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Superfamily SCOLIOIDEA
Family MUTILLIDAE

Females are entirely wingless and have a greatly modified thorax. Males are normally fully
winged, but are brachypterous or apterous in a few species. Some females are called cowkillers
or mulekillers because of their size and painful sting, or velvet ants because of the appearance of
the dense pile covering the body.

Hosts are known for very few of the North American species. The stages of host attacked are
limited to diapausing larvae or pupae. Most recorded North American hosts are wasps or bees
which nest in the soil, in borings in wood or which build mud or resin cells or paper nests. How-
ever, there is one authenticated record of coleopterous pupae as a host. ExtraHmital genera have
been recorded as parasitizing Lepidoptera, Coleoptera and Diptera, as well as aculeate
Hymenoptera.

Brothers' important contribution on phylogeny of the Mutillidae was received too late to adopt




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his classification but it will be used in the next edition. He places the Myrmosinae here rather
than in the Tiphiidae. He raises the exotic Bradynobaeninae to family rank and transfers to it as
subfamilies the Typhoctinae and Chyphotinae (here considered a tribe of Apterogyninae). He as-
signs the other North American genera as follows: Sphaeropthalminae, Pseudomethocina, Myr-
ryiilloides, Pseudometlwca; Sphaeropthalminae, Sphaeropthalmina, Acanthopliotopsis,
Acrophotopsis, Dasymufilla, Dilophotopsis, Lomachaeta, Morsyma, Odontophotopsis,
PItotomorphus, Protophotopsis, Sniicwnnitilla, Spliaeropthahna; Mutillinae, Mutillini, Smicro-
myrmina, Timulla; and Mutillinae, Ephutini, Ephuta.

Taxonomy: Blake, 1886. Amer. Ent.. Soc, Trans. 13: 179-286, 21 figs. (N. A. spp.).

— Ashmead, 1899. N. Y. Ent. Soc, Jour. 7: 52-60 (gen. of world). —Fox, 1899. Amer. Ent.
Soc, Trans. 25: 219-292 (N. A. spp.). —Andre, 1903. In Wytsman, Gen. Ins., fasc. 11, pp.
1-77, 3 pis. (gen. of world). — Melander, 1903. Amer. Ent. Soc, Trans. 29: 291-330, pi. 4
(taxonomic notes and some revised keys). —Bradley, 1916. Amer. Ent. Soc, Trans. 42:
309-336 (spp. of east. U. S.). —Schuster, 1946. Ent. Soc. Amer., Ann. 39: 700-703 (subfam.
of New World). —Schuster, 1949. Ent. Amer. (n. s.) 29: 61-64 (key to Neotrop. subfam.).
— Mickel, 1970. Minn. Univ., Agr. Expt. Sta., Tech. Bui. 271: 1-77 (annotated bibliography
of world literature). —Brothers, 1975. Kans. Univ. Sci. Bui. 50: 483-648, 101 figs., 7 tabs,
(phylogeny).

Morphology: Reid, 1941. Roy. Ent. Soc. London, Trans. 91: 375-387, figs. 1-10, 15-20 (female,
male thorax). — Hinton, Gibbs and Silberglied, 1969. Jour. Ins. Physiol. 15: 549-552, 10 figs,
(stridulatory file). —Sheldon, 1970. Ent. News 81: 57-61 (male adaptations for female
carriage). — Debolt, 1973. Ent. Soc. Amer., Ann. 66: 100-108, 15 figs, (felt line and felt line
organ).

Subfamily APTEROGYNINAE

The North American species are all nocturnal.

Tribe CHYPHOTINI

Genus CHYPHOTES Blake

Revision: Buzicky, 1941. Ent. Amer. (n. s.) 21: 201-243. —Mickel, 1967. Amer. Ent. Soc,
Trans. 93: 125-234, 40 figs., 26 maps.

Genus CHYPHOTES Subgenus CHYPHOTES Blake

Chyphotes Blake, 1886. Amer. Ent. Soc, Trans. 13: 276.

Type-species: Chyphotes elevatus Blake. Orig. desig.
Blaketa Pate, 1948. Ent. News 59: 41. N. name for Chyphotes Blake, not Cyphotes
Burmeister 1835.

Chyphotes aenigmus Mickel
N. Mex. to Calif.; Mexico (Sonora).

Chyphotes {Chyphotes) aenigmus Mickel, 1967. Amer. Ent. Soc, Trans. 93: 147, fig. 2, map
5. ♂.

Chyphotes atriceps Mickel
Wash., Idaho, Oreg., Calif., Nev., Utah, Ariz., N. Mex., west. Tex.; Mexico
(Sonora, Chihuahua).
Chyphotes (Chyphotes) atriceps Mickel, 1967. Amer. Ent. Soc, Trans. 93: 134, fig. 1, map 1.
S.

Chyphotes auripilus Buzicky
Calif., Ariz.

Chyphotes auripilus Buzicky, 1941. Ent. Amer. (n. s.) 21: 232. ♀.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 174.

Chyphotes belfragei (Blake)
Wyo., Nebr., Colo., Utah, Ariz., N. Mex.; Mexico (Baja California, Chihuahua
to Nuevo Leon, south to Jalisco and Puebla).
Agama Belfragei Blake, 1871. Amer. Ent. Soc, Trans. 3: 263. ♂.
Sphaerophthalmail) frugala Cameron, 1896. Biol. Cent.-Amer., Hym. 2: 394. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 151, fig. 12, map 7. ♂, ♀.

Morphology: Reid, 1941. Roy. Ent. Soc London, Trans. 91: 386-387, figs. 19-20 (male thorax).




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Chyphotes bicolor Schuster
Calif. (San Diego).

Chyphotes bicolor Schuster, 1945. Pan-Pacific Ent. 21: 89. ♀.

Chyphotes calexicensis Bradley
Ariz., Calif.; Mexico (Baja California, Sonora).

Chyphotes calexicensis Bradley, 1917. Amer. Ent. Soc, Trans. 43: 284. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 144, fig. 7, map 4.

Chyphotes californicus Baker
Ariz., Nev., Calif.

Chyphotes californicus Baker, 1905. Invertebrata Pacifica 1: 117. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 171, fig. 23, map 14.

Chyphotes capitatus Mickel
Ariz.

Chyphotes (Chyphotes) capitatus Mickel, 1967. Amer. Ent. Soc, Trans. 93: 166, fig. 19, map
11. ♂.

Chyphotes cooki Mickel
Calif.

Chyphotes (Chyphotes) cooki Mickel, 1967. Amer. Ent. Soc, Trans. 93: 159, fig. 17, map 9.

i.

Chyphotes dubius Mickel
Calif., Ariz.; Mexico (Chihuahua).

Chyphotes (Chyphotes) dubius Mickel, ^967. Amer. Ent. Soc, Trans. 93: 175. ♀.

Chyphotes elevatus Blake
N. Mex., Ariz.; Mexico (Sonora).

Chyphotes elevatus Blake, 1886. Amer. Ent. Soc, Trans. 13: 276. ♀.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 155, fig. 11, map 8. ♂, ♀.

Chyphotes incredulus Mickel
Southwest Tex., south. N. Mex.; Mexico (Chihuahua, Nuevo Leon,
Durango).
Chyphotes (Chyphotes) incredulus Mickel, 1967. Amer. Ent. Soc, Trans. 93: 149, fig. 15,
map 6. ♂.

Chyphotes knulli Mickel
Tex. (Davis Mts.).

Chyphotes (Chyphotes) knulli Mickel, 1967. Amer. Ent. Soc, Trans. 93: 181. ♀.

Chyphotes mandibularis Mickel
Calif.

Chyphotes (Chyphotes) mandibularis Mickel, 1967. Amer. Ent. Soc, Trans. 93: 139, fig. 3,
map 2. ♂, ♀.

Chyphotes melaniceps (Blake)
Utah, Nev., Ariz., Calif.; Mexico (Baja California).

Photopsis melaniceps Blake, 1886. Amer. Ent. Soc, Trans. 13: 264. ♂.

Chyphotes piceiceps Baker, 1905. Invertebrata Pacifica 1: 116. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 141, fig. 6, map 3.

Chyphotes mexicanus Mickel
West. Tex., Ariz.; Mexico (Chihuahua, Sinaloa, Nayarit).

Chyphotes (Chyphotes) mexicanus Mickel, 1967. Amer. Ent. Soc, Trans. 93: 168, fig. 20,
map 12. ♂.

Chyphotes minusculus Mickel
Ariz., Nev., Calif.; Mexico (Baja California).

Chyphotes (Chyphotes) minusculus Mickel, 1967. Amer. Ent. Soc, Trans. 93: 163, fig. 18,
map 10. ♂.

Chyphotes peninsularis Fox
Calif.; Mexico (Baja California).

Chyphotes peninsularis Fox, 1899. Amer. Ent. Soc, Trans. 25: 277. cj.
Chyphotes jugatus Buzicky, 1941. Ent. Amer. (n. s.) 21: 222. ♂.
Chyphotes pilosus Buzicky, 1941. Ent. Amer. (n. s.) 21: 233. ♀.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 170, figs. 4, 21, map 13.

Chyphotes petiolatus Fox
Tex., Ariz., Nev., Calif.

Chyphotes petiolatus Fox, 1899. Amer. Ent. Soc, Trans. 25: 277. ♀.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 174.

Chyphotes pimus Mickel
Ariz. (Pima Co.).

Chyphotes (Chyphotes) pimus Mickel, 1967. Amer. Ent. Soc, Trans. 93: 181, fig. 5. ♀.

Chyphotes rugosus Mickel
Calif.

Chyphotes (Chyphotes) rugosus Mickel, 1967. Amer. Ent. Soc, Trans. 93: 177. ♀.

Chyphotes rugulosus Mickel
Ariz. (Pima Co.).

Chyphotes (Chyphotes) rugulosus Mickel, 1967. Amer. Ent. Soc, Trans. 93: 173. ♀.




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Chyphotes testaceipes Fox
Ariz.

Chyphotes testaceipes Fox, 1899. Amer. Ent. Soc, Trans. 25: 277. ♀.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 173.

Morphology: Reid, 1941. Roy. Ent. Soc. London, Trans. 91: 386-387, figs. 17-18 (thorax).

Chyphotes wasbaueri Mickel
Calif. (San Bernardino Co.).

Chyphotes (Chyphotes) wasbaueri Mickel, 1967. Amer. Ent. Soc, Trans. 93: 180. ♀.

Genus CHYPHOTES Subgenus PITANTA Pate

Pitanta Pate, 1947. Notulae Nat. 192: 1.

Type-species: Pitanta mojave Pate. Orig. desig.

Species Group Similis

Chyphotes similis Baker
Calif.; Mexico (Baja California).

Chyphotes similis Baker, 1905. Invertebrata Pacifica 1: 117. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 188, fig. 8, map 15.

Species Group Nubeculus

Chyphotes fergusoni Mickel
Oreg., Idaho, Utah, Nev., Calif.

Chyphotes (Pitanta) fergusoni Mickel, 1967. Amer. Ent. Soc, Trans. 93: 193, fig. 24, map
17. ♂.

Chyphotes nubeculus (Cresson)
Mont., Wyo., Nebr., Kans., Colo., Tex., N. Mex., Ariz.; Mexico (Sonora).

Mutilla nubecula Cresson, 1865. Ent. Soc Phila., Proc 4: 440. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 191, fig. 22, map 16.

Species Group Heathii

Chyphotes bruscus Buzicky
Ariz., Calif.

Chyphotes bruscus Buzicky, 1941. Ent. Amer. (n. s.) 21: 214. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 195, fig. 26, map 19.

Chyphotes heathii Melander
Calif. (Pacific Grove).

Chyphotes Heathii Melander, 1903. Amer. Ent. Soc, Trans. 29: 326. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 194, fig. 25, map 18.

Species Group Pallidus

buzickyi Mickel. Calif.; Mexico (Baja California).

Chyphotes (Pitanta) buzickyi Mickel, 1967. Amer. Ent. Soc, Trans. 93: 200, fig. 14. ♂, ♀.

Chyphotes marginalis Mickel
Calif. (Imperial Co.).

Chyphotes (Pitanta) marginalis Mickel, 1967. Amer. Ent. Soc, Trans. 93: 196, fig. 27. ♂.

Chyphotes minimus Mickel
Ariz.; Mexico (Sonora).

Chyphotes yninimus Mickel, 1963. Pan-Pacific Ent. 39: 186. ♂.

Chyphotes mojave (Pate)
Calif.

Pitanta mojave Pate, 1947. Notulae Nat. 192: 3. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 202, fig. 29. ♂, ♀.

Chyphotes nitidus Mickel
Calif. (Winterhaven, Glamis).

Chyphotes nitidus Mickel, 1963. Pan-Pacific Ent. 39: 188. ♀.

Taxonomy: Mickel, 1974. Ent. Soc Amer., Ann. 67: 461, fig. 1. ♂.

Chyphotes pallidus Buzicky
Ariz., Calif.

Chyphotes pallidus Buzicky, 1941. Ent. Amer. (n. s.) 21: 226. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 198, fig. 28, map 20.

Species Group Albipes

Chyphotes albipes (Cresson)
Alta., B. C, Idaho, Wash., Oreg., Calif., Nev., Utah, Ariz.
Agama albipes Cresson, 1874. Amer. Ent. Soc, Trans. 5: 99. ♂.
Chyphotes nevadensis Baker, 1905. Invertebrata Pacifica 1: 118. ♂.




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Taxonomy: Buzicky, 1941. Ent. Amer. (n. s.) 21: 215-217. 9. — Mickel, 1967. Amer. Ent. Soc,
Trans. 93: 206, fig. 34, map 21.

Chyphotes mickeli
***authority mismatch
inyoensis Mickel. Calif. (Inyo Co.).

Chyphotes (Pitanta) mickeli inijoensis Mickel, 1967. Amer. Ent. Soc, Trans. 93: 213, fig.
33. ♂.

Chyphotes mickeli
***authority mismatch
mickeli Buzicky. N. Mex., Ariz., Calif.; Mexico (Baja California).
Chyphotes mickeli luickeli Buzicky, 1941. Ent. Amer. (n. s.) 21: 217. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 209, fig. 32, map 22. ♂, ♀.

Chyphotes mickeli
***authority mismatch
polingi Buzicky. Tex., N. Mex.

Chyphotes inickeli polingi Buzicky, 1941. Ent. Amer. (n. s.) 21: 219. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 214.

Species Group Stramineus

Chyphotes stramineus Mickel
Ariz., Nev., Calif.; Mexico (Baja California).

Chyphotes (Pitanta) stramineus Mickel, 1967. Amer. Ent. Soc, Trans. 93: 215, fig. 30, map
23. ♂.

Species Group Mellipes

Chyphotes epedaphus Buzicky
Ariz., Calif.

Chyphotes epedaphus Buzicky, 1941. Ent. Amer. (n. s.) 21: 231. ♀.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 232.

Chyphotes evansi Mickel
N. Mex.

Chyphotes (Pitanta) evansi Mickel, 1967. Amer. Ent. Soc, Trans. 93: 231. ♀.

Chyphotes gracilis Mickel
Tex.; Mexico (Nuevo Leon).

Chyphotes (Pitanta) gracilis Mickel, 1967. Amer. Ent. Soc, Trans. 93: 225, fig. 37, map 26.
6.

Chyphotes mellipes (Blake)
Wash., Oreg., Idaho, Calif., Nev., Utah, Colo., west. Kans., Tex., N. Mex.,
Ariz.; Mexico (Chihuahua, Durango, Zacatecas, San Luis Potosi, Jalisco).
Agama attenuata Blake, 1872. Amer. Ent. Soc, Trans. 4: 76. ♂.
Photopsis mellipes Blake, 1886. Amer. Ent. Soc, Trans. 13: 262. ♂.
Photopsis picus Cockerell, 1895. Amer. Ent. Soc, Trans. 22: 289. ♂.
Mutilla picas (!) Dalla Torre, 1897. Cat. Hym., v. 8, p. 73.
Mutilla tenula Dalla Torre, 1897. Cat. Hym., v. 8, p. 91. N. name for Agama attenuata

Blake, not Mutilla attenuata Spinola.
Chyphotes punctatus Fox, 1899. Amer. Ent. Soc, Trans. 25: 276. ♀.
Chyphotes striatus Buzicky, 1941. Ent. Amer. (n. s.) 21: 228. ♀.
Chyphotes segregatus Buzicky, 1941. Ent. Amer. (n. s.) 21: 229. ♀.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 98: 216, fig. 31, map 24.

Chyphotes pixus Buzicky
Ariz. (Gila Bend).

Chyphotes pixus Buzicky, 1941. Ent. Amer. (n. s.) 21: 227. ♀.

Chyphotes sonorus Mickel
Ariz.; Mexico (Sonora).

Chyphotes (Pitayita) sonorus Mickel, 1967. Amer. Ent. Soc, Trans. 93: 223, fig. 35, map 25
S.

Chyphotes subulatus Buzicky
Ariz.

Chyphotes subulatus Buzicky, 1941. Ent. Amer. (n. s.) 21: 224. ♂.

Taxonomy: Mickel, 1967. Amer. Ent. Soc, Trans. 93: 226, fig. 39.

Subfamily TYPHOCTINAE
Revision: Krombein and Schuster, 1957. Ent. Soc Wash., Proc 59: 209-232, 2 pis.
All known species are diurnal insects.

Genus TYPHOCTES Ashmead

Typhoctes Ashmead, 1899. N. Y. Ent. Soc, Jour. 7: 53.

Type-species: Mutilla peculiaris Cresson. Orig. desig.




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Avonimiifilla Mickel, 1936. Ent. Soc. Amer., Ann. 29: 295.

Type-species: Anomtnutilla difficilis Mickel. Orig. desig.

Taxonomy: Brothers, 1970. Kans. Ent. Soc, Jour. 43: 305 (generic redescription, female).

Morphology: Reid, 1941. Roy. Ent. Soc. London, Trans. 91: 385-386, figs. 15-16 (female

thorax).

Typhoctes peculiaris
***authority mismatch
mirabilis (Cockerell). Kans., Colo., Tex., N. Mex., Ariz.; Mexico (Chihuahua).

Chijphotes mirabilis Cockerell, 1896. Canad. Ent. 28: 285. ♀.

Typhoctes peculiaris
***authority mismatch
peculiaris (Cresson). Wash., Idaho, Utah, Ariz., Calif.

Mutilla peculiaris Cresson, 1875. Amer. Ent. Soc, Trans. 5: 119. ♀.

Anommutilla difficilis Mickel, 1936. Ent. Soc Amer., Ann. 29: 295. ♂.

Morphology: Hermann, 1975. Ent. Soc. Amer., Ann. 68: 882-884, 5 figs, (sting).

Typhoctes striolatus Krombein and Schuster
Ariz, to western Tex.

Typhoctes striolatus Krombein and Schuster, 1957. Ent. Soc. Wash., Proc 59: 223, fig. 2. ♀.
6.

Typhoctes williamsi Krombein and Schuster
Southern Calif.

Typhoctes williamsi Krombein and Schuster, 1957. Ent. Soc. Wash., Proc. 59: 220, figs. 1, 4.
6, 9.

Subfamily MUTILLINAE
Tribe MUTILLINI

Blake (1886. Amer. Ent. Soc, Trans. 13) described three species o{ Mutilla, mislabeled as com-
ing from Florida, from the Harris collection. Fox, 1898 (Ent. News 9: 14) stated that M. ajax was
a synonym of the African Psammothervia flabellata {¥.), M. floridana a synonym of the Eu-
ropean Dasylabris maura (L.), and M. trisignata a synonym of the European D. arenaria {¥.).

Genus TIMULLA Ashmead

Genus TIMULLA Subgenus TIMULLA Ashmead

Timulla Ashmead, 1899. N. Y. Ent. Soc, Jour. 7: 55.

Type-species: Mutilla dubitata Smith. Orig. desig.

Revision: Mickel, 1937. Ent. Amer. (n. s.) 17: 1-119 (N. Amer. spp.).

Only the typical subgenus occurs in the New World. Most species are diurnal, but a few spe-
cies with enlarged ocelli, such as ocellaria Mick., are crepuscular or nocturnal.

Timulla barbata (Fox)
Mo., La.

Mutilla barbata Fox, 1899. Amer. Ent. Soc, Trans. 25: 272. ♂.

Timulla barbigera
***authority mismatch
barbigera (Bradley). Va. south to Fla., west to S. Dak. and Tex. in U. and L. Austr.
Zones.

Mutilla (Timulla) barbigera Bradley, 1916. Amer. Ent. Soc, Trans. 42: 206. ♂.

Timulla barbigera
***authority mismatch
rohweri Mickel. Kans., Okla. in U. Sonor. Zone.

Timulla (Timulla) barbigera rohweri Mickel, 1937. Ent. Amer. (n. s.) 17: 58. ♂.

Timulla compressicornis Mickel
N. C. to Tex. in L. Austr. Zone.

