. ♀.

Perdita halictoides Smith
Wis., Mich, and Ind., west to N. Dak., Colo., and Tex.; Fla. Pollen: Unknown,

but visits flowers of Euphorbia nuttallii, Gaillardia, Monarda punctata var.

occidentalis, Physalis heterophylla, P. lanceolata, Solidago.
Perdita halictoides Smith, 1853. Cat. Hym. Brit. Mus., v. 1, p. 128. ♀.
Perdita sexmaculata var. punctata Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 71. ♀.
Perdita maura Cockerell, 1901. Entomologist 34: 191. cJ, 9.
Perdita bisignata Cockerell, 1922. Amer. Mus. Novitates 33: 11. ♀.

Taxonomy: Cockerell, 1904. Canad. Ent. 36: 303. —Timberlake, 1928. Hawaii. Ent. Soc, Proc.
7: 155. —Cockerell, 1922. Ann. and Mag. Nat. Hist. (9) 10: 268 (tax. characters, as maura).
—Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 366, 367, 370-371, figs. 448, 449, 508
(redescription, key, geogr. and floral records, as maura). —Timberlake, 1958. Calif. Univ.
Pubs. Ent. 14: 366, 371. 9 (key, redescription). -Timberlake, 1960. Calif. Univ. Pubs. Ent.
17: 135 (synonymy, geogr. records). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141:
318-319, figs. 76-79, table 9 (redescription, synonymy). —Timberlake, 1968. Calif. Univ.
Pubs. Ent. 49: 57, 58 (key).




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Perdita lenis Timberlake
Tex., N. Mex., Ariz., Calif. (Indio). Pollen: Unknown, but visits flowers of
Baileya viultiradiata, Chamaesaracha conioides, Physalis heterophylla, Verbesina
enceUoides.
Perdita leiiis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 372, figs. 452, 453, 510. ♀, ♂.

Taxonomy: Rozen, 1966. Amer. Mas. Novitates 2259: 19, figs. 47-49 (larva). —Timberlake,
1968. Calif. Univ. Pubs. Ent. 49: 58, 59 (key). —Timberlake, 1971. Calif. Univ. Pubs. Ent.
66: 22 (geogr. and floral records).

Perdita levigata Timberlake
Colo. (Fremont County). Pollen: Unknown, but visits flowers of Physalis.
Perdita levigata Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 60, figs. 1225, 1226, 1316. ♂.

Perdita missionis Timberlake
Tex.; Mexico (Tamaulipas, San Luis Potosi). Pollen: Unknown, but visits
flowers of Chamaesaracha conioides, Lesquerella, Physalis lobata.
Perdita ynissionis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 369, figs. 446, 447, 507. ♂, ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 58 (key).

Perdita physalidis Timberlake
Calif. (Riverside County), Ariz. (Continental). Pollen: Unknown, but
visits flowers of Physalis, including P. crassifolia.
Perdita physalidis Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 374, figs. 454, 455, 511. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 58, 59, 61 (key, geogr. and floral
records).

Perdita rozeni Timberlake
Calif. (Colorado Desert); Mexico (Baja California and Sonora). Pollen:
Unknown, but visits flowers of Boerhaavia, Dalea mollis.
Perdita rozeni Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 61, figs. 1227, 1228, 1317. ♀, ♂.

Perdita sexmaculata
***authority mismatch
sexmaculata Cockerell. Ariz., N. Mex., Tex., Kans., Colo.; Mexico (Chihuahua,

Durango, Hidalgo). Pollen: Unknown, but visits flowers of Astragalus, Chamaesaracha
conioides, C. coronopus, Phacelia piperi, Physalis lobata, Prosopis, Scutellaria,
Sphaeralcea albicaulis, Thelesperma gracile. Another subspecies, Perdita sexmaculata
octonaria Timberlake, occurs in Baja California, Mexico.
Perdita sexmaculata Cockerell, 1895. Acad. Nat. Sci. Phila., Proc. 47: 12. ♀.
Perdita anograe Cockerell, 1902. Amer. Nat. 36: 812. ♂.

Taxonomy: Cockerell and Porter, 1899. Ann. and Mag. Nat. Hist. (7) 4: 416. ♀, ♂.

—Timberlake, 1929. N. Y. Ent. Soc, Jour. 37: 118. c^. —Timberlake, 1951. In Muesebeck,

Krombein and Townes, U. S. Dept. Agr., Agr. Monog. 2: 1095 (synonymy). —Timberlake,

1958. Calif. Univ. Pubs. Ent. 14: 366, 367-369, figs. 444, 445, 506 (key, redescription).

—Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 101 (tax. gtatus). —Rozen, 1966. Amer.

Mus. Novitates 2259: 20-21, fig. 51 (larva, as sexmaculataO. —Timberlake, 1968. Calif.

Univ. Pubs. Ent. 49: 58 (key).
Biology: Rozen, 1967. Amer. Mus. Novitates 2297: 38-40, figs. 16, 18, table 1 (nest

architecture, life history).

Species Group Octomaculata

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 1-22 (tax. characters, key to

included spp.). —Timberlake, 1968. CaHf. Univ. Pubs. Ent. 49: 62-67 (suppl. key to included
spp.). -Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 22-25 (suppl. key to included spp.).

Perdita abdominalis Timberlake
South. Calif. (Colorado Desert); Mexico (Baja California). Pollen:
Unknown, but visits flowers of Aplopapjms, Heliotropium curassavicum, Pectis

papposa.
Perdita abdominalis Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 38, figs. 550, 551, 667. ♀, ♂.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 26 (geogr. and floral records).

Perdita affinis Cresson
Colo., N. Mex., Ariz. Pollen: Unknown, but visits flowers of Aplopappus
gracilis, Grindelia inomata, G. squarrosa, Helianthus petiolaris, Solidago.
Perdita affiyiis Cresson, 1878. Amer. Ent. Soc, Trans. 7: 69. ♀, ♂.




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Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 11, 21, 108-109, figs. 636, 637, 720
(redescription, geogr. and floral records).

Perdita albipes Timberlake
Tex. (Big Bend National Park).

Perdita albipes Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 102, figs. 837, 838, 897. ♂.

Perdita apacheorum Timberlake
N. Mex. (Hidalgo County), Wyo. (Johnson County), Ariz. (Cochise
and Mohave Counties). Pollen: Unknown, but visits flowers of Asclepias, Baccharis,
Euphorbia, Grindelia squarrosa, Gutierrezia sarothrae, Heteroiheca.
Perdita apacheorum Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 99. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 67-68. 6 (geogr. and floral records).

Perdita aperta Timberlake
N. Mex. (Lincoln County), Ariz. (Navajo County). Pollen: Unknown, but
visits flowers of Gutierrezia microcephala, Lepidium montanum.
Perdita aperta Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 68, figs. 9, 1231, 1232, 1319. ♂.

Perdita aplopappi Timberlake
Ariz., Utah (Beaver and Washington Counties). Pollen: Unknown, but
visits flowers of Aplopappus gracilis, Eriogonum, Gutierrezia microcephala.
Perdita aplopappi Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 98, figs. 626, 627, 715. ♀.
6.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49; 69 (geogr. and floral record).
aridelia Timberlake. Utah, Ariz. (Navajo County). Pollen: Unknown, but visits flowers of
Gutierrezia sarothrae, Helianthus.
Perdita aridelia Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 81, figs. 600, 601, 702. ♀, ♂.

Perdita atriventris Timberlake
Tex. (Calvert). Pollen: Unknown, but visits flowers of Heterotheca
subaxillaris.
Perdita atriventris Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 111, figs. 640, 641, 722. ♂.

Perdita beatula Timberlake
Nev. (Churchill, Humboldt and Washoe Counties). Pollen: Unknown, but
visits flowers of Sphaeralcea.
Perdita beatula Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 22, figs. 522, 523, 662. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 62, 69-70. 9 (geogr. and floral
records).

Perdita bigeloviae Cockerell
Tex. (El Paso County), N. Mex., Colo. (Bent and Logan Counties). Pollen:
Unknown, but visits flowers of Aplopappus heterophyllus, A. pleuriflorus.
Perdita bigeloviae Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 87. ♂, ♀.
Perdita fasciata Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 57. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 7, 15, 71-72, figs. 588, 589, 696
(redescription). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 70 (synonymy, geogr. and
floral records).

Perdita biparticeps Cockerell
N. Mex. Pollen: Unknown, but visits flowers of Pedis papposa.
Perdita biparticeps Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 84. ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 121 (tax. characters). —Timberlake,
1968. Calif. Univ. Pubs. Ent. 49: 66, 70. 6 (type, redescription).

Perdita bridwelli Timberlake
Ariz. (Yuma County), south. Calif. (Colorado Desert). Pollen: Unknown,
but visits flowers of Sphaeralcea including S. emoryi, S. orcuttii.
Perdita bridwelli Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 34, figs. 542, 543, 673. ♀, ♂.

Perdita bruneri Cockerell
Tex. to Man. and Alta. east of Rocky Mts. Pollen: Unknown, but visits
flowers oi Aplopappus divaricatus, Boltonia asteroides, Cleome serrulata, Grindelia
squarrosa, Solidago altissima, S. rigida.
Perdita bruneri Cockerell, 1897. Ent. News 8: 23. (J ( 9 misdet.).
Perdita Cockerelli Crawford, 1906. Canad. Ent. 38: 282. ♂, ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 11, 21, 101-103, figs. 628, 629, 716
(redescription, geogr. and floral records).

Perdita butleri Timberlake
Ariz. (Gila Bend and Salome). Pollen: Unknown, but visits flowers of
Hymenothrix wislizeni. Pedis papposa.
Perdita butleri Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 96, figs. 624, 625, 714. ♀, ♂.




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Perdita chloris Timberlake
Ariz. (Yuma County), Nev. (Churchill and Lyon Counties), Calif. (Imperial
and Inyo Counties). Pollen: Unknown, but visits flowers of Dalea polyadenia,
Sphaemlcea.
Perdita chloris Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 64, figs. 576, 577, 690. ♀, ♂.

Perdita claripennis Timberlake
N. Mex. (Dona Ana County).

Perdita claripennis Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 71. ♀.

Perdita cognata Timberlake
Wyo. (Sweetwater County), Utah (Grand County). Pollen: Unknown, but
visits flowers of Chrysothannius.

Perdita cognata Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 93, figs. 616, 617, 710. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 67, 71-72. 9, 6 (redescription, key,
tax. status, geogr. and floral records).

Perdita congrua Timberlake
Tex. (Marathon). Pollen: Unknown, but visits flowers of Gutierrezia.

Perdita congrua Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 26, figs. 1373, 1374, 1405. ♂.

Perdita consobrina
***authority mismatch
consobrina Timberlake. N. C, S. C. Pollen: Unknown, but visits flowers of
Chrysopsis, Gerardia, Haplopappus, Heterotheca, Kuhnistera.
Perdita consobrina Timberlake, 1928. Amer. Mus. Novitates 321: 3. ♀.

Taxonomy: Timberlake, 1952. Ent. Soc. Wash., Proc. 54: 202-203. i. —Timberlake, 1960.
Calif. Univ. Pubs. Ent. 17: 12, 21, 109-111, figs. 638, 639, 721. 9, S (redescription).
—Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 315, figs. 76-79, table 9
(redescription).

Perdita consobrina
***authority mismatch
lepida Timberlake. Fla. (Dunedin, Lake Worth).

Perdita consobrina lepida Timberlake, 1952. Ent. Soc. Wash., Proc. 54: 203. ♀, ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 12, 21, 111. -Mitchell, 1960. N. C.
Agr. Expt. Sta. Tech. Bui. 141: 315-316 (redescription).

Perdita croceipes Timberlake
N. Mex. (Eddy and Socorro Counties), Ariz. (Mohave and Yavapai
Counties), Utah (Kane County). Pollen: Unknown, but visits flowers oi Baileya,
Gutierrezia including G. microcephala.
Perdita croceipes Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 72, figs. 590, 591, 697. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 64, 66, 72-73. 9 (key, geogr. and
floral records).

Perdita crotonis
***authority mismatch
caerulea Timberlake. Utah (St. George), Nev. (Glendale). Pollen: Apparently an
oligolege of Croton including C. longipes.
Perdita crotonis caerulea Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 47. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 73 (key).

Perdita crotonis
***authority mismatch
crotonis Cockerell. N. Mex., Ariz. (Maricopa County); Mexico (Chihuahua). Pollen:
Apparently an oligolege of Croton.
Perdita crotonis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 64. ♀, ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 5, 17, 43-44, figs. 558, 559, 681

(redescription, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 73
(key).

Perdita crotonis
***authority mismatch
cucullata Timberlake. Calif. (Anza). Pollen: Apparently an oligolege of Croton
including C. californicus.
Perdita crotonis cucullata Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 46. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 73 (key).

Perdita crotonis
***authority mismatch
decipiens Timberlake. Tex. (Victoria).

Perdita crotonis decipiens Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 48. ♀, ♂.

Perdita crotonis
***authority mismatch
dilucida Timberlake. Ariz., possibly Kans. and Nebr. Pollen: Presumably an oligolege
of Croton, but visits flowers of Euphorbia.
Perdita crotonis dilucida Timberlake, 1960. Cahf. Univ. Pubs. Ent. 17: 47. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 73 (key).

Perdita crotonis
***authority mismatch
juabensis Timberlake. Utah (Juab County). Pollen: Apparently an oligolege of Croton,
visits flowers of C. texensis.
Perdita crotonis juabensis Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 103. ♀, ♂.




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Perdita crotonis
***authority mismatch
leucoptera Timberlake. South. Calif. (Riverside, San Diego and San Bernardino

Counties). Pollen: Apparently an oligolege of Croton califomicus, but also visits flowers
of Hoffmanseggia presumably for nectar.
Perdita crotonis leucoptera Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 45. ♀, ♂.

Perdita crotonis
***authority mismatch
perpicta Timberlake. Nebr., N. Mex. (Arriba County). Pollen: Presumably an
oligolege of Croton, but visits flowers of Eriogonum, Euphorbia marginata.
Perdita crotonis perpicta Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 74. ♀, ♂.

Perdita crotonis
***authority mismatch
subnitens Timberlake. Kans. (Reno County). Pollen: Apparently an oligolege of
Crotoyi, visits flowers of C. monanthogynus.
Perdita crotonis subnitens Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 48. ♀.

Perdita crotonis
***authority mismatch
titusi Timberlake. Calif. (Chino).

Perdita crotonis titusi Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 46. ♀, ♂.

Perdita crotonis
***authority mismatch
undecimalis Cockerell. N. Mex. (San Miguel County). Pollen: Presumably an oligolege
of Croton, visits flowers of Croton sp.
Perdita crotonis undecimalis Cockerell, 1916. Ann. and Mag. Nat. Hist. (8) 17: 281. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 5, 45. 9 (key, redescription).
daiyi Timberlake. Tex. (El Paso). Pollen: Unknown, but visits flowers of Haplopappus
heterophyllus.
Perdita dalyi Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 119. ♀.

Perdita dilecta Timberlake
Utah (Kane County), Tex. (El Paso).

Perdita dilecta Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 40, figs. 554, 555, 679. ♀, ♂.

Perdita discreta Timberlake
N. C. (coastal plains and sand dunes). Pollen: Unknown, but visits flowers
of Haplopappus.
Perdita discreta Timberlake, 1954. Ent. News 65: 14. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 12, 22, 115-116, figs. 646, 647, 725. ♂.
9 (key, redescription, geogr. and floral records). —Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 76, 77 table 9 (redescription).

Perdita dolichocephala Swenk and Cockerell
Nebr., Kans., Colo. (Weld County). Pollen: Unknown, but
visits flowers of Amorpha canescens, Boltonia asteroides, Euphorbia, Helianthus
petiolaris, H. subrhomboideus, Heterotheca subaxillaris.
Perdita dolichocephala Swenk and Cockerell, 1907. Ent. News 18: 54. ♀.
Perdita wuiideri Cockerell, 1922. Amer. Mus. Novitates 33: 10. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 12, 19, 116-117, figs. 648, 649, 726. ♀.
6 (redescription, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49:
74 (synonymy, tax. characters).

Perdita electa Timberlake
Wyo (Sweetwater County). Pollen: Unknown, but visits flowers of
Chrysothamnus 7iauseosus.
Perdita electa Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 86. ♀.

Perdita elegans Timberlake
Calif. (Colorado Desert), Ariz. (Yuma). Pollen: Unknown, but visits
flowers o{ Abro7iia, Larrea tridentata, Palafoxia linearis. Pedis papposa.
Perdita elegans Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 55, figs. 568, 569, 686. ♀, ♂.

Perdita fallax Cockerell
Tex. to Ariz., north to Nebr. and Mont. Pollen: Unknown, but visits flowers of
Baileya, Eriogonum, Grindelia, Gutierrezia, Helianthus including H. petiolaris,
Heterotheca, Verbesina encelioides. Predator: Philanthus crabroniformis Smith, P.
pacificus arizonae Dunning.
Perdita fallax Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 90. ♀.
Perdita erigeronis Cockerell, 1897. Ann. and Mag. Nat. Hist. (6) 19: 898. ♂.
Perdita fallax fontis Cockerell, 1922. Amer. Mus. Novitates 33: 9. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 11, 20, 106-108, figs. 634, 635, 719
(key, redescription, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent.
49: 65, 74-75 (key, synonymy, variation, geogr. and floral records).

Perdita flavicauda Timberlake
Calif. (Colorado Desert). Pollen: Unknown, but visits flowers of Larrea
tridentata, Melilotus, Noma hispidum, Sphaeralcea.
Perdita flavicauda flavicauda Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 39. 9
(Misspelled flavicauda flavicanda in key, p. 4).




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Perdita flavicauda formosa Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 40. 9 (Misspelled
Jlavicanda formosa in key, p. 4).

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 63, 66, 75-76, figs. 1233, 1234, 1320.
6, 9 (key, synonymy, tax. characters, geogr. and floral records).

Perdita flaviceps Timberlake
Nev. (Las Vegas).

Perdita Jlaviceps Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 37, figs. 548, 549, 676. ♂.
flavifrons Timberlake. N. Mex. (Carrizozo), Utah (Lucia), south. Calif. (Colorado Desert).
Pollen: Unknown, but visits flowers of Chrysothamnus, Gutierrezia microcepliala.
Perdita flavifrons Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 76, figs. 1235, 1236, 1321.
S.

Perdita fuscipes Timberlake
N. Mex. (Arch in Roosevelt County).

Perdita fuscipes Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 77. ♀.

Perdita gerardiae Crawford
N. C, Fla., Miss. Pollen: Apparently an oligolege of Gerardia including G.
fasciculata, G. purpurea.
Perdita gerardiae Crawford, 1932. Ent. See. Wash., Proc. 34: 75. cJ, 9.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 13, 18, 68, figs. 582, 583, 693. 9, 6
(redescription). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 317, figs. 76-79, table
9 (redescription).