Timulla (Tiiriulla) compressicornis Mickel, 1937. Ent. Amer (n. s.) 17: 59. ♂.

Timulla contigua Mickel
Tex. (Brownsville).

Timulla (Timulla) contigua Mickel, 1937. Ent. Amer. (n. s.) 17: 107. ♀.

Timulla dubitata
***authority mismatch
dubitata (Smith). N. Y. south to Fla., west to Okla. and Tex. in L. Austr. Zone.

Mutilla dubitata Smith, 1855. Cat. Hym. Brit. Mus., v. 3, p. 60. ♀.

Mutilla dubiatta (!) Blake, 1886. Amer. Ent. Soc, Trans. 13: 201. ♀.

Biology: Sheldon, 1970. Ent. News 81: 59-60 (mating behavior).

Timulla dubitata
***authority mismatch
fugitiva Mickel. N. J. south to Ga., west to Nebr. and Tex. in U. Austr. Zone.

Timulla (Timulla) dubitata fugitiva Mickel, 1937. Ent. Amer. (n. s.) 17: 39. ♂.

Timulla dubitatiformis Mickel
Mass. south to Fla., west to Mont, and Tex.

Timulla (Timulla) dubitatiformis Mickel, 1937. Ent. Amer. (n. s.) 17: 102. ♀.




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Timulla euphrosyne Mickel
Fla., Tex.

Timulla (Timulla) euphrosyne Mickel, 1937. Ent. Amer. (n. s.) 17: 64. ♀.

Timulla euterpe (Blake)
N. C, Fla., La., Tex.

Mutilla Euterpe Blake, 1879. Amer. Ent. Soc, Trans. 7: 249. ♀.

Timulla ferrugata (Fabricius)
N. J. south to Fla., west to Okla. and Tex. in U. and L. Austr. Zones.
Host: Eumenes fraternus Say.

Mutilla ferrugata Fabricius, 1805. Systema Piezatorum, p. 438. ♀.

Mutilla rufa Lepeletier, 1845. Hist. Nat. Ins. Hym., v. 3, p. 631. ♂.

Mutilla Promethea Blake, 1871. Amer. Ent. Soc, Trans. 3: 229. ♂.

Biology: Fattig, 1943. Emory Univ. Mus. Bui. 1: 3 (host).

Timulla floridensis (Blake)
S. C. to Tex. in L. Austr. Zone.

Mutilla floridensis Blake, 1879. Amer. Ent. Soc, Trans. 7: 249. ♂.

Timulla grotei (Blake)
Colo., Utah, Tex., N. Mex., Ariz.; Mexico (Jalisco, Guanajuato).

Mutilla Grotei Blake, 1871. Amer. Ent. Soc, Trans. 3: 228. ♂.

Mutilla fulviventris Gerstaecker, 1874. Arch. Naturgesch. 40: 323. ♂.

Taxonomy: Mickel, 1937. Ent. Amer. (n. s.) 17: 78.

Timulla hollensis
***authority mismatch
hollensis (Melander). Mass., N. Y., Md.

Mutilla Sayi var. hollensis Melander, 1903. Amer. Ent. Soc, Trans. 29: 324. ♂.

Timulla hollensis
***authority mismatch
melanderi Mickel. N. J., Pa., Md., Va.

Timulla (Timulla) hollensis melanderi Mickel, 1937. Ent. Amer. (n. s.) 17: 101. ♂.

Timulla huntleyensis Mickel
Mont. (Huntley).

Timulla {Timulla) hniitleyeusis Mickel, 1937. Ent. Amer. (n. s.) 17: 82. ♂.

Timulla kansana Mickel
Kans.

Timulla (Timulla) kansayia Mickel, 1937. Ent. Amer. (n. s.) 17: 94. ♂.

Timulla leona (Blake)
Wis. south to Fla., west to Nebr. and Tex. in U. and L. Austr. Zones. Host:
Bembix troglodytes Handl.?

Mutilla Leona Blake, 1871. Amer. Ent. Soc, Trans. 3: 230. ♀.

Biology: Evans, 1957. Studies in Compar. Ethology Bembix, p. 134, fig. 36 (entering burrows
of B. troglodytes).

Timulla navasota
***authority mismatch
coahuila Krombein. Tex. west of 100th Meridian, N. Mex., Ariz.

Timulla (Timulla) navasota nebulosa Mickel, 1937. Ent. Amer. (n. s.) 17: 29. ♂. Preocc. by

Mickel, 1935.
Timulla (Timulla) navasota coahuila Krombein, 1951. U. S. Dept. Agr., Monog. 2: 771. N.
name.

Timulla navasota
***authority mismatch
navasota (Bradley). Tex. east of 100th Meridian.

Mutilla (Timulla) navasota Bradley, 1916. Amer. Ent. Soc, Trans. 42: 213. ♂.

Timulla neobule Mickel
Ariz.

Timulla (Timulla) neobule Mickel, 1937. Ent. Amer. (n. s.) 17: 44. ♂.

Timulla nicholi Mickel
Ariz.

Timulla (Timulla) nicholi Mickel, 1937. Ent. Amer. (n. s.) 17: 47. ♀.

Timulla nitela Mickel
Ariz. (Douglas).

Timulla (Timulla) nitela Mickel, 1937. Ent. Amer. (n. s.) 17: 43. ♂.

Timulla oajaca (Blake)
Tex., N. Mex., Ariz.; Mexico (Morelos, Jalisco, Colima).

Mutilla oajaca Blake, 1871. Amer. Ent. Soc, Trans. 3: 228. ♂ (♀ misdet.).

Mutilla ardens Gerstaecker, 1874. Arch. Naturgesch. 40: 323. ♂.

Mutilla mazatlanae Cameron, 1894. Biol. Cent.-Amer., Hym., v. 2, p. 294. ♂.

Mutilla nestor Fox, 1899. Amer. Ent. Soc, Trans. 25: 271. ♂.

Mutilla omata Howard, 1901. Insect Book, pi. 8, fig. 21. ♀.

Taxonomy: Mickel, 1938. Roy. Ent. Soc. London, Trans. 87: 641.

Biology: Linsley, 1960. Pan-Pacific Ent. 36: 36 (mating behavior).

Timulla ocellaria
***authority mismatch
ocellaria Mickel. Ohio, Ind., Ky., Tenn., Ga. Host: Ligyrus gibbosus (Deg.), pupae.
Timulla (Ti7nulla) ocellaria ocellaria Mickel, 1937. Ent. Amer. (n. s.) 17: 91. ♂.

Biology: Fattig, 1943. Emory Univ. Mus. Bui. 1: 5 (host record).




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Timulla ocellaria
***authority mismatch
rufidorsa Mickel. Miss., Ark., Mo., Kans., Okla., Tex.

TiniuUa (TimuUa) ocellaria ntfidorna Mickel, 1937. Ent. Amer. (n. s.) 17: 93. ♂.

Timulla ornatipennis (Bradley)
N. J. south to Fla. west to Tenn. and Miss.

Mutilla (Timnlla) ornatipennis Bradley, 1916. Amer. Ent. Soc, Trans. 42: 205. ♀, ♂.

Timulla rufosignata (Bradley)
N. C, Ga., Fla. Host: Ground-nesting eumenid wasp.

Mutilla {Timulla) rufosignata Bradley, 1916. Amer. Ent. Soc, Trans. 42: 212. ♂.

Taxonomy: Krombein, 1953. Ent. Soc. Wash., Proc. 55: 128. ♀.

Biology: Fattig, 1943. Emory Univ. Mus. Bui. 1: 6 (ground-nesting host misdet. as Odynerus
erin7iys (!) Lep.).

Timulla sayi (Blake)
Tex. (Piano).

Mutilla Sayi Blake, 1871. Amer. Ent. Soc, Trans. 3: 229. ♂.

Timulla subhyalina Mickel
111., Iowa, Minn., N. Dak., S. Dak., Nebr., Kans., Mont., Oreg., B. C.

Timulla {Timulla) subhyalina Mickel, 1937. Ent. Amer. (n. s.) 17: 97. ♂.

Timulla suspensa
***authority mismatch
jonesi Mickel. Tex., Ariz.

Timulla (Timulla) suspensa jonesi Mickel, 1937. Ent. Amer. (n. s.) 17: 90. ♂.

Timulla suspensa
***authority mismatch
sonora Mickel. Kans. and Colo, south to Tex. and Ariz.

Timulla (Timulla) suspensa sonora Mickel, 1937. Ent. Amer. (n. s.) 17: 84. ♀, ♂.

Timulla suspensa
***authority mismatch
suspensa (Gerstaecker). Ariz.

Mutilla suspensa Gerstaecker, 1874. Arch. f. Naturgesch. 40: 299. ♀.

Timulla tolerata Mickel
Mo., Ga.

Timulla (Timulla) tolerata Mickel, 1937. Ent. Amer. (n. s.) 17: 109. ♂.

Timulla tyro Mickel
Ariz., Cahf.

Timulla (Timulla) tyro Mickel, 1937. Ent. Amer. (n. s.) 17: 25. ♀, ♂.

Timulla vagans
***authority mismatch
rufinota Mickel. D. C, S. C, Ga., Fla.

Timulla (Timulla) vagans rufinota Mickel, 1937. Ent. Amer. (n. s.) 17: 78. ♀, ♂.

Timulla vagans
***authority mismatch
vagans (Fabricius). Vt. and Ont. south to Ala., west to B. C. and N. Mex.

Mutilla vagans Fabricius, 1798. Sup. Ent. System., p. 282. ♀.

Mutilla (Mutilla) hexagona Say, 1836. Boston Jour. Nat. Hist. 1: 295. ♂.

Mutilla oniativentris Cresson, 1865. Ent. Soc. Phila., Proc 4: 438. ♀.

Mutilla Briaxus Blake, 1871. Amer. Ent. Soc, Trans. 3: 227. ♂.

Mutilla Canadensis Provancher, 1887. Addit. Corr. Faune Ent. Canada Hym., p. 250. ♂.
Preocc.

Mutilla secunda Dalla Torre, 1897. Cat. Hym., v. 8, p. 84. N. name for Mutilla Canadensis
Provancher, not Mutilla ca7iade7isis Blake.

Biology: Fattig, 1936. Ent. News 47: 51-52 (mating behavior). — Shappirio, 1948. Brooklyn
Ent. Soc, Bui. 42: 163 (mating behavior).

Timulla wileyae Mickel
Ark., La., Okla., Tex.

Timulla (Timulla) wileyae Mickel, 1937. Ent. Amer. (n. s.) 17: 41. ♀.

Tribe EPHUTINI

Genus EPHUTA Say

Revision: Bradley, 1916. Amer. Ent. Soc, Trans. 42: 187-198. —Schuster, 1951. N. Y. Ent.
Soc, Jour. 59: 1-43, 5 pis. (Part I containing only gen. diag. and key to spp.). —Schuster,
1957. N. Y. Ent. Soc, Jour. 64: 7-84 (Part II of revis. treating Species Group Grisea).

All of the species are diurnal.

Genus EPHUTA Subgenus EPHUTA Say

Ephuta Say, 1836. Boston Jour. Nat. Hist. 1: 297.

Type-species: Mutilla (Ephuta) scrupea Say. Desig. by Ashmead, 1899.
Rhoptromutilla Andre, 1903. In Wytsman, Gen. Ins., fasc. 11, p. 43.

Type-species: Mutilla chrysodora Perty. Orig. desig.
Ephutopsis Ashmead, 1904. Canad. Ent. 36: 6.

Type-species: Mutilla odontophora Cameron. Desig. by Mickel, 1928.




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The few authenticated host records indicate that species of this subgenus have various species
of Pompilidae as hosts. The spider wasp hosts have diverse nesting habits. Some nest in borings
in wood (Dipogoyi), some construct mud cells beneath stones {Phanagenia), and some nest in the
ground (? Anoplius fratemus (Bks.)).

Ephuta albiceps Schuster
Tex. (Tyler).

Ephiita albiceps Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 31 (in key). 9.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 75 (female description).

Ephuta argenticeps Schuster
Calif.

Ephuta caiifornica Schuster, 1951. N. Y. Ent. Soc, Jour. 59: footnote on p. 25. d (On p. 15
Schuster states that there is only one sp. in Calif., and in key to females, argenticeps is
the only sp. stated to be from Calif.).
Ephuta argenticeps Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 26, 32 (in key). ♂, ♀.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 16 (male, female description).

Ephuta auricapitis Schuster
Tex. (Edinburgh). Possibly the opposite sex of ecarinata ecarinata
Schuster.
Ephuta auricapitis Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 31 (in key). 9.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 76 (female description).

Ephuta baboquivari Schuster
Ariz. (Baboquivari Mts.)

Ephuta baboquivari Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 31 (in key), figs. 39, 41. ♀.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 79 (female description).

Ephuta battlei
***authority mismatch
battlei Bradley. Coastal N. C, Ga., Fla.

Ephuta battlei Bradley, 1916. Amer. Ent. Soc, Trans. 42: 195. ♂.

Taxonomy: Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 29, figs. 5, 44 (in key; reduced to subsp.
rank).

Ephuta battlei
***authority mismatch
confusa Schuster. Piedmont and west, coastal plain, N. C, Ga., Ala.

Ephuta battlei confusa Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 29 (in key), fig. 44. ♂.

Ephuta battlei
***authority mismatch
microcellaria Schuster. N. J. (Cape May).

Ephuta battlei microcellaria Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 29 (in key), fig. 44.
6.

Ephuta battlei
***authority mismatch
transitionalis Schuster. Transit, and U. Austr. Zones, Mass. to Va., Ohio.

Ephuta battlei transitionalis Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 28 (in key), fig. 44.
d.

Ephuta cephalotes Schuster
N. Dak., S. Dak., Wyo., Ariz., west. Tex. The opposite sex is possibly
m inula Schuster.
Ephuta cephalotes Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 24 (in key), figs. 10, 20, 46. ♂.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 68 (male description).
coioradella Schuster. Colo. Possibly the opposite sex of grisea Bradley.

Ephuta coioradella Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 32 (in key). 9.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 73 (female description).

Ephuta conchate Mickel
N. Y., Mich., 111., Iowa, Minn., S. Dak., Nebr., Kans.
Ephuta conchate Mickel, 1928. Minn. State Ent, Rpt. 19: 111. ♂.

Taxonomy: Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 32 (in key).

Ephuta copano (Blake)
Tex.; Mexico.

Mutilla Copano Blake, 1871. Amer. Ent. Soc, Trans. 3: 232. ♂.
Mutilla susura Melander, 1903. Amer. Ent. Soc, Trans. 29: 324. ♂.

Taxonomy: Schuster, 1958. hi Krombein, U. S. Dept. Agr., Monog. 2, Sup. 1: 105 (synonymy).

Ephuta ecarinata
***authority mismatch
ecarinata Schuster. Tex.; Mexico. The opposite sex is possibly sudatrix (Melander)
or auricapitis Schuster.
Ephuta ecarinata ecarinata Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 26 (in key). ♂.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 49 (male description).




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Ephuta ecarinata
***authority mismatch
neomexicana Schuster. N. Mex.

Ephuta ecarinata neomexicana Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 26 (in key). ♂.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 52 (male description).

Ephuta ecarinata
***authority mismatch
pima Schuster. Ariz.

Ephuta ecarinata pima Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 26 (in key). ♂.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 51 (male description).

Ephuta eurygnathus Schuster
N. C. (Southern Pines).

Ephuta eurygnathus Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 29 (in key), fig. 11. ♂.

Ephuta floridana
***authority mismatch
dietrichi Schuster. Ala., Miss.

Ephuta floridana dietrichi Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 32 (in key), figs. 35,
36. ♀.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 40 (female description).

Ephuta floridana
***authority mismatch
floridana Schuster. Fla.

Ephuta floridana floridana Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 25, 32 (in key). ♂, ♀.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 34 (male, female description).

Ephuta grisea
***authority mismatch
fuscosericea Schuster. Alta., Mont., Utah, N. Dak.

Ephuta grisea fuscosericea Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 27 (in key), figs. 14,
15,42. ♂.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 14 (male description).

Ephuta grisea
***authority mismatch
grisea Bradley. Colo. The opposite sex is possibly coloradella Schuster.
Ephuta grisea Bradley, 1916. Amer. Ent. Soc, Trans. 42: 194. ♂.

Taxonomy: Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 27, fig. 21 (reduced to subsp. rank).
—Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 13 (male description).

Ephuta margueritae
***authority mismatch
margueritae Schuster. Fla.

Ephuta margueritae margueritae Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 25, 30 (in key),
figs. 17, 46. ♂, ♀.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 56 (male, female description).

Ephuta margueritae
***authority mismatch
xanthocephala Schuster. Pa., N. C, Tenn. Host: "cocoons under stones",

presumably mud cells oi Phanagenia bombycina (Cr.) or of a species of Auplopus.
Ephuta margueritae xanthocephala Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 25, 30 (in
key), figs. 25, 46. cj, 9.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 63 (male, female description).
Biology: Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 14 (rearing note).

Ephuta minuta
***authority mismatch
minuta Schuster. Tex. (Brownsville, Fedor, College Sta.). Possibly the opposite sex of

Ephuta cephalotes Schuster

Ephuta minuta minuta Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 33 (in key). 9.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 71 (tentative sex association).

Ephuta minuta
***authority mismatch
modesta Schuster. Ariz. (Tempe).

Ephuta minuta modesta Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 32 (in key), figs. 31, 32.
9.

Ephuta pauxilla
***authority mismatch
pauxilla Bradley. Minn, to Ala., east to Maine and Fla. Host: Dipogon s. sayi Bks.
Almost certainly the male of puteola (Blake).
Ephuta ixtuxilla Bradley, 1916. Amer. Ent. Soc, Trans. 42: 197. ♂.

Taxonomy: Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 29, figs. 1, 12, 37, 43 (in key; reduced to
subsp. rank).

Biology: Evans and Yoshimoto, 1962. Ent. Soc. Amer., Misc. Pub. 3: 104 (host record).

Ephuta pauxilla
***authority mismatch
texanella Schuster. Tex., La., Miss.

Ephuta pauxilla texanella Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 29 (in key), figs. 13,
43. ♂.

Ephuta psephenophila Schuster
Ga. (Stone Mt.) The opposite sex is possibly tentativa Schuster.
Ephuta psephenophila Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 25 (in key). ♂.




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Epiiuta ocellaria Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 36, fig. 18. MS name changed to

Ephuta psephenophila
***authority mismatch
(teste R. M. Schuster).

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 44 (male description; tentative sex
association).

Ephuta puteola (Blake)
Mass. south to Fla., west to Iowa, Tex. Possibly the opposite sex of p.

Ephuta pauxilla
***authority mismatch
Brad.
Mutilla puteola Blake, 1879. Amer. Ent. Soc, Trans. 7: 252. ♀.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 40 (female description in key; putative
sex association).

Ephuta rufisquamis (Andre)
Ariz., south Calif. The opposite sex is possibly timiacacori Schuster.
RhoptromutUla rufisquamis Andre, 1905. Ztschr. System. Hym. Dipt. 5: 366. ♂.

Taxonomy: Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 24 (in key). —Schuster, 1957. N. Y. Ent.
Soc, Jour. 64: 65 (male description; tentative sex association).

Ephuta sabaliana
***authority mismatch
fattigi Schuster. Ga. (Tifton).

Ephuta sabaliana fattigi Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 28 (in key). ♂.

Ephuta sabaliana
***authority mismatch
sabaliana Schuster. Fla. Host: Anoplius fraternus (Bks.)?

Ephuta sabaliayia sabaliana Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 28 (in key). ♂.

Biology: Krombein, 1955. Ent. Soc. Wash. Proc 57: 225-226.

Ephuta scrupea (Say)
N. J. south to Ga. and Ala., Mich., Wis., Ohio, W. Va., Mo., Ark., Tex. Host:
Phanagenia bombycina (Cr.).
Mutilla (Ephuta) scrupea Say, 1836. Boston Jour. Nat. Hist. 1: 297. ♂.

Taxonomy: Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 25, 34, figs. 2-4, 27-30, 42 (male, female
in key). —Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 26 (male, female description).

Ephuta slossonae
***authority mismatch
monochroa Schuster. Tex. (Rock Island).

Ephuta slossonae monochroa Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 34 (in key). 9.

Ephuta slossonae
***authority mismatch
slossonae (Fox). Fla.

Mutilla Slossonae Fox, 1899. Amer. Ent. Soc, Trans. 25: 273. ♂.

Taxonomy: Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 34 (in key; reduced to subsp. rank).

Ephuta spinifera Schuster
N. J. to Ga.

Ephuta spi7iifera Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 27, 34 (in key). ♂, ♀.

Ephuta stenognatha
***authority mismatch
psephenophora Schuster. North. Fla.

Ephuta stenognatha psephenophora Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 28. ♂.

Ephuta stenognatha
***authority mismatch
stenognatha Schuster. Cent, and south. Fla.

Ephuta stenognatha stenognatha Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 28 (in key), figs.
7,8. ♂.

Ephuta sudatrix (Melander)
Tex. (Fedor). Possibly the opposite sex of ecarinata ecarinata Schuster.

Mutilla sudatrix Melander, 1903. Amer. Ent. Soc, Trans. 29: 325. ♀.

Taxonomy: Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 30 (in key). —Schuster, 1957. N. Y. Ent.
Soc, Jour. 64: 54 (female description; tentative sex association).

Ephuta tegulicia Bradley
Tex., Ariz. Possibly a syn. or subsp. of idiasfa (Cameron) from Mexico.
Ephuta tegulicia Bradley, 1916. Amer. Ent. Soc, Trans. 42: 193. ♂.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 47 (male description).

Ephuta tentativa Schuster
Ga. (Atlanta, Ochlochnee). Possibly the opposite sex of psephenophila
Schuster.
Ephuta tentativa Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 34 (in key). 9.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 42 (female description; tentative sex
association).

Ephuta tumacacori Schuster
Ariz. (Tumacacori Mts.). Possibly the opposite sex of rufisqua^nis
(Andre).
Ephuta tumacacori Schuster, 1951. N. Y. Ent. Soc, Jour. 59: 30 (in key). 9.

Taxonomy: Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 82 (female description; tentative sex
association).




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Genus EPHUTA Subgenus XENOCHILE Schuster

Ephuta subg. Xenochile Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 8.

Type-species: Ephuta (Xenochile) kromheini Schuster. Orig. desig.

Ephuta krombeini Schuster
Ariz. (Ramsey Canyon in Huachuca Mts.).

Ephuta (Xenochile) krombeini Schuster, 1957. N. Y. Ent. Soc, Jour. 64: 9. ♂.

Subfamily SPHAEROPTHALMINAE

Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci. BuL, Biol. Ser. 8, no. 4: 1-26, 7 figs,
(males of Nevada Test Site).

Tribe SPHAEROPTHALMINI

Taxonomy: Schuster, 1958. Ent. Amer. (n. s.) 37: 1-130, 7 pis. (Part II of a tribal revision of
males containing keys to genera and subgenera and revisions of Acrophotopsis Schuster,
Dilophotopsis Schuster and Acanthophotopsis Schuster).

The majority of species in this tribe are nocturnal, but all members of Morsyma Fox,
Sphaeropthahna subg. Sphaeropthahna Blake, and Protophotopsis Schuster, and most species of
Photomorphus subg. Photomorphus Viereck, are diurnal.

Genus MORSYMA Fox

Morsyma Fox, 1899. Amer. Ent. Soc, Trans. 25: 287.

Type-species: Morsyma Ashyneadii Fox. Orig. desig.

ashmeadii Fox. Calif.

Morsyma Ashmeadii Fox, 1899. Amer. Ent. Soc, Trans. 25: 287. "♀"=♂.

Taxonomy: Krombein, 1940. Amer. Ent. Soc, Trans. 65: 420.

Genus ACROPHOTOPSIS Schuster

Acrophotopsis Schuster, 1958. Ent. Amer. (n. s.) 37: 4 (in key), 61.

Type-species: Acrophotopsis eurygnathtis Schuster. Orig. desig.

Acrophotopsis campylognathus Schuster
Calif. (Riverside Co.); Mexico (Baja California).

Acrophotopsis campylognathus Schuster, 1958. Ent. Amer. (n. s.) 37: 11 (in key), 69. ♂.

Acrophotopsis eurygnathus Schuster
Ariz., Nev.

Acrophotopsis eurygnathus Schuster, 1958. Ent. Amer. (n. s.) 37: 10 (in key), 65, pi. 1, figs.
1,2. ♂.

Genus DILOPHOTOPSIS Schuster

Dilophotopsis Schuster, 1958. Ent. Amer. (n. s.) 37: 5 (in key), 71.
Type-species: Mutilla concolor Cresson. Orig. desig.

Dilophotopsis concolor
***authority mismatch
concolor (Cresson). Kans. to Colo, and Wyo., Tex., N. Mex., Ariz.; Mexico (Zacatecas,
Durango, Jalisco).
Mutilla concolor Cresson, 1865. Ent. Soc. Phila., Proc 4: 890. ♂.
Odontophotopsis alamonis Viereck, 1904. Amer. Ent. Soc, Trans. 30: 87. ♂.

Taxonomy: Schuster, 1958. Ent. Amer. (n. s.) 37: 84 (reduction to subsp. rank and syn.).

Dilophotopsis concolor
***authority mismatch
crassa (Viereck). B. C, Mont., southwest. Idaho, west. Colo., Nev., Utah, Ariz., Calif.,
Wash.
Odontophotopsis crassus Viereck, 1924. Canad. Ent. 56: 112. ♂.
Dilophotopsis concolor utahensis Schuster, 1958. Ent. Amer. (n. s.) 37: 84 (in key), 87. ♂.

Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci. Bui., Biol. Ser. 8, no. 4: 8 (synonymy).

Dilophotopsis concolor
***authority mismatch
laredo Schuster. Tex. (Winterhaven, Laredo, Cotulla).

Dilophotopsis concolor laredo Schuster, 1958. Ent. Amer. (n. s.) 37: 84 (in key), 86. ♂.

Dilophotopsis concolor
***authority mismatch
paron (Cameron). Nev., Ariz., Calif.; Mexico (Sonora, Baja California).

Sphaerophthalma (!) paron Cameron, 1896. Biol. Cent.-Amer., Hym., v. 2, p. 381. ♂.

Dilophotopsis concolor sonorensis Schuster, 1958. Ent. Amer. (n. s.) 37: 84 (in key), 88. ♂.




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Taxonomy: Mickel, 1965. Ent. Soc. Wash., Proc. 67: 1 (synonymy).

Dilophotopsis stenognatha Schuster
Ariz. (Tucson).

Dilophotopsis stenognatha Schuster, 1958. Ent. Amer. (n. s.) 37: 11 (in key), 74. ♂.

Taxonomy: Mickel, 1963. Pan-Pacific Ent. 39: 184 (female description).

Genus ACANTHOPHOTOPSIS Schuster

Acanthophotopsis Schuster, 1958. Ent. Amer. (n. s.) 37: 5 (in key), 88.

Type-species: Acantliopliotopsis falciforviis Schuster. Orig. desig.

Acanthophotopsis bequaertii Schuster
Ariz. (Hereford).

Acanthophotopsis bequaertii Schuster, 1958. Ent. Amer. (n. s.) 37: 12 (in key), 101. ♂.

Acanthophotopsis bifurca Schuster
Okla., Tex., N. Mex.

Acanthophotopsis bifurca Schuster, 1958. Ent. Amer. (n. s.) 37: 13 (in key), 98, pi. 3, fig. 3.
6.

Acanthophotopsis dorophora Schuster
Ariz. (Tucson, Yuma).

Acanthophotopsis dorophora Schuster, 1958. Ent. Amer. (n. s.) 37: 13 (in key), 104, pi. 2,
fig. 1, pi. 3, fig. 2. ♂.

Acanthophotopsis evansii Schuster
Tex. (Alpine and Chisos Mts.); Mexico (Durango).

Acanthophotopsis evansii Schuster, 1958. Ent. Amer. (n. s.) 37: 12 (in key), 93, pi. 3, fig. 1.
6.

Acanthophotopsis falciformis
***authority mismatch
falciformis Schuster. Nev., Calif.; Mexico (Zacatecas).

Acanthophotopsis falcifonnis falcifomiis Schuster, 1958. Ent. Amer. (n. s.) 37: 13 (in key),
108, pi. 2, figs. 2, 3. ♂.

Acanthophotopsis falciformis
***authority mismatch
furcisterna Schuster. Ariz. (Tucson, Phoenix, Ajo).

Acanthophotopsis falciformis furcistema Schuster, 1958. Ent. Amer. (n. s.) 37: 14 (in key),
111. ♂.

Genus SPHAEROPTHALMA Blake

In addition to the host records cited under several species in this genus, Parker and Bohart,
1968 (Pan-Pacific Ent. 44: 2) record an unidentified species of Sphaeropthalma from a twig nest
oi Leptochilus washo Parker, and Hurd and Powell, 1958 (Pan-Pacific Ent. 34: 152-153) record an
unidentified female of this species as leaving a burrow of Colletes stepheni Timb.

Genus SPHAEROPTHALMA Subgenus MICROMUTILLA Ashmead

Micromutilla Ashmead, 1899. N. Y. Ent. Soc, Jour. 7: 59.

Type-species: Photopsis nanus Ashmead. Orig. desig.

Sphaeropthalma acontia (Fox)
N. Mex., Ariz., Nev.

Photopsis nanus Ashmead, 1896. Amer. Ent. Soc, Trans. 23: 181. ♂. Preocc.
Mutilla acontius Fox, 1899. Amer. Ent. Soc, Trans. 25: 266. ♂.
Mutilla Ashmeadii Fox, 1899. Amer. Ent. Soc, Trans. 25: 289. N. name.

Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci. Bui., Biol. Ser. 8, no. 4: 9 (synonymy).

Sphaeropthalma becki Ferguson
Nev., Calif.

Sphaeropthalma (Micromutilla) becki Ferguson, 1967. Brigham Young Univ. Sci. BuL,
Biol. Ser. 8, no. 4: 9, fig. 5. ♂.

Sphaeropthalma bellerophon (Fox)
Tex., N. Mex., Ariz.

Mutilla bellerophon Fox, 1899. Amer. Ent. Soc, Trans. 25: 254. ♂.

Sphaeropthalma brachyptera Schuster
Calif., Nev., Ariz.; Mexico (Sonora).

Photopsis brachyptera Schuster, 1945. Pan-Pacific Ent. 21: 149. ♂.

Sphaeropthalma (Micromutilla) yavapai Schuster, 1958. Ent. Amer. (n. s.) 37: 19. ♂.

Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci. BuL, Biol. Ser. 8, no. 4: 11 (synonymy).

Sphaeropthalma difficilis (Baker)
Calif., Nev., Ariz., N. Mex.; Mexico (Baja California). Host: Auplopus
architect/us metallicus (Bks.).
Photopsis difficilis Baker, 1905. Invertebrata Pacifica 1: 114. ♂.

Sphaeropthahna (Micromutilla) maricopella purism.ella Schuster, 1958. Ent. Amer. (n. s.)
37: 17. ♂.




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Sphaeropthahiia {Mkrontutilla) maricopella maricopella Schuster, 1958. Ent. Amer. (n. s.)

37: 17. ♂.
Sphaeropthalma (M icromtitilla) maricopella castanea Schuster, 1958. Ent. Atner. (n. s.)

37: 17. ♂.
Sphaeropthahna (MkrovintUla) californiense califortiiense Schuster, 1958. Ent. Amer. (n.

s.)37: 18. ♂.
Sphaeropthahna (Micronuifilla) californiense fnscatella Schuster, 1958. Ent. Amer. (n. s.)

37: 18. ♂.
Sphaeropthahna (Micromutilla) quijotoa quijotoa Schuster, 1958. Ent. Amer. (n. s.) 37: 18.

6.
Sphaeropthahna (Mcroimitilla) quijotoa parrasia Schuster, 1958. Ent. Amer. (n. s.) 37: 18.

cJ.
Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci. Bui., Biol. Ser. 8, no. 4: 11 (synonymy).
Biology: Ferguson, 1962. Univ. Calif. Pubs. Ent. 27: tab. 10 (host record). jjj

Sphaeropthalma hyalina (Blake)
Tex.

Agama hyalina Blake, 1871. Amer. Ent. Soc, Trans. 3: 263. ♂.

Agama minuta Blake, 1872. Amer. Ent. Soc, Trans. 4: 76. ♂.

Sphaeropthalma macswaini Ferguson
Nev., Calif.

Sphaeropthahna (Micromutilla) macswaini Ferguson, 1967. Brigham Young Univ. Sci.
Bui, Biol. Ser. 8, no. 4: 12, fig. 6. ♂.

Sphaeropthalma mesillensis (Cockerell)
N. Mex. (Mesilla).

Photopsis mesilleiisis Cockerell, 1897. Entomologist 30: 137. ♂.

Sphaeropthalma pallida (Blake)
Tex., N. Mex., Ariz., Nev.

Agama pallida Blake, 1871. Amer. Ent. Soc, Trans. 3: 263. ♂.

Sphaeropthahna {Micromutilla) arizonae Schuster, 1958. Ent. Amer. (n. s.) 37: 16. ♂.

Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci. Bui., Biol. Ser. 8, no. 4: 13 (synonymy).

Sphaeropthalma parapenalis Ferguson
Wash., Oreg., Idaho, Calif., Nev., Utah, Ariz., Tex.; Mexico (Chihuahua,
Coahuila, Durango, Zacatecas).
Sphaeropthahna (Micromutilla) parapenalis Ferguson, 1967. Brigham Young Univ. Sci.
Bui, Biol. Ser. 8, no. 4: 14, fig. 7. ♂.

Sphaeropthalma pateli Schuster
Ariz. (Hereford).

Sphaeropthahna (Micromutilla) pateli Schuster, 1958. Ent. Amer. (n. s.) 37: 15. ♂.

Sphaeropthalma reducta Schuster
Ariz.

Sphaeropthahna (Micromutilla) reducta Schuster, 1958. Ent. Amer. (n. s.) 37: 18. ♂.

Sphaeropthalma sabino Schuster
Ariz.

Sphaeropthahna (Micromutilla) sabino Schuster, 1958. Ent. Amer. (n. s.) 37: 19. ♂.

Sphaeropthalma sonora Schuster
Ariz., Nev., Calif.

Sphaeropthahna (Micromutilla) sonora Schuster, 1958. Ent. Amer. (n. s.) 37: 16. ♂.

Sphaeropthalma sublobata Schuster
Idaho.

Sphaeropthahna (Micromutilla) sublobata Schuster, 1958. Ent. Amer. (n. s.) 37: 16. ♂.

Sphaeropthalma uvaldella Schuster
Tex.

Sphaeropthahna (Micromutilla) uvaldella Schuster, 1958. Ent. Amer. (n. s.) 37: 19. ♂.

Sphaeropthalma yumaella Schuster
Nev., Ariz., Calif.; Mexico (Baja California).

Sphaeropthalma (Micromutilla) yumaella Schuster, 1958. Ent. Amer. (n. s.) 37: 19. ♂.

Genus SPHAEROPTHALMA Subgenus PHYSETAPSIS Schuster

Sphaeropthalma subg. Physetapsis Schuster, 1958. Ent. Amer. (n. s.) 37: 9 (in key), 20.
Type-species: Sphaeropthalma (Physetapsis) papaga Schuster. Orig. desig.

Sphaeropthalma borealis Schuster
Sask.

Sphaeropthahna (Physetapsis) borealis Schuster, 1958. Ent. Amer. (n. s.) 37: 20. ♂.

Sphaeropthalma ecarinata Schuster
Calif.

Sphaeropthahna (Physetapsis) ecarinata Schuster, 1958. Ent. Amer. (n. s.) 37: 20. ♂.




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Sphaeropthalma papaga
***authority mismatch
ephysetos Schuster. Southwestern Ariz., southern Calif.

Sphaeropthalma (Physetapsis) papaga ephysetos Schuster, 1958. Ent. Amer. (n. s.) 37: 20.
6.

Sphaeropthalma papaga
***authority mismatch
papaga Schuster. Southeastern Ariz., Calif. (Imperial Co.).

Sphaeropthalma (Physetapsis) papaga papaga Schuster, 1958. Ent. Amer. (n. s.) 37: 20, pi.
6, fig. 3. ♂.

Sphaeropthalma subcarinata Schuster
N. Mex.

Sphaeropthalma (Physetapsis) subcarinata Schuster, 1958. Ent. Amer. (n. s.) 37: 20. ♂.

Genus SPHAEROPTHALMA Subgenus PHOTOPSIS Blake

Agarna Blake, 1871. Amer. Ent. Soc, Trans. 8: 258. Preocc.

Type-species: Agama iviperialis Blake. Automatic desig. by Ashmead, 1899.
Photopsis Blake, 1886. Amer. Ent. Soc, Trans. 13: 179. N. name.

Type-species: Agavia iinperialis Blake. Desig. by Ashmead, 1899.
Pyrrhomutilla Ashmead, 1899. N. Y. Ent. Soc, Jour. 7: 56.

Type-species: Sphaerophthalma (!) anthophorae Ashmead. Orig. desig.
Neophotopsis Ashmead, 1903. Canad. Ent. 35: 306.

Type-species: Mutilla pluto Fox. Orig. desig.

In addition to the host records cited under several species in this subgenus, Parker and
Bohart, 1968 (Pan-Pacific Ent. 44: 2, 3 and 5) record several unidentified species of
Sphaeropthalma (Photopsis) from the following hosts from nests in twigs: Ancistrocerus cat-
skill (Sauss.); Odynerus erythrogaster Bohart; Pisonopsis birkmanni Rohwer; Anthocopa
hypostomalis Michener; Hoplitis fulgida (Cresson); H. sayubnci Titus; and Proteriades bullifa-
cies (Michener). Ferguson, 1962 (Univ. Calif. Pubs. Ent. 27: tab. 9) states that Davidson reared
an unidentified species of Sphaeropthalma (Photopsis) from a nest of Hoplitis producta
(Cresson) in a hollow plant stem.

Biology: Ferguson, 1962. Univ. Calif. Pubs. Ent. 27: 1-92, 7 pis., 2 text figs, (life history,

hosts).

Sphaeropthalma albopilosa (Blake)
Tex. (Comal Co.).

Mutilla (Sphaeropthalma) albopilosa Blake, 1872. Amer. Ent. Soc, Trans. 4: 74. ♀.

Sphaeropthalma anaspasia (Cockerell and Rohwer)
Colo. (Boulder).

Photopsis anaspasia Cockerell and Rohwer, 1908. Psyche 15: 5. ♀.

Sphaeropthalma angulifera Schuster
Calif., Nev.

Sphaeropthalma (Photopsis) angulifera Schuster, 1958. Ent. Amer. (n. s.) 37: 32. ♂.

Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci. Bui., Biol. Ser. 8, no. 4: 17
(distribution).

Sphaeropthalma arenicola Schuster
Ariz.

Sphaeropthalma (Photopsis) arenicola Schuster, 1958, Ent. Amer. (n. s.) 37: 25. ♂.

Sphaeropthalma arota (Cresson)
Calif. (San Diego).

Mutilla Arota Cresson, 1875. Amer. Ent. Soc, Trans. 5: 120. ♀.

Sphaeropthalma baboquivari Schuster
Ariz.

Sphaeropthalma (Photopsis) baboqtiivari Schuster, 1958. Ent. Amer. (n. s.) 37: 24, pi. 4,
figs. 1-6. ♂.

Sphaeropthalma bisetosa Schuster
Tex., N. Mex., Ariz.

Sphaeropthalma (Photopsis) bisetosa Schuster, 1958. Ent. Amer. (n. s.) 37: 36. ♂.

Sphaeropthalma blakeii (Fox)
Nev., Ariz., N. Mex., Calif.; Mexico (Baja California). Host: Diadasia vallicola
Timb.
Photopsis Blakeii Fox, 1893. Calif. Acad. Sci., Proc (2) 4: 6. ♂.
Mutilla Gautschii Dalla Torre, 1897. Cat. Hym. 8: 43. N. name erroneously proposed for

Sphaeropthalma blakeii
***authority mismatch
Fox, thought to be preocc. by Cameron, 1894.
Mutilla ceyx Fox, 1899. Amer. Ent. Soc, Trans. 25: 262. ♂.

Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci. Bui., Biol. Ser. 8, no. 4: 17 (synonymy).

Biology: Ferguson, 1962. Univ. Calif Pubs. Ent. 27: 10-15, pis. 2, 3, 4, 5, 7, tab. 9 (life
history).




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Sphaeropthalma capricornis (Rohwer)
Colo. (Boulder).

Photopsis capricornis Rohwer, 1909. Amer. Ent. Soc, Trans. 35: 131. ♀.

Sphaeropthalma ceres (Fox)
Ariz., Colo., Utah.

Mutilla ceres Fox, 1899. Amer. Ent. Soc, Trans. 25: 257. ♀.

Sphaeropthalma ceyxoides Schuster
Ariz.

Sphaeropthalma (Photopsis) ceyxoides Schuster, 1958. Ent. Amer. (n. s.) 37: 25. ♂.

Sphaeropthalma clara
***authority mismatch
clara (Cresson). N. Mex., Colo, to Wyo. and Idaho. This subsp. is transitional to
Sphaeropthalma subg. Physetapsis Schuster.
Mutilla clara Cresson, 1865. Ent. Soc. Phila., Proc. 4: 439. ♂.

Taxonomy: Schuster, 1958. Ent. Amer. (n. s.) 37: 27.

Sphaeropthalma clara
***authority mismatch
cleomella Schuster. Mont. This subsp. is transitional to Sphaeropthalma subg.
Physetapsis Schuster.
Sphaeropthalma (Photopsis) clara cleomella Schuster, 1958. Ent. Amer. (n. s.) 37: 27. ♂.