Perdita gerhardi
***authority mismatch
dallasiana Cockerell. Tex. Pollen: Unknown, but visits flowers of Aster

tanacetifolius, Dalea aurea, Helianthus ayinuus, Monarda citriodora, M. punctata.
Perdita dallasiana Cockerell, 1906. Entomologist 39: 178. ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 13, 14, 15, 28-29. 9, 6
(redescription, tax. status, geogr. and floral records).

Perdita gerhardi
***authority mismatch
gerhardi Viereck. Wis., 111., Ind. Pollen: Unknown, but visits flowers of Monarda
punctata.
Perdita gerhardi Viereck, 1904. Ent. News 15: 21. ♀, ♂.
Perdita gerhardi arenicola Timberlake, 1929. N. Y. Ent. Soc, Jour. 37: 119. ♀, ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 2, 3, 13, 27-28 (redescription,
synonymy, geogr. range). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 317-318
(redescription, as gerhardi gerhardi and g. arenicola).

Perdita gerhardi
***authority mismatch
monardae Viereck. N. J., Va., Fla. Pollen: Unknown, but visits flowers of Monarda
punctata.
Perdita monardae Viereck, 1904. Ent. News 15: 22. ♀, ♂.

Taxonomy: Timberlake, 1952. Ent. Soc. Wash., Proc. 54: 204 (tax. status). —Timberlake, 1960.
Calif. Univ. Pubs. Ent. 17: 7, 15, 28 (redescription). —Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 318, figs. 76, 77, table 9 (redescription).

Perdita gutierreziae Cockerell
Colo., Tex., N. Mex., Ariz., Calif. (Mojave Desert). Pollen: Unknown,
but visits flowers of Baileya pleniradiata, Chrysothamnus, Gutierrezia lucida, G.
microcephala, Haplopappus acradenius, H. heterophyllus, Solidago confinis,
Sphaeralcea.
Perdita gutierreziae Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 85. ♂.

Taxonomy: Swenk and Cockerell, 1907. Ent. News 18: 56. cJ, 9. —Timberlake, 1960. Calif.
Univ. Pubs. Ent. 17: 8, 16, 74-76, figs. 594, 595, 699. 9, S (key, redescription, geogr. and
floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 65, 77-78 (tax. status, geogr.
and floral records).
halli Timberlake. Calif. (Palm Springs).

Perdita halli Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 89, figs. 612, 613, 708. ♀, ♂.

Perdita hirsuta Cockerell
N. Mex., Ariz. (Willcox). Pollen: Unknown, but visits flowers of Dalea
including D. scoparia.
Perdita hirsuta Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 79. ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 8, 18, 66, figs. 580, 581, 692. 9, 6
(redescription).




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Perdita hirtella Timberlake
Utah (Wasatch County).

Perdita hirtella Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 78, fig. 10. ♀.

Perdita hirticeps
***authority mismatch
apicata Timberlake. Calif. (Colorado and Mojave Deserts), Ariz. (Mohave County),
Utah (Cornish and Oak City). Pollen: Collects pollen from flowers oi Stephanovieria
including S. exigua, S. pauciflora, but visits these and other flowers for nectar including
Malacotkrix glabrata.
Perdita hirticeps apicata Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 59. ♀, ♂.
Perdita hirticeps candidipennis Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 60. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 79 (tax. characters, as hirticeps

Perdita apicata
***authority mismatch
and h. candidipennis). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 28
(synonymy, geogr. and floral record).

Perdita hirticeps
***authority mismatch
hirticeps Timberlake. Calif, (cismontane). Pollen: Collects pollen from the flowers of
Stephanomeria including S. exigua, S. e. var. coronaria, S. virgata, but visits these and
other flowers for nectar including Coreopsis lanceolata, Hemizonia.
Perdita hirticeps hiHiceps Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 58, figs. 570, 571,
687. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 79 (tax. characters, variation).

Perdita hirticeps
***authority mismatch
luteocincta Timberlake. Calif. (Antioch). Pollen: Unknown, but visits flowers of
Gutierrezia califomica.
Perdita hirticeps luteocincta Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 60. ♀.

Perdita indioensis Timberlake
Calif. (Indio). Parasite: Neolarra vigilans (Ckll.). Pollen: Unknown, but
visits flowers of Haplopappus acradenius.
Perdita indioensis Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 54, figs. 566, 567, 685. ♀.
6.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 63 (key).

Perdita idonea Timberlake
Utah, Ariz. (Coconino County). Pollen: Unknown, but visits flowers of
Gutierrezia.
Perdita idonea Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 79, figs. 1237, 1238, 1322. ♀.
6.

Perdita jonesi Cockerell
Tex., Kans. Pollen: Unknown, but visits flowers oi Dalea aurea, Monarda
citriodora, M. punctata, Petalostemon multiflorum.
Perdita jonesi Cockerell, 1906. Entomologist 39: 177. ♀, ♂.
Perdita birkmanni Cockerell, 1916. Ann. and Mag. Nat. Hist. (8) 17: 282. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 12, 22, 114-115, figs. 644, 645, 724
(redescription, synonymy, geogr. and floral records).

Perdita knowltoni Timberlake
Utah (Moab). Pollen: Unknown, but visits flowers of Chrysothamnus.
Perdita knowltoni Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 51, figs. 562, 563, 683. 9
(cJ misdet.).

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 63, 80 (key, tax. status, geogr. and
floral record).

Perdita labergei Timberlake
Kans. (Hutchinson). Pollen: Unknown, but visits flowers of Euphorbia.

Perdita labergei Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 41, figs. 556, 557, 680. ♀, ♂.

Perdita lasiogastra Timberlake
Tex. (Cameron County), N. Mex. (San Miguel County), Colo. (Moffat
County). Pollen: Unknown, but visits flowers of Pedis jxipposa.
Perdita lasiogastra Timberlake, 1929. N. Y. Ent. Soc, Jour. 37: 115. ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 9, 20, 84-86, figs. 606, 607, 705. 9, 6
(key, redescription, geogr. and floral records).

Perdita laticincta Swenk and Cockerell
N. Dak., Nebr., Kans., Wyo., N. Mex. Pollen: Unknown, but
visits flowers of Aniorpha canescens, Haplopappus ciliatns, Helianthus petiolaris,
Melilotus.
Perdita laticincta Swenk and Cockerell, 1907. Ent. News 18: 52. ♀, ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 8, 20, 76, figs. 596, 597, 700. 9, 6
(key, redescription, geogr. and floral records).




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Perdita luteiceps Cockerell
Colo. (Conejos, Garfield and Routt Counties), Wyo. (Fremont County).
Pollen: Unknown, but visits flowers of Gutierrezia.
Perdita Luteiceps Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 74. ♂.

Taxonomy: Timberlake, 1928. N. Y. Ent. Soc, Jour. 37: 112. d, 9. -Timberlake, 1960. Calif.
Univ. Pubs. Ent. 17: 3, 14, 36-37, figs. 544, 545, 674. 9 , 6 (key, redescription, geogr
records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 81 (geogr. and floral records).

Perdita luteola Cockerell
Nebr., Colo., N. Mex., Ariz. Pollen: Unknown, but visits flowers of
Chrysothamnus, Gutierrezia sarothrae, Haplopappiis heterophyllus.
Perdita luteola Cockerell, 1894. Ent. News 5: 328. ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 2, 13, 23-24, figs. 526, 527, 664 (key,
geogr. and floral records).

Perdita maculigera
***authority mismatch
var. bilineata Timberlake. 111., Mo., Kans., Nebr., S. Dak.

Perdita maculipeyinis var. bilineata Timberlake, 1929. N. Y. Ent. Soc, Jour. 37: 121. ♀, ♂.

Taxonomy: Timberlake, 1952. Ent. Soc. Wash., Proc. 54: 204 (tax. status). —Timberlake, 1960.
Calif. Univ. Pubs. Ent. 17: 3, 14, 33 (tax. status, tax. characters, key, geogr. records).
—MitcheU, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 320, figs. 76, 77, table 9 (tax.
characters).

Perdita maculigera
***authority mismatch
maculigera Cockerell. Tex. (Southmost), N. Mex. (Las Cruces). Pollen: Unknown,
but visits flowers of Salix.
Perdita maculigera Cockerell, 1896. Ent. News 7: 255. ♂.

Taxonomy: Timberlake, 1952. Ent. Soc. Wash., Proc. 54: 204 (tax. status). —Timberlake, 1960.
Calif. Univ. Pubs. Ent. 17: 3, 14, 32-33 (key, redescription, geogr. and floral records).

Perdita maculigera
***authority mismatch
maculipennis Graenicher. Minn., Wis., Mich., 111., Iowa, Kans., Tex. (Anderson and
Robertson Counties). Parasite: Eupliyto pollinaris Reinhard. Pollen: Collects pollen
from flowers of Salix nigra although has been listed as collecting pollen from flowers of
Melilot2is alba, visits other flowers presumably for nectar including Erigeron
philadelphicus.
Perdita maculipennis Graenicher, 1910. Canad. Ent. 42: 102. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 9, 14, 33-34, figs. 540, 541, 671, 672.
9, 6 (key, redescription, geogr. and floral records). —Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 320-321, figs. 75, 77, table 9 (redescription). — Rozen, 1966. Amer. Mus.
Novitates 2259: 19-20 (larva). —Yager and Rozen, 1966. Amer. Mus. Novitates 2265: 12
(pupa).

Biology: Michener and Ordway, 1963. Kans. Ent. Soc, Jour. 36: 34-45, 10 figs, (nest
architecture, life history, immature stages, flight behavior, pollen source, parasite).

Perdita maculipes Cockerell
N. Mex. (Las Cruces). Pollen: Unknown, but visits flowers of
Haplopappiis heterophyllus.
Perdita maculipes Cockerell, 1896. Acad. Nat. Sci. Phila., Proc 48: 87. ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 121 (tax. characters). —Timberlake,
1968. Calif. Univ. Pubs. Ent. 49: 65, 81-82 (key, redescription of type).

Perdita media Timberlake
Calif. (Colorado and Mojave Deserts). Pollen: Unknown, but visits flowers
of Gutierrezia lucida, Haplopappus acradenius, Solidago confinis.
Perdita media Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 73, figs. 592, 593, 698. ♀, ♂.

Perdita melanostoma
***authority mismatch
albocincta Timberlake. N. Mex., Ariz. Pollen: Unknown, but visits flowers of
Baileya, Gutierrezia, Haplopappus spinulosus.
Perdita albocincta Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 78. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 63, 83-84 (geogr. and floral records).

Perdita melanostoma
***authority mismatch
melanostoma Swenk and Cockerell. Idaho, Utah, Colo., Nebr., N. Mex., Ariz.
Pollen: Unknown, but visits flowers of Chrysothamnus, Gutierrezia including G.
sarothrae.
Perdita melanostoma Swenk and Cockerell, 1907. Ent. News 18: 57. ♀.




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Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 8, 78 (key, redescription, geogr. and
floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 63, 65, 82-83, figs. 11, 1241,
1242, 1324. 6 (key, redescription, geogr. and floral records).

Perdita mesillensis Timberlake
N. Mex. (Mesilla Park). Pollen: Unknown, but visits flowers of
Gutierrezia lucida, Pedis papposa.
Perdita mesillensis Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 84, figs. 1243, 1244, 1325.
6.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 28 (tax. characters, synonymy).

Perdita microsticta Timberlake
Kans. (Scott County).

Perdita microsticta Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 89. ♀.

Perdita mimula Timberlake
Calif. (Colorado Desert). Pollen: Unknown, but visits flowers of
Haplopappus acradenius.
Perdita mimula Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 53. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 63 (key).

Perdita nuda Cockerell
N. Mex., Ariz., Colo., Utah, Idaho (Franklin County). Parasite: Megaselia,
Sphecodes sp. near fragariae Ckll. Pollen: Apparently collects pollen only from vernal
and autumnal flowering Compositae including Chrysothamnus, Erigeron, Grindelia
including G. squarrosa.
Perdita nuda Cockerell, 1896. Acad. Nat. Sci. PhUa., Proc. 48: 93. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 12, 119. 9 (redescription, key).
—Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 104-105, figs. 839, 840, 898. 6, 9 (geogr.
and floral records). — Torchio, 1975. Kans. Ent. Soc, Jour. 48: 266-275, 277-278, figs. 1-18,
26, 27 (immature stages).

Biology: Torchio, 1975. Kans. Ent. Soc, Jour. 48: 257-279, 27 figs, (nest architecture, life
history, nest associates).

Perdita numerata
***authority mismatch
hesperia Timberlake. South. Calif. (Colorado Desert); Mexico (Baja California).

Pollen: Unknown, but visits flowers of Geraea canescens, Larrea tridentata, Melilotus,
Prosopis, Tamarix.
Perdita mimerata hesperia Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 62. ♀, ♂.

Perdita numerata
***authority mismatch
numerata Cockerell. Tex., N. Mex., Ariz. (Phoenix), ?Calif. (Needles). Pollen:
Unknown, but visits flowers of Actinea, Prosopis, Salix.
Perdita numerata Cockerell, 1895. Amer. Ent. Soc, Trans. 22: 296. ♀.

Taxonomy: Cockerell, 1916. Ann. and Mag. Nat. Hist. (8) 17: 281. 6. —Timberlake, 1960.
Calif. Univ. Pubs. Ent. 17: 6, 15, 60-62, figs. 572, 573, 688 (key, redescription, tax. status,
geogr. and floral records).

Perdita occidua Timberlake
Ariz. (Oak Creek Canyon).

Perdita occidua Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 120. ♀.

Perdita occlusa Timberlake
Ariz. (Walnut Canyon near Flagstaff).

Perdita occlusa Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 85, figs. 1245, 1246, 1326. ♂.

Perdita octomaculata
***authority mismatch
octomaculata (Say). N. B. south to Ga., west to Minn., 111. and Miss. Pollen:
Unknown, but visits flowers of Aster ericoides, A. puniceus, Bidens, Boltonia,
Eupatorium, Euthamia, Lycopus, Monarda, Solidago.
Panurgus 8-maculatus Say, 1824. In Keating, Narr. Long's 2nd Exped., v. 2, p. 350. cJ, 9.
Perdita octomaculata Dalla Torre, 1896. Cat. Hym., v. 10, p. 173. Emend.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 12, 22, 111-113, figs. 642, 643, 723
(key, redescription, geogr. and floral records). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech.
Bui. 141: 321-322, figs. 76-79, table 9 (redescription, geogr. range, floral records).

Perdita octomaculata
***authority mismatch
terminata Cockerell. N. Dak., Nebr., Kans., Iowa, lU., Ark., Miss. Pollen:

Unknown, but visits flowers of Aster azureus, A. miiltiflonis, A. saliciflorus, Solidago
canadensis, S. rigida.
Perdita octomaculata terminata Cockerell, 1922. Amer. Mus. Novitates 33: 8. ♂, ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 12, 22, 113-114 (key, tax. characters,
geogr. and floral records). —Mitchell, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 322 (tax.
characters, geogr. range).




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Perdita ordinata Timberlake
Ariz. (Cochise and Navajo Counties), N. Mex. (Hidalgo and Socorro

Counties), Utah (Cache and Washington Counties). Pollen: Unknown, but visits flowers
of Baileya pleniradiata, Dithyrea including D. wislizenii, Heterotheca subaxillaris,
Pedis papposa.
Perdita ordinata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 105, figs. 841, 842, 899. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 67, 86, fig. 12 (key, geogr. and floral
records). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 28-29. d, 9 (geogr. and floral
records).

Perdita paroselae Timberlake
South. Calif. (Colorado Desert). Pollen: Possibly an oligolege of Dalea,
visits flowers of Dalea califoniica, D. emoryi.
Perdita paroselae Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 65, figs. 578, 579, 691. ♀, ♂.

Perdita pectidis Cockerell
N. Mex., Ariz. (Salome). Pollen: Unknown, but visits flowers of
Kallstroemia, Pedis papposa, Sesuviitm verrucosum, Wedeliella.
Perdita pedidis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 83. ♀, ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 9, 19, 91-92, figs. 614, 615, 709. 9, S
(key, tax. characters, geogr. and floral records).

Perdita percincta Timberlake
Utah (Skull Rock Pass, 6,100ft.), Ariz. (Coconino County). Pollen:

Unknown, but visits flowers of Aster, Gutierrezia microcephala, Wislizenia refrada.
Perdita percincta Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 86. ♀.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 30-31, figs. 1379, 1380, 1408. 6, 9
(key, redescription, geogr. and floral records).

Perdita perixantha Timberlake
Ariz. (Navajo County).

Perdita perixantha Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 23, figs. 524, 525, 663. ♂.

Perdita perpallida
***authority mismatch
citrinella Graenicher. Wis., Minn., N. Dak., Nebr., Alta. Pollen: Unknown, but visits
flowers o{ Petalostemo7i purpureum, P. villosum.
Perdita citrinella Graenicher, 1910. Canad. Ent. 42: 103. ♀.

Taxonomy: Crawford, 1912. Canad. Ent. 44: 359. 6,9. —Timberlake, 1952. Ent. Soc. Wash.,
Proc. 54: 204 (tax. status). — Timberiake, 1960. Calif. Univ. Pubs. Ent. 17: 3, 14, 26-27 (key,
tax. characters, geogr. and floral records).

Perdita perpallida
***authority mismatch
perpallida Cockerell. Man., Minn., N. Dak., Nebr., Kans., N. Mex. Pollen:

Apparently an oHgolege of vernal and autumnal flowering Compositae, visits flowers of
Amorpha canescens, Heleniiim autumnale, Petalostemon candidum, P. oligophyllum,
P. villosum, Solidago occidentalis.
Perdita perpallida Cockerell, 1901. Entomologist 34: 190. ♀, ♂.

Taxonomy: Swenk and Cockerell, 1907. Ent. News 18: 57. —Stevens, 1919. Canad. Ent. 51:
207. 9 , 6 (tax. characters, floral records). —Timberlake, 1952. Ent. Soc. Wash., Proc. 54:
204 (tax. status). -Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 2, 13, 25-26, figs. 532, 533,
667. 9, 6 (key, redescription, geogr. and floral records). —Mitchell, 1960. N. C. Agr. Expt.
Sta. Tech. Bui. 141: 322 (tax. status).

Perdita phymatae Cockerell
N. Mex., Ariz., Utah. Pollen: Unknown, but visits flowers of Baliia

absinthifolia, Baileya pleniradiata, Gutierrezia lucida, G. microcephala, Haplopappus

heteropiiyllus, Pedis papposa. Predator: Phymata sp.
Perdita phymatae Cockerell, 1895. Acad. Nat. Sci. Phila., Proc. 47: 12. ♀.
Perdita nitidella Cockerell, 1895. Acad. Nat. Sci. Phila., Proc. 47: 16. ♂.

Taxonomy: Timberiake, 1960. Calif. Univ. Pubs. Ent. 17: 9, 92-93 (key, redescription,
geographic, floral and predator records). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17:
70-71, figs. 586, 587, 695. 6 (as nitidella). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49:
64, 85. 9 (key, redescription, as nitidella). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49:
65' (key). -Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 31 (synonymy, geogr. range, tax.
characters).