Sphaeropthalma coaequalis Cameron
Mont, and Oreg. to N. Mex. and north. Ariz., Kans., Tex.; north. Mexico.
Sphaerophthalma (!) coaequalis Cameron, 1896. Biol. Cent.-Amer., Hym., v. 2, p. 379. ♂.
Mutilla albicincta Fox, 1899. Amer. Ent. Soc, Trans. 25: 255. ♂.
Sphaeropthalma (Photopsis) albicincta piceoceps Schuster, 1958. Ent. Amer. (n. s.) 37: 35.

6.
Sphaeropthalma (Photopsis) albicincta flavipes Schuster, 1958. Ent. Amer. (n. s.) 37: 36.

S.

Taxonomy: Mickel, 1965. Ent. Soc. Wash., Proc. 67: 1 (synonymy).

Sphaeropthalma danaus (Blake)
Kans., Tex., N. Mex.; Mexico.

Agama Danaiis Blake, 1871. Amer. Ent. Soc, Trans. 3: 261. ♂.

Sphaeropthalma dentifera Schuster
S. Dak., Minn.

Sphaeropthalma (Photopsis) dentifera Schuster, 1958. Ent. Amer. (n. s.) 37: 24. ♂.

Sphaeropthalma diomeda (Fox)
Tex.

Mutilla diomeda Fox, 1899. Amer. Ent. Soc, Trans. 25: 257. ♀.

Sphaeropthalma dirce (Fox)
Ariz., Utah, Tex.

Mutilla dirce Fox, 1899. Amer. Ent. Soc, Trans. 25: 258. ♀.

Sphaeropthalma edwardsii
***authority mismatch
edwardsii (Cresson). Oreg., Idaho.

Mutilla Edwardsii Cresson, 1875. Amer. Ent. Soc, Trans. 5: 119. ♂.

Sphaeropthalma edwardsii
***authority mismatch
flammifcra Schuster. Calif.

Sphaeropthalma (Photopsis) edwardsii flammifera Schuster, 1958. Ent. Amer. (n. s.) 37:
36. ♂.

Sphaeropthalma erato (Blake)
Tex.

Mutilla Erato Blake, 1879. Amer. Ent. Soc, Trans. 7: 251. ♀.

Sphaeropthalma erigone (Fox)
Colo.

Mutilla erigone Fox, 1899. Amer. Ent. Soc, Trans. 25: 268. ♀.

Sphaeropthalma facilis (Cameron)
N. Mex., Ariz.; Mexico (Durango).

Sphaerophthahna (\) facilis Cameron, 1896. Biol. Cent.-Amer., Hym., v. 2, p. 392. ♂.

Mutilla Hubbardii Fox, 1899. Amer. Ent. Soc, Trans. 25: 256. ♂.

Sphaeropthalma (Photopsis) piceogaster Schuster, 1958. Ent. Amer. (n. s.) 37: 27. ♂.

Sphaeropthalma ferruginea (Blake)
Nev., Calif. Host: Isodontia etegans (Smith).

Agama ferruginea Blake, 1879. Amer. Ent. Soc, Trans. 7: 254. ♂.

Mutilla ferruginosa Dalla Torre, 1897. Cat. Hym., v. 8, p. 40. N. name for Agama

Sphaeropthalma ferruginea
***authority mismatch
Blake, not M utilla feirruginea Smith.

Biology: Davidson, 1899. Ent. News 10: 180 (host).

Sphaeropthalma ferruginopsis Schuster
Ariz., Calif.

Sphaeropthalma (Photopsis) ferruginopsis Schuster, 1958 Ent. Amer. (n. s.) 37: 29. ♂.

Sphaeropthalma fuscipes Schuster
Ariz.; Mexico (Durango). This sp. is transitional to Sphaeropthalma subg.
Physetapsis Schuster.
Sphaeropthalma (Photopsis) fuscipes Schuster, 1958. Ent. Amer. (n. s.) 37: 21. ♂.

Sphaeropthalma halcyone (Fox)
Kans. (Hamilton Co.).

Mutilla halcyone Fox, 1899. Amer. Ent. Soc, Trans. 25: 257. ♀.




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Sphaeropthalma helicaon (Fox)
Nev., Ariz., Calif.; Mexico (Baja California, Coahuila).
Mutilla helicaon Fox. 1899. Amer. Ent. Soc, Trans. 25: 254. ♂.
Photopsis lingulatus Viereck, 1903. Acad. Nat. Sci. Phila., Proc. 54: 737. ♂.
Sphaeropthahna {Photopsis) carinata Schuster, 1958. Ent. Amer. (n. s.) 37: 34. ♂.
Sphaeropthalma (Photopsis) helicaon coahuilae Schuster, 1958. Ent. Amer. (n. s.) 37: 34.

6.
Sphaeropthalma (Photopsis) helicaon diegueno Schuster, 1958. Ent. Amer. (n. s.) 37: 35. ♂.

Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci. BuL, Biol. Ser. 8, no. 4: 17 (synonymy).

Sphaeropthalma hypermnestra (Fox)
Calif. (Poway).

Mutilla hypermnestra Fox, 1899. Amer. Ent. Soc, Trans. 25: 268. ♀.
ignacio Schuster. Southern Calif.; Mexico (Baja California).

Sphaeropthalma (Photopsioides) amphion ignacio Schuster, 1958. Ent. Amer. (n. s.) 37: 38.
6.

Sphaeropthalma imperialiformis
***authority mismatch
imperialiformis (Viereck). Tex. to Kans. and Colo.

Mutilla (Photopsis) imperialiformis Viereck, 1906. Amer. Ent. Soc, Trans. 32: 189. ♂.

Sphaeropthalma imperialiformis
***authority mismatch
maricopae Schuster. Ariz, to Wash.

Sphaeropthalma (Photopsis) imperialiformis maricopae Schuster, 1958. Ent. Amer. (n. s.)
37:34. ♂.

Sphaeropthalma imperialis (Blake)
Tex.

Agama imperialis Blake, 1871. Amer. Ent. Soc, Trans. 3: 260. ♂.

Sphaeropthalma insignis (Baker)
Calif. (Claremont); Mexico (Baja California).

Photopsis insig7iis Baker, 1905. Invertebrata Pacifica 1: 115. ♂.

Sphaeropthalma jacala Schuster
Ariz.; Mexico (Jacala).

Sphaeropthalma (Photopsis) jacala Schuster, 1958. Ent. Amer. (n. s.) 37: 21. ♂.

Sphaeropthalma juxta (Blake)
Iowa, Kans., Tex., N. Mex., Ariz.

Agama juxta Blake, 1872. Amer. Ent. Soc, Trans. 4: 76. ♂.

Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci. BuL, ser. 8, no. 4: 15.

Sphaeropthalma laodamia (Fox)
Ariz.

Mutilla laodamia Fox, 1899. Amer. Ent. Soc, Trans. 25: 258. ♀.

Sphaeropthalma luiseno Schuster
Calif.

Sphaeropthalma (Photopsis) luiseno Schuster, 1958. Ent. Amer. (n. s.) 37: 23. ♂.

Sphaeropthalma marpesia (Blake)
Kans., Utah.

Mutilla Marpesia Blake, 1879. Amer. Ent. Soc, Trans. 7: 247. ♀.

Sphaerophthalma (!) luteola Blake, 1886. Amer. Ent. Soc, Trans. 13: 235. ♀.

Sphaeropthalma megagnathos
***authority mismatch
aurifera Schuster. Calif., Ariz.

Sphaeropthalma (Photopsis) megagnathos auriferus Schuster, 1958. Ent. Amer. (n. s.) 37:
36. ♂.

Sphaeropthalma megagnathos
***authority mismatch
megagnathos Schuster. Calif., Ariz.

Sphaeropthalma (Photopsis) megagnathos megagnathos Schuster, 1958. Ent. Amer. (n. s.)
37:36. ♂.

Sphaeropthalma militaris Schuster
Ariz., Calif.

Sphaeropthalma (Photopsis) militaris Schuster, 1958. Ent. Amer. (n. s.) 37: 27, pi. 5, fig. 2.
S.

Sphaeropthalma minutella (Mickel)
Calif. (Lassen Co.).

Photopsis minutella Mickel, 1938. Pan-Pacific Ent. 14: 183. ♀.

Sphaeropthalma nanula (Dalla Torre)
Tex., Colo., Nev., Idaho. Host: Tachysphex sp.

Mutilla pygmea Blake, 1879. Amer. Ent. Soc, Trans. 7: 250. ♀. Preocc.

Mutilla nanula Dalla Torre, 1897. Cat. Hym., v. 8, p. 65. N. name.

Mutilla pygmaea (!) Dalla Torre, 1897. Cat. Hym., v. 8, p. 65.

Biology: Krombein, 1958. U. S. Dept. Agr., Monog. 2, Sup. 1: 102 (host).

Sphaeropthalma neomexicana Schuster
N. Mex.

Sphaeropthalma (Photopsis) neomexicana Schuster, 1958. Ent. Amer. (n. s.) 37: 30. ♂.

Sphaeropthalma noctivaga (Melander)
Nebr. and Wyo. south to Tex. and Ariz.

Mutilla noctivaga Melander, 1903. Amer. Ent. Soc, Trans. 29: 318. tj.




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Sphaeropthalma {Photopsis) noctivaga infuscata Schuster, 1958. Ent. Amer. (n. s.) 37: 32.
6.

Sphaeropthalma nokomis
***authority mismatch
nokomis (Blake). Ariz.

Agama Nokomis Blake, 1871. Amer. Ent. Soc, Trans. 3: 260. ♂.

Sphaeropthalma nokomis
***authority mismatch
ruficeps Schuster. Ariz.

Sphaeropthalma {Photopsis) nokomis ruficeps Schuster, 1958. Ent. Amer. (n. s.) 37: 35. ♂.

Sphaeropthalma ordae Schuster
Tex.

Sphaeropthalma (Photopsis) ordae Schuster, 1958. Ent. Amer. (n. s.) 37: 23. ♂.

Sphaeropthalma orestes (Fox)
Wash, to Calif., Nev.; Mexico (Baja California). Host: Anthidium c. collectum
Huard; Euodynerus a auranus (Cam.); Tachysphex tenuipunctus Fox.

Mutilla orestes Fox, 1899. Amer. Ent. Soc, Trans. 25: 256. ♂.

Mutilla Pattersonae Melander, 1903. Amer. Ent. Soc, Trans. 29: 309. ♂.

Photopsis indigens Baker, 1905. Invertebrata Pacifica 1: 112. ♂.

Photopsis uniformis Baker, 1905. Invertebrata Pacifica 1: 113. ♂.

Photopsis pedatus Baker, 1905. Invertebrata Pacifica 1: 115. ♂.

Photopsis inge^iuiis Baker, 1905. Invertebrata Pacifica 1: 116. ♂.

Photopsis salmani Mickel, 1938. Pan-Pacific Ent. 14: 178. ♀, ♂.

Sphaeropthahna {Photopsis) salmani fresnoensis Schuster, 1958. Ent. Amer. (n. s.) 37: 30.

6.
Sphaeropthalma (Photopsis) salmani oregana Schuster, 1958. Ent. Amer. (n. s.) 37: 31. ♂.

Taxonomy: Schuster, 1958. Ent. Amer. (n. s.) 37: 30 (synonymy). —Ferguson, 1962. Univ.

Calif. Pubs. Ent. 27: 10 (synonymy).
Biology: Ferguson, 1962. Univ. Calif. Pubs. Ent. 27: 6-10, pis. 1, 6, tabs. 1, 9 (life history).

Sphaeropthalma pallidipes Schuster
Ariz.

Sphaeropthalma (Photopsis) pallidipes pallidipes Schuster, 1958. Ent. Amer. (n. s.) 37: 27.

6.
Sphaeropthalma (Photopsis) pallidipes gila Schuster, 1958. Ent. Amer. (n. s.) 37: 28. ♂.

Sphaeropthalma parkeri Schuster
Ariz., Calif.

Sphaeropthalma (Photopsis) parkeri Schuster, 1958. Ent. Amer. (n. s.) 37: 28. ♂.

Sphaeropthalma pervaga (Melander)
Tex. (Fedor).

Mutilla pervaga Melander, 1903. Amer. Ent. Soc, Trans. 29: 321. ♀.

Sphaeropthalma pinalea Schuster
N. Dak. to Tex., Ariz.; Mexico (Coahuila, Durango).

Sphaeropthalma (Photopsis) pinalea pinalea Schuster, 1958. Ent. Amer. (n. s.) 37: 32. ♂.

Sphaeropthalma (Photopsis) pinalea texanella Schuster, 1957. Ent. Amer. (n. s.) 37: 32. ♂.

Sphaeropthalma pinales Schuster
Ariz.

Sphaeropthalma (Photopsis) pinales Schuster. 1958. Ent. Amer. (n. s.) 37: 36. ♂.

Sphaeropthalma pluto (Fox)
Tex., N. Mex., Ariz.

Mutilla pluto Fox, 1899. Amer. Ent. Soc, Trans. 25: 255. ♂.
Mutilla palamedes Fox, 1899. Amer. Ent. Soc, Trans. 25: 264. ♂.

Sphaeropthalma rubriventris Schuster
Tex., N. Mex., Ariz.; Mexico (Tamaulipas).

Sphaeropthalma (Photopsis) faciiis rubriventris Schuster, 1958. Ent. Amer. (n. s.) 37: 26.
S.

Sphaeropthalma sanctaefeae (Cockerell)
N. Mex. (Santa Fe).

Mutilla sanctae-feae Cockerell, 1897. In Cockerell and Fox, Acad. Nat. Sci. Phila., Proc 49:
137. ♀.

Sphaeropthalma scudderi Schuster
Ariz.

Sphaeropthahna (Photopsis) scudderi Schuster, 1958. Ent. Amer. (n. s.) 37: 23. ♂.

Sphaeropthalma seminanula Rohwer
Colo. (Boulder).

Photopsis seminanula Rohwer, 1909. Amer. Ent. Soc, Trans. 35: 130. ♀.

Sphaeropthalma similis Schuster
Calif.

Sphaeropthalma (Photopsis) siinilis Schuster, 1958. Ent. Amer. (n. s.) 37: 30. ♂.

Sphaeropthalma spinifera Schuster
Kans., Okla., Colo.

Sphaeropthalma (Photopsis) spinifera Schuster, 1958. Ent. Amer. (n. s.) 37: 23. ♂.

Sphaeropthalma tetracuspis Schuster
Calif. (Riverside Co.); Mexico (Baja California).

Sphaeropthalma (Photopsis) tetracuspis Schuster, 1958. Ent. Amer. (n. s.) 37: 31. ♂.




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Sphaeropthalma triangularis (Blake)
Tex., N. Mex., Ariz., Nev., Calif.; Mexico (Baja California).
Agama triangularis Blake, 1871. Amer. Ent. Soc, Trans. 3: 262. ♂.

Sphaeropthalma tuberculifera Schuster
Calif. (?).

Sphaeropthalvia (Plwtopsis) tuberculifera Schuster, 1958. Ent. Amer. (n. s.) 37: 25. ♂.

Sphaeropthalma unicolor (Cresson)
U. S. west of Rocky Mts.; Mexico (Baja California del Norte). Host:

Isodontia elegans (Sm.); Anthidiuyn c. collectum Huard, Callanthidiuni illustre (Cr.),
Ashmeadiella c. califoniica (Ashm.); Anthophora abruptella Ckll., A. bomboides
neomexicayia Ckll., A. linsleyi Timb., A. montana Cr.?, A. occidentalis Cr., Diadasia
bituberculata (Cr.), Melissodes robustior Ckll., Xeromelecta califomica (Ashm.).

Mutilla unicolor Cresson, 1865. Ent. Soc. Phila., Proc. 4: 389. ♂.

Agama mendica Blake, 1871. Amer. Ent. Soc, Trans. 3: 259. ♂.

Mutilla auraria Blake, 1879. Amer. Ent. Soc, Trans. 7: 248. ♀.

Mutilla Aspasia Blake, 1879. Amer. Ent. Soc, Trans. 7: 250. ♀.

Mutilla Phaedra Blake, 1879. Amer. Ent. Soc, Trans. 7: 251. ♀.

Agama rustica Blake, 1879. Amer. Ent. Soc, Trans. 7: 252. ♂.

Photopsis nebulosus Blake, 1886. Amer. Ent. Soc, Trans. 13: 275. ♂.

Sphaeroplithalmia (!) anthophora Ashmead, 1897. In Davidson, South. Calif. Acad. Sci.,
Proc 1: 5. ♀, ♂.

Mutilla monochroa Dalla Torre, 1897. Cat. Hym., v. 8, p. 63. N. name for Mutilla unicolor
Cresson, not Myrmosa unicolor Say.

Dasymutilla sumneriella Cockerell, 1915. Entomologist 48: 249. ♀.

Sphaeropthalma (Photopsis) rustica ocellaria Schuster, 1958. Ent. Amer. (n. s.) 37: 32. ♂.

Biology: Davidson, 1894. Ent. News 5: 170 (host records). —Davidson, 1897. South. Calif.

Acad. Sci., Proc. 1: 4-5 (host records). —Davidson, 1899. Ent. News 10: 180 (host record).

— Linsley and MacSwain, 1942. Amer. Midland Nat. 27: 409 (host records). —Ferguson,

1962. Univ. Calif. Pubs. Ent. 27: 15-22, pis. 3, 5, 7, tab. 9 (life history, host records).

Sphaeropthalma virguncula (Blake)
N. Mex.

Sphaerophthalma (!) virguncula Blake, 1886. Amer. Ent. Soc, Trans. 13: 253. ♀.

Sphaeropthalma zenobia (Blake)
Calif., ? Tex.

Mutilla zenobia Blake, 1879. Amer. Ent. Soc, Trans. 7: 250. ♀.

Sphaeropthalma zephyritis (Fox)
Calif. (Los Angeles).

Mutilla zephyritis Fox, 1899. Amer. Ent. Soc, Trans. 25: 269. ♀.

NoMEN Nudum in Sphaeropthalma Subgenus Photopsis Blake

Sphaeropthalma (Photopsis) nixonensis Parker and Bohart, 1966. Pan-Pacific Ent. 42: 98. Nev.
Host: Proteriades sp.

Genus SPHAEROPTHALMA Subgenus PHOTOPSIOIDES Schuster

Sphaeropthalma subg. Photopsioides Schuster, 1958. Ent. Amer. (n. s.) 37: 10 (in key), 36.
Type-species: Agama uro Blake. Orig. desig.

In addition to the host records cited below under uro (Blake), Ferguson, 1962 (Univ. Calif.
Pubs. Ent. 27: tab. 10) records Sceliphron caementarium (Dru.) as the host of an unidentified
species of this subgenus.

Sphaeropthalma abdominalis (Blake)
Kans., Tex., Colo., Ariz., Utah. Host: Trypargilum t. tridentatum (Pack.).
Photopsis abdominalis Blake, 1886. Amer. Ent. Soc, Trans. 13: 275. ♂.
Mutilla coloradensis Dalla Torre, 1897. Cat. Hym., v. 8, p. 25. N. name for Photopsis

Sphaeropthalma abdominalis
***authority mismatch
Blake, not Mutilla abdomiyialis Westwood.

Sphaeropthalma amphion (Fox)
Ariz., Nov., Calif., Oreg.; Mexico (Baja California). Host: Sapyga aculeata Cr.;

Leptochilus chiricahua Prkr., Ancistrocerus c. catskill (Sauss.); Trypargilum t.

tridentatum (Pack.); Hoplitis bullifacies Mich., H . f. fulgida (Cr.), H. g. grijmelli Ckll.,

H. hypostomalis Mich., H. sambuci Titus, Anthocopa copelandica (Ckll.), Ashmeadiella

m. meliloti (Ckll.), Stelis spp.
Mutilla amphion Fox, 1899. Amer. Ent. Soc, Trans. 25: 263. ♂.
Photopsis absfrusa Baker, 1905. Invertebrata Pacifica 1: 113. ♂.
Photopsis nudata Baker, 1905. Invertebrata Pacifica 1: 114. ♂.




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Taxonomy: Ferguson, 1967. Brigham Young Univ., Sci. Bui., ser. 8, no. 4: 20 (synonymy).

Biology: Parker, 1975. Pan-Pacific Ent. 51: 116 (host).

Sphaeropthalma contracta (Blake)
Nev., Oreg.

Agama contmcta Blake, 1879. Amer. Ent. Sec, Trans. 7: 253. ♂.
Mutilla contrahenda Dalla Torre, 1897. Cat. Hym., v. 7, p. 27. N. name for Agama
covtracta Blake, not Mutilla coiitracta Say.

Sphaeropthalma uro (Blake)
Kans., Tex., N. Mex., Ariz. Host: Pachodynerns astraeus (Cam.), eumenid sp.;
Trypargilum t. tridentatum (Pack.); Dianthidixm cnrvatum sayi Ckll.
Agama uro Blake, 1879. Amer. Ent. Soc, Trans. 7: 253. ♂.
PJiotopsis melanderi Baker, 1905. Invertebrata Pacifica 1: 112. ♂.

Sphaeropthalma (Photopsioides) uro stenognatha Schuster, 1958. Ent. Amer. (n. s.) 37: 38.
6.

Biology: Fischer, 1951. Kans. Ent. Soc, Jour. 24: 49 (host record). —Ferguson, 1962. Univ.
Calif. Pubs. Ent. 27: tab. 10 (host records). —Parker and Bohart, 1966. Pan-Pacific Ent. 42:
93 (host record). — Krombein, 1967. Trap-nesting wasps and bees, pp. 478-479 (life history).

Genus SPHAEROPTHALMA Subgenus SPHAEROPTHALMA Blake

Sphaeropthalma Blake, 1871. Amer. Ent. Soc, Trans. 3: 232.

Type-species: Mutilla (Sphaeropthalma) scaeva Blake. Automatic desig. by
Ashmead, 1899 (=Sphaeropthalma pensylvanica scaeva (Blake)).
Sphaerophthalma (!) Blake, 1886. Amer. Ent. Soc, Trans. 13: 179. Emend.

Type-species: Mutilla (Sphaeropthahna) scaeva Blake. Desig. by Ashmead, 1899
{=Sphaeropthalma peiisylvanica scaeva (Blake)).

Revision: Schuster, 1944. Brooklyn Ent. Soc, Bui. 39: 141-147.

Sphaeropthalma auripilis
***authority mismatch
auripilis (Blake). Kans., Okla., Tex. Host: Chalyhion califomicum (Sauss.),

Sceliphron caementarium (Drury), Trypoxylon sp.; solitary eumenid in old clay cell of
Sceliph ron.
Mutilla (Sphaeropthalma) auripilis Blake, 1871. Amer. Ent. Soc, Trans. 3: 233. ♂.
Sphaerophthalma (!) albiplumosa Schuster, 1944. Brooklyn Ent. Soc, Bui. 39: 145. ♀.

Biology: Rau, 1940. Ent. Soc Amer., Ann. 33: 594 (host).

Sphaeropthalma auripilis
***authority mismatch
fasciventris Schuster. Tex. (Edinburg).

Sphaeropthalma (Sphaeropthalma) auripilis fasciventris Schuster, 1958. Ent. Amer. (n. s.)
37:39. ♂.

Taxonomy: Mickel, 1974. Ent. Soc. Amer., Ann. 67: 466. ♀.

Sphaeropthalma boweri Schuster
Tex. (College Station).

Sphaerophthalma (!) boweri Schuster, 1944. Brooklyn Ent. Soc, Bui. 39: 143. ♂.

Sphaeropthalma pensylvanica
***authority mismatch
floridensis Schuster. South. Fla.

Sphaerophthalma (!) pennsylvanica(\) floridensis Schuster, 1944. Brooklyn Ent. Soc, Bui.
39: 142. ♀.

Taxonomy: Mickel, 1974. Ent. Soc. Amer., Ann. 67: 466. ♂.

Sphaeropthalma pensylvanica
***authority mismatch
pensylvanica (Lepeletier). N. C, Ga., cent. Fla., La., Mo., Kans., Tex. Host:
Sceliphron caementarium (Drury), Trypargilmn politum (Say), T. clavatum (Say);
anthidiine sp.
Mutilla pensylvanica Lepeletier, 1845. Hist. Nat. Ins. Hym., v. 3, p. 628. ♂.
Mutilla (Sphaeropthalma) pennsylvanica (!) Blake, 1871. Amer. Ent. Soc, Trans. 3: 233.

S.
Mutilla (Sphaeropthalma) balteola Blake, 1871. Amer. Ent. Soc, Trans. 3: 248. ♀.

Taxonomy: Bradley, 1916. Amer. Ent. Soc, Trans. 42: 331.

Biology: Rau, 1922. Acad. Sci. St. Louis, Trans. 24: 7 (host). —Rau, 1928. Acad. Sci. St. Louis,
Trans. 25: 437 (host).

Sphaeropthalma pensylvanica
***authority mismatch
scaeva (Blake). Mass. to N. C, Ohio, 111., Mo., Kans., Tex. Host: Auplopus a.
architectus (Say), A. m. mellipes (Say); Trypargilum clavatum (Say), T. collinum




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rubwcinctu))) (Pack.), T. politioii (Say), T. lactitarse (Sauss.), Sceliphron caementarium
(Dru.). Parasite: Melittobia chalijbii Ashm.
Mitfilla (Sphaewpthahua) scaeva Blake, 1871. Amer. Ent. Soc, Trans. 3: 232. ♂.

Taxonomy: Bradley, 1916. Amer. Ent. Soc, Trans. 42: 831.

Biology: Rau and Rau, 1918. Wasp studies afield, pp. 88-89 (host). — Rau, 1922. Acad. Sci. St.
Louis, Trans. 24, no. 7: 8 (host). — Krombein, 1951. U. S. Dept. Agr., Monog. 2: 752 (hosts).
— Krombein, 1967. Trap-nesting wasps and bees, pp. 476-478, figs. 135-139 (life history).

Genus PHOTOMORPHUS Viercck

Taxonomy: Schuster, 1944. Brooklyn Ent. Soc, Bui. 39: 147-151 (east spp. only). —Krombein,
1954. Amer. Ent. Soc, Trans. 80: 6-7 (key to eastern females).

Genus PHOTOMORPHUS Subgenus PHOTOMORPHINA Schuster

Photomorphus subg. Photomorphina Schuster, 1952. Brooklyn Ent. Soc, Bui. 47: 53.
Type-species: Photomorphus {Photomorphina) aurifera Schuster. Orig. desig.

Photomorphua subg. Photomorphina Schuster, 1958. Ent. Amer. (n.s.) 37: 8 (in key), 39.
Preocc. by Photomorphus subg. Photomorphina Schuster, 1952, and a syn. of it.
Type-species: Photomorphus {Photontorphina) quadriangulata Schuster. Orig.
desig.

Photomorphus aurifer Schuster
Ariz.

Photomorphus (Photomorphina) aurifera Schuster, 1952. Brooklyn Ent. Soc, Bui. 47: 53.
6.

Photomorphus (Photomorphina) aurifera Schuster, 1958. Ent. Amer. (n. s.) 37: 44. S-
Preocc. by and a syn. oi Photomorphus (Photomorphina) aurifera Schuster, 1952.

Photomorphus auriventris Schuster
Tex.

Photomorphus (Photomorphina) auriventris Schuster, 1958. Ent. Amer. (n. s.) 37: 40. ♂.

Photomorphus bequaertii (Schuster)
n. comb. (W. E. Ferguson). Tex.

Odontophotopsis (Odontophotopsis) bequaertii Schuster, 1958. Ent. Amer. (n. s.) 37: 52. ♂.

Photomorphus californicus Schuster
Calif.

Photomorphus (Photomorphina) califomica Schuster, 1958. Ent. Amer. (n. s.) 37: 44. ♂.

Photomorphus clandestinus (Viereck)
Tex., N. Mex.

Odontophotopsis clandestinus Viereck, 1903. Acad. Nat. Sci. Phila., Proc. 54: 740. ♂.

Photomorphus cobabi
***authority mismatch
Schuster. Ariz.; Mexico (Sonera, Baja California).

Photomorphus (Photomorphina) cobabi Schuster, 1958. Ent. Amer. (n. s.) 37: 41. ♂.

Photomorphus cochisae Schuster
Ariz.; Mexico.

Photomorphus (Photomorphina) cochisae Schuster, 1958. Ent. Amer. (n. s.) 37: 43. ♂.

Photomorphus coloradellus Schuster
Colo.

Photomorphus (Photomorphina) coloradella Schuster, 1958. Ent. Amer. (n. s.) 37: 44. ♂.

Photomorphus comobabi Schuster
Ariz.; Mexico.

Photomorphus (Photomorphina) comobabi Schuster, 1958. Ent. Amer. (n. s.) 37: 45. ♂.

Photomorphus crepusculus (Viereck)
Kans. (Morton Co.).

Mutilla (PhotomorphusO crepuscula Viereck, 1906. Amer. Ent. Soc, Trans. 32: 188. ♂.

Photomorphus dichrous Schuster
Ariz.

Photomorphus (Photomorphina) dichrous Schuster, 1958. Ent. Amer. (n. s.) 37: 41. ♂.

Photomorphus hebes (Melander)
Tex., N. Mex., Ariz.; Mexico (Sonora, Coahuila, Tamaulipas).

Mutilla hebes Melander, 1903. Amer. Ent. Soc, Trans. 29: 311. ♂.

Photomorphus (Photomorphina) taeniatus Schuster, 1958. Ent. Amer. (n. s.) 37: 42. ♂.

Photomorphus (Photomorphina) taeniatus parksi Schuster, 1958. Ent. Amer. (n. s.) 37: 42.

i.

Photomorphus imperialoides Schuster
Ariz.

Photomorphus (Photomorphina) imperialoides Schuster, 1958. Ent. Amer. (n. s.) 37: 40. ♂.

Photomorphus jason (Fox)
Tex., Okla.

Mutilla jason Fox, 1899. Amer. Ent. Soc, Trans. 25: 250. ♂.




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Photomorphus juanita Schuster
Tex.; Mexico (Tamaulipas).

Photoinorphufi (Photomorphina) juanita Schuster, 1952. Brooklyn Ent. Soc, Bui. 47: 57. ♂.
PhotomorpliHfi {Pliotoniorphina) juanita Schuster, 1958. Ent. Amer. (n. s.) 37: 43. ♂.
Preocc. by and a syn. o{ Photoniorplius (Photomorphina) juanita Schuster, 1952.

Photomorphus minimus Schuster
Ariz. (Wellton).

Photomorphu.'i (Photomorphina) minima Schuster, 1958. Ent. Amer. (n. s.) 37: 43. ♀.

Photomorphus myrmicoides (Cockerell)
Ala., Tex., Ill, Iowa, Kans., S. Dak.

Mutilla panmla Blake, 1886. Amer. Ent. Soc, Trans. 13: 206. ♀. Preocc.
Sphaerophthalnia (!) mip-micoides Cockerell, 1895. Ent. News 6: 62. ♀.
Mutilla unpar Melander, 1903. Amer. Ent. Soc, Trans. 29: 321. ♀.
Odontophotopsis subtenuis Viereck, 1904. Amer. Ent. Soc, Trans. 30: 85. ♂.

Taxonomy: Bradley, 1916. Amer. Ent. Soc, Trans. 42: 336. 6. — Mickel, 1934. Ent. Soc
Amer., Ann. 27: 611. 9, cJ. —Mickel, 1965. Ent. Soc. Wash., Proc 67: 3-4 (synonymy).

Photomorphus nanullus Schuster
Ariz.

Photomorphuii (Photomorphina) itanulla Schuster, 1958. Ent. Amer. (n. s.) 37: 44. ♂.

Photomorphus obscurus Schuster
Calif., Ariz.

Photomorpkus (Photomorphina) obscura Schuster, 1958. Ent. Amer. (n. s.) 37: 45. ♂.

Photomorphus quadriangulatus Schuster
Ariz.; Mexico (Sonora).

Photomorphus (Photomorphina) quadriangulata Schuster, 1958. Ent. Amer. (n. s.) 37: 42.
6.

Photomorphus sarpedon (Fox)
Tex.; Mexico (Coahuila).

Mutilla sarpedon Fox, 1899. Amer. Ent. Soc, Trans. 25: 267. ♂.

Photomorphus spinci (Bradley)
N. C. to Fla., Ala.

Photopsis (Odontophotopsis) spinci Bradley, 1916. Amer. Ent. Soc, Trans. 42: 335. ♂.

Photopsis (Odontophotopsis) spinci floridensis Schuster, 1944. Brooklyn Ent. Soc, Bui. 39:
155. ♂.

Photomorphus spinciformis Schuster
Tex.

Photomorphus (Photomorphina) spinciformis Schuster, 1958. Ent. Amer. (n. s.) 37: 44. ♂.

Photomorphus thamyras (Fox)
Tex.

Mutilla thamyras Fox, 1899. Amer. Ent. Soc, Trans. 25: 267. ♂.

Photomorphus trunculus (Viereck)
Kans., Okla., Tex., Ariz.

Mutilla simpliciventris Melander, 1903 (Oct.). Amer. Ent. Soc, Trans. 29: 315. 3. Preocc
by Andre, July, 1903.

Odontophotopsis trunculus Viereck, 1904. Amer. Ent. Soc, Trans. 30: 85. ♂.

Odontophotopsis melanderella Krombein, 1951. U. S. Dept. Agr., Monog. 2: 757. N. name.

Photomorphus wheeleri (Melander)
Tex. (Austin).

Mutilla Wheeleri Melander, 1903. Amer. Ent. Soc, Trans. 29: 316. ♂.

Photomorphus vegas Schuster
N. Mex., Ariz., Utah.

Photomorphus (Photomorphina) vegas Schuster, 1958. Ent. Amer. (n. s.) 37: 42. ♂.

Genus PHOTOMORPHUS Subgenus PHOTOMORPHUS Viereck

Photomorphus Viereck, 1903. Ent. News 14: 249.

Type-species: Photomorphus Johnsoni Viereck. Orig. desig.

Photomorphus alogus Viereck
Ga., Fla., Miss.

Photomorphus alogus Viereck, 1903. Ent. News 14: 251. ♂.

Taxonomy: Krombein, 1954. Amer. Ent. Soc, Trans. 80: 3. ♀.

Photomorphus banksi (Bradley)
N. J. south to Fla., Ala.

Sphaerophthaima (!) (Photomorphus) banksi Bradley, 1916. Amer. Ent. Soc, Trans. 42:
331. ♂.

Photomorphus bradleyi (Schuster)
S. C. (Clemson).

Photopsis bradleyi Schuster, 1944. Brooklyn Ent. Soc, Bui. 39: 152. ♀.

Photomorphus johnsoni
***authority mismatch
johnsoni Viereck. N. Y. to Ga., Okla.

Photomorphus Johnsoni Viereck, 1903. Ent. News 14: 249. ♂.




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Photomorplius jolmso7ii var. argentipilis Schuster, 1944. Brooklyn Ent. Soc, Bui. 39: 148.
6.

Photomorphus johnsoni
***authority mismatch
natchitoches Schuster. La.

Plwtomorphus {Pliotomorphus) johnsoni natchitoches Schuster, 1958. Ent. Amer. (n. s.) 37:
46. ♂.

Photomorphus paulus (Bradley)
Ga., Fla.

Photopsis (Odontophotopsis) paula Bradley, 1916. Amer. Ent. Soc, Trans. 42: 333. ♂.

Taxonomy: Schuster, 1944. Brooklyn Ent. Soc, Bui. 39: 154. ♂.

Photomorphus quintilis (Viereck)
Iowa, Kans., La.

Mutilla iPhotonwrphus) qui^itilis Viereck, 1906. Amer. Ent. Soc, Trans. 32: 187. ♂.

Photomorphus rubroscutellatus (Bradley)
Va. (Falls Church).

Sphaerophfhalma (!) (Photomorphus) rubroscutellatus Bradley, 1916. Amer. Ent. Soc,
Trans. 42: 332. ♂.

Genus ODONTOPHOTOPSIS Viereck

Revision: Viereck, 1904. Amer. Ent. Soc, Trans. 30: 81-92.

Genus ODONTOPHOTOPSIS Subgenus ODONTOPHOTOPSIS Viereck

Odontophotopsis Viereck, 1903. Acad. Nat. Sci. Phila., Proc 54: 738.

Type-species: Odontophotopsis exogyrus Viereck. Orig. desig.
Tetraphotopsis Ashmead, 1903. Canad. Ent. 35: 305.

Type-species: Tetraphotopsis Hubbardi Ashmead. Orig. desig. (Not Mutilla
Hubbardii Fox as cited by Mickel, 1928. U. S. Natl. Mus., Bui. 143: 37).

Odontophotopsis acmaea Viereck
Ariz., south. Calif.; Mexico (Sonora).

Odontophotopsis aanaeus Viereck, 1904. Amer. Ent. Soc, Trans. 30: 84. ♂.

Odontophotopsis adonis (Fox)
N. Mex. (Las Cruces).

Mutilla adonis Fox, 1899. Amer. Ent. Soc, Trans. 25: 265. ♂.

Odontophotopsis alemon (Fox)
Tex., N. Mex.

Mutilla aletnon Fox, 1899. Amer. Ent. Soc, Trans. 25: 266. ♂.

Mutilla trita Melander, 1903. Amer. Ent. Soc, Trans. 29: 317. ♂.

Odontophotopsis crucis Viereck, 1904. Amer. Ent. Soc, Trans. 30: 86. ♂.

Odontophotopsis annulata Baker
Calif. (Claremont).

Odontophotopsis a7mulatus Baker, 1905. Invertebrata Pacifica 1: 94. ♂.

Odontophotopsis anomala Schuster
Tex.

Odontophotopsis (Odontophotopsis) anomala Schuster, 1958. Ent. Amer. (n. s.) 37: 60. ♂.

Odontophotopsis appacheorum
***authority mismatch
appacheorum Schuster. Tex.

Odontophotopsis (Odontophotopsis) appacheorum appacheorum Schuster, 1958. Ent.
Amer. (n. s.) 37: 55. ♂.

Odontophotopsis appacheorum
***authority mismatch
rostrata Schuster. Ariz.

Odontophotopsis (Odontophotopsis) appacheorum rostratus Schuster, 1958. Ent. Amer. (n.
s.)37: 55. ♂.

Odontophotopsis argentipilis (Provancher)
Published type locality Fla. is erroneous.

Sphaerophthalma (!) argentipilis Provancher, 1887. Addit. Corr. Faune Ent. Canada Hym.,
p. 251. ♂.

Odontophotopsis armata Schuster
Nev.

Odontophotopsis (Odontophotopsis) armata Schuster, 1958. Ent. Amer. (n. s.) 37: 60. ♂.

Odontophotopsis augusta Viereck
N. Mex. (St. Augustine).

Odontophotopsis augustus Viereck, 1904. Amer. Ent. Soc, Trans. 30: 90. ♂.

Odontophotopsis bicolor (Blake)
Tex., Ariz.

Agama bicolor Blake, 1879. Amer. Ent. Soc, Trans. 7: 252. ♂.

Mutilla Madejskii Dalla Torre, 1897. Cat. Hym., v. 8, p. 56. N. name for Agajna bicolor
Blake, not Mutilla bicolor Pallas.

Taxonomy: Fox, 1899. Amer. Ent. Soc, Trans. 25: 259-260. 6 (in key to group imperialis).




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Odontophotopsis biramosa Schuster
Calif. (Holtville).

Odontophotopsis (Odontopliotopsis) biramosa Schuster, 1952. Brooklyn Ent. Sec, Bui. 47:
43, fig. I. 6.

Odontophotopsis braccata Schuster
Tex., N. Mex.

Odontophotopsis (Odontophotopsis) braccatus Schuster, 1958. Ent. Amer. (n. s.) 37: 54. ♂.

Odontophotopsis clypeata Schuster
Ariz., Nev., Calif.

Odontophotopsis (Odontophotopsis) clypeatns Schuster, 1958. Ent. Amer. (n. s.) 37: 59. ♂.

Odontophotopsis cockerelli (Melander)
Tex. (Cuero).

Mutilla Cockerelli Melander, 1903. Amer. Ent. See, Trans. 29: 307. ♂.

Odontophotopsis conifera Schuster
Ariz.

Odontophotopsis (Odontophotopsis) coniferus Schuster, 1958. Ent. Amer. (n. s.) 37: 48. ♂.

Odontophotopsis cookii Baker
Calif., Nev.

Odontophotopsis cookii Baker, 1905. Invertebrata Pacifica 1: 93. ♂.

Odontophotopsis delodonta Viereck
Ariz.

Odontophotopsis delodontus Viereck, 1904. Amer. Ent. Sec, Trans. 30: 91. ♂.

Odontophotopsis edentata
***authority mismatch
distans Schuster. Tex. (Devils River).

Odontophotopsis (Odontophotopsis) edentata dista^is Schuster, 1958. Ent. Amer. (n. s.) 37:
53. ♂.

Odontophotopsis edentata
***authority mismatch
edentata Schuster. Ariz.

Odontophotopsis (Odontophotopsis) edentata edentata Schuster, 1958. Ent. Amer. (n. s.) 37:
53. ♂.

Odontophotopsis erebus (Melander)
Kans., Okla., Tex., N. Mex., Ariz., Utah; Mexico (Tamaulipas).
Mutilla Erebus Melander, 1903. Amer. Ent. Soc, Trans. 29: 312. ♂.
Odoyitophotopsis avellanus Viereck, 1904. Amer. Ent. Soc, Trans. 30: 88. ♂.