Perdita picturata Timberlake
Tex. (Victoria and Aransas Counties).

Perdita picturata Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 50, figs. 560, 561, 682. ♀, ♂.




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Perdita plucheae Timberlake
Calif. (Colorado Desert), Ariz.; Mexico (Baja California). Pollen:
Unknown, but visits flowers of Pluchea sericea, Tamarix gallica.
Perdita plucheae Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 69, figs. 584, 585, 694. ♀, ♂.

Perdita polygonellae Timberlake
N. C. (Coastal Plain). Pollen: Unknown, but visits flowers of
Polygonella polygama.
Perdita polygonellae Timberlake, 1954. Ent. News 65: 12. ♀, ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 4, 16, 40, figs. 552, 553, 678 (key, tax.
characters). — MitcheU, 1960. N. C. Agr. Expt. Sta. Tech. Bui. 141: 323, figs. 76-79, table 9
(redescription).

Perdita prionopsidis Timberlake
Kans. (Reno and Montgomery Counties). Pollen: Unknown, but visits
flowers of Boltonia asteriodes, Haplopappus ciliatus, Helianthus petiolaris, Heterotheca
subaxillaris.
Perdita prionopsidis Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 104, figs. 632, 633, 718.
9,

Perdita rectangulata Cockerell
Colo., Idaho, Ariz. (Holbrook). Pollen: Unknown, but visits flowers of
Chrysothamnus, Gutierrezia sarothrae, Solidago.
Perdita rectangulata Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 72. ♀ (♂ misdet.).

Taxonomy: Stevens, 1919. Canad. Ent. 51: 207. cJ, 9 (as martini). —Timberlake, 1960. Calif.
Univ. Pubs. Ent. 17: 3, 9, 16, 82-83, figs. 602, 603, 703. 9, 6 (key, redescription, geogr. and
floral records, in part martini). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 64, 65,
87-88 (key, tax. characters and tax. status).

Perdita reperta Timberlake
N. Mex. (Quemado), Ariz. (Navajo County), Colo. (Alamosa County).
Pollen: Unknown, but visits flowers of Chrysothavinus including C. viscidiflonis.
Predator: Philanthus sp.
Perdita repeHa Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 88, figs. 13, 1247, 1248, 1327.
9,cJ.

Perdita retusa Timberlake
Ariz. (Mohave and Yuma Counties), Calif. (Colorado and Mojave Deserts).
Pollen: Unknown, but visits flowers of Chrysothamnus paniculatus, Haplopappus.
Perdita retiisa Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 86, figs. 608, 609, 706. ♀, ♂.

Perdita rhodura Cockerell
Ariz., N. Mex., Nebr. (Sioux County). Pollen: Unknown, but visits flowers
of Gutierrezia sarothrae, Haplopappus, Zinnia grandiflora.
Perdita rhodura Cockerell, 1897. Ann. and Mag. Nat. Hist. (6) 20: 511. ♀, ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 6, 15, 52-53, figs. 564, 565, 684. 9, 6
(key, redescription, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent.
49: 63, 66 (key).

Perdita scitula
***authority mismatch
antiochensis Timberlake. Calif. (Antioch, Oakley). Pollen: Unknown, but visits flowers
of Eriogonum, Gutierrezia califomica, Heterotheca grandijlora, Lessingia glandnlifera.
Perdita scitula antiochensis Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 80, figs. 598, 599,
701. ♀, ♂.

Perdita scitula
***authority mismatch
scitula Timberlake. Calif. (Inyo County). Pollen: Unknown, but visits flowers of
Chrysothamnus nauseosus var. gnaphalodes.
Perdita scitula scitula Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 79. ♀, ♂.

Perdita sedulosa Timberlake
N. Mex. (Socorro County). Pollen: Unknown, but visits flowers of
Baileya pleniradiata.
Perdita sedulosa Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 31. ♀.

Perdita sejuncta Timberlake
N. Mex. (Socorro County), Utah (Wah Wah Mts.); Mexico (Chihuahua).
Pollen: Unknown, but visits flowers of Gutierrezia microcephala.
Perdita sejuncta Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 90. ♀.

Taxonomy: Timberiake, 1971. Calif. Univ. Pubs. Ent. 66: 32 (geogr. record).

Perdita senecionis Cockerell
N. Mex. (Las Cruces). Pollen: Unknown, but visits flowers of Senecio
douglasii.
Perdita senecionis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 94. ♀.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 11, 100-101. 9 (redescription, key).




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Perdita separata Timberlake
Tex. (Zapata County).

Perdita separata Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 106. ♀.

Perdita snowii Cockerell
Colo. (Boulder, Costilla and Moffat Counties), Ariz. (Holbrook). Pollen:
Unknown, but visits flowers o{ Aster, Grindelia, Gutierrezia.

Perdita snowii Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 73. ♀.

Taxonomy: Cockerell, 1910. Psyche 17: 244. 6. —Timberlake, 1960. Calif. Univ. Pubs. Ent.
17: 9, 20, 83-84, figs. 604, 605, 704. 9, 6 (key, redescription, geogr. and floral records).
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 64, 91 (tax. characters, key).

Perdita solidaginis Cockerell
Colo. (White Rocks, near Boulder). Pollen: Unknown, but visits flowers
of Solidago.
Perdita solidaginis Cockerell, 1922. Amer. Mus. Novitates 33: 12. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 316-317 (tax. characters).
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 66, 91-92. 6 (key, redescription of type
specimen).

Perdita stepheni Timberlake
Tex. (Dimmit County). Pollen: Unknown, but visits flowers of Monarda
punctata.
Perdita stepheni Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 29, figs. 536, 537, 669. ♂.

Perdita submedia Timberlake
Ariz. (Apache County). Pollen: Unknown, but visits flowers of
Chrysothaniniis.
Perdita submedia Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 32. ♀.

Perdita swenki Crawford
N. Y. (Long Island), Mich., 111., Wis., Minn., N. Dak., Nebr., Alta. Pollen:
Unknown, but visits flowers of Chrysopsis, Grindelia, Helianthus maximillianii,
Liatris, Solidago juncea, S. rigida.
Perdita swenki Crawford, 1915. Insecutor Inscitiae Menstruus 3: 109. tJ, 9.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 10, 19, 103-104, figs. 630, 631, 717. ♀.
(J (key, redescription, geogr. and, floral records). —Mitchell, 1960. N. C. Agr. Expt. Sta.
Tech. Bui. 141: 324, figs. 76-78, table 9 (redescription). —Timberlake, 1971. Calif. Univ.
Pubs. Ent. 66: 33 (geogr. records).

Perdita tacita Timberlake
Idaho (Owyhee County).

Perdita tacita Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 93. ♀.

Perdita translineata Timberlake
Tex. (Brazos and Van Zandt Counties), N. Mex. (Socorro County).
Pollen: Unknown, but visits flowers of Baileya pleniradiata, Eriogonum midtijlorum,
Haplopappus divaricatus, Heterotheca subaxillaris.
Perdita translineata Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 93, figs. 618, 619, 711. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 65, 67, 94. 9, (J (key, redescription,
geogr. and floral records).

Perdita tridentata Stevens
N. Dak., Nebr., Kans. Pollen: Unknown, but visits flowers of Haplopappus,
Helianthus petiolaris.
Perdita tridentata Stevens, 1919. Canad. Ent. 51: 206. ♂.

Taxonomy: Stevens, 1921. Canad. Ent. 53: 66. 9. —Timberlake, 1960. Calif. Univ. Pubs. Ent.
17: 12, 21, 118-119, figs. 650, 651, 727. 9, S (key, redescription, geogr. and floral records).

Perdita trifida Timberlake
Calif. (Mojave Desert). Pollen: Unknown, but visits flowers of Baccharis
emoryi, Haplopappus acradenius.
Perdita trifida Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 37, figs. 546, 547, 675. ♂.

Perdita trimaculata Timberlake
South. Calif. (Colorado and Mojave Deserts), Ariz. (Mohave County).
Pollen: Unknown, but visits flowers of C hrysothamnus nauseosus, C. paniculatus,
Gutierrezia lucida.
Perdita trimacxdata Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 62, figs. 574, 575, 689. ♀.
6.

Perdita truncatella Timberlake
Ariz. (Snowflake).

Perdita truncatella Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 94, figs. 1251, 1252, 1329.
6.




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Perdita variegata
***authority mismatch
pura Timberlake. Tex., Kans. (Garden City). Pollen: Unknown, but visits flowers of
Monarda punctata var. occidentalis.
Perdita variegata jnira Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 31. ♀, ♂.

Perdita variegata
***authority mismatch
variegata Timberlake. Kans. Pollen: Unknown, but so far has been collected
exclusively at the flowers of Monarda including M. punctata var. occidentalis.
Perdita variegata variegata Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 30, figs. 538, 539,
670. ♀, ♂.

Perdita versuta Timberlake
Tex. (Kingsville).

Perdita versuta Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 105, figs. 843, 844, 900. ♂.

Perdita xanthochroa Timberlake
Colo. (Moffat County), Utah (Juab and Uinta Counties), Idaho
(Owyhee County). Pollen: Unknown, but visits flowers of Sphaeralcea ambigua,
Stanleya pinnata.
Perdita xanthochroa Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 24, figs. 530, 531, 668. ♀.
S.

Perdita xanthodes Timberlake
Calif. (Mohave Desert). Pollen: Unknown, but visits flowers of
Gutierrezia lucida, Haplopappus cooperi.
Perdita xanthodes Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 24, figs. 528, 529, 665. ♀.
6.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 63 (key). —Timberlake, 1971. Calif.
Univ. Pubs. Ent. 66: 33 (geogr. and floral records).

Perdita xanthops Timberlake
Wyo. (Carbon County).

Perdita xanthops Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 33, figs. 1381, 1382, 1409. ♀.
6.

Species Group Ventralis

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 1, 2-14 (key, tax. characters, key to
included spp.). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 95-97 (suppl. key to included
spp.).

Perdita acaciae Timberlake
South. Calif. (Colorado Desert). Pollen: Unknown, but visits flowers of
Acacia greggii. Agave consociata, A. deserti.
Perdita acaciae Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 61, figs. 799, 800, 878. ♀, ♂.

Perdita albata Timberlake
Ariz. (Navajo County). Pollen: Unknown, but visits flowers of Mentzelia
pumila.
Perdita albata Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 97. ♀.

Perdita amplipennis Timberlake
Calif, mts. (Inyo and Mono Counties). Pollen: Unknown, but visits
flowers of Eriogonum fasciculatum, Stanleya pinnata, and presumably Dalea.
Perdita amplipennis Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 30, figs. 751, 752, 854. ♀.
6.

Perdita austini Cockerell
N. Mex. Pollen: Unknown, but visits flowers of Baileya pleniradiata,
Gutierrezia microcephala, G. sarothrae, Haplopappus.
Perdita austini Cockerell, 1895. Acad. Nat. Sci. PhUa., Proc. 47: 13. ♂.
Perdita aeneifrons Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 91. ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 10, 40-41, figs. 769, 770, 863. 6
(redescription). —Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 125. 6, 9 (synonymy).
—Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 96, 97, 98-99 9, 6 (key, redescription,
geogr. and floral records).

Perdita bicuspidariae Timberlake
South. Calif. (Colorado Desert). Pollen: Collects pollen from the
flowers of Mentzelia involucrata, but also visits flowers of Hyptis enioryi presumably
for nectar.
Perdita bicuspidariae Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 20, figs. 741, 742, 849.
6, 9.

Perdita brevihirta Timberlake
Ariz. (Maricopa County), Wyo. (Fremont County). Pollen: Unknown,
but visits flowers of Chrysothamnus nauseosus, Pedis papposa.
Perdita brevihirta Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 41, figs. 771, 772, 864. ♀.
6.




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Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 34 (geogr. and floral records).

Perdita claypolei
***authority mismatch
australior Timberlake. Calif. (San Diego and Los Angeles Counties). Pollen:

Unknown, but visits flowers oi Adenostoma fasciculatum, Eriogonuni fasciculatum,
Hemizonia fasciculata, Heteromeles arbutifolia, Opuntia.
Perdita claypolei australior Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 68. ♀, ♂.

Perdita claypolei
***authority mismatch
claypolei Cockerell. South. Calif, (cismontane). Pollen: Collects pollen from the

flowers oi Eriogonum including E. elongatum, E. fasciculatum, E. gracile, E. wrightii
var subscaposuvi, E. umbellatum, but visits these and other flowers for nectar
including Eriastruvi virgatum.
Perdita claypolei Cockerell, 1901. Canad. Ent. 33: 281. ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 6, 13, 66-68, figs. 805, 806, 881. 9, 6
(key, redescription, geogr. and floral records).

Perdita claypolei
***authority mismatch
limatula Timberlake. Calif. (Kern County) to Oreg., and Nev. (Douglas County).
Pollen: Collects pollen from the flowers of Eriogonum including E. elatuvt, E.
fasciculatum, E. gracile, E. heennannii, E. inflatum, E. latifolium, E. nudum, E.
vimiyieum, E. virgatum, E. wrightii var. subscaposum, but visits these and other
flowers for nectar including Achillea millefolium, Brickellia, Cirsium, Solidago.
Perdita claypolei limatula Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 68. ♀, ♂.

Perdita clypeata
***authority mismatch
clypeata Timberlake. South. Calif. (Colorado Desert). Pollen: Unknown, but visits
flowers of Eriogonum inflatum.
Perdita clypeata clypeata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 79, figs. 817, 818,
887. ♀, ♂.

Perdita clypeata
***authority mismatch
immaculata Timberlake. Ariz, (near Mesa), south. Calif. (Desert Center and
Hayfield). Pollen: Unknown, but visits flowers of Eriogonum inflatum.
Perdita clypeata immaculata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 80. ♀, ♂.

Perdita colei Timberlake
Calif. (San Bernardino Mts. and vicinity). Pollen: Unknown, but visits
flowers oi Chorizanthe staticoides, Chrysothamnus nauseosus, Gutierrezia lucida,
Lepidospartum squamatum, Solidago confinis.
Perdita colei Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 48, figs. 783, 784, 870. ♀, ♂.

Perdita crandalli Timberlake
Ariz. (Pima and Pinal Counties). Pollen: Unknown, but visits flowers of
Petalonyx thurberi.
Perdita craiidalli Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 38. ♀.

Perdita dasylirii Cockerell
Western Tex. to Ariz. Pollen: Unknown, but visits flowers of Acacia,

Agave palmeri, Amorpha fruticosa, Asclepias galioides, Ceanothus, Dasylirion wheeleri.
Euphorbia albomarginata, Mertensia franciscana, Nolina microcarpa, Rudbeckia
laciniata, Salix taxifolia, Sapindus sapo7iaria.
Perdita dasylirii Cockerell, 1907. Ann. and Mag. Nat. Hist. (7) 19: 539. "9" = cJ (d

misdet.).
Perdita nolinae Cockerell, 1922. Amer. Mus. Novitates 33: 11. ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 5, 12, 55-57, figs. 793, 794, 875. 9, (key, redescription, geogr. and floral records).

Perdita dimidiata Timberlake
Ariz. (Pinal County); Mexico (Baja California). Pollen: Unknown, but
visits flowers oi Sesuvium verrucosuyn.
Perdita dimidiata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 39, figs. 767, 768, 862. ♂.

Perdita distans Timberlake
South. Calif. (Colorado and Mojave Deserts). Pollen: Unknown, but visits
flowers oi Eriogonum including E. reniforme, E. thomasii.
Perdita distans Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 73, figs. 811, 812, 884. ♀, ♂.
erythropyga Timberlake. South. Calif. (Colorado Desert). Pollen: Collects pollen of Dalea
including D. califoniica, D. emoryi, but also visits other flowers for nectar including
Cryptantha angustifolia, Isomeris arborea, Larrea tridentata.
Perdita erythropyga Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 42, figs. 773, 774, 865. ♀.
6.

Perdita exilis Timberlake
South. Calif. (Colorado and Mojave Deserts). Pollen: Collects pollen from
Petalonyx thurberi.
Perdita exilis Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 36, figs. 763, 764, 860. ♂, ♀.




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Perdita fulvicauda Timberlake
South. Calif. (Colorado Desert); Mexico (Baja California). Pollen:

Unknown, but visits flowers of Baileya plenimdiata, Coldenia plicata, Dalea viollis,
Larrea tridentata, Melilotus, Nama hispidum, Phacelia distayis, Prosopis juliflora.
Verbena.
Perdita fulvicauda Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 43, figs. 775, 776, 866. ♀.
6.

Perdita gentilis Timberlake
Nev. (Churchill and Washoe Counties), Ariz. (Vermillion Cliffs). Pollen:
Unknown, but visits flowers of Eriogoniim.
Perdita gentilis Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 78. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 96, 97, 109-110. 9, 6 (key, geogr.
and floral record).

Perdita glabrella Timberlake
Ari2. (Continental).

Perdita glahrella Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 104. ♀.

Perdita glabrescens Timberlake
Calif. (Inyo County).

Perdita glabrescens Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 38, figs. 765, 766, 861. ♀.

S.

Perdita grandiceps Cockerell
N. Mex. (Alamogordo, La Cueva and Las Cruces). Pollen: Unknown, but
visits flowers of Fallugia paradoxa, Solidago.
Perdita grandiceps Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 96. ♂.

Taxonomy: Cockerell, 1899. Canad. Ent. 31: 256. 9. —Timberlake, 1962. Calif. Univ. Pubs.
Ent. 28: 3, 10, 32-33, figs. 757, 758, 857. 9, S (key, redescription, geogr. and floral records).

Perdita holoxantha Timberlake
Utah (Grand and Washington Counties), Ariz. (Holbrook). Pollen:
Unknown, but visits flowers of Mentzeiia piiniila, Poleomintha incana.
Perdita holoxantha Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 17, figs. 737, 738, 847. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 95, 96, 100. 9, 6 (key, geogr. and
floral records).

Perdita inornata Timberlake
Calif. (Clark Mt. and Panamint Mts.), Nev. (Charleston Mts.), Ariz.
(Grand Canyon, south rim). Pollen: Unknown, but visits flowers of Agave deserti,
Cowania stansburiana, Eriodictyon.
Perdita inoniata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 49, figs. 787, 788, 872. ♀, ♂.

Perdita jucunda Timberlake
Calif., Nev. (Churchill County), Ariz. (Painted Desert). Pollen: Collects

pollen from the flowers of Eriogonum, including E. aiireum, E . fasciculatum, E. gi'acile,
E. inflatum, E. molestum var. davidsonii, E. nitduni, but also visits other flowers for
nectar including Chorizanthe staticoides, Gutierrezia lucida, Lepidospartnui
squaiuatum.
Perdita jucimda Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 64, figs. 803, 804, 880. ♀, ♂.

Perdita kanabensis Timberlake
Utah (Kanab), Ariz. (Coconino County). Pollen: Unknown, but visits
flowers of cleomella, Eriogonum corymbosuni.
Perdita kanabensis Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 34, figs. 1383, 1384, 1410.