Odontophotopsis eubule (Cameron)
N. Mex.; Mexico (Sonora). Host: Probably Evaniella neomexicana (Ashm.)
in ootheca oi Arenivaga genitalis Caud., or. less likely, the ootheca itself.
Sphaerophthalma (!) eubule Cameron, 1896. Biol. Cent.-Amer., Hym., v. 2, p. 383. ♂.
Mutilla hamata Melander, 1903. Amer. Ent. Soc, Trans. 29: 314. ♂.

Taxonomy: Mickel, 1965. Ent. Soc. Wash., Proc 67: 1 (synonymy).

Biology: Mickel, 1974. Ent. Soc. Amer., Ann. 67: 468 (host).

Odontophotopsis exogyra Viereck
Calif. (La Jolla).

Odontophotopsis exogyrus Viereck., 1903. Acad. Nat. Sci. Phila., Proc. 54: 738. ♂.

Odontophotopsis fallax Viereck
Nev.

Odontophotopsis fallax Viereck, 1904. Amer. Ent. Soc, Trans. 30: 89. ♂.
Odontophotopsis ocellatus Baker, 1905. Invertebrata Pacifica 1: 98. ♂.

Odontophotopsis grandiceps Schuster
Ariz. (Globe).

Odontophotopsis (Odontophotopsis) grandiceps Schuster, 1958. Ent. Amer. (n. s.) 37: 53. ♂.

Odontophotopsis grata (Melander)
N. Mex. (La Cueva).

Mutilla grata Melander, 1903. Amer. Ent. Soc, Trans. 29: 308. ♂.

Odontophotopsis hexadonta Schuster
Calif.

Odontophotopsis (Odontophotopsis) hexadontus Schuster, 1958. Ent. Amer. (n. s.) 37: 51. ♂.

Odontophotopsis inconspicua
***authority mismatch
brunnea Schuster. Ariz.

Odontophotopsis (Odontophotopsis) inconspicua brunnea Schuster, 1958. Ent. Amer. (n. s.)
37: 54. ♂.

Odontophotopsis inconspicua
***authority mismatch
inconspicua (Blake). Calif., Nev.

Photopsis inconspicuus Blake, 1886. Amer. Ent. Soc, Trans. 13: 272. ♂.
Mutilla infelix Dalla Torre, 1897. Cat. Hym., v. 8, p. 50. N. name for Photopsis
inconspicuus Blake, not Mutilla inconspicua Smith.

Odontophotopsis lamellifera Schuster
Ariz.

Odontophotopsis (Odontophotopsis) lamellifera Schuster, 1958. Ent. Amer. (n. s.) 37: 56. ♂.

Odontophotopsis melicausa
***authority mismatch
melicausa (Blake). B. C, Mont, Calif., Ariz., Tex.

Agama melicausa Blake, 1871. Amer. Ent. Soc, Trans. 3: 261. ♂.
Mutilla brevicornis Fox, 1899. Amer. Ent. Soc, Trans. 25: 255. ♂.
Odontophotopsis mellicomis Baker, 1905. Invertebrata Pacifica 1: 96. ♂.




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Taxonomy: Schuster, 1958. Ent. Amer. (n. s.) 37: 58 (synonymy).

tnelicausa piceipes Schuster. Mont.

Odontophotopsis (Odontophotopsis) mellicausa (!) piceipes Schuster, 1958. Ent. Amer. (n.
s.)37:58. ♂.
tnelicausa westcottii (Melander). Tex., N. Mex.

Mutilla Westcottii Melander, 1903. Amer. Ent. Soc, Trans. 29: 310. ♂.
Odontophotopsis indotatus Viereck, 1904. Amer. Ent. Soc, Trans. 30: 89. ♂.

Odontophotopsis microdonta Ferguson
Nev., Calif.

Odontophotopsis (Odontophotopsis) microdonta Ferguson, 1967. Brigham Young Univ. Sci.,
Bui., Ser. 8, no. 4: 22. ♂.

Odontophotopsis obliqua Viereck
B. C, Mont., Idaho, Wash., Nev., Calif.; Mexico (Baja California).
Odontophotopsis obliquus Viereck, 1925. Canad. Ent. 56: 112. ♂.

Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci., Bui., Ser. 8, no. 4: 23 (redescription).

Odontophotopsis parksiana Schuster
Tex.

Odontophotopsis (Odontophotopsis) parksiana Schuster, 1958. Ent. Amer. (n. s.) 37: 57. ♂.

Odontophotopsis parva Schuster
Ariz. (Arlington).

Odontophotopsis (Odontophotopsis) parva Schuster, 1958. Ent. Amer (n. s.) 37: 55. ♂.

Odontophotopsis pudica (Melander)
Wash., Calif.

Mutilla pudica Melander, 1903. Amer. Ent. Soc, Trans. 29: 309. ♂.
Odontophotopsis atripes Mickel, 1938. Pan-Pacific Ent. 14: 182. ♂.

Odontophotopsis quadridentata Schuster
Calif.

Odontophotopsis (Odontophotopsis) quadridentata Schuster, 1958. Ent. Amer. (n. s.) 37: 51.
6.

Odontophotopsis quadrispinosa Schuster
Nev., Calif.; Mexico (Baja California).

Odontophotopsis (Odontophotopsis) quadrispinosa Schuster, 1958. Ent. Amer. (n. s.) 37: 51.
^■
Taxonomy: Ferguson, 1967. Brigham Young Univ. Sci., Bui., Ser. 8, no. 4: 24 (redescription).

Odontophotopsis serca Viereck
Nev., Calif.; Mexico (Baja California). Host: Anthophora linsleyi Timberlake.
Odontophotopsis sercus Viereck, 1904. Amer. Ent. Soc, Trans. 30: 87. ♂.

Biology: Linsley and MacSwain, 1942. Amer. Midland Nat. 27: 409 (host).

Odontophotopsis setifera Schuster
South. Calif., Nev., Ariz.

Odontophotopsis (Odontophotopsis) setifera Schuster, 1952. Brooklyn Ent. Soc, Bui. 47: 47.
i.

Odontophotopsis succinea Viereck
Calif.

Odontophotopsis succineus Viereck, 1903. Acad. Nat. Sci. Phila., Proc 54: 741. ♂.

Odontophotopsis tapajos
***authority mismatch
aulus (Blake). Tex.

Agama Aulus Blake, 1872. Amer. Ent. Soc, Trans. 4: 75. ♂.

Odontophotopsis tapajos
***authority mismatch
tapajos (Blake). Tex., Ariz.

Agama Tapajos Blake, 1871. Amer. Ent. Soc, Trans. 3: 262. ♂.

Agama Astyanax Blake, 1879. Amer. Ent. Soc, Trans. 7: 254. ♂.

Odontophotopsis tenuiptera Schuster
Ariz.; Mexico.

Odontophotopsis (Odontophotopsis) tenuiptera Schuster, 1958. Ent. Amer. (n. s.) 37: 47. ♂.

Odontophotopsis territa
***authority mismatch
obscura Schuster. Ariz.

Odontophotopsis (Odontophotopsis) territa obscura Schuster, 1958. Ent. Amer. (n. s.) 37:
59. ♂.

Odontophotopsis territa
***authority mismatch
territa (Cockerell). N. Mex. (Las Cruces).

Photopsis territus Cockerell, 1894. Ent. News 5: 200. ♂.

Odontophotopsis unicornis Schuster
Ariz., Calif.

Odontophotopsis (Odontophotopsis) u7iicor7iis Schuster, 1958. Ent. Amer. (n. s.) 37: 52. ♂.

Odontophotopsis venusta (Blake)
Ariz.

Photopsis venustus Blake, 1886. Amer. Ent. Soc, Trans. 13: 270. ♂.

Mutilla pretiosissivia Dalla Torre, 1897. Cat. Hym., v. 8, p. 74. N. name for Photopsis
venustus Blake, not Mutilla venusta Smith.

Tetraphotopsis Hubbardi Ashmead, 1903. Canad. Ent. 35: 305. ♂.




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Taxonomy: Schuster, 1958. Ent. Amer. (n. s.) 37: 54 (synonymy).

Odontophotopsis viereckii Baker
Nev. (Ormsby Co.).

Odontophotopsis viereckii Baker, 1905. Invertebrata Pacifica 1: 97. ♂.

Genus ODONTOPHOTOPSIS Subgenus PERIPHOTOPSIS Schuster

Odontophotopsis subg. Periphotopsis Schuster, 1958. Ent. Amer. (n. s.) 37: 8 (in key), 60.
Type-species: Odontophotopsis (Periphotopsis) inamatus Schuster. Grig, desig.

Odontophotopsis mamata Schuster
Ariz., Nev., Calif.

Odontophotopsis (Periphotopsis) mamatus Schuster, 1958. Ent. Amer. (n. s.) 37: 60. ♂.

Genus PROTOPHOTOPSIS Schuster

Genus PROTOPHOTOPSIS Subgenus PROTOPHOTOPSIS Schuster

Protophotopsis Schuster, 1947. Ent. Soc. Amer., Ann. 39: 693.

Type-species: Protophotopsis scudderi Schuster. Orig. desig.

Only the typical subgenus occurs in North America.

Taxonomy: Schuster, 1949. Ent. Amer. (n. s.) 29: 82-85.

Protophotopsis scudderi Schuster
Kans., Tex., Colo.

Protophotopsis scudderi Schuster, 1947. Ent. Soc. Amer., Ann. 39: 694. ♂.

Protophotopsis venenaria (Melander)
Kans., Tex., Calif. Possibly the opposite sex of scudderi Schuster.

Mutilla venenaria Melander, 1903. Amer. Ent. Soc, Trans. 29: 320. ♀.

Taxonomy: Schuster, 1949. Ent. Amer. (n. s.) 29: 83.

Tribe PSEUDOMETHOCINI

All species are diurnal. The authenticated host records suggest that members of this tribe are
parasitic on various kinds of ground-nesting bees, principally halictids belonging to the subfami-
lies Halictinae and Nomiinae. Fattig, 1943 (Emory Univ. Mus. Bui. 1: 4) records frigida (Smith)
as having been reared from mud cells of the eumenid wasp Ancistrocenis iutojiidns Boh.; this
may be based on a misidentification.

Revision: Mickel, 1924. U. S. Natl. Mus., Proc. 64(15): 1-51, pis. 1-4. — Mickel, 1935. Amer.
Ent. Soc, Trans. 61: 383-398.

Genus PSEUDOMETHOCA Ashmead

Pseudomethoca Ashmead, 1896. Amer. Ent. Soc, Trans. 23: 181.

Type-species: Photopsis Cressoni Fox. Orig. desig. (= Pseudomethoca frigida
(Smith)).
Nomiaephagus Ashmead, 1899. N. Y. Ent. Soc, Jour. 7: 56.

Type-species: Mutilla (Sphaeropthalma) Sanbomii Blake. Orig. desig.

Pseudomethoca albicoma Mickel
Tex. (Rio Grande in Brewster Co.).

Pseudomethoca albicoma Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 29. ♂.

Pseudomethoca anthracina (Fox)
Oreg., Calif.

Sphaerophthalma (!) anthracina Fox, 1892. Ent. News 3: 172. ♂.
Mutilla anthracicolor Dalla Torre, 1897. Cat. Hym., v. 8, p. 9. N. name for

Sphaerophthalma (!) anthracina Fox, not Mutilla anthracina Gerstaecker.
Mutilla harpalyce Fox, 1899. Amer. Ent. Soc, Trans. 25: 227. ♀.

Taxonomy: Kurd, 1951. Pan-Pacific Ent. 27: 156 (synonymy).

Pseudomethoca athamas (Fox)
Oreg., Calif.

Mutilla athamas Fox, 1899. Amer. Ent. Soc, Trans. 25: 225. cj.

Pseudomethoca aureovestita Bradley
N. Mex., Ariz.

Pseudomethoca aureovestita Bradley, 1924. In Mickel, U. S. Natl. Mus., Proc. 64 (15): 22.
9,6.

Pseudomethoca bequaerti Mickel
Tex., N. Mex., Ariz., Colo., Mont.

Pseudomethoca bequaerti Mickel, 1924. U. S. Natl. Mus., Proc 64 (15): 13. ♀.




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Pseudomethoca brazoria (Blake)
Okla., Tex.

Mutilla (Sphaeropthalma) brazoria Blake, 1871. Amer. Ent. Soc, Trans. 3: 255. ♀.

Pseudomethoca carbonaria Mickel
Tex.

Pseudomethoca carbonaria Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 43. ♂.

Pseudomethoca cephalargia Mickel
Ariz. (Santa Catalina Mts.).

Pseudomethoca cephalargia Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 17. ♀.

Pseudomethoca connectens (Cresson)
Calif.

Mutilla co7inectens Cresson, 1865. Ent. Soc. Phila., Proc. 4: 387. ♀.
contumax (Cresson). Iowa, S. Dak., Nebr., Kans., Tex., Colo., Ariz.; Mexico.

Mutilla contu77iax Cresson, 1865. Ent. Soc. Phila., Proc. 4: 437. ♀.

Mutilla 7nicrophthaima Gerstaecker, 1874. Arch. Naturgesch. 40: 64. ♂.

Pseudomethoca contumeliosa Mickel
Utah, Idaho, Calif.

Pseudomethoca coiitumeliosa Mickel, 1935. Amer. Ent. Soc, Trans. 61: 393. ♀.

Pseudomethoca damia (Cameron)
Ariz.; Mexico (Guerrero).

Sphaerophthahna (!) datnia Cameron, 1894. Biol. Cent.-Amer., Hym., v. 2, p. 383. ♂.

Mutilla sphaerophthalmica Dalla Torre, 1897. Cat. Hym., v. 8, p. 87. N. name.

Pseudomethoca dentifrontalis Bradley
Oreg., Calif.

Pseudoviethoca doitifrontalis Bradley, 1924. /«, Mickel, U. S. Natl. Mus., Proc. 64 (15): 11.
9.

Pseudomethoca dentigula Mickel
Tex. (Trans-Pecos).

Pseudomethoca dentigula Mickel, 1935. Amer. Ent. Soc, Trans. 61: 391. ♀.

Pseudomethoca donaeanae (Cockerell)
Tex., N. Mex., Ariz., Calif.

Sphaerophthalma (!) donae-anae Cockerell, 1897. In Cockerell and Fox, Acad. Nat. Sci.
Phila., Proc 49: 127. ♀.

Pseudomethoca flammigera Mickel
Ariz.

Pseudomethoca flammigera Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 21. ♀.

Pseudomethoca flavida (Blake)
Tex.

Mutilla (Sphaeropthalma) flavida Blake, 1871. Amer. Ent. Soc, Trans. 3: 249. ♂.

Pseudomethoca frigida (Smith)
Southern Canada and U. S. east of Rocky Mts. Host: Diaiictus zephyrus (Sm.),
D. coeruleus (Robt.), D. rohioeri (Ellis)?, D. laevissiynus (Sm.)?, D. versatus (Robt.), D.
i7)iitatus (Sm.)?, Evylaeus cinctipes (Prov.), Augochlorella striata (Prov.)?

Mutilla frigida Smith, 1855. Cat. Hym. Brit. Mus., v. 3, p. 60. ♀.

Mutilla (.Sphaeropthahna) canadensis Blake, 1871. Amer. Ent. Soc, Trans. 3: 252. ♀.

Photopsis Cressoni Fox, 1890. Ent. News 1: 138. ♂.

Sphaerophthalma (!) alveolata Provancher, 1895. Nat. Canad. 22: 110. ♀.

Mutilla neojerseiensis Dalla Torre, 1897. Cat. Hym., v. 8, p. 65. N. name for Phytopsis (!)
Cressoni Fox, not Mutilla Cressoni Blake.

Taxonomy: Brothers, 1972. Kans. Univ. Sci. Bui. 50: 25-31, figs. 11-40 (egg, larvae, pupae).

Biology: Melander and Brues, 1903. Biol. Bui. 5: 4-7, fig. 4 (behavior in host nest).

— Krombein, 1938. Brooklyn Ent. Soc, Bui. 33:14-15 (behavior in host nest). — Fattig, 1943.
Emory Univ. Mus. Bui. 1: 4 (questionable host record). — Shappirio, 1947. Brooklyn Ent.
Soc, Bui. 42: 163 (mating behavior). — Michener and Wille, 1961. Kans. Univ. Sci. Bui. 42:
1129-1130 (behavior in host nests). —Lin, 1964. Ins. Sociaux 11: 187-192 (behavior in host
nests). — Batra, 1965. Kans. Ent. Soc, Jour. 38: 383-386, figs. 6 a-c, 7 a-b (life history,
behavior). —Michener, 1966. Kans. Ent. Soc, Jour. 39: 196-197 (host record). — Knerer and
Atwood, 1967. Ent. Soc. Ontario, Proc 97: 106 (host record). —Brothers, 1972. Kans. Univ.
Sci. Bui. 50: 1-25, figs. 1-10 (life history, behavior).

Pseudomethoca gila (Blake)
Tex. (Brownsville).

Mutilla (Sphaeropthalwa) Gila Blake, 1871. Amer. Ent. Soc, Trans. 3: 250. ♂.

Pseudomethoca ilione (Fox)
Tex., Okla.

Mutilla ilione Fox, 1899. Amer. Ent. Soc, Trans. 25: 268. ♀.
Mutilla aprica Melander, 1903. Amer. Ent. Soc, Trans. 29: 322. ♀.

Taxonomy: Mickel, 1965. Ent. Soc Wash., Proc. 67: 2 (synonymy).

Pseudomethoca klotsi Mickel
Colo. (Hugo).

Pseudomethoca klotsi Mickel, 1935. Amer. Ent. Soc, Trans. 61: 389. ♀.




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Pseudomethoca manca Mickel
Utah.

Paeudoniethoca manca Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 45. ♂.

Pseudomethoca meritoria Mickel
Tex.

P>

Pseudomethoca mulaiki Mickel
Tex. (Cameron Co.).

Pseudomethoca Diiilaiki Mickel, 1938. Ent. Soc. Amer., Ann. 31: 150. ♀.

Pseudomethoca nephele (Fox)
Tex.

Mufilla nephele Fox, 1899. Amer. Ent. Soc, Trans. 25: 223. ♀.

Pseudomethoca nigricula Mickel
Ariz.

Pseudomethoca nigricula Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 26. ♂.

Pseudomethoca nudula Mickel
N. Mex. (Pecos). Host: Possibly Calliopsis sp.

Pseudomethoca nudula Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 15. ♀.

Pseudomethoca oceola (Blake)
N. J. south to Fla., west to S. Dak. and Ariz.

Mutilla (Sphaewpthalma) Oceola Blake, 1871. Amer. Ent. Soc, Trans. 3: 248. ♂.

Mutilla hippodamia Fox, 1899. Amer. Ent. Soc, Trans. 25: 227. ♀.

Pseudomethoca oculata (Banks)
N. C, Ga., Fla.

Nomiaephagun oculatus Banks, 1921. Ent. Soc Amer., Ann. 14: 24. ♀.

Pseudomethoca oculissima Mickel
N. Mex. (Pecos).

Pseudomethoca oculissima Mickel, 1924. U. S. Natl. Mus., Proc 64 (15): 14. ♀.

Pseudomethoca paludata Mickel
Nebr., Kans., Wyo., Colo., N. Mex., Ariz.

Pseudomethoca paludata Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 29. ♀.
Pseudomethoca scrupulosa Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 31. ♂.

Pseudomethoca pigmentata Mickel
Tex.

Pseudomethoca pigmentata Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 24. ♀.

Pseudomethoca praeclara (Blake)
Ariz., Calif.

Sphaerophthalma (!) praeclara Blake, 1886. Amer. Ent. Soc, Trans. 13: 252. ♀.

Mutilla aegaeon Fox, 1899. Amer. Ent. Soc, Trans. 25: 229. ♂.

Nomiaephagus acuum Cockerell, 1915. Entomologist 48: 250. ♀.

Pseudomethoca (Nomiaephagus) aegeon (!) Bradley, 1916. Amer. Ent. Soc, Trans. 42: 321.
6.

Pseudomethoca propinqua (Cresson)
Minn, south to Tex., west to Alta. and Ariz. Host: Noinia m. melanderi
Ckll.

Mutilla propinqua Cresson, 1865. Ent. Soc Phila., Proc. 4: 433. ♂.

Mutilla montivaga Cresson, 1865. Ent. Soc. Phila., Proc. 4: 436. ♀.

Pseudomethoca sacatona Caldwell, 1939. Pan-Pacific Ent. 15: 94. ♀.

Biology: Krombein, 1958. U. S. Dept. Agr., Monog. 2, Sup. 1: 103 (host record).

Pseudomethoca quadrinotata Mickel
Ariz. (Tucson).

Pseudomethoca quadrinotata Mickel, 1938. Ent. Soc. Amer., Ann. 31: 149. ♀.

Pseudomethoca russeola Mickel
Tex. (San Diego).

Pseudomethoca russeola Mickel, 1924. U. S. Natl. Mus., Proc 64 (15): 44. ♂.