Perdita labrata Timberlake
South. Calif. (Colorado and Mojave Deserts). Pollen: Evidently collects
pollen from the flowers of Eriogonum including E.fasciculatum var. polifolium, but
also visits other flowers presumably for nectar including Larrea tridentata.
Perdita labrata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 81, figs. 821, 822, 889, 901. ♀.
3.

Perdita lateralis
***authority mismatch
daleae Timberlake. Calif. (Inyo and Panamint Mts.). Pollen: Apparently an oligolege
of Larrea tridentata, but visits other flowers presumably for nectar including
Chaenactis, D.fremontii, D.f. war. johnsonii, Eiiogonum fasciculatum, E. inflatum.
Perdita lateralis daleae Timberlake, i962. Calif. Univ. Pubs. Ent. 28: 29. 9, cJ. "

Perdita lateralis
***authority mismatch
lateralis Timberlake. Calif. (Colorado Desert), Ariz. (Maricopa, Pinal and Yuma
Counties). Pollen: Apparently an oligolege of La n-ea tridentata, but also visits other
flowers presumably for nectar including Chaenactis, Dalea califomica, Eriogonum
fasciculatum, Hyptis emoryi, Prosopis juliflora.
Perdita lateralis lateralis Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 27, figs. 749, 750,
853. ♀, ♂.

Biology: Hurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 25 (floral relationships).




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Perdita lucens Timberlake
South. Calif. (Mojave Desert), Nev. (Churchill County). Pollen: Possibly an
oHgolege of Eriogonum, visits flowers of E.fasciculatum, E. mflatum, E. rnohavense,
E. nodosum.
Perdita lucens Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 71, figs. 809, 810, 883. ♀, ♂.

Perdita lunulata Timberlake
Ariz. (Apache and Navajo Counties). Pollen: Collects pollen from the
flowers of Mentzelia puynila, but also visits the flowers oi Eriogonum aureum
presumably for nectar.
Perdita lunulata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 22. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 96, 101. 9, d (tax. status, key,
geogr. and floral relationships).

Perdita melanops Timberlake
South. Calif. (Santa Rosa Mts.). Pollen: Unknown, but visits flowers of
Physalis crassifolia.

Perdita melanops Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 105. ♀.

Perdita melanura Timberlake
Ariz. (Grand Canyon).

Perdita melanura Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 52. ♀.

Perdita mentzeliae Cockerell
Colo. (Antonita and Trinidad), N. Mex., Ariz., (Cochise, Greenlee and
Navajo Counties). Pollen: Collects pollen from the flowers of Mentzelia ptimila.

Perdita mentzeliae Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 60. ♂, ♀.

Perdita pallidior Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 61. ♂, ♀.

Perdita pulchrior Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 62. ♂.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 3, 10, 24-26, figs. 745, 746, 851. 9, 3
(key, redescription, synonymy, geogr. and floral records). —Timberlake, 1968. Calif. Univ.
Pubs. Ent. 49: 101 (geogr. and floral records).

Biology: Linsley and Hurd, 1959. Ent. News 70: 64, 65, 67 (floral relationships).

Perdita mentzeliarum Cockerell
West. Tex. to Ariz.; Mexico (Baja California and Chihuahua). Pollen:
Apparently an oligolege of Mentzelia pumila, but visits flowers of Bahia absinthifolia
for nectar.
Perdita mentzeliarum Cockerell, 1897. N. Mex. Agr. Expt. Sta., Bui. 24: 43. ♀, ♂.
Perdita mentzeliarum var. lauta Cockerell, 1905. Entomologist 38: 145. ♀, ♂.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 3, 9, 23-24, figs. 743, 744, 850. 9, S
(key, redescription, synonymy, geogr. and floral records). —Timberlake, 1968. Calif. Univ.
Pubs. Ent. 49: 101 (geogr. and floral records).

Perdita michelbacheri Timberlake
Ariz. (Maricopa County); Mexico (Baja California). Pollen:
Unknown, but visits flowers of Acacia, Camegiea gigantea.
Perdita michelbacheri Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 60, figs. 797, 798, 877.
9, 6.

Perdita nasuta
***authority mismatch
galacticoptera Timberlake. Calif. (Inyo and Panamint Mts.). Pollen: Apparently an
oligolege of Eriogonum, visits flowers of E. inflatum.
Perdita nasuta galacticoptera Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 76. ♀, ♂.

Perdita nasuta
***authority mismatch
nasuta Timberlake. South. Calif. (Colorado and Mojave Deserts), Nev. (Clark County),
Ariz. (Mohave, Pinal and Yuma Counties). Pollen: Apparently an oligolege of
Eriogonum, visits flowers of E. trichopes.
Perdita nasuta nasuta Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 74, figs. 813, 814, 885.
9,6.

Perdita nasuta
***authority mismatch
obscurescens Timberlake. Ariz., N. Mex. (Las Cruces). Pollen: Apparently an oligolege
of Eriogonum, visits flowers of E. thomasii, E. trichopes.
Perdita nasuta obscurescens Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 76. ♀, ♂.

Perdita nigridia Timberlake
Calif. (Colorado and Mojave Deserts), Ariz. (Mohave County); Mexico

(Baja California). Pollen: Apparently collects pollen only from the flowers of Mentzelia
including M. albicaulis, M. involucraia, M. pumila, M. tricuspis, but visits these and
apparently other flowers for nectar including Encelia farinosa, Eriogonum
fasciculatum, Eucnide urens.
Perdita nigridia Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 18, figs. 777, 778, 867. cJ, 9.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 101 (geogr. and floral record).




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Perdita nodosicornia Timberlake
Calif. (Coalinga).

Perdita nodosicomia Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 110, fig. 1360. ♂.

Perdita pectoralis Timberlake
Ariz. (Vermillion Cliffs). Pollen: Unknown, but visits flowers of
Eriogoniim.
Perdita pectoralis Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 111, figs. 1265, 1266, 1336.
6.

Perdita perplexa Timberlake
N. Mex. (Catron County). Pollen: Unknown, but visits flowers of
Mentzelia.
Perdita perplexa Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 19, figs. 739, 740, 848. 9 (6
misdet.).

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 101 (tax. status).

Perdita polytropica
***authority mismatch
obsoleta Timberlake. Ariz. (Maricopa, Pima and Pinal Counties). Pollen: Polylectic,
visits flowers of Acacia, Camegiea gigantea, Cercidium, Echinocactus.
Perdita polytropica obsoleta Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 59. ♀, ♂.

Perdita polytropica
***authority mismatch
polytropica Timberlake. South. Calif. (Colorado and Mojave Deserts). Pollen:
Polylectic, visits a wide variety of flowers including Acacia greggii, Agave deserti,
Asclepias subulata, Camegiea gigantea, Croton califomicus, Dalea spiyiosa, Encelia
farinosa, Echinocactus, Eriogonum inflatum, Ferrocactus acanthodes, Hyptis emoryi,
Larrea tridentata, Olneya tesota, Opuntia, Prosopis juliflora.
Perdita polytropica polytropica Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 57, figs. 795,
796,876. ♀, ♂.

Perdita punctifera Cockerell
Ariz., south. Calif. (Blythe); Mexico (Baja California). Pollen: Collects
pollen from the flowers of Mentzelia pumila, but also visits the flowers of Gossypium
thurberi presumably for nectar.
Perdita punctifera Cockerell, 1914. Ent. Soc. Wash., Proc. 16: 32. ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 3, 9, 26-27, figs. 747, 748, 852. 9, 6
(key, redescription, geogr. and floral records).

Perdita rehni
***authority mismatch
praeclara Timberlake. Ariz. (Southwestern Research Station).

Perdita rehni praeclara Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 106, fig. 14. ♀.

Perdita rehni
***authority mismatch
rehni Cockerell. N. Mex., western Tex.; Mexico (Chihuahua). Pollen: Unknown, but visits
flowers of Dasylirion wheeleri, Nolina.
Perdita Rehni Cockerell, 1907. Ann. and Mag. Nat. Hist. (7) 20: 131. 9 (= male).

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 8, 12, 50-52, figs. 789, 790, 873. 9, S
(key, redescription, geogr. and floral records).

Perdita rhois
***authority mismatch
reducta Cockerell. Calif. (South Coast Ranges). Pollen: Apparently polylectic, visits a
wide variety of flowers including Adeyiostoma fasciculatum, Eriodictyon parryi,
Eriogonum fasciculatum, Heteromeles arbutifolia, Rhus laurina.
Perdita rhois mut. reducta Cockerell, 1901. Canad. Ent. 33: 283. ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 6, 54-55. 9, 6 (key, redescription,

geogr. and floral records).

Perdita rhois
***authority mismatch
rhois Cockerell. South. Calif, (chiefly cismontane); Mexico (Baja California). Pollen:
Polylectic, visits a wide variety of flowers for pollen and nectar including Acacia
greggii, Adenostoma fasciculatum, A. sparsifolimn, Agave deserti, Alyssum
maritimum, Baccharis emoryi, B. viminea. Chrysanthemum, Eriogonum fasciculatum,
E.f var. polifolium, E. gracile. Euphorbia albomarginata, Gutierrezia califomica,
Hapiopappus linearifolius, Heliotropium oculatum, Heteromeles arbutifolia, Lotus
scoparius, Mentha, Nolina parryi. Polygonum lapanthifolium, Prunus ilicifolia,
Rhamnus califomicus, Rhus laurina, Rhus ovata, Schinus molle.

Perdita rhois Cockerell, 1901. Canad. Ent. 33: 282. ♀.

Perdita hypoxantha Cockerell, 1914. Ann. and Mag. Nat. Hist. (8) 13: 424. ♂.

Taxonomy: Timberlake, 1951. In Muesebeck, Krombein and Townes, U. S. Dept. Agr., Agr.
Monog. 2: 1094 (synonymy). —Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 6, 12, 53-54,
figs. 791, 792, 874. 9, 6 (key, redescription, geogr. and floral records).




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Perdita semicrocea Cockerell
N. Mex. (Dona Ana, Otero and San Miguel Counties), Ariz. (Willcox).

Pollen: Unknown, but visits flowers of Aster, Bahia ahsinthifolia, Giitierrezia sarothrae,
Haplopappus, Solidago.
Perdita semicrocea Cockerell, 1895. Acad. Nat. Sci. Phila., Proc. 47: 13. ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 4, 9, 35-36, figs. 761, 762, 859. 9, S
(key, redescription, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent.
49: 96, 101 (key, geogr. and floral record).

Perdita semilutea Timberlake
Calif. (Box Canyon in Riverside County). Pollen: Unknown, but. visits
flowers oi Eriogonum thomasii.
Perdita semilutea Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 81, figs. 819, 820, 888. ♂.

Perdita snellingi Timberlake
South. Calif. (Kern and Inyo Counties). Pollen: Presumably an oligolege
of vernal and autumnal flowering Compositae, visits flowers of Chrysothamnus
nauseosus co7isi7nilis, Cleome, Eriogonum heerniannii, Euphorbia albomarginata,
Gutierrezia lucida, G. microcephala, Rosa califortiica, Solidago spectabilis.
Perdita snellingi Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 46, figs. 781, 782, 869. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 106-107 (tax. characters, geogr. and
floral records).

Perdita sodalis Timberlake
South. Calif. (Santa Rosa Mts.). Pollen: Unknown, but visits flowers of
Hyptis emoryi.
Perdita sodalis Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 101, figs. 1255, 1256, 1331. ♂.

Perdita stabilis Timberlake
Ariz. (Douglas).

Perdita stabilis Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 102, figs. 1257, 1258, 1332. ♂.

Perdita subfasciata Cockerell
N. Mex. Ariz., Utah (Juab County), Nev. (White Pine County), Calif.
(Inyo County Mts.). Pollen: Apparently an oligolege of vernal and autumnal flowering
Compositae, visits flowers of Chrysothamnus nauseosus, C. viscidiflorus, Gutierrezia
lucida, G. microcephala.
Perdita subfasciata Cockerell, 1897. Ann. and Mag. Nat. Hist. (6) 20: 512. ♀.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 4, 11, 45-46, figs. 779, 780, 868. 9, 6
(key, redescription, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent.
49: 107 (geogr. and floral records).
thertnophila thermophila Timberlake. South. Calif. (Colorado and Mojave Deserts). Pollen:
Possibly an oligolege of Eriogonum, visits flowers of E. deserticola, E. hirtella, E.
inflatum, E. renifonne, E. thomasii.
Perdita thertnophila thermophila Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 77, figs.
815,816,886. ♀, ♂.

Perdita thermophila
***authority mismatch
trilobata Timberlake. Ariz., south. Calif. (Mojave Desert). Pollen: Possibly an

oligolege of Eriogonum, visits flowers of Baileya, Eriogonum deflexuyn, E. densum, E.
deserticola, E. trichopes.
Perdita thertuophila trilobata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 78. ♀, ♂.

Perdita timberlakei Cockerell
South. Calif, (cismontane and adjacent desert areas). Pollen: Evidently
depends mostly upon the pollen and nectar of Eriogonum gracile, but also visits flowers
of Brassica, Eriogonum fasciculatum, E. mohavense, E. virgatuni.
Perdita timberlakei Cockerell, 1925. Calif. Acad. Sci., Proc. (4) 14: 194. ♀, ♂.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 6, 13, 70-71, figs. 807, 808, 882. 9, 6
(key, redescription, geogr. and floral records).

Perdita varleyi
***authority mismatch
niveipennis Timberlake. Calif. (San Benito County). Pollen: Apparently an oligolege of
Eriogonum including E. inflatum.
Perdita varleyi niveipennis Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 86. ♀, ♂.

Perdita varleyi
***authority mismatch
varleyi Timberlake. South. Calif. (Mojave Desert, western margin). Pollen: Apparently
an oligolege of Eriogonum, visits flowers of E . fasciculatum var. polifolium, E.
inflatum.
Perdita varleyi varleyi Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 85, figs. 825, 826, 891,
904. ♂, ♀.




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Perdita vicina Timberlake
Nev. (Charleston Mts.). Pollen: Unknown, but visits flowers of Falliigia
paradoxa.
Perdita vicma Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 31, figs. 753, 754, 855. ♂.

Perdita viridinotata Timberlake
N. Mex. (Alamogordo).

Perdita viridinotata Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 16, figs. 735, 736, 846. ♀.
6.

wheeler! Timberlake. Ariz. (Grand Canyon). Pollen: Unknown, but possibly may visit flowers
of Mentzelia.
Perdita wheeleri Timberlake, 1928. Amer. Mus. Novitates 321: 5. ♂.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 10, 32, figs. 755, 756, 856. 6
(redescription, possible pollen source).

Perdita wootonae Cockerell
N. Mex., Colo., Kans., Nebr. Pollen: Apparently an oligolege of Mentzelia
including M. decapetala, M. nuda, but visits other flowers presumably for nectar
including Eriogonum effusuyn, Tragopogon porrifolius.
Perdita wootonae Cockerell, 1898. Ent. News 9: 215. ♀, ♂.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 2, 8, 15-16, figs. 733, 734, 845. 9, 6
(key, redescription, geogr. and floral records).
xanthoxyii Timberlake. Tex. (Corpus Christi State Park); Mexico (Puebla). Pollen: Unknown,
but visits flowers of Xanthoxy Ion.
Perdita xanthoxyii Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 107, figs. 15, 1261, 1262,
1334. ♀, ♂.

Perdita xerophila
***authority mismatch
discrepans Timberlake. Nev. (Clark County). Pollen: Possibly an oligolege of
Eriogonum, visits flowers oi E.fasciculatum, E. trichopes.
Perdita xerophila discrepaiis Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 84, figs. 823,
824,890. ♀, ♂.

Perdita xerophila
***authority mismatch
fuscicornis Timberlake. Calif. (Inyo County Mts.). Pollen: Possibly an oligolege of
Eriogonum, visits flowers of E. inflatum, Lan-ea tridentata.
Perdita xerophila fuscicornis Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 84, fig. 903. ♀.
6.

Perdita xerophila
***authority mismatch
xerophila Timberlake. Ariz. (Yuma County), south. Calif. (Colorado Desert). Pollen:
Possibly an oligolege of Eriogonum, visits flowers of ^. inflatum, E. trichopes.
Perdita xerophila xerophila Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 82, fig. 902. ♀, ♂.

Perdita yosemitensis Timberlake
Calif. (Sierra Nevada Mts.). Pollen: Possibly an oligolege of
Eriogonum, visits flowers of E. nud^im, E. wrightii.
Perdita yosemitensis Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 63, figs. 801, 802, 879.
6, 9.

Species Group Sphaeralceae

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 1 (key). —Timberlake, 1964. Calif.
Univ. Pubs. Ent. 28: 126-158 (key to included spp.). —Timberlake, 1971. Calif. Univ. Pubs.
Ent. 66: 38-42 (suppl. key to included spp.).

Perdita adustiventris Timberlake
South. Calif. (Parker Dam); Mexico (Baja California: Angel de
la Guardia Island). Pollen: Unknown, but visits flowers of Eucnide urens, Mentzelia
hirsutissima.
Perdita adustiventris Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 166. ♀.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 42-43, figs. 5, 1385, 1386, 1411. 6
(geogr. and floral records).

Perdita albiventris Timberlake
Tex. (Pecos County). Pollen: Unknown, but the type specimen bears a
load of fine, mealy fulvous pollen, probably leguminaceous.
Perdita albiventris Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 294, fig. 60. ♀.

Perdita albofasciata Timberlake
N. Mex. (Santa Fe). Pollen: Unknown, but visits flowers of Tamarix
gallica.
Perdita albofasciata Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 152. ♀.




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Perdita ambigua Timberlake
Nev. (Lander and Washoe Counties), Calif. (Siskiyou County). Pollen:
Unknown, but visits flowers of Chrysotlmmyius nanseosus var. consiniilis, Cirshirn
lanceolatian.
Perdita ambigua Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 311. ♀.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 54-55. 6 (geogr. and floral records).

Perdita amicula Timberlake
Ariz. (Willcox). Pollen: Unknown, but visits flowers of Aplopappus
liartwegi.
Perdita amicula Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 295. ♀.

Perdita ancoralis Timberlake
South. Calif. (Boyd Desert Research Center).

Perdita ancoralis Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 155. ♀.
ashmeadi ashmeadi Cockerell. Tex. (Brownsville); Mexico (Sonora). Pollen: Unknown, but
visits flowers of Prosopis.
Perdita ashmeadi Cockerell, 1899. Ann. and Mag. Nat. Hist. (7) 8: 492. ♀.
Perdita exclamans atramentata Cockerell, 1923. Calif. Acad. Sci., Proc. (4) 12: 96. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 141, 158, 324-325, figs. 1069, 1070,
1178. 9, 6 (key, redescription, synonymy, geogr. and floral records).
ashmeadi simulans Timberlake. South. Calif. (Colorado Desert), ?Ariz. (Sentinel); Mexico
(Baja California). Pollen: Unknown, but visits flowers of Geraea canescens, Larrea
tridentata, Prosopis jiiliflora.
Perdita ashmeadi simulans Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 326, figs. 1071,
1072, 1179. ♂.
ashmeadi vierecki Cockerell. West. Tex., N. Mex. (Alamogordo), Ariz. Pollen: Unknown, but
visits flowers of Mimosa, Prosopis.
Perdita Vierecki Cockerell, 1907. Ann. and Mag. Nat. Hist. (7) 20: 129. ♀, ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 141, 158, 325-326. 9, S (key,
redescription, tax. status, geogr. and floral records).

Perdita assimilis Timberlake
Calif. (Fresno County). Pollen: Unknown, but visits flowers of Melilotus.

Perdita assimilis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 228, figs. 37, 987, 988, 1137.
9, 6.

Perdita atrata Timberlake
Calif. (Colorado and Mojave Deserts). Pollen: Unknown, but visits flowers
of Camissonia brevipes, C. dentata var. parishii, Chaenactis stevioides, Mentzelia
affinis, M. albicaiilis, M. veatchiana, Sphaeralcea ambigua.
Perdita atrata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 163, figs. 3, 911, 912, 1009. ♀.
6.

Perdita barri Timberlake
Idaho (Midvale). Pollen: Unknown, but visits flowers of Phacelia.

Perdita barri Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 251, figs. 1013, 1014, 1150. ♂.

Perdita blanda Timberlake
N. Mex. (Albuquerque).

Perdita blayida Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 358. ♀.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 137, 262 (key, tax. status).

Perdita calloleuca
***authority mismatch
calloleuca Cockerell. N. Mex., Colo., Utah, Ariz. Pollen: Unknown, but visits

flowers of Artemisia filifolia, Chrysothamnus, Cleome lutea, Eriogonum, Salsola kali,
Tamarix, Trifolium, Wislizenia refracta.
Perdita calloleuca Cockerell, 1922. Amer. Mus. Novitates 33: 12. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 140, 154, 298-300, figs. 63, 1059,
1060, 1173. 9, 6 (key, redescription, geogr. and floral records). —Timberlake, 1966. Calif.
Univ. Pubs. Ent. 66: 43 (geogr. and floral records).

Perdita calloleuca
***authority mismatch
convergens Timberlake. N. Mex. (Virden), Ariz. (Graham and Yuma Counties), Nev.
(Clark County). Pollen: Unknown, but visits flowers of Atriplex canescens,
Chenopodium ohlongifolium, Euphorbia, Suaeda, Wislizenia refracta.
Perdita calloleuca convergens Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 300. ♀, ♂.

Perdita chihuahua Timberlake
N. Mex. (Rodeo); Mexico (Chihuahua and Zacatecas).

Perdita chihuahua Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 203, figs. 955, 956, 1121.
9,6.




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Perdita chionostoma Timberlake
Calif. (Fresno County). Pollen: Unknown, but visits flowers of
Melilotiis.
Perdita chionostoma Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 240, figs. 43, 1001, 1002,
1144. ♀, ♂.

Perdita cleomellae Cockerell
Calif. (Mojave Desert). Pollen: Apparently an oligolege of Cleomella
including C. obtusifolia.
Perdita cleomellae Cockerell, 1925. Calif. Acad. Sci., Proc. (4) 14: 193. ♀, ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 309, 315, 353-354, figs. 430, 431, 499.
9 , 6 (key, redescription, geogr. and floral records).

Perdita cochiseana Timberlake
Ariz. (Douglas). Pollen: Unknown, but visits flowers of Euphorbia.

Perdita cochiseana Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 43. ♀.

Perdita compta Timberlake
South. Cahf. (Colorado and Mojave Deserts). Pollen: Unknown, but most
likely source is Eriastnim; visits flowers oi Cryptantha intermedia, Eriastrum
virgatnm, Eriogonum reniforme, E. trichopes, Lepidium fremontii, Malacothrix
glabrata, Stilli^igia paucidenfata.
Perdita compta Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 236, figs. 41, 997, 998, 1142.
9, d.

Perdita confusa Timberlake
West. Tex., N. Mex., Ariz.; Mexico (Chihuahua). Pollen: Apparently an
oligolege of Lepidium including L. alyssoides, L. montanum, L. thurberi.
Perdita confusa Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 247, figs. 46, 1007, 1008, 1147.
9, 6.

Taxonomy: Rozen, 1966. Amer. Mus. Novitates 2259: 19, fig. 50 (larva, as questionably this
species).

Biology: Rozen, 1967. Amer. Mus. Novitates 2297: 38, fig. 15, table 1 (nest architecture, life
history, floral association).

Perdita covilleae Timberlake
Ariz., Utah (St. George), Nev. (Las Vegas), south. Calif. (Colorado and
Mojave Deserts); Mexico (Baja California). Pollen: Oligolege oi Larrea tridentata, also
visits other flowers for nectar including Heliotropium curassavicum, Prosopis.
Perdita covilleae Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 361, figs. 438, 439, 503. ♀.
S.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 139, 262 (key).

Biology: Hurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 24, fig. 5 (intrafloral ecology).

Perdita cruciferarum Timberlake
Nev. (Battle Mountain). Pollen: Unknown, but visits flowers of an
unidentified Cruciferae.
Perdita cruciferarum Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 140, figs. 24, 25, 1291,
1292, 1349. ♀, ♂.

Perdita cushmani Timberlake
Tex. (Brewster and Hudspeth Counties); Mexico (Chihuahua). Pollen:
Unknown, but visits flowers of Gutierrezia, Prosopis juliflora.
Perdita cushmani Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 222, figs. 979, 980, 1133. ♀.
6.

Perdita cuspidata Timberlake
South. Calif. (Colorado and Mojave Deserts). Pollen: Collects pollen
from the flowers of Nama demissum, Phacelia distans, but visits other flowers for
nectar including Oenothera.
Perdita cuspidata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 231, figs. 39, 991, 992, 1139.
9, 6.

Perdita dammersi Timberlake
South. Calif. (Colorado and Mojave Deserts). Pollen: Collects pollen
from flowers of Malacothrix including M. glabrata, but visits these and other flowers
for nectar including Chaenactis fremontii, Stephanorneria exigua.
Perdita dammersi Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 169, figs. 7, 919, 920, 1103.
9, cJ.

Perdita davidsoni Timberlake
South. Calif.

Perdita davidsoni Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 235, figs. 995, 996, 1141. ♂.




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Perdita deltophora Timberlake
Tex. Pollen: Unknown, but visists flowers of Cha7naesaracha
co7iioides, Physalis lobata, Prosopis.
Perdita deltophora Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 207, figs. 25, 959, 960,
1123. ♀, ♂.

Perdita dentata Timberlake
South. Calif. (Colorado Desert). Pollen: Possibly an oligolege of Phacelia
including P. distans, but visits other flowers presumably for nectar including Acacia
greggii, Geraea ca^tescens, Kmmeria canescens, Larrea tridentata.
Perdita dentata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 165, figs. 4, 913, 914, 1100. ♀.

S.

Perdita difficilis Timberlake
Tex. (Brewster County), N. Mex. (Dona Ana and Eddy Counties), Ariz.,
south. Calif. (Colorado and Mojave Deserts); Mexico (Baja California). Pollen:
Apparently an oligolege of Prosopis including P.juliflora, but visits flowers of other
desert shrubs presumably for nectar including Cercidium floridum, Larrea tridentata.
Mimosa and also has been collected at the flowers of Melilotns.
Perdita difficilis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 323, figs. 1067, 1068, 1177.
6, 9.

Perdita digna Timberlake
CaHf. (Quincy).

Perdita digna Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 310. ♀.

Perdita discors Timberlake
South. Calif. (Thousand Palms).

Perdita discors Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 361. ♀.

Perdita dispar Timberlake
Tex. (Starr and Webb Counties); Mexico (Chihuahua). Pollen: Unknown,
but visits flowers of Condalia lycioides, Croton, Prosopis juliflora.
Perdita dispar Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 220^ figs. 33, 997, 998, 1132. ♀.
6.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 43: 136 (geogr. and floral record).

Perdita distincta Timberlake
Oreg. (Baker County), Calif. (Sierra County). Pollen: Unknown, but
visits flowers of Eriogonum.
Perdita distincta Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 174, figs. 10, 925, 926, 1106.

6.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 45-46. ♀.

Perdita drymariae Timberlake
N. Mex. (Dona Ana and Hidalgo Counties), Mich. (Baraga County);

Mexico (San Luis Potosi). Pollen: Unknown, but visits flowers of Dryntaria holosteoides,
Tidestro7)tia lanuginosa.
Perdita drymariae Timberlake, 1960. In Mitchell, N. C. Agr. Expt. Sta. Tech. Bui. 141: 324.
9, 6.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 129, 148, 201-202, figs. 22, 951, 952,
1119. 9, 6 (key, tax. characters, geogr. and floral records). —Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 112, 134 (key, geogr. and floral record).

Perdita emarginata Timberlake
Ariz. (Tucson), south. Calif. (Desert Center). Pollen: Unknown, but
visits flowers of Dalea.
Perdita emarginata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 304. ♀.

Perdita eremica Timberlake
Ariz. (Maricopa and Pinal Counties), south. Calif. (Colorado and Mojave
Deserts). Pollen: Apparently an oligolege of Larrea tridentata, but visits other flowers
including Dalea californica, D.fremontii var. jolinsoyiii, D.f. var. sau7idersii, D. schottii,
Eriogo7iuni inflatum, Funastruni hirtellu7n, Hyptis emoryi, Nama de7nissum.
Perdita eremica Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 268, figs. 54, 1033, 1034, 1160.
9,6.

Biology: Hurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 25 (floral relationships).

Perdita eremophila Timberlake
South. Calif. (Mojave Desert). Pollen: Possibly an oligolege of
Phacelia including P. distans, P. fremontii, but visits other flowers including
Haplopappus cooperi, Lepidium fremontii.
Perdita eremophila Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 238, figs. 42, 999, 1000,
1143. ♀, ♂.




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Perdita eriastri Timberlake
South. Calif, (cismontane). Pollen: Collects pollen from the flowers of
Eriastrnm including E. virgatum, but visits other flowers for nectar including
Cryptantha intermedia, Eriogonum fasciculatum, Gilia.
Perdita eriastri Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 233, figs. 40, 993, 994, 1140.
9,6.

Perdita eriastri
***authority mismatch
var. fusciventris Timberlake. South. Calif. (Palm Springs). Pollen: Collects pollen from
flowers o{ Eriastnon virgatum. A specimen similar to the holotype has been collected
in Arroyo Seco, Los Angeles County at the flowers oi Eriodictyon parryi.
Perdita eriastri var. fusciventris Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 235. ♀.

Perdita eucnides
***authority mismatch
eucnides Timberlake. Calif. (Inyo County), Nev. (Churchill County). Pollen:

Polylectic, collects pollen from the flowers of Cowania mexicana var. stansburiana,
Dalea polyadenia, Eucnide urens, Larrea tridentata, but also visits other flowers
including Eriogonum inflatum possibly for nectar.
Perdita eucnides eucnides Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 301, figs. 64, 1061,
1062, 1174. ♂, ♀.

Perdita eucnides
***authority mismatch
platyzona Timberlake. Ariz. (Yuma County), south. Calif. (Colorado Desert); Me.xico
(Baja California). Pollen: Polylectic, visits flowers of Acacia greggii, Lan-ea tridentata,
Pluchea sericea, Sesuviu^n verrucosum.
Perdita euc7iides platyzona Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 304. ♀, ♂.

Perdita euzonata Timberlake
Ariz. (Painted Desert). Pollen: Unknown, but visits flowers of
Eriogonum aureum.
Perdita euzonata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 295. ♀.

Perdita exclamans Cockerell
West. Tex. to south. Calif. (Colorado and Mojave Deserts) and south

Nev. (Clark County); Mexico (Baja California and Sonora). Pollen: Collects pollen from
the flowers of Prosopis, but also visits other flowers presumably for nectar including
Baccharis, Cercidium florid um, Dithyrea wislizeni, Lan-ea tridentata, Melilotus,
Mimosa, Phoradendron.
Perdita nitidella var. exclamans Cockerell, 1895. Psyche 1 (sup.): 5. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 141, 157, 158, 320-323, figs. 70, 1065,
1066, 1176. 6, 9 (key, redescription, geogr. and floral records).

Perdita exigua
***authority mismatch
becki Timberlake. Nev. (Nye and White Pine Counties).

Perdita exigua becki Timberlake, 1971 Calif. Univ. Pubs. Ent. 66: 44. ♀, ♂.

Perdita exigua
***authority mismatch
exigua Timberlake. Calif. (Hallelujah Junction).

Perdita exigua Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 278, figs. 1043, 1044, 1165. ♀.
6.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 44 (tax. status).

Perdita eximia Timberlake
Ariz. (Maricopa and Yuma Counties), south. Calif. (Colorado and Mojave
Deserts), Nev. (Esmeralda County). Pollen: Possibly an oligolege of Phacelia including
P. crenulata, but visits a variety of other flowers including Acacia greggii, Camisso7iia
clavaeformis, Cryptantha angustifolia, Cercidium florid um, Dalea schottii, Encelia,
Eriogonum inflatum, E. thomasii, Heliotropium curassaricutn, Iso7neris arborea,
Larrea tridentata, Prosopis juliflora.
Perdita eximia Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 255, figs. 49, 1019, 1020, 1153.
9,6.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 145 (key).

Perdita eysenhardtiae Timberlake
Tex. (Alpine), Ariz. (Cochise and Santa Cruz Counties); Mexico
(Distrito Federal, Durango and Hidalgo). Pollen: Unknown, but visits flowers of
Baccharis glutinosa, Eysenhardtia polystachya.
Perdita eysenhardtiae Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 215. ♀.

Perdita falcata Timberlake
Calif. (Inyo County). Pollen: Unknown, but visits flowers of Dalea
fremontii, Encelia farinosa, Gilia, Sphaeralcea.
Perdita falcata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 161, figs. 2, 909, 910, 1098. ♀.
6.




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Perdita flavipes Timberlake
South. Calif. (Colorado and Mojave Deserts); Mexico (Baja California).
Pollen: Apparently an oligolege of Larrea tridentata, but visits other flowers including
Heliotropium cumssavicum, Prosopis juliflora.
Perdita flavipes Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 259, figs. 50, 1023, 1024, 1155.

6, 9.

Biology: Hurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 24 (floral relationships).

Perdita florissantella Cockerell
Colo., N. Mex. (Taos County). Pollen: Collects pollen from the flowers
of Lepidium jonesi, but visits other flowers presumably for nectar including
Chenopodinm albu7n, Eriogoimm umbellatum, Phacelia.
Perdita florissantella Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 440. ♂, ♀.
Perdita lepidii Cockerell, 1907. Entomologist 40: 266. ♀.
Perdita opacifrons Timberlake, 1929. N. Y. Ent. Soc, Jour. 37: 113. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 137, 151, 242-244, figs. 44, 1003,
1004, 1145. 9, 6 (key, redescription, synonymy, geogr. and floral records). —Timberlake,
1968. Calif. Univ. Pubs. Ent. 49: 139 (key).

Perdita foxi Cockerell
N. Mex. (Santa Fe).

Perdita foxi Cockerell, 1895. Acad. Nat. Sci. Phila., Proc. 47: 18. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 152, 317. 6 (key, redescription).

Perdita gemella Timberlake
Calif. (Apple Valley and Deep Creek on Mojave Desert). Pollen:
Unknown, but visits flowers oi Eriodictyon trichocalyx.
Perdita gemella Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 200, figs. 949, 950, 1118. ♂.

Perdita geminata Timberlake
N. Mex. (Otero County). Pollen: Unknown, but visits flowers of Larrea
tridentata.
Perdita geminata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 251, figs. 1015, 1016, 1151.
9, 6.

Perdita genalis
***authority mismatch
genalis Timberlake. South. Calif. (Colorado and Mojave Deserts). Pollen: Unknown,
but visits flowers of Acacia greggii, Cercidium floridum, Larrea tridentata, Prosopis
juliflora.
Perdita genalis genalis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 359, figs. 76, 1093,
1094, 1190. ♀, ♂.

Perdita genalis
***authority mismatch
panamintensis Timberlake. Calif. (Panamint Mts.). Pollen: Unknown, but visits flowers
of Larrea tridentata, Prosopis juliflora, Stanleya pinnata.
Perdita genalis panamintensis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 360. ♀, ♂.

Perdita gracilis Timberlake
Calif. (Lone Pine). Pollen: Unknown, but visits flowers of Malacothrix.

Perdita gracilis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 309. ♀.

Perdita greggiae Timberlake
Tex. (Reeves County), N. Mex. (Eddy County). Pollen: Unknown, but
visits flowers of Lepidium montanum, Lesquerella gordonii, Nerisyi-enia campoi^utn.
Perdita greggiae Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 142, figs. 1293, 1294, 1350. ♀.
6.

Perdita heliotropii
***authority mismatch
heliotropii Cockerell. Tex. (El Paso) west to south. Calif. (Colorado and Mojave
Deserts), Nev. (Lyon County), Utah (St. George).; Mexico (Baja California, Sonora,
Chihuahua). Pollen: Collects pollen from the flowers of Heliotropium including H.
curassavicuni, but visits these and other flowers for nectar including Larrea tridentata,
Salix, Tamarix.
Perdita heliotropii Cockerell, 1900. Entomologist 33: 63. ♀, ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 309, 314, 364-365, figs. 442, 443, 505,
521. 9, 6 (key, tax. characters, tax. status, geogr. and floral records). —Timberlake, 1964.
Calif. Univ. Pubs. Ent. 28: 139, 262 (key, tax. characters). —Timberlake, 1968. Calif. Univ.
Pubs. Ent. 49: 116, 148 (key, tax. characters, geogr. record).

Perdita heliotropii
***authority mismatch
perducta Timberlake. Central CaHf. Pollen: Collects pollen from the flowers of

Heliotropium curassavicuni, but also visits flowers of Stanleya pinnata presumably for
nectar.
Perdita heliotropii perducta Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 365. ♀, ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 139, 262 (key, geogr. and floral
record).




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Perdita humilis Timberlake
N. Mex. (Mesilla Park). Pollen: Unknown, but visits flowers of Dithyrea
wislizeni.
Perdita humilis Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 156. ♀.

Perdita imberbis Timberlake
Calif. (Sierra and Siskiyou Counties). Pollen: Unknown, but visits
flowers of Erigeron.
Perdita imberbis Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 139, fig. 23. ♀.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 46-47, figs. 1387, 1388, 1412. 6, 9
(key, geogr. and floral record).

Perdita impigra Timberlake
Tex. (San Jacinto County).

Perdita impigra Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 157, fig. 31. ♀.

Perdita impressa Timberlake
Ariz. (Apache County). Pollen: Unknown, but visits flowers of
Eriogonum aureum.
Perdita impressa Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 153. ♀.

Perdita incompta Timberlake
Tex. (Southmost). Pollen: Unknown, but visits flowers of Lippia.