Pseudomethoca sanbornii
***authority mismatch
acetis (Fox). ?Conn., N. C. south to Fla., west to Tex. in L. Austr. Zone.
Mutilla aeetis Fox, 1899. Amer. Ent. Soc, Trans. 25: 228. ♀.

Pseudomethoca {Nomiaephagus) aetis (!) Bradley, 1916. Amer. Ent. Soc, Trans. 42: 312,
316. ♀.
Taxonomy: Krombein, 1953. Wasmann Jour. Biol. 10: 308 (reduced to subsp. rank).

Pseudomethoca sanbornii
***authority mismatch
sanbornii (Blake). Mass. south to Fla., west to Nebr. and Tex. in Transition and U.
Austr. Zones. Host: Noinia pattoni Cockerell.
Mutilla (Sphaeropthalma) Sanboniii Blake, 1871. Amer. Ent. Soc, Trans. 3: 248. ♂.

Biology: Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 38 (host).

Pseudomethoca scaevolella (Cockerell and Casad)
Tex., N. Mex.

Sphaerophthalma (!) scaevolella Cockerell and Casad, 1895. Amer. Ent. Soc, Trans. 22:
298. ♀.

Pseudomethoca simillima (Smith)
Mass. south to Fla., west to Nebr. and Tex.

Mutilla simillima Smith, 1855. Cat. Hym. Brit. Mus., v. 3, p. 62. ♀.




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M lit ilia gerijon Fox, 1899. Amer. Ent. Soc, Trans. 25: 255. ♂.

Mutilla Henshawi Melander, 1903. Amer. Ent. Soc, Trans. 29: 303. ♂.

Mutilla daeckei Rohwer, 1910. Ent. Soc. Wash., Proc. 12: 49. ♂.

Pseudomefhoca swiillinia Mickel, 1924. U. S. Natl. Mus., Proc. 64 (15): 32. ♀ (♂ misdet.).

Taxonomy: Krombein, 1948. Ent. News 59: 187-189.

Biology: Shappirio, 1947. Brooklyn Ent. Soc, Bui. 42: 163 (mating behavior).

Pseudomethoca torrida Krombein
Fla.

Pseudometlioca frigida torrida Krombein, 1954. Amer. Ent. Soc, Trans. 80: 7, fig. 7. ♀, ♂.

Pseudomethoca toumeyi (Fox)
Tex., Ariz.; Mexico (Sonora).

Sphaerophthahna (!) toumeyi Fox, 1894. Ent. News 5: 297. ♀.

Taxonomy: Mickel, 1974. Ent. Soc. Amer., Ann. 67: 468. ♂.

Pseudomethoca vanduzei Bradley
Va., S. C, Ga., Fla., Tex. Host: Nomia maneei Ckll.

Pseudomethoca (Nomiaephagus) vanduzei Bradley, 1916. Amer. Ent. Soc, Trans. 42: 321.

6.
Pseudomefhoca fattigi Mickel, 1938. Ent. Soc. Amer., Ann. 31: 147. ♀.
Pseudomefhoca vanduzeei (!) Krombein, 1951. U. S. Dept. Agr., Monog. 2: 761. Emend.

Biology: Fattig, 1943. Emory Univ. Mus. Bui. 1: 6 (host record).
wickhami (Cockerell and Casad). Tex. (Fedor).

Sphaerophfhalvia (!) wickhami Cockerell and Casad, 1895. Amer. Ent. Soc, Trans. 22: 297.
9.

Genus MYRMILLOIDES Andre

Myrmilloides Andre, 1903. In Wytsman, Gen. Ins., fasc. 11, p. 26.

Type-species: Mutilla (Sphaeropfhahna) grandiceps Blake. Orig. desig.

Myrmilloides grandiceps (Blake)
Iowa, Nebr., Colo., Kans., Mo., Okla., Ark., Tex., La., Tenn. Host: Dialictus
zephyrus (Sm.)?, Augocklorella sfi'iafa (Prov.)?, A. persiinilis (Vier.)?
Mufilla (Sphaeropthalma) grandiceps Blake, 1872. Amer. Ent. Soc, Trans. 4: 74. ♀, ♂.

Biology: Melander, 1903. Amer. Ent. Soc, Trans. 29: 293 (host record). — Ordway, 1964. Kans.
Ent. Soc, Jour. 37: 149 (host records).

Tribe DASYMUTILLINI

Members of this tribe are diurnal except for a few species of Dasymutilla.

Genus DASYMUTILLA Ashmead

Dasymutilla Ashmead, 1899. N. Y. Ent. Soc, Jour. 7: 57.

Type-species: Mufilla (Sphaeropthalma) Gorgon Blake. Orig. desig.
Bruesia Ashmead, 1903. Canad. Ent. 35: 306.

Type-species: Mufilla harmonia Fox. Orig. desig.
Pycnomufilla Ashmead, 1904. Canad. Ent. 36: 8.

Type-species: Mufilla (Sphaeropthalma) Waco Blake. Orig. desig.

Revision: Mickel, 1928. U. S. Natl. Mus., Bui. 143: 1-351, 5 pis., 28 text figs. —Mickel, 1936.
Ent. Soc. Amer., Ann. 29: 29-60, 3 figs, (revised key to spp.).

Taxonomy: Hurd, 1951. Calif. Ins. Survey Bui. 1: 89-118, 1 pi. (Calif, spp.)
Morphology: Snodgrass, 1941. Smithsn. Inst., Misc. Collect. 99, no. 14, p. 43, pi. 14 A-F (male
genitalia).

Dasymutilla albiceris Mickel
Calif. (San Bernardino Co.); Mexico (Baja California).

Dasymutilla albiceris Mickel, 1936. Pan-Pacific Ent. 12: 91. ♀.

Dasymutilla alesia Banks
Mass. south to Fla., west to 111. and Okla.

Dasymutilla alesia Banks, 1921. Ent. Soc. Amer., Ann. 14: 24. ♀.

Dasymutilla angulata Krombein
Mich., 111., Iowa, Mo., S. Dak., Nebr., Kans.

Mufilla anguliceps Fox, 1899. Amer. Ent. Soc, Trans. 25: 240. 9. Preocc by Andre, 1897.

Dasymutilla angulata Krombein, 1951. U. S. Dept. Agr., Monog. 2: 762. N. name.




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Dasymutilla apicalata (Blake)
Tex., Ariz.; Mexico (Guerrero).

Mntilla {Sphaeropthahna) apicalata Blai Splmerophtliaima (!) perfidiosa Cameron, 1895. Biol. Cent.-Amer., Hym., v. 2, p. 368. ♂.

Preocc. in Mntilla.
Splmerophtliaima (!) giicrreroensis Cameron, 1895. Biol. Cent.-Amer., Hym., v. 2, p. 369. ♂.
Mntilla perfida Dalla Torre, 1897. Cat. Hym., v. 8, p. 72. N. name.

Taxonomy: Mickel, 1965. Ent. Soc. Wash., Proc. 67: 2 (synonymy).

Dasymutilla arcana Mickel
Nebr., Okla., Tex., N. Mex., Colo.

Dasymntilla arcana Mickel, 1928. U. S. Natl. Mus., Bui. 143: 217. ♂.

Dasymutilla arenivaga Mickel
Ariz., Calif. D. megaloplithalvia Mick, may be the male.

Dasymntilla arenivaga Mickel, 1928. U. S. Natl. Mus., Bui. 143: 278. ♀.

Dasymutilla arenivaga
***authority mismatch
var. unicolor Mickel. Ariz. (Ft. Mojave).

Dasymntilla arenivaga var. nnicolor Mickel, 1936. Ent. Soc. Amer., Ann. 29: 59. ♀.

Dasymutilla asopus
***authority mismatch
asopus (Cresson). Man., Minn., N. Dak., Mont., S. Dak., Nebr., Kans., Colo., Tex. Host:
Dianthidiuin sp.?

Mutilla Asopns Cresson, 1865. Ent. Soc. Phila., Proc. 4: 435. ♂.

Ephuta sparsiformis Cockerell and Rohwer, 1908. Psyche 15: 4. ♀.

Biology: Hicks, 1926. Colo. Univ., Studies, p. 249. -Mickel, 1928. U. S. Natl. Mus., Bui. 143:
61.

Dasymutilla asopus
***authority mismatch
bexar (Blake). Mass. south to Fla., Miss., Tex., Mo., Ind. Host: Anthophora abrnpta

Say, A. nrsina Cr.; Dianthidiuvi cnrvatnm sayi Ckll., Paranthidium j. jugatorinm Say.
Mntilla {Sphaeropthahna) Bexar Blake, 1871. Amer. Ent. Soc, Trans. 3: 238. ♂.
Mutilla hannonia Fox, 1899. Amer. Ent. Soc, Trans. 25: 229. ♀.

Biology: Fattig, 1943. Emory Univ. Mus. Bui. 1: 4-5 (host records). — Krombein, 1958. U. S.
Dept. Agr., Monog. 2, Sup. 1: 103 (host record). — Michener, 1975. Kans. Ent. Soc, Jour. 48:
199 (host).

Dasymutilla asopus
***authority mismatch
Cassandra Mickel. Fla.

Dasymntilla cassa^idra Mickel, 1928. U. S. Natl. Mus., Bui. 143: 64. ♂.
asteria Mickel. Ariz. (Pima Co.). Host: Microbembex nigrifrons (Prov.).

Dasymntilla asteria Mickel, 1936. Ent. Soc Amer., Ann. 29: 57. ♀.

Biology: Haddock, 1967. hi Krombein, U. S. Dept. Agr., Monog. 2, Sup. 2: 339 (host record).
atricauda Mickel. Calif., Ariz.

Dasymntilla atricanda Mickel, 1936. Pan-Pacific Ent. 12: 92. ♀.

Dasymutilla atrifimbriata Mickel
Kans., Ark., Okla., Tex., Ariz.

Dasymutilla atrifimbriata Mickel, 1928. U. S. Natl. Mus., Bui. 143: 169. ♀.

Dasymutilla atrifulva Mickel
Ariz.

Dasymntilla atrifnlva Mickel, 1928. U. S. Natl. Mus., Bui. 143: 290. ♂.

Dasymutilla aureola (Cresson)
Nev., Oreg., Calif. Host: Anthophora stanfordiana Ckll., Melissodes sp.

Mntilla anreola Cresson, 1865. Ent. Soc. Phila., Proc. 4: 386. ♀.

Sphaerophthalma (!) pannosa Blake, 1886. Amer. Ent. Soc, Trans. 13: 210. ♀.

Sphaerophthalma (!) mollissima Blake, 1886. Amer. Ent. Soc, Trans. 13: 215. ♂.

Biology: Krombein, 1958. U. S. Dept. Agr., Monog. 2, Sup. 1: 103 (host records).

Dasymutilla aureola
***authority mismatch
var. pacifica (Cresson). B. C, Calif.

Mntilla pacifica Cresson, 1875. Amer. Ent. Soc, Trans. 5: 120. ♀.

Dasymutilla biguttata (Cockerell)
Miss., La., Tex., Ark., Kans., Ariz.

Sphaerophthalma (!) qnadrignttata var. biguttata Cockerell, 1895. Ent. News 6: 63. ♀.

Dasymutilla bioculata (Cresson)
Man. south to La., west to B. C. and N. Mex. Host: Bembix pruinosa Fox,
B. americana spinolae Lep.?, Microbembex monodonta (Say).

Mntilla biocnlata Cresson, 1865. Ent. Soc Phila., Proc. 4: 431. ♂.

Mntilla Aegina Cresson, 1865. Ent. Soc. Phila., Proc 4: 435. ♀.

Mntilla chlamydata Melander, 1903. Amer. Ent. Soc, Trans. 29: 299. ♀.




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Biology: Mickel, 1924. Ent. News 35: 236-242. — Krombein, 1958. U. S. Dept. Agr., Monog. 2,

Sup. 1: 103 (host record). — Cottrell, 1972. In Brothers, Kans. Univ. Sci. Bui. 50: 31-36, figs.

41-42 (life history).

Dasymutilla birkmani (Melander)
Nebr., Wyo., Kans., Okla., Tex.

Mutilla Birkmani Melander, 1903. Amer. Ent. Soc, Trans. 29: 313. ♀.

Dasymutilla bollii (Fox)
Miss., La., Tex., Okla., Kans.

Mutilla Bollii Fox, 1899. Amer. Ent. Soc, Trans. 25: 242. ♀.

Dasymutilla blawana Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 461. ♀.

Dasymutilla bonita Mickel
Ariz.

Dasymutilla boyiita Mickel, 1928. U. S. Natl. Mus., Bui. 143: 208. ♀.

Dasymutilla poliothrix Mickel, 1928. U. S. Natl. Mus., Bui. 143: 285. ♂.

Dasymutilla californica (Radoszkowski)
B. C, Oreg., Utah, Calif.

Mutilla Californica Radoszkowski, 1861. Soc. Ent. Rossica, Horae 1: 86. ♀.

Dasymutilla abdita Mickel, 1928. U. S. Natl. Mus., Bui. 143: 293.

Dasymutilla californica
***authority mismatch
var. clio (Blake). B. C, Idaho, Nev., Calif.

Mutilla Clio Blake, 1879. Amer. Ent. Soc, Trans. 7: 251. ♀.

Dasymutilla calorata Mickel
Kans., Okla., Tex., N. Mex., Colo.

Dasymutilla calorata Mickel, 1928. U. S. Natl. Mus., Bui. 143: 227. ♀.

Dasymutilla campanula Mickel
Kans., Colo.

Dasymutilla campanula Mickel, 1928. U. S. Natl. Mus., Bui. 143: 178. ♀.
Candida Mickel. Ariz (Douglas).

Dasymutilla Candida Mickel, 1928. U. S. Natl. Mus., Bui. 143: 296. ♂.
canella (Blake). Mass. to Pa., west to Man. and Tex. Host: Dialictus pruinosus (Robt.)?

Mutilla (Sphaeropthalma) canella Blake, 1871. Amer. Ent. Soc, Trans. 3: 239. ♂.

Mutilla nigulosa Fox, 1899. Amer. Ent. Soc, Trans. 25: 240. ♀.

Mutilla infensa Melander and Brues, 1903. Biol. Bui. 5: 24. ♀.

Biology: Viereck, 1902. Ent. News 13: 72. —Melander and Brues, 1903. Biol. Bui. 5: 25

(putative host).

Dasymutilla caneo (Blake)
Minn., N. Dak., south to Tex. and N. Mex., west to Alta. and B. C.

Mutilla caneo Blake, 1879. Amer. Ent. Soc, Trans. 7: 250. ♀.

Mutilla mixtura Blake, 1879. Amer. Ent. Soc, Trans. 7: 251. ♀.

Mutilla myirha Fox, 1899. Amer. Ent. Soc, Trans. 25: 258. ♀.

Dasymutilla castor (Blake)
S. C, Ga., Fla., Ala., Miss., La., Tex. Host: Polistes fuscatus (F.).

Mutilla (Sphaeropthalma) castor Blake, 1871. Amer. Ent. Soc, Trans. 3: 237. ♂.

Dasymutilla mediatoria Mickel, 1938. Ent. Soc. Amer., Ann. 31: 152. ♀.

Biology: Fattig, 1943. Emory Univ. Mus. Bui. 1: 6 (host record).

Dasymutilla chattahoochei Bradley
N. C. south to Fla.

Dasymutilla {Dasymutilla) chattahoochei Bradley, 1916. Amer. Ent. Soc, Trans. 42: 324.

9.
Dasymutilla {Dasymutilla) arenerronea Bradley, 1916. Amer. Ent. Soc, Trans. 42: 324. ♀.

Taxonomy: Krombein, 1954. Amer. Ent. Soc, Trans. 80: 9, fig. 5 (description male).

Dasymutilla chiron (Blake)
Tex.

Mutilla {Sphaeropthalma) Chiron Blake, 1872. Amer. Ent. Soc, Trans. 4: 72. ♂.

Mutilla Ursula var. texana Cresson, 1875. Amer. Ent. Soc, Trans. 5: 120. ♂.

Dasymutilla chiron
***authority mismatch
var. ursuIa (Cresson). B. C. south to Ariz, and Calif, and southeast to Tex.

Mxitilla Ursula Cresson, 1875. Amer. Ent. Soc, Trans. 5: 120. ♂.

Dasymutilla chisos Mickel
Tex.

Dasymutilla chisos Mickel, 1928. U. S. Natl. Mus., Bui. 143: 284. ♂.

Dasymutilla clotho (Blake)
Okla., Tex.

Mutilla {Sphaeropthalma) Clotho Blake, 1872. Amer. Ent. Soc, Trans. 4: 72. ♀.

Dasymutilla clytemnestra (Fox)
Oreg., Calif.

Mutilla clytemnestra Fox, 1899. Amer. Ent. Soc, Trans. 25: 246. ♀.

Dasymutilla coccineohirta (Blake)
Idaho, Nev., Calif., Oreg., Wash.

Mutilla (Sphaeropthalma) coccineohirta Blake, 1871. Amer. Ent. Soc, Trans. 3: 235. ♂.




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Mutilla ochracea Blake, 1879. Amer. Ent. Soc, Trans. 7: 247. ♂.
Sphaerophthalma (!) venifica Blake, 1886. Amer. Ent. Soc, Trans. 13: 210. ♀.
Mutilla progne Fox, 1899. Amer. Ent. Soc, Trans. 25: 247. ♀.
Dasymntilla aletina Cockerell, 1915. Entomologist 48: 249. ♂.

Columbiana Mickel. B. C. (Nicola).

Dasymufilla columbiana Mickel, 1928. U. S. Natl. Mus., Bui. 143: 119. ♂.

Dasymutilla connectens (Cameron)
Ariz.; Mexico (Chihuahua, Jalisco, Baja California).

Sphaerophtliaima (!) connectens Cameron, 1895. Biol. Cent.-Amer., Hym., v. 2, p. 362. ♀.

Secondary homonym in Mutilla.
Mutilla eggeri Dalla Torre, 1897. Cat. Hym., v. 8, p. 33. N. name.
Dasymutilla helva Mickel, 1928. U. S. Natl. Mus., Bui. 143: 259. ♀.

Taxonomy: Mickel, 1965. Ent. Soc. Wash., Proc 67: 2 (synonymy).

Dasymutilla corcyra Mickel
Tex.

Dasymutilla corcyra Mickel, 1928. U. S. Natl. Mus., Bui. 143: 180. ♀.

Dasymutilla cotulla Mickel
Tex.

Dasyviutilla cotulla Mickel, 1928. U. S. Natl. Mus., Bui. 143: 75. ♂.

Dasymutilla creon (Blake)
Tex., Ark.

Mutilla {Sphaeropthalma) Creon Blake, 1872. Amer. Ent. Soc, Trans. 4: 73. ♂.

Dasymutilla creusa (Cresson)
Ark., Tex., Okla., Kans., Nebr., Colo., N. Mex.

Mutilla Creusa Cresson, 1865. Ent. Soc. Phila., Proc 4: 431. ♀.

Dasymutilla creusa
***authority mismatch
var. bellona (Cresson). Ark., Kans., Nebr., Wyo., Colo. Host: Bembix anioena Handl.
Possibly a synonym of bioculata (Cr.).

Mutilla Bellona Cresson, 1865. Ent. Soc. Phila., Proc. 4: 434. ♀.

Biology: Evans, 1966. Compar. ethology and evolution of sand wasps, p. 288 (host record).

Dasymutilla curialis Mickel
N. Mex. (Jemez Springs).

Dasymutilla curialis Mickel, 1928. U. S. Natl. Mus., Bui. 143: 274. ♂.

Dasymutilla curticeps Mickel
Kans., Tex.

Dasymutilla curticeps Mickel, 1928. U. S. Natl. Mus., Bui. 143: 173. ♀.

Dasymutilla cypris (Blake)
N. J., W. Va., Ga., Fla., Miss., Mo., Minn. Host: Bembix cinerea Handl.

Mutilla (Sphaeropthalma) Cypris Blake, 1871. Amer. Ent. Soc, Trans. 3: 246. ♀.

Dasymufilla mutata miamensis Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 462. ♀.

Dasymutilla hora Mickel, 1928. U. S. Natl. Mus., Bui. 143: 196. ♂.

Biology: Fattig, 1943. Emory Univ. Mus. Bui. 1: 4, 7 (host record, mating behavior).

Dasymutilla dammersi Mickel
Calif. (Palm Springs).

Dasymutilla damynersi Mickel, 1936. Pan-Pacific Ent. 12: 94. ♀.

Dasymutilla dawsoni Mickel
Nebr. (Halsey).

Dasymutilla dawsoni Mickel. 1936. Ent. Soc. Amer., Ann. 29: 40. ♀.

Dasymutilla digressa Mickel
N. Mex. (Pecos).

Dasymutilla digressa Mickel, 1928. U. S. Natl. Mus., Bui. 143: 273. ♂.