Perdita incompta Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 208, figs. 26, 961, 962, 1124.
9, 6.

Perdita infelix Timberlake
Nebr. (Harrison).

Perdita infelix Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 314. ♀.

Perdita innotata Timberlake
Ariz. (Maricopa and Yuma Counties), south. Calif. (Colorado and Mojave
Deserts). Pollen: Possibly an oligolege of Prosopis including P. juliflora, but also visits
flowers of Na7na hispidum.
Perdita innotata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 333, figs. 1081, 1082, 1184.
9, 6.

Perdita insequens Timberlake
South. Calif. (Piute Butte in Mojave Desert). Pollen: Unknown, but
visits flowers of Chaenactis brachypappa.
Perdita insequens Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 146, fig. 27. ♀.

Perdita koebelei
***authority mismatch
concinna Timberlake. South. Calif. (Colorado and Mojave Deserts), Ariz. (Mohave
and Yuma Counties). Pollen: Apparently an oligolege of Mentzelia including M.
involucrata, M. tricuspis, but also visits flowers of Eucnide iirens, Hyptis emoryi.
Perdita koebelei concinna Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 161, figs. 907, 908.

6, 9.
Perdita masoni Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 306. ♀.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 47 (synonymy, geogr. and floral
records).

Perdita koebelei
***authority mismatch
koebelei Timberlake. Calif. (Inyo County). Pollen: Collects pollen from the flowers of
Eucnide urens, Mentzelia, but also visits other flowers presumably for nectar including
Argemone platyceras, Encelia farinosa, Eriogonum inflatum, Phacelia caltliifolia,
Stepkanomeria.
Perdita koebelei koebelei Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 159, figs. 1, 905, 906,
1097. cJ, 9.

Perdita krombeini Timberlake
Fla. (Lee County).

Perdita krombeini Timberlake, 1960. In Mitchell, N. C. Agr. Expt. Sta. Tech. Bui. 141: 327.
6.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 144, 167, figs. 915, 916, 1101. S (key,
tax. characters).

Perdita leucogastra Timberlake
Tex. (Terrell County). Pollen: Unknown, but visits flowers of Gilia
acerosa.
Perdita leucogastra Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 280, figs. 1045, 1046,
1166. ♀, ♂.

Perdita leucosticta Timberlake
South. Calif. (San Jacinto Mts. and environs). Pollen: Possibly an
oligolege of Eriastnim virgatum, but visits other flowers including Eriogonum
fasciculatum, Gilia exilis, Layia platyglossa.
Perdita leucosticta Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 224, figs. 34, 981, 982,
1134. ♀, ♂.




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Perdita linsleyi Timberlake
South. Calif. (Los Angeles and Ventura Counties). Pollen: Unknown, but
visits flowers of Chaenactis, Euphorbia albomargmata, Lasthenia chrysostoma.
Perdita linsleyi Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 199, figs. 21, 947, 948, 1117.
9,6.

Perdita luciae
***authority mismatch
decora Timberlake. Ariz. (Maricopa, Pima and Yuma Counties), Nev. (Clark County),
south. Calif. (Colorado Desert); Mexico (Baja California and Sonora). Pollen: Polylectic,
especially flowers of Larrea tridentata and Prosopis juliflora, but also visits a wide
variety of other flowers including Acacia greggii, Baileya pleniradiata, Cercidium
floridum, Cryptantha angustifolia, C. intermedia, Naina hispidum, Psilostrophe
cooperi, Sphaeralcea emoryi.
Perdita luciae decora Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 336, fig. 73. ♀, ♂.

Biology: Hurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 25 (floral relationships).

Perdita luciae
***authority mismatch
luciae Cockerell. Tex. (Pecos County), N. Mex. (Las Cruces), Ariz. (Cochise, Maricopa
and Pima Counties); Mexico (Sonora). Pollen: Unknown, but visits flowers of Medicago
sativa, Mimosa, Prosopis.
Perdita luciae Cockerell, 1899. Ann. and Mag. Nat. Hist. (7) 8: 494. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 141, 156, 157, 334-336, figs. 72, 1083,
1084, 1185 (key, tax. status, redescription, geogr. and floral records).

Perdita lycii Timberlake
Ariz. (Pima County). Pollen: Unknown, but visits flowers of Lycium.

Perdita lycii Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 305, fig. 65. ♀.

Perdita macneilli Timberlake
Calif. (Santa Clara County).

Perdita macneilli Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 312. ♀.

Perdita macswaini Timberlake
Tex. (Cameron and Harligen Counties), N. Mex. (Eddy County).
Pollen: Unknown, but visits flowers of Coreopsis douglasii, Gaillardia, Monarda
citriodora, Prosopis.
Perdita macswaini Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 289, figs. 1053, 1054, 1170.
9,6.

Perdita macswaini
***authority mismatch
var. flavolineata Timberlake. Tex. (Hidalgo County).

Perdita macswaini var. flavolineata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 290. ♂.

Perdita martini Cockerell
West. Tex., N. Mex. (Dona Ana County). Pollen: Unknown, but visits
flowers of Sesuvium vemicosum.
Perdita martini Cockerell, 1895. Acad. Nat. Sci. Phila., Proc. 47: 14. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 140, 155, 287, figs. 58, 1051, 1052,
1169. 9, 6 (key, tax. status, redescription, geogr. and floral records).

Perdita melanogastra Timberlake
Calif. (Tulare County).

Perdita melanogastra Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 313, fig. 67. ♀.

Perdita mimosae Timberlake
Tex., N. Mex. (Eddy County). Pollen: Unknown, but visits flowers of
Mimosa, Phacelia popei, Prosopis.
Perdita mimosae Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 270, figs. 1035, 1036, 1161.
9,6.

Perdita modestissima Timberlake
Nev. (Battle Mountain). Pollen: Unknown, but holotype is labeled
from an undetermined species of Cruciferae.
Perdita modestissima Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 147, fig. 28. ♀.

Perdita munita Timberlake
Ariz. (Cochise County); Mexico (Durango). Pollen: Unknown, but visits
flowers of Chamaesaracha coronopns, Eriogonum, Physalis.
Perdita munita Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 205, figs. 24, 957, 958, 1122.
9, 6.

Perdita nigricornis Timberlake
Ariz. (Cochise and Pima Counties); Mexico (Sonora). Pollen:
Apparently may collect pollen from the flowers of Prosopis.
Perdita nigricomis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 331, figs. 1077, 1078, 1182.
6.

Perdita nigroaenea Timberlake
Utah (Kane County). Pollen: Unknown, but visits flowers of
Gutierrezia microcephala.
Perdita nigroaenea Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 158. ♀.




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Perdita nigroclypeata Timberlake
Colo. (Alamosa and Rio Grande Counties).

Perdita nigroclypeata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 314, fig. 69. ♀.

Perdita obliqua Timberlake
Ark. (Garland County), Ariz., south. Calif. (Colorado Desert); Mexico
(Baja California and Sonora). Pollen: Apparently polylectic, visits flowers of Acacia,
Carnegiea gigantea, Cercidium, Dalea spinosa, Eriogonum trichopes, Medicago sativa,
Prosopis, Wislizenia refracta.
Perdita obliqua Timberlake, 1928. Pan-Pacific Ent. 5: 26. ♀.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 134, 144, 148, 218-219, figs. 975, 976,
1131. 9, 6 (key, redescription, geogr. and floral records). —Timberlake, 1971. Calif. Univ.
Pubs. Ent. 66: 48 (geogr. record).

Perdita obscurella Timberlake
N. Mex. (Hidalgo and Socorro Counties), Ariz. (Cochise and Pima
Counties); Mexico (Chihuahua). Pollen: Unknown, but visits flowers of Baileya
pleniradiata, Boerhaavia, Euphorbia albornarginata.
Perdita obscurella Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 202, figs. 23, 593, 594,
1120. ♀, ♂.

Perdita omani Timberlake
Calif. (Bakersfield).

Perdita omani Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 296, fig. 62. ♀.

Perdita oreophila Timberlake
Calif. (Tuolumne County). Pollen: Unknown, but visits flowers of a
Compositae.
Perdita oreophila Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 244. ♀.

Perdita ovaliceps Timberlake
N. Mex. (Eddy County), Ariz. (Maricopa and Mohave Counties), Nev.

(Clark County), south. Calif. (Colorado and Mojave Deserts). Pollen: Unknown, but visits
flowers oi Chaenactis carphoclinia, C. sfevioides var. brachypappa, Encelia farinosa,
Gerrea canescens, Larrea tridentata.
Perdita ovaliceps Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 265, figs. 52, 1029, 1030,

1158. ♂.
Perdita iuctuosa Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 266, fig. 53. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 150-151 (synonymy, geogr. records).

Perdita pallidipes Timberlake
Ariz. (Maricopa and Pima Counties), south. Calif. (Colorado Desert);
Mexico (Sonora). Pollen: Unknown, but visits flowers of Prosopis juliflora.
Perdita pallidipes Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 332, figs. 1079, 1080, 1183.
6.

Perdita panocheana Timberlake
Calif. (Fresno County). Pollen: Unknown, but visits flowers of
Heliotropium oculatum.
Perdita panocheana Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 274, figs. 1039, 1040,
1163. ♀, ♂.

Perdita parryellae Timberlake
Ariz. (Navajo County). Pollen: Unknown, but visits flowers of
Parryella fiiifolia.
Perdita parryellae Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 48, figs. 1389, 1390, 1413.
9, 6.

Perdita placens Timberlake
Ariz. (Willcox).

Perdita placens Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 124, figs. 1273, 1274, 1340. ♂.

Perdita planifrons Timberlake
Calif. (Mono County).

Perdita planifrons Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 148. ♀.

Perdita propinqua Timberlake
Calif. (Ventura County). Pollen: Unknown, but visits flowers of
Linanthus aureus, Phacelia douglasii.
Perdita propinqua Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 226, figs. 35, 983, 984,
1135. ♀, ♂.

Perdita prosopidis Timberlake
Ariz. (Yuma County), south. Calif. (Colorado and Mojave Deserts);
Mexico (Baja California). Pollen: Possibly an oligolege of Prosopis juliflora, but visits
other flowers for nectar including Melilotus.
Perdita prosopidis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 318, figs. 69, 1063, 1064,
1175. ♀, ♂.

Perdita puncticeps Timberlake
Calif. (Palm Springs).

Perdita puncticeps Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 49. ♀.




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Perdita punctosignata
***authority mismatch
flava Timberlake. Tex. west, to Ariz.; Mexico (Baja California, Chihuahua,
Coahuila, Sonora and Sinaloa). Pollen: Apparently an oligolege of Prosopis, but also
visits other flowers presumably for nectar including Gutierrezia, Medicago, Sapiyidus
saponaria.

Perdita punctosigyiata flava Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 354. ♀, ♂.

Perdita punctosignata
***authority mismatch
punctosignata Cockerell. N. Mex. (Dona Ana and Otero Counties), Ariz., Calif.
(Colorado Desert); Mexico (Baja California and Sonora). Pollen: Apparently an oligolege
of Prosopis.

Perdita punctosignata Cockerell, 1895. Psyche 7 (supp.): 6. ♂.

Perdita howardi Cockerell, 1899. Ann. and Mag. Nat. Hist. (7) 3: 492. ♀, ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 142, 157, 351-353, figs. 75, 1087,
1088, 1187. 9, 6 (key, redescription, synonymy, geogr. and floral records).

Perdita punctosignata
***authority mismatch
sulphurea Timberlake. Ariz., south. Calif. (Colorado and Mojave Deserts, also
San Benito County), Nev. (Clark County), Utah (Washington County); Mexico (Baja
California). Pollen: Apparently an oligolege of Prosopis, but also visits other flowers
presumably for nectar including Acacia greggii, Cercidium floridum, Larrea tridentata,
Melilotus.
Perdita punctosignata sulphurea Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 353. ♀, ♂.

Perdita punctulata Timberlake
South. Calif. (Colorado and Mojave Deserts), Ariz. (Yuma); Mexico

(Baja California and Sonora). Pollen: OUgolege of Larrea tridentata, but also visits other
flowers for nectar including Bacckaris emoryi, C ercidium floridum, Lasthenia
chrysostoma, Melilotus, Prosopis juliflora.
Perdita punctulata Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 362, figs. 440, 441, 504. ♂.
9.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 139, 262 (key).

Biology: Hurd and Linsley, 1975. Calif. Univ. Pubs. Ent. 193: 24-25, fig. 6 (floral
relationships).

Perdita pusilla Timberlake
South. Calif, (western margin of Mojave Desert).

Perdita pusilla Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 257, figs. 1021, 1022, 1154. ♀.
6.

Perdita quadraticeps Timberlake
Ariz. (Patagonia).

Perdita quadraticeps Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 213, figs. 29, 969, 970,
1128. ♂.

Perdita replicans Timberlake
Calif. (San Luis Obispo County).

Perdita replicans Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 149, fig. 29. ♀.

Perdita richardsi Timberlake
California (Apple Valley in the Mojave Desert).

Perdita richardsi Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 227. ♂.

Perdita salicis
***authority mismatch
coloradana Timberlake. Colo. (Delta and Garfield Counties).

Perdita salicis coloradana Timberlake, 1929. N. Y. Ent. Soc, Jour. 37: 111. ♀.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 340, 341. 9 (key, redescription,
geogr. records).

Perdita salicis
***authority mismatch
euxantha Timberlake. Idaho (Idaho and Nez Perce Counties), Oreg. (near Corvallis).

Perdita salicis euxantha Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 349. ♀, ♂.

Perdita salicis
***authority mismatch
hirsutior Timberlake. Calif. (Olancha). Pollen: Presumably an oligolege of Salix, visits
the flowers of S. exigua.
Perdita salicis hirsutior Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 346. ♀, ♂.

Perdita salicis
***authority mismatch
imperialis Cockerell. Wash. (Asotin, Spokane, WaUa Walla and Whitman Counties),
Idaho (Blaine, Canyon, Nez Perce and Twin Falls Counties), Oreg. (Umatilla and
Walloun Counties), Utah (Cache, Iron, Juab and Washington Counties), south. Calif.
(Colorado and Mojave Deserts), Ariz. (Maricopa and Pima Counties), Colo. (Archuleta
County). Pollen: Presumably an oligolege of Salix, including S. gooddingii, but visits
other flowers apparently for nectar including Acacia greggii. Asparagus, Daucus carota,
Prosopis, Sisymbrium.
Perdita exclamans imperialis Cockerell, 1925. Calif. Acad. Sci., Proc. (4) 14: 193. ♀.




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Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 340, 342-343. 9, 3 (key,
redescription, geogr. and floral records).

Perdita salicis
***authority mismatch
laeta Timberlake. South. Calif. (Needles). Pollen: Presumably an oligolege of Salix,
visits flowers of Prosopis juliflora.
Perdita salicis laeta Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 347. ♀, ♂.

Perdita salicis
***authority mismatch
monoensis Timberlake. Calif. (Eldorado and Mono Counties). Pollen: Presumably an
oligolege of Salix, visits flowers of S. sessilifolia.
Perdita salicis ynonoejisis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 345. ♀, ♂.

Perdita salicis
***authority mismatch
occidentalis Timberlake. South. Calif, (cismontane, except one population at Helendale
on Mojave Desert). Pollen: Apparently an oligolege of Salix including S. exigua, S.
gooddingii, S. nigra, but visits other flowers presumably for nectar including Alyssum
maritimum, Cryptantha microphylla var. lepida, Funastrum heterophyllum,
Ligustrum, Pyracantha.
Perdita salicis occidentalis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 343. ♀, ♂.

Perdita salicis
***authority mismatch
personata Timberlake. South. Calif. (San Diego County). Pollen: Presumably an
oligolege of Salix, visits flowers of S. laevigata.
Perdita salicis personata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 344. ♀, ♂.

Perdita salicis
***authority mismatch
salicis Cockerell. Colo., N. Hex. (Las Cruces); Mexico (Chihuahua). Pollen: Presumably
an oligolege of Salix.
Perdita salicis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 80. ♀, ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 138, 141, 157, 338-341, figs. 1085,
1086, 1186. 9, 6 (key, redescription, geogr. and floral records),

Perdita salicis
***authority mismatch
sublaeta Timberlake. Oreg. (Hood River and The Dalles).

Perdita salicis sublaeta Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 348. ♀, ♂.

Perdita salicis
***authority mismatch
subtristis Cockerell. Colo. (Archuleta and Chaffee Counties), Utah (Grand County).
Pollen: Apparently an oligolege oi Salix.
Perdita subtristis Cockerell, 1933. Canad. Ent. 65: 234. ♀.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 135, 339, 341-342. 9, 6 (key,
redescription, geogr. and floral records).

Perdita salicis
***authority mismatch
tristis Timberlake. Calif. (San Joaquin Valley). Pollen: Apparently an oligolege of Salix,
visits flowers of S. hindsiana and also Populus.
Perdita salicis tristis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 345, fig. 74. ♀, ♂.

Perdita salviae Timberlake
South. Calif. (Colorado Desert). Pollen: Collects pollen from the flowers of
Salvia vaseyi.
Perdita salviae Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 184, figs. 935, 936, 1111. ♀, ♂.

Perdita sandhouseae Timberlake
Ariz. (Apache County). Pollen: Unknown, but visits flowers of
Gutierrezia.
Perdita sandhouseae Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 171, figs. 8, 921, 922,
1104. ♂.

Perdita schlingeri Timberlake
Nev. (Emigrant Pass).

Perdita schlingeri Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 258. ♀.

Perdita semicaerulea Cockerell
Tex. to Ariz.; Mexico (Chihuahua and Coahuila). Pollen: Oligolege of
Larrea tridentata, visits both the spring and late summer and fall blooms of this plant
from which it obtains pollen and nectar, but also visits flowers of Prosopis presumably
for nectar only.
Perdita semicaerulea Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 64. ♀.
Perdita quadraiigularis Cockerell, 1907. Ann. and Mag. Nat. Hist. (7) 20: 129. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 131, 150, 186-188, figs. 14, 937, 938,
1112. 9, 6 (key, redescription, synonymy, geogr. and floral records).

Biology: Hurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 25 (floral relationships).

Perdita sexnotata Timberlake
South. Calif. (Colorado Desert). Pollen: Unknown, but visits flowers of
Coldenia pahneri, Eschscholzia parishii.
Perdita sexnotata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 307, fig. 66. ♀.




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Perdita sidae Cockerell
N. Mex. (Mesilla); Mexico (Durango). Pollen: Unknown, but visits flowers of
Sida hederacea.
Perdita sidae Cockerell, 1897. Acad. Nat. Sci. Phila., Proc. 49: 353. ♀, ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 128, 132, 148, 263-264, figs. 51, 1025,
1026, 1156. 6 (key, redescription). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 50. 9
(redescription, geogr. record).

Perdita sonorensis Cockerell
Ariz. (Yuma County), south. Calif. (Colorado and Mojave Deserts);

Mexico (Baja California and Sonora). Pollen: Oligolege o{ Prosopis including P.juliflora,
P. pubescens, but also visits flowers of Encelia farinosa, Eucnide urens, Heliotropium
curassavicuin, Melilotus, PeucephyUum schottii, Pluchea sericea.
Perdita sonorensis Cockerell, 1899. Ann. and Mag. Nat. Hist. (7) 3: 493. ♀.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 136, 138, 152, 272-274, figs. 55, 1037,
1038, 1162. 9, 6 (key, redescription, geogr. and floral records).

Perdita speciosa Timberlake
Ariz. (Coconino County). Pollen: Unknown, but visits flowers of
Wislizeyiia refracta.
Perdita speciosa Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 51, figs. 1391, 1392, 1414. ♂.

Perdita sphaeralceae
***authority mismatch
alticola Cockerell. N. Mex., Ariz. (Cochise and Coconino Counties). Pollen:

Possibly an oligolege of Spliaeralcea, visits flowers of S. lobata, but also visits flowers
of Heterotheca subaxillaris.
Perdita sphaeralceae race alticola Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 68. ♀.

Perdita sphaeralceae race alticola mut. suffiisa Cockerell, 1896. Acad. Nat. Sci. Phila.,
Proc. 48: 68. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 154, 178-179, fig. 11. 9,6 (key,

geogr. and floral records).

Perdita sphaeralceae
***authority mismatch
sphaeralceae Cockerell. Tex. (Hudspeth County), N. Mex. (Dona Ana and Luna
Counties); Mexico (Chihuahua, San Luis Potosi and Zacatecas). Pollen: Possibly an
oligolege o{ Spliaeralcea including S. angiistifolia, but visits other flowers including
Cologania, Gutieii-ezia lucida, Haplopappus heterophyllus, Mentzelia, Solidago
canadensis.

Perdita sphaeralceae Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 66. ♀, ♂.

Perdita pellucida Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 88. ♂.

Perdita sphaeralceae var. ridens Cockerell, 1922. Ann. and Mag. Nat. Hist. (9) 10: 547. ♂.

Taxonomy: Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 121 (tax. status, as pellucida).
—Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 127, 154, 176-178. 9, 6 (key, redescription,
synonymy, geogr. and floral records).

Perdita stathamae
***authority mismatch
eluta Timberlake. Ariz. (Salt River Mts.).

Perdita stathamae eliita Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 358. ♀, ♂.

Perdita stathamae
***authority mismatch
stathamae Timberlake. N. Mex. (Alamogordo), Ariz. (Chihuahua and Sonora

Deserts), south. Calif. (Colorado Desert); Mexico (Baja California and Sonora). Pollen:
Unknown, but visits flowers of Acacia greggii, Prosopis jtiliflora.
Perdita stathamae stathamae Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 365, figs. 1091,
1092, 1189. ♀, ♂.

Perdita stenopyga Timberlake
Calif. (Inyo County). Pollen: Unknown, but visits flowers of
Chaenactis, Mentzelia affinis.
Perdita stenopyga Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 229, figs. 38, 989, 990, 1138.

9, 6.

Perdita sternalis Timberlake
South. Tex. (Cameron and Galveston Counties). Pollen: Unknown, but
visits flowers of Coreopsis douglasii, Heliotropium, Rubus.
Perdita stemalis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 291, figs. 1055, 1056, 1171.
9, 6.

Perdita sulphuripes Timberlake
Calif, (near Palm Springs, also Slate Mts., Inyo County). Pollen:
Unknown, but visits flowers of Malacothrix.
Perdita sulphuripes Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 267, figs. 1031, 1032,
1159. ♂.




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Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 151. 6 (geogr. and floral record).

Perdita tarda Cockerell
N. Mex. (Las Cruces). Pollen: Unknown, but visits flowers of Gutierrezia
microcephala.
Perdita tarda Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 85. ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 146, 151, 279-280 (redescription of
type specimen).

Perdita tenebrosa Timberlake
Ariz. (Chiricahua Mts.).

Perdifa tenebrosa Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 131. ♀.

Perdita tessellata Timberlake
Idaho (Elmore County), Nev. (Battle Mountain). Pollen: Unknown, but
visits flowers oi Chaenactis, Tetradymia.
Perdita tessellata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 264, figs. 1027, 1028, 1157.
6.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 113, 117, 151-152, fig. 30. 9, S (key,
redescription, geogr. and floral records).

Perdita thelypodii Timberlake
Calif. (Inyo and Los Angeles Counties). Pollen: Unknown, but visits
flowers of Cleomella obtusifolia, Thelypodium brack ycarpiivi.
Perdita thelypodii Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 354, figs. 432, 433, 500. ♀.
6.

Perdita tortifoliae
***authority mismatch
fremonti Timberlake. South. Calif. (Mohave Desert), Nev. (Clark County), Utah
(Washington County). Pollen: Collects pollen from the flowers oi Lepidium fremontii.
Perdita fremonti Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 250, figs. 1011, 1012, 1149.
9, S.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 143 (tax. status, geogr. and floral
records).

Perdita tortifoliae
***authority mismatch
tortifoliae Cockerell. Colo. (Florissant and Wilkerson Pass), N. Mex. (Rio Arriba
County). Pollen: Presumably collects pollen from the flowers of Lepidium jonesii, but
also visits flowers of Bigelovia tortifoliae, Eriogonum umbellatiim, Ranunculus
eremogenes.
Perdita tortifoliae Cockerell, 1906. Amer. Mus. Nat. Hist., Bui. 22: 440. ♀.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 132, 144, 245-247, figs. 45, 1005,
1006, 1146. 9, 6 (key, redescription, geogr. and floral records). —Timberlake, 1968. Calif.
Univ. Pubs. Ent. 49: 143 (tax. status, geogr. and floral record).

Perdita triangulifera Timberlake
West. Tex. to south. Calif., deserts. Pollen: Apparently an oligolege
of Prosopis, but visits other flowers presumably for nectar including Acacia greggii,
Cercidium floridum, Dithyrea wislizeni, Melilotus, Mimosa, Phoradendron, Tamarix.
Perdita triangulifera Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 328, figs. 71, 1075-1076,
1181. ♀, ♂.

Perdita trinotata Timberlake
Tex. Pollen: Unknown, but visits flowers of Lesqtierella gordonii, Nama
hispidum, Nerisyrenia ca7npo7'U7n.
Perdita trinotata Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 249, figs. 47, 1009, 1010,
1148. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 115, 116, 144-145, fig. 26. 9,6 (key,
redescription, geogr. and floral records).

Perdita veris Timberlake
Ariz. (Pima County). Pollen: Collects pollen from the flowers of Nama

hispidum, but visits other flowers for nectar including Teucrium cubense depressum.
Perdita veris Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 137, figs. 21, 1289, 1290, 1348. 9
6.

Perdita vidua Timberlake
Tex. (El Paso County). Pollen: Unknown, but visits flowers of Gossypium.

Perdita vidua Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 309. ♀.
vittata conformis Timberlake. Nev. (Lincoln County).

Perdita vittata conformis Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 284. ♀, ♂.




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vittata tricolor Timberlake. South. Calif. (Mojave Desert and south. San Joaquin Valley).

Pollen: Unknown, but visits flowers of Cleomella obtusifolia, WislizeJiia refracta. The
typical subspecies, Perdita vittata vittata Cockerell, occurs in Mexico (Baja California).
Perdita vittata tricolor Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 283. ♂, ♀.

Perdita werneri Timberlake
Ariz. (Springerville).

Perdita wenieri Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 154, figs. 1297, 1298, 1352. ♂.

Perdita wilmattae
***authority mismatch
miricornis Cockerell. Wyo. (Green River), Colo. (Mesa and Moffat Counties), Utah
(Carbon County), Ariz. (Grand Canyon). Pollen: Unknown, but visits flowers of Cleome,
Engehnannia pinnatifida, Phacelia, Stanleya pinnata.
Perdita miricornis Cockerell, 1922. Amer. Mus. Novitates 33: 9. ♂, ♀.
Perdita miricornis var. leucorhina Cockerell, 1922. Amer. Mus. Novitates 33: 10. ♀.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 310, 314, 360. 6, 9 (key,
redescription, tax. status, geogr. and floral records). —Timberlake, 1964. Calif. Univ. Pubs.
Ent. 28: 245 (tax. status). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 145 (key).

Perdita wilmattae
***authority mismatch
stanleyae Timberlake. Utah (Grand, Uintah and Washington Counties), Nev., Calif.
(Inyo County). Pollen: Unknown, but visits flowers of Asclepias, Cleome lutea,
Mentzelia, Pninus, Stanleya pinnata, Thelypodium laciniatum.
Perdita stanleyae Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 253, figs. 48, 1017, 1018,
1152. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 145 (key, tax. status, geogr. record).

Perdita wilmattae
***authority mismatch
wilmattae Cockerell. Colo., Utah, Nev. (Humboldt County). Pollen: Unknown, but
visits flowers of Cleome lutea, Engehnannia, Phacelia alba, Stanleya pinnata.
Perdita wilmattae Cockerell, 1906. Amer. Mus. Nat. Hist, Bui. 22: 441. ♂, ♀.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 310, 314, 359-360, figs. 436, 437, 502.
6, 9 (key, redescription, tax. status, geogr. and floral records). —Timberlake, 1964. Calif.
Univ. Pubs. Ent. 28: 137, 245 (key, floral records). —Timberlake, 1968. Calif. Univ. Pubs.
Ent. 49: 145 (key).

Perdita wislizeniae Timberlake
N. Mex. (Luna County), Ariz. (Cochise County); Mexico (Chihuahua).
Pollen: Apparently an oligolege of Wislizeriia refracta, but also visits flowers of Cleome,
Eriogonuyn thomasii, Lepidium.
Perdita wislizeniae Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 284, figs. 57, 1049, 1050,
1168. ♀, ♂.

Perdita zebrata
***authority mismatch
flavens Timberlake. Nev., Oreg., Idaho, Wyo., Utah, Colo., Calif. (Inyo County). Pollen:
Apparently collects pollen only from the flowers of Cleome including C. lutea, C.
serrnlata, but visits other flowers for nectar including Engehnannia pinnatifida,
Phacelia, Salsola kali.
Perdita zebrata flavens Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 357. ♀, ♂.

Taxonomy: Timberlake, 1964. Calif. Univ. Pubs. Ent. 28: 138, 153, 262 (key, geogr. and floral
record).

Perdita zebrata
***authority mismatch
zebrata Cresson. N. Dak., Nebr. and N. Mex., west to Idaho, Nev. and Ariz. Parasite:
Neolaira pi'uinosa Ashmead. Pollen: Collects pollen from the flowers of Cleome
including C. serrnlata, but visits other flowers presumably for nectar including
Haplopappus heterophyllus, Helianthus petiolaris, Nolina microcarpa, Solidago.
Perdita zebrata Cresson, 1878. Amer. Ent. Soc, Trans. 7: 69. 9 (part).
Perdita caimia Cockerell, 1895. Acad. Nat. Sci. Phila., Proc. 47: 17. ♂.
Perdita bakerae Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 69. ♂, ♀.

Taxonomy: Cockerell, 1903. In Viereck, Amer. Ent. Soc, Trans. 29: 51. —Timberlake, 1958.
CaHf. Univ. Pubs. Ent. 14: 310, 313, 356-357, figs. 434, 435, 501 (key, tax. characters, tax.
status, synonymy, geogr. and floral records). —Timberlake, 1964. Calif. Univ. Pubs. Ent.
28: 138, 153, 262 (key, geogr. range). — Rozen, 1966. Amer. Mus. Novitates 2259: 17-19, figs.
41-46 (larva). -Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 118, 150, figs. 1295, 1296,
1351. 6 (key, variation, geogr. records).

Biology: Custer, 1929. Canad. Ent. 61: 49 (nesting habits with description of larva). —Rozen,
1967. Amer. Mus. Novitates 2297: 34-38, figs. 2, 4, 11, 14, tables 1-2 (nest architecture, life
history, pollen source, parasite).




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Species Group Valida

This species may be better placed in a new genus related to Perdita.

Taxonomy: Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 106 (tax. characters).

Perdita valida Timberlake
N. Mex. (Mesilla).

Perdita valida Timberlake, 1962. Calif. Univ. Pubs. Ent. 28: 106. ♀.

Genus PERDITA Subgenus PERDITELLA Cockerell

Perdita subg. Perditella Cockerell, 1899. Psyche 8: 312.

Type-species: Perdita larreae Cockerell. Orig. desig. (=Perdita laneae Cockerell).

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 348 (key). —Timberlake, 1956. Calif.
Univ. Pubs. Ent. 11: 267-268 (redescription, key to included spp.).

Perdita cladothricis Cockerell
West. Tex., N. Mex., Ariz.; Mexico (Baja California, San Luis Potosi,
Sonora and Zacatecas). Pollen: Possibly an oligolege of Tidestromia including T.
lanuginosa, but visits other flowers presumably for nectar including Gutierrezia
microcephala, Haplopappus wrightii, Medicago sativa, Pedis papposa.
Perdita cladothricis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 82. ♀, ♂.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 268, 270, figs. 207, 208, 305 (male

genitalia, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 30

(geogr. records).

Perdita larreae Cockerell
N. Mex., Ariz., South. Calif.; Mexico (Baja California, Chihuahua and

Coahuila). Pollen: Collects pollen from the flowers of Larrea tridentata, but visits these
and other flowers for nectar including Aplopappus heterophylius, Asclepias, Baccharis
glutinosa, Croton califomicus, Dalea, Dicraurus, Eriogonum reniforme. Euphorbia
polycarpa, Pedis papposa, Tamarix.

Perdita larreae Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 62. ♂.

Perdita larrearum Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 63. ♀.

Perdita larreae var. modesta Cockerell, 1897. Acad. Nat. Sci. Phila., Proc. 49: 355. ♂.

Perdita laneae Cockerell, 1899. Psyche 8: 312. Error for larreae.

Taxonomy: Timberlake, 1928. Amer. Mus. Novitates 321: 11. 9. —Timberlake, 1956. Calif.
Univ. Pubs. Ent. 11: 268, figs. 203, 204, 304 (tax. characters, synonymy, geogr. and floral
records). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17: 134 (variation, geogr. and floral
records).

Biology: Kurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 25-26, fig. 7 (floral
relationships).

Perdita marcialis Cockerell
West. Tex. to south. Calif., Nev.; Mexico (Coahuila, Sonora). Pollen:
Collects pollen from the flowers of Larrea tridentata and possibly also from other
flowers including Aloysia wrightii, Chrysothanmus paniculatus, Eriogonum
fasciculatum var. polifolium, Phacelia, Salvia vaseyi.
Perdita marcialis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 62. ♂.
Perdita phaceliae Cockerell, 1898. Ann. and Mag. Nat. Hist. (7) 2: 450. ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 268, 269-270, figs. 205, 206 (key,
redescription, geogr. and floral records). —Timberlake, 1960. Calif. Univ. Pubs. Ent. 17:
134-135 (synonymy).

Biology: Hurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 26 (floral relationships).

Perdita minima Cockerell
Ariz., South. Calif.; Mexico (Baja California). Pollen: Collects pollen from
the flowers of Euphorbia including E. albomarginata, E. polycarpa, E. p. var. hirtella,
but visits these and other flowers for nectar including Croton califomicus, Eriogonum
gracile, Gutierrezia.
Perdita minima Cockerell, 1923. Amer. Mus. Novitates 66: 4. ♀.

Taxonomy: Cockerell, 1925. Pan-Pacific Ent. 1: 179. 6. —Timberlake, 1956. Calif. Univ. Pubs.
Ent. 11: 268, 270, figs. 209, 210 (key, redescription, geogr. and floral records).




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Genus PERDITA Subgenus PROCOCKERELLIA Timberlake

Perdita subg. Procockerellia Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 402.
Type-species: Perdita alboyiotata Timberlake. Orig. desig.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 5-7 (key to spp. of subgenera
Procockerellia, Allomacrotera).

Perdita albonotata Timberlake
South. Calif. (Colorado and Mojave Deserts), Ariz. (Mohave Cou\nty),
Utah (Cache and Washington Counties); Mexico (Sonora). Pollen: Collects pollen from
the flowers of Stephano7neria including S. exigua, S. pauciflora, but visits other flowers
presumably for nectar including Agave deserti, Chaenactis glahriiiscula, Eriogonum
plumatella.
Perdita albonotata Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 403, figs. 91, 92, 161. ♀, ♂.

Perdita brachyglossa Timberlake
Ariz. (Coconino County). Pollen: Unknown, but visits flowers of
Thelesperma.
Perdita brachyglossa Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 6. ♀.
exceliens Timberlake. Utah (Grand and Washington Counties), Ariz. (Coconino County).
Pollen: Unknown, but visits flowers of Cleome lutea.
Perdita exceliens Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 384. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 22. ♂.

Perdita moabensis Timberlake
Utah (Moab).

Perdita moabensis Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 7. ♀.

Genus PERDITA Subgenus PSEUDOMACROTERA Timberlake

Perdita subg. Pseudomacrotera Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 349.
Type-species: Perdita turgiceps Timberlake. Orig. desig.

Perdita turgiceps Timberlake
Calif. (Colorado and Mojave Deserts), Ariz., (Maricopa Mts.); Mexico

(Baja California). Pollen: Apparently an oligolege oi Larrea tridentata, but visits a wide
variety of other flowers for nectar including Acacia greggii, Cercidium floridiim,
Cowania stansburiana, Cryptantha angustifolia, C. barbigera, Dalea califoniica, D.
fremontii, D. schottii, Datura meteloides, Encelia farinosa, Eriodictyon crassifoliuni,
Eriogonum inflatum, E. trichopes, Eschscholzia, Eucnide urens, Hyptis emoryi, Larrea
tridentata, Lasthenia, Nama hispida, Prosopis juliflora.

Perdita turgiceps Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 350, figs. 3, 4, 116, 117. ♂, ♀.

Biology: Kurd and Linsley, 1975. Smithsn. Contrib. Zool. 193: 26 (floral relationships).

Genus PERDITA Subgenus PYGOPERDITA Timberlake

Perdita subg. Pygoperdita Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 275.
Type-species: Perdita interrupta Cresson. Orig. desig.

Although several of the species visit flowers of the Compositae, especially Layia and
Malacotkrix, most members of this subgenus visit flowers of other famihes, such as the
Rosaceae, Papaveraceae, Rhamnaceae and Liliaceae.

Taxonomy: Timberlake, 1956. CaHf. Univ. Pubs. Ent. 11: 275-284 (key to included spp.).

Species Group Interrupta

Perdita associata Timberlake
Calif. (San Benito County).

Perdita associata Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 32. ♀.

Perdita aureovittata
***authority mismatch
aureovittata Cockerell. South. Calif, (cismontane, south of San Gabriel Mts.).
Pollen: Unknown, but visits flowers oi Layia, Malacotkrix califoniica var. glabrata.
Perdita (Cockerellia) aureovittata Cockerell, 1916. Canad. Ent. 48: 391. ♀.
Taxonomy Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 391 (tax. status). -Timberlake, 1956.
Calif Univ. Pubs. Ent. 11: 276, 281, 285, figs. 221, 222, 311, 337. 9, c5 (redescription, new
status, geogr. and floral records). -Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 33 (geogr.
range).




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Perdita aureovittata
***authority mismatch
maderensis Timberlake. Calif. (Madera County). Pollen: Unknown, but
presumably visits flowers of Malacothrix califoniica.
Perdita aureovittata maderensis Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 286. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 33 (geogr. range).

Perdita aureovittata
***authority mismatch
soluta Timberlake. Calif. (South Coast and Peninsular Ranges). Pollen: Unknown,
but visits flowers of Agoseris auriantiaca, A. heterophylla, Encelia actoni, Linanthus
aureus, Malacothrix califomica.
Perdita soluta Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 394, fig. 520. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 33 (tax. status, geogr. range).

Perdita aureovittata
***authority mismatch
stenozona Timberlake. Calif. (San Benito and San Luis Obispo Counties). Pollen:
Unknown, but visits flowers of Chaenactis glabriuscula, Malacothrix califomica.
Perdita aureovittata stenozona Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 286. ♀, ♂.

Perdita fieldi Timberlake
South. Calif. (Monterey to San Diego County). Pollen: Unknown, but visits
flowers of Adenostoma fasciculatum, Calochortus splendens, Eriogonum fasciculatum.
Perdita fieldi Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 291, figs. 227, 228, 315, 340. ♀.
6.

Perdita inflexa Timberlake
Calif. (Riverside County). Pollen: Unknown, but visits flowers of
Eschscholzia multiflora var. darwinensis.
Perdita inflexa Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 298, figs. 237, 238, 345, 346. ♂.
interrupta interrupta Cresson. South. Calif, (cismontane). Pollen: Collects pollen from the
flowers of Eschscholzia califomica, but visits other flowers for nectar including
Baccharis emoryi. Coreopsis califomica, Cryptantha intermedia, Lasthenia
chrysostoma, Lepidospartum squamatum. Sisymbrium irio, Solidago occidentalis,
Tamarix gallica; also has been taken at the flowers of Yucca, but since these flowers
are not attractive to bees, the plant in question was presumably Nolina parryi.
Perdita interrupta Cresson, 1878. Amer. Ent. Soc, Trans. 7: 70. ♂.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 277, 282, 292, figs. 229, 230, 316, 341.
9, 6 (redescription, tax. status, geogr. and floral records).
interrupta kernensis Timberlake. Calif. (Kern County). Pollen: Unknown, but visits flowers of
Cryptantha.
Perdita interrupta kernensis Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 293. ♀, ♂.
interrupta vernalis Timberlake. Calif. (San Diego County).

Perdita interrupta vemalis Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 293. ♀, ♂.

Perdita layiae
***authority mismatch
excisa Timberlake. South. Calif, (cismontane). Pollen: Unknown, but visits flowers of
Chaenactis glabriuscula, Layia platyglossa.
Perdita layiae excisa Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 288. ♀, ♂.

Perdita layiae
***authority mismatch
layiae Cockerell. Calif. (San Miguel Island). Pollen: Unknown, but visits flowers of
Layia platyglossa.
Perdita layiae Cockerell, 1938. Ann. and Mag. Nat. Hist. (11) 2: 152. ♀, ♂.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 277, 281, 288, figs. 223, 224, 313, 338
(redescription).

Perdita malacothricis Timberlake
South. Calif, deserts, Nev. (Clark County), Ariz. (Tucson). Pollen:
Unknown, but visits flowers o{ Aster, Baileya multiradiata, B. pleniradiata,
Calycoseris parryi, Chaenactis fremontii, Hymenoclea salsola, Larrea tridentata,
Malacothrix califomica var. glabrata, Stephanomeria exigua, S. pauciflora.
Perdita malacothricis Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 289, figs. 225, 226, 314,
339. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 36 (geogr. and floral records).

Perdita malacothricis
***authority mismatch
var. unica Timberlake. CaHf. (Riverside County).

Perdita malacothricis var. unica Timberlake, 1962. CaHf. Univ. Pubs. Ent. 28: 96. ♀.

Perdita micheneri
***authority mismatch
micheneri Timberlake. South. Calif, (cismontane); Mexico (Baja California). Pollen:
Unknown, but visits flowers of Ceanothus cordulatus, C. greggii, C. leucodermis, C.




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orcuttii, Cryptantha intermedia, C. micrantha, C. muricata, Eriophyllum
confertiflorum, Eschscholzia califomica, Rhaynnus crocea, Rhus ovata.
Perdita micheneri micheneri Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 294, figs. 231,
232,317,342. ♀, ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 36 (geogr. record).

Perdita micheneri
***authority mismatch
rhamnophila Timberlake. Calif. (Rock Creek in San Gabriel Mts.). Pollen:
Unknown, but visits flowers oi Rliamnus crocea.
Perdita micheneri rhamnophila Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 295. ♂.

Perdita mucronata Timberlake
Calif. (Riverside County); desert. Pollen: Unknown, but visits flowers
oi Camissonia brevipes, Eschscholzia minutiflora var. darwinensis.
Perdita mucronata Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 297, figs. 235, 236, 318,
344. ♂.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 36. ♀.

Perdita nitens Timberlake
Calif. (Mendocino, San Benito, Santa Barbara and Stanislaus Counties).

Pollen: Unknown, but visits flowers oi Eriogonum, Eschscholzia califomica, Mimulus
fremontii, Phacelia.
Perdita nitens Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 295, figs. 233, 234, 343. ♀, ♂.

Perdita quadrisignata Timberlake
Calif. Pollen: Unknown, but visits flowers oi Baccharis, Ceanothus,
Clarkia, Coreopsis bigelovii, Cryptantha muricata, Eriophyllum confertiflorum,
Eschscholzia califomica, Layia platyglossa, Plagiobothrys nothofulvus, Pnmus
subcordata, Rhamnus crocea, Salix.
Perdita quadrisignata Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 296. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 39. ♂.

Perdita transversa Timberlake
Oreg. (Jackson County).

Perdita transversa Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 287, figs. 312. ♂.

Perdita vandykei Timberlake
Calif. (Monterey County).

Perdita vandykei Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 286. ♀, ♂.

Species Group Californica

Biology: Linsley, 1958. Hilgardia 27: 562, table 5 (oligolecty).

Perdita argemones Timberlake
Ariz. (Maricopa, Pima and Pinal Counties). Pollen: Unknown, but visits
flowers of Argemone.
Perdita argemones Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 309, figs. 261, 262, 327,
358. ♀, ♂.

Perdita arizonica Timberlake
Ariz. (Oak Creek Canyon). Pollen: Apparently collects pollen from the
flowers of Calochortus.
Perdita aHzonica Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 310, figs. 263, 264, 328, 359.

9,6.

Perdita bilobata Timberlake
Calif. (Inyo and San Bernardino Counties). Pollen: Unknown, but visits
flowers oi Calochortus aureus, C. kennedyi, Haplopappus cooperi.
Perdita bilobata Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 305, figs. 253, 254, 354. ♀, ♂.

Perdita bispinata Timberlake
Calif. (San Bernardino and Riverside Counties). Pollen: Unknown, but
visits flowers oi Calochortus kennedyi, Haplopappus cooperi.
Perdita bispinata Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 303, figs. 249, 250, 323, 352.
9, 6.

Perdita bohartorum Timberlake
Calif. (Mono County).

Perdita bohartorum Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 319, figs. 275, 276, 365.

9,6.

Perdita californica
***authority mismatch
californica (Cresson). Coastal Calif. (Mt. Diablo, southward); Mexico (Baja

California). Pollen: Apparently collects pollen from the flowers oi Calochortus including
C. concolor, C. kennedyi, C. luteus, C. plummerae, C. splendens, C. venusius, C. weedu,
but visits other flowers for nectar including Cryptantha, Eriogonum fasciculatum,
Eriophyllum confertiflorum, Grindelia, Opuntia.
Macrotera califomica Cresson, 1878. Amer. Ent. Soc, Trans. 7: 71. ♂.




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Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 280, 282, 301-302. 9. -Timberlake,
1968. Calif. Univ. Pubs. Ent. 49: 34 (tax. status, geogr. range).

Perdita californica
***authority mismatch
inopina Timberlake. Calif. (Inyo County).

Perdita californica inopina Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 33. ♀.

Perdita calochorti Timberlake
Calif. (San Bernardino and Inyo Counties). Pollen: Unknown, but visits
flowers of Calochortus nuttallii.
Perdita calochorti Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 299, figs. 239, 240, 320, 347.
9,6.

Perdita coalingensis Timberlake
Calif. (Coalinga). Pollen: Unknown, but visits flowers of Eschscholzia
californica.
Perdita coalingensis Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 313. ♀.

Perdita cowaniae Timberlake
Calif. (Inyo County). Pollen: Unknown, but visits flowers of Calochortus
kennedyi var. munzii, Cowania stansburiana.
Perdita cowaniae Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 315, figs. 269, 270, 331, 362.
9,

Perdita digressa Timberlake
Ariz. (Tucson), Calif. (Granite Mts., San Bernardino County). Pollen:
Unknown, but visits flowers of Baileya multiradiata, Chaenactis.
Perdita digressa Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 34, figs. 1211, 1212, 1309,
1354. ♂, ♀.

Perdita distropica Timberlake
Calif. (South Coast Ranges). Pollen: Collects pollen from the flowers of
Calochortus including C. clavatus, C. luteiis, C. splendens, C. venustus and Eschscholzia
californica, but visits other flowers for nectar including Adenostoma, Eriogonum.
Perdita distropica Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 312, figs. 265, 266, 329, 360.
9, 6.

Perdita duplonotata Timberlake
Utah (Garfield County), Nev. (Charleston Mts.), Cahf. (San
Bernardino County). Pollen: Unknown, but visits flowers oi Fallugia paradoxa,
Isomeris arhorea.
Perdita duplonotata Timberlake, 1956. Cahf. Univ. Pubs. Ent. 11: 307, figs. 257, 258, 356.
9,6.

Perdita eriogoni Cockerell
Colo. Pollen: Unknown, but visits flowers of Eriogonum umbellatum.
Perdita eriogoni Cockerell, 1925. Ann. and Mag. Nat. Hist. (9) 16: 621. ♀, ♂.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 279, 284, 322-323, figs. 279, 280, 367
(tax. characters). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 35-36. 6 (color variation).

Perdita fallugiae Timberlake
Nev. (Charleston Mts.). Pollen: Unknown, but visits flowers of
Calochortus flexuosus, Fallugia paradoxa, Hymenoxys cooperi.
Perdita fallugiae Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 314, figs. 267, 268, 330, 361.
9,6.

Perdita leucostoma Timberlake
Calif. (Kern and Inyo Counties). Pollen: Unknown, but visits flowers
of Calochortus leichtlinii, Haplopappus linearifolius.
Perdita leucostoma Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 300, figs. 241, 242, 321,
348. ♀, ♂.

Perdita macrostoma Cockerell
Calif. Pollen: Apparently collects pollen from the flowers of
Calochortus.
Perdita macrostoma Cockerell, 1922. U. S. Natl. Mus., Proc. 60 (18): 18. cJ, 9.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 278, 283, 304, figs. 251, 252, 324, 353

(tax. characters).

Perdita mohavensis
***authority mismatch
mohavensis Timberlake. South. Calif, deserts. Pollen: Probably collects pollen
mainly from the flowers of Eschscholzia, but visits flowers of Amsinckia intermedia,
Argemone platyceras, Calochortus aureus, Calycoseris wrightii, Chaenactis stevioides
var. brachypappa, Encelia farinosa, Eschscholzia californica, E. glyptosperyna, E.
minutiflora var. darwinensis, E. parishii, Haplopappus cooperi, Hyptis emoryi, Larrea
tridentata, Monardella exilis, Salix, Salvia.

Perdita mohavensis Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 306, figs. 255, 256, 325,
355. ♀, ♂.




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Perdita mohavensis
***authority mismatch
pimana Timberlake. Ariz. (Pima County). Pollen: Unknown, but visits flowers of
Baileya.
Perdita mohavensis pimana Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 36. ♀.

Perdita montereyensis Timberlake
Calif. (Monterey, San Benito and San Luis Obispo Counties).
Pollen: Collects pollen from the flowers of Calochortus splendens, C. veiiustus and
Eschscholzia califomica, but visits other flowers for nectar including Eriogonum,
Rhamnus crocea.
Perdita montereyensis Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 302, figs. 245, 246, 350.
9, (J.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 36 (color variation, geogr. and floral
record).

Perdita mormonica Timberlake
Utah (Garfield and Tooele Counties). Pollen: Unknown, but visits
flowers of Sphaeralcea.
Perdita mormonica Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 311. ♀.

Taxonomy: Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 10 (color variation).

Perdita nebrascensis Swenk and Cockerell
Nebr., (Neligh, Antelope County). Pollen: Unknown, but
visits flowers of Helianthus.
Perdita nebrascensis Swenk and Cockerell, 1907. Ent. News 18: 53. ♀, ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 395-396 (key, tax. status).

Perdita nevadensis
***authority mismatch
culbertsoni Timberlake. Calif. (Tulare, Santa Cruz Counties, Yosemite Natl. Pk.).

Perdita nevadensis culbertsoni Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 318. 9, tJ.

Perdita nevadensis
***authority mismatch
molina Timberlake. Calif. (San Bernardino Mts.).

Perdita nevadensis molina Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 318, figs. 273, 274,
364. ♀, ♂.

Taxonomy: Timberlake, 1958. Calif. Univ. Pubs. Ent. 14: 396 (key).

Perdita nevadensis
***authority mismatch
nevadensis Cockerell. B. C. to north. Calif. Pollen: Unknown, but visits flowers of
Eriogonum.
Perdita nevadensis Cockerell, 1896. Acad. Nat. Sci. Phila., Proc. 48: 58. ♀.
Perdita nigricollis Timberlake, 1929. Pan-Pacific Ent. 6: 54. ♀.

Taxonomy: Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 279, 284, 316-318, figs. 271, 272, 332,
363. 9 , 6 (redescription, synonymy).

Perdita obtusa Timberlake
Calif. (Tulare County).

Perdita obtusa Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 37. ♀.

Perdita pulliventris Timberlake
Calif. (Tuolumne County).

Perdita pulliventris Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 38. ♀.

Perdita robustula Timberlake
South. Calif. Pollen: Unknown, but visits flowers of Acacia greggii,
Agave deserti, Argemone corymbosa, A. platyceras, Cryptantha utahensis, Encelia
farinosa, Eriogonum inflatum, Eschscholzia minutiflora var. darwinensis, Prosopis
juliflora, Stanleya pinnata.
Perdita robustula Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 308, figs. 259, 260, 326, 357.
9,S.

Perdita santaclarensis Timberlake
Calif. (Santa Clara County).

Perdita santaclarensis Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 314. ♀.

Perdita tularensis Timberlake
Calif. (Alpine, Tuolumne, and Mariposa Counties, south to Tehachapi
Pass). Pollen: Unknown, but visits flowers of Calochortus luteus, C. venustus.
Perdita tularensis Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 303, figs. 247, 248, 351. ♀.
S.

Perdita wyomingensis
***authority mismatch
sculleni Timberlake. Oreg., Wash.

Perdita sculleni sculleni Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 320, figs. 277, 278,
333,366. ♀, ♂.
Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 40 (tax. status, key).

Perdita wyomingensis
***authority mismatch
segona Timberlake. Utah, Wyo., Nev. Pollen: Unknown, but visits flowers of
Calochortus nuttallii.
Perdita sculleni segona Timberlake, 1956. Calif. Univ. Pubs. Ent. 11: 322. ♀, ♂.




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Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 40 (tax. status, key).

Perdita wyomingensis
***authority mismatch
wyomingensis Cockerell. Wyo. (Teton County). Predator: Philanthus pulcher
Dalla Torre.
Perdita wyoyriingensis Cockerell, 1922. Amer. Mus. Novitates 33: 13. ♀.

Taxonomy: Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 40 (type, tax. status, key).

Genus PERDITA Subgenus XEROMACROTERA Timberlake

Perdita subg. Xeromacrotera Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 412.
Type-species: Macrotera cephalotes Cresson. Orig. desig.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 347 (key).

Perdita cephalotes (Cresson)
Nev., Calif. (Kern and San Bernardino Counties), Ariz. (Coconino

County). Pollen: Unknown, but visits flowers of Clirysotliavmus, Gutierrezia lucida.
Macrotera cephalotes Cresson, 1878. Amer. Ent. Soc, Trans. 7: 71. ♂.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 413, figs. 109, 110, 170 (male
genitalia, geogr. and floral records). —Timberlake, 1968. Calif. Univ. Pubs. Ent. 49: 22
(geogr. and floral record). —Timberlake, 1971. Calif. Univ. Pubs. Ent. 66: 7 (geogr. and
floral record).

Genus PERDITA Subgenus XEROPHASMA Cockerell

Xerophasma Cockerell, 1923. Amer. Mus. Novitates 66: 1.

Type-species: Perdita beqiiaertiana Cockerell. Monotypic and orig. desig.
(=Xerophasvia bequaerti Cockerell).

These species are apparently crepuscular and obtain pollen only from the flowers of
Oenothera.

Taxonomy: Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 346, 348-349 (redescription, key to
included spp.).

Perdita bequaertiana Cockerell
Tex. (El Paso and Howard Counties), N. Mex. (Albuquerque). Pollen:
Collects pollen from the flowers oi Oenothera.
Xerophasma bequaerti Cockerell, 1923. Amer. Mus. Novitates 66: 2. ♀. Preocc.
Perdita bequaertiana Cockerell, 1951. In Michener, In Muesebeck, Krombein and Townes,
U. S. Dept. Agr., Agr. Monog. 2: 1089. N. name.

Taxonomy: Timberlake, 1953. Kans. Univ. Sci. Bui. 35: 962-963. 6. —Timberlake, 1954. Calif.

Univ. Pubs. Ent. 9: 349, figs. 1, 2, 115 (key). —Timberlake, 1958. Calif. Univ. Pubs. Ent. 14:

375 (geogr. record).

Perdita pallida Timberlake
Calif. (Colorado Desert), Ariz. (Yuma). Ecology: Attracted to artificial
lights near dusk through early evening. This species was readily observed in flight at
Borrego and Hopkins Well on the Colorado Desert by shining automobile headlights on
the patches of Oenothera being visited by this species. Commencing about nightfall
many males and some females were attracted to the lights of gasoline lanterns. Activity
at the flowers continued well into the darkness of early evening. These observations
were made in sand dune areas and suggest the possibility that this species may be
arenophilous in its choice of nesting sites. Pollen: Collects pollen from the flowers of
Oenothera deltoides, 0. trichocalyx, but visits these and other flowers for nectar
including Trichoste7)ia.

Perdita pallida Timberlake, 1954. Calif. Univ. Pubs. Ent. 9: 349. ♀, ♂.

NoMEN Nudum in Perdita
Perdita excisa Timberlake, 1928. Amer. Mus. Novitates 321: 10.


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