Dasymutilla dilucida Mickel
Ariz.

Dasymutilla dilucida Mickel, 1928. U. S. Natl. Mus., Bui. 143: 267. ♀.

dionysia Mickel. Ariz.

Dasymutilla dionysia Mickel, 1928. U. S. Natl. Mus., Bui. 143: 104. ♀.

Dasymutilla dorippa Mickel
Nev. (Ormsby Co.).

Dasymutilla dorippa Mickel, 1928. U. S. Natl. Mus., Bui. 143: 297. ♂.

Dasymutilla dugesii (Cockerell and Casad)
Kans., Colo., Tex., Ariz. Host: Antlwplwra occidentalis Cr.

Sphaerophthalma (!) dugesii Cockerell and Casad. 1894. Ent. News 5: 294. ♀.

Ephata (Ephuta) Dugesi (!) Andre, 1903. In Wytsman, Gen. Ins., fasc. 11, p. 59.

Biology: Krombein, 1967. U. S. Dept. Agr., Monog. 2, Sup. 2: 339 (host record).

Dasymutilla electra (Blake)
Tex., Okla.

Mutilla Electra Blake, 1872. Amer. Ent. Soc, Trans. 4: 75. ♀.
eminentia Mickel. Ariz., Calif.; Mexico (Baja California).

Dasymutilla eminentia Mickel, 1928. U. S. Natl. Mus., Bui. 143: 79. ♂.




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Dasymutilla errabunda Mickel
Ariz., Calif.

Dasymutilla errabunda Mickel, 1928. U. S. Natl. Mus., Bui. 143: 269. ♂.

Dasymutilla eurynome Mickel
Tex. (Valentine in Presidio Co.).

Dasymutilla eurynome Mickel, 1928. U. S. Natl. Mus., Bui. 143: 209. ♀.

Dasymutilla fasciventris Mickel
Ariz. (Globe).

Dasyiuutilla fasciveritris Mickel, 1938. Ent. See. Amer., Ann. 31: 155. ♂.

Dasymutilla ferruginea (Smith)
Ariz.; Mexico (Puebla, Oaxaca).

Mutilla fe)-ruginea Smith, 1879. Descr. New Species Hym. Brit. Mus., p. 226. ♀.
Dasymutilla chrysocoma Mickel, 1928. U. S. Natl. Mus., Bui. 143: 266. ♀.

Taxonomy: Mickel, 1965. Ent. Soc. Wash., Proc. 67: 3 (synonymy).

Dasymutilla flammifera Mickel
Idaho, Oreg., Calif., Ariz.

Dasymutilla flammifera Mickel, 1928. U. S. Natl. Mus., Bui. 143: 240. ♀.

Dasymutilla foxi (Cockerell)
N. Mex., Ariz. Host: Diadasia di7)iinuta (Cr.).

Sphaerophthalma (I) foxi Cockerell, 1894. Ent. News 5: 199. ♂.

Sphaerophthalma (!) heterochroa Cockerell and Casad, 1894. Ent. News 5: 298. ♀.

Sphaerophthalma (I) foxi var. arizonica Cockerell, 1900. Entomologist 33: 65. ♀.

Biology: Cockerell, 1896. Nature 54: 461.

Dasymutilla fulgida Mickel
Ariz. Possibly a synonym oi erythrina (Say) from Mexico.
Dasymutilla fulgida Mickel, 1928. U. S. Natl. Mus., Bui. 143: 76. ♀.

Taxonomy: Mickel, 1974. Ent. Soc. Amer., Ann. 67: 470. ♂.

Dasymutilla furina Mickel
Ariz. (Douglas).

Dasymutilla furiua Mickel, 1928. U. S. Natl. Mus., Bui. 143: 82. ♀.

Dasymutilla gentilis Mickel
Kans., Tex., Rocky Mts.

Dasymutilla gentilis Mickel, 1928. U. S. Natl. Mus., Bui. 143: 199. ♂.

Dasymutilla gibbosa (Say)
N. H. south to Ga., west to Minn, and Mo.

Mutilla (Ephuta) gibbosa Say, 1836. Boston Jour. Nat. Hist. 1: 298. ♂.

Mutilla cariniceps Fox, 1899. Amer. Ent. Soc, Trans. 25: 241. ♀.

Dasymutilla scrobinata Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 462. ♀.

Dasymutilla camiceps (!) Fattig, 1943. Emory Univ. Mus. Bui. 1: 9, 16. ♀.

Dasymutilla gloriosa (Saussure)
Tex., Utah, Nev., Ariz., Calif.; Mexico (Baja California). Host:
Microbembex nigrifrons (Prov.)

Mutilla gloriosa Saussure, 1868 (1867). Soc. Ent. France, Ann. (4) 7: 359. ♀.

Mutilla tecta Cresson, 1875. Amer. Ent. Soc, Trans. 5: 119. ♀.

Dasymutilla reperticia Mickel, 1928. U. S. Natl. Mus., Bui. 143: 287. ♂.

Taxonomy: Mickel, 1936. Ent. Soc. Amer., Ann. 29: 56 (synonymy).

Biology: Haddock, 1967. In Krombein, U. S. Dept. Agr., Monog. 2, Sup. 2: 339 (host record).

Dasymutilla glycera Mickel
Ariz. (Florence).

Dasyynutilla glycera Mickel, 1928. U. S. Natl. Mus., Bui. 143: 299. ♂.

Dasymutilla gorgon (Blake)
La., Okla., Tex., N. Mex.

Mutilla (Sphaeropthalma) Gorgon Blake, 1871. Amer. Ent. Soc, Trans. 3: 233. ♀.

Mutilla Tisiphone Blake, 1879. Amer. Ent. Soc, Trans. 7: 249. ♀.

Dasymutilla gorgons (!) Ashmead, 1899. N. Y. Ent. Soc, Jour. 7: 57, 60. ♀, ♂.

Dasymutilla hector (Blake)
Iowa, Nebr., Kans., Colo.

Mutilla (Sphaeropthalma) Hector Blake, 1871. Amer. Ent. Soc, Trans. 3: 237. ♂.

Dasymutilla heliophila (Cockerell)
Ariz., Calif.

Sphaerophthalma (!) heliophila Cockerell, 1900. Entomologist 33: 65. ♀.

Dasymutilla welltonensis Bradley and Bequaert, 1923. Amer. Mus. Novitates 82: 1. ♀.

Dasymutilla hersilia Mickel
Tex. (Bexar Co.).

Dasymutilla hersilia Mickel, 1936. Ent. Soc. Amer., Ann. 29: 51. ♂.

Dasymutilla hispidaria Mickel
Tex. (Hondo).

Dasymutilla hispidaria Mickel, 1936. Ent. Soc. Amer., Ann. 29: 55. ♀.

Dasymutilla homole Mickel
N. Mex. (Organ Mts.).

Dasymutilla homole Mickel, 1928. U. S. Natl. Mus., Bui. 143: 72. ♀.




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intermixta Mickel. N. Mex., Ariz.

DasyviufiUa intermixta Mickel, 1928. U. S. Natl. Mus., Bui. 143: 256. ♂.
interrupta Banks. Mass. south to Ga., west to N. Dak. and Colo.

Dasyniutilla cypris var. interrupta Banks, 1921. Ent. See. Amer., Ann. 14: 24. ♀.

Taxonomy: Krombein, 1953. Wasmann Jour. Biol. 10: 310. ♂.

Dasymutilla klugii (Gray)
Kans. south to Tex., west to Utah and Ariz. Host: Sphecius grandis (Say).

Mutilla Klugii Gray, 1872. In Griffith, Cuvier's Animal Kingdom 15 (Insecta 2), p. 516. ♀.
Mutilla OrcHs Cresson, 1865. Ent. Soc. Phila., Proc. 4: 428. ♀, ♂.

Biology: Bradley, 1920. Ent. News 31: 112-113 (host record).

Dasymutilla klugioides Mickel
Okla., Tex.

Dasyniutilla klugioides Mickel, 1936. Ent. Soc. Amer., Ann. 29: 54. ♀.

Dasymutilla leda (Blake)
S. Dak. south to Tex., Colo., N. Mex.

Mutilla (Sphaeropthalma) Leda Blake, 1872. Amer. Ent. Soc, Trans. 4: 72. ♀.

Mutilla prognoides Viereck, 1906. Amer. Ent. Soc, Trans. 32: 186. ♀.

Dasymutilla lepeletierii (Fox)
Mass. south to Fla., west to Iowa and Tex.

Mutilla fenestrata Lepeletier, 1845. Hist. Nat. Ins. Hym., v. 3, p. 627. ♂. Preocc.

Mutilla Lepeletierii Fox, 1899. Amer. Ent. Soc, Trans. 25: 244. N. name.

Mutilla vierecki Rohwer, 1910. Ent. Soc Wash., Proc 12: 49. ♀.

Dasymutilla ferrugata var. balabetei Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 456. ♀.

Dasymutilla georgiana Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 456. ♀.

Dasymutilla plesia Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 456. ♀.

Dasymutilla (Dasymutilla) ferrugata var. ballabetei (!) Bradley, 1916. Amer. Ent. Soc,
Trans. 42: 327. ♀.

Biology: Shappirio, 1948. Brooklyn Ent. Soc, Bui. 42: 163 (mating). jlj

Dasymutilla macilenta (Blake)
Ga., Fla., Tex. '''

Mutilla (Sphaeropthalma) macilenta Blake, 1871. Amer. Ent. Soc, Trans. 3: 239. ♂.

Dasymutilla macra (Cresson)
N. Dak. south to Ark., west to Alta. and N. Mex.

Mutilla macra Cresson, 1865. Ent. Soc. Phila., Proc. 4: 429. ♂.

Sphaerophthalma (!) hispida Blake, 1886. Amer. Ent. Soc, Trans. 13: 226. ♂.

Dasymutilla magna (Cresson)
Ariz., Calif.; Mexico (Baja California).

Mutilla magna Cresson, 1865. Ent. Soc Phila., Proc. 4: 385. ♀.
magnifica Mickel. N. Mex., Ariz., Nev., Calif.; Mexico (Baja California).

Dasymutilla magnifica Mickel, 1928. U. S. Natl. Mus., Bui. 143: 234. ♀, ♂.

Dasymutilla medea (Cresson)
Colo., N. Mex. Host: Microbembex monodonta (Say)?

Mutilla Medea Cresson, 1865. Ent. Soc Phila., Proc 4: 432. ♀.

Biology: Melander, 1903. Amer. Ent. Soc, Trans. 29: 297 (host ?).

Dasymutilla megalophthalma Mickel
Ariz., Calif. D. arenivaga Mick, may be the female.

Dasymutilla megalophthalma Mickel, 1928. U. S. Natl. Mus., Bui. 143: 282. ♂.

Dasymutilla melanippe Mickel
Tex.

Dasymutilla melanippe Mickel, 1928. U. S. Natl. Mus. Bui. 143: 152. ♀.
Dasymutilla lauta Mickel, 1928. U. S. Natl. Mus., Bui. 143: 154. ♀.

Taxonomy: Mickel, 1936. Ent. Soc. Amer., Ann. 29: 46 (synonymy).

Dasymutilla melanippe
***authority mismatch
var. conformis Mickel. Tex.

Dasymutilla melanippe var. confonnis Mickel, 1928. U. S. Natl. Mus., Bui. 143: 153. ♀.

Dasymutilla meracula Mickel
Kans., Okla., Tex.

Dasymutilla meracula Mickel, 1928. U. S. Natl. Mus., Bui. 143: 201. ♂.

monticola (Cresson). Minn, south to Kans., west to B. C. and Ariz.
Mutilla monticola Cresson, 1865. Ent. Soc Phila., Proc. 4: 430. ♂.
Mutilla eximia Blake, 1886. Amer. Ent. Soc, Trans. 13: 200. ♂.
Ephuta boulderensis Rohwer, 1909. Amer. Ent. Soc, Trans. 35: 133. ♂.

Dasymutilla montivagoides (Viereck)
Kans. (Hamilton Co.).

Mutilla (Timulla) montivagoides Viereck, 1906. Amer. Ent. Soc, Trans. 32: 185. ♀.




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Dasymutilla mutata (Blake)
N. Y. south to Fla., west to Mo. and Tex. Host: Mijzinum obsciinim (F.).
MutUla (Sphaeropthalma) ynufata Blake, 1871. Amer. Ent. Soc, Trans. 3: 247. ♀.
Dasymutilla allardi Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 463. ♀.
Dasymntilla ruhricosa Mickel, 1928. U. S. Natl. Mus., Bui. 143: 192. ♂.

Taxonomy: Mickel, 1938. Ent. Soc. Amer., Ann. 31: 152. ♀.

Biology: Fattig, 1943. Emory Univ. Mus. Bui. 1: 4, 7 (host record, mating).

Dasymutilla myrice Mickel
Nebr., Kans., Okla., Tex., Colo.

Dasymufilla myrke Mickel, 1928. U. S. Natl. Mus., Bui. 143: 121. ♂.

Dasymutilla nigricauda (Viereck)
Kans., Okla., Tex.

MutUla (Timulla) iiigricauda Viereck, 1906. Amer. Ent. Soc, Trans. 32: 187. ♀.

Dasymutilla nigripes (Fabricius)
Mass. to Fla., west to Alta. and Ariz. Host: Philanthus gibbosus (F.)?,
Cerceris f. flavofasciata Sm.

MutUla nigripes Fabricius, 1787. Mantissa Insectorum, v. 1, p. 313. ♀.

MutUla sparsa Fox, 1899. Amer. Ent. Soc, Trans. 25: 240. ♀.

Dasymutilla blawa Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 457. ♀.

Dasymutilla segregata Rohwer, 1912. U. S. Natl. Mus., Proc 41: 459. ♀.

Dasymutilla segregata finni Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 459. ♀.

Dasymutilla bruneri Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 460. ♀.

Dasymutilla champlaini Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 461. ♀.

Dasymutilla hirticula Mickel, 1923. Minn. State Ent. Rpt. 19: 107. ♂.

Taxonomy: Mickel, 1936. Ent. Soc. Amer., Ann. 29: 43-44 (synonymy).

Biology: Shappirio, 1948. Brooklyn Ent. Soc, Bui. 42: 163 (mating behavior). — Shappirio,
1949. Brooklyn Ent. Soc, Bui. 43: 158 (host record). — Krombein, 1959. Ent. Soc Wash.,
Proc. 61: 198 (host record). —Evans and Miller, 1969. Mich. Ent. 2: 74 (adult female
overwintering). —Evans, Miller and Bartlett, 1973. Kans. Ent. Soc, Jour. 46: 343-346 (host
searching range).

Dasymutilla nitidula Mickel
Kans., Okla., Tex.

Dasymutilla nUidula Mickel, 1928. U. S. Natl. Mus., Bui. 143: 174. ♀.

Dasymutilla nitidula
***authority mismatch
var. nigridia Mickel. Tex.

Dasymutilla nitidula var. nigridia Mickel, 1936. Ent. Soc. Amer., Ann. 29: 47. ♀.

Dasymutilla nocturna Mickel
Calif., Nev.

DasymutUla nocturna Mickel, 1928. U. S. Natl. Mus., Bui. 143: 279. ♀.

Dasymutilla nogalensis Mickel
Ariz.

DasymutUla nogalensis Mickel, 1928. U. S. Natl. Mus., Bui. 143: 258. ♀.

Dasymutilla nupera Mickel
Tex., N. Mex.

Dasymutilla nupera Mickel, 1928. U. S. Natl. Mus., Bui. 143: 103. ♀.

Dasymutilla occidentalis
***authority mismatch
comanche (Blake). Iowa and S. Dak., south to Tex., Colo., Ariz., west to Idaho
and Oreg.
MutUla (Sphaeropthahna) Comanche Blake, 1871. Amer. Ent. Soc, Trans. 3: 234. ♀, ♂.

Dasymutilla occidentalis
***authority mismatch
occidentalis (Linnaeus). Conn, south to Fla., west to Mo. and Tex. Host: Bombus
fraternus Smith.
MutUla occidentalis Linnaeus, 1758. Syst. Nat, ed. 10, v. 1, p. 582. ♀.
MutUla bifasciata Swederus, 1787. Svensk. Vetensk. Akad. Handl. 8: 285. ♀.
MutUla coccinea Fabricius, 1793. Ent. System. 2: 366. ♀.

Biology: Fattig, 1943. Emory Univ. Mus. Bui. 1: 5 (host).

Dasymutilla ocydrome Mickel
Ariz.

DasymutUla ocydrome Mickel, 1928. U. S. Natl. Mus., Bui. 143: 292. ♂.

Dasymutilla paenulata Mickel
Ariz., Calif.

DasymutUla paenulata Mickel, 1928. U. S. Natl. Mus., Bui. 143: 206. ♀.

Dasymutilla paranocturna Barr and Hurd
Calif.

Dasymutilla parayiocturna Barr and Hurd, 1947. Pan-Pacific Ent. 23: 88. ♀.
parks! Mickel. Tex. (Bexar Co.).

Dasymutilla parksi Mickel, 1936. Ent. Soc. Amer., Ann. 29: 50. ♀.




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Dasymutilla perilla Mickel
Tex.

Dasymxtilla perilla Mickel, 1928. U. S. Natl. Mus., Bui. 143: 300. ♂.

Dasymutilla perilla
***authority mismatch
var. gentilicia Mickel. Kans., Colo.

Dasyviutilla perilla var. gentilicia Mickel, 1928. U. S. Natl. Mus., Bui. 143: 301. ♂.

Dasymutilla phaon (Fox)
Tex., N. Mex., Ariz., Utah.

Mutilla phaon Fox, 1899. Amer. Ent. Soc, Trans. 25: 243. ♂.

Dasymutilla phaon
***authority mismatch
var. fimbrialis Mickel. Utah, Ariz., Calif.

Dasymutilla phaon var. fimbrialis Mickel, 1928. U. S. Natl. Mus., Bui. 143: 302. ♂.

Dasymutilla phoenix (Fox)
Ariz., Calif. Host: Ptilothrix sumichrasti (Cr.).

Mutilla phoenix Fox, 1899. Amer. Ent. Soc, Trans. 25: 247. ♀.

Biology: Krombein, 1958. U. S. Dept. Agr., Monog. 2, Sup. 1: 104 (host record).

Dasymutilla polia Mickel
Ariz.

Dasyinutilla polia Mickel, 1928. U. S. Natl. Mus., Bui. 143: 215. ♂.

Dasymutilla praegrandis Mickel
Tex.

Dasymutilla praegrandis Mickel, 1928. U. S. Natl. Mus., Bui. 143: 148. ♀, ♂.

Dasymutilla praegrandis
***authority mismatch
var. russata Mickel. Tex.

Dasymutilla praegratidis var. russata Mickel, 1928. U. S. Natl. Mus., Bui. 143: 152. ♂.

Dasymutilla pseudopappus (Cockerell)
Colo., N. Mex., Ariz., Calif.

Sphaerophthalma (!) gloriosa var. pseudopappus Cockerell, 1895. Psyche 7(sup.): 6. ♀.

Dasymutilla pyrrhus (Fox)
Ga., Fla. Host: Sp. of Bembicini.

Mutilla pyrrhus Fox, 1899. Amer. Ent. Soc, Trans. 25: 243. ♂.

Biology: Krombein, 1964. Amer. Mus. Novitates 2201: 2-4, fig. 1 (host record).

Dasymutilla quadriguttata (Say)
111., Minn., and S. Dak. south to Miss, and Tex. Host: Myzinum
quinquecinctum (F.).
Mutilla quadriguttata Say, 1823. West. Quart. Rptr. 2: 74. ♀.
Dasymutilla permista Mickel, 1923. Minn. State Ent. Rpt. 19: 108. ♂.

Taxonomy: Remington, 1944. Ent. Soc. Amer., Ann. 37: 198-200 (synonymy).

Biology: Swezey, 1915. Hawaii Ent. Soc, Proc 3: 72 (host record; misdet. as castor Blake.)
—Hayes, 1924. Brooklyn Ent. Soc, Bui. 19: 153 (host).

Dasymutilla reclusa Mickel
S. Dak. south to Tex., Mont., Wyo.

Dasymutilla reclusa Mickel, 1928. U. S. Natl. Mus., Bui. 143: 219. ♂.

Dasymutilla rubicunda Bradley
Ga., Fla.

Dasymutilla [Dasymutilla) rubicunda Bradley, 1916. Amer. Ent. Soc, Trans. 42: 325. ♀.

Dasyynutilla beameri Mickel, 1936. Ent. Soc. Amer., Ann. 29: 48. ♀.

Dasymutilla sackenii (Cresson)
Oreg., Calif., Nev.; Mexico (Baja California). Host: Bembix occidentalis
beutenynulleri Fox.

Mutilla Sackenii Cresson, 1865. Ent. Soc. Phila., Proc 4: 385. ♀.

Mutilla erudita Cresson, 1875. Amer. Ent. Soc, Trans. 5: 120. ♀.

Biology: Bohart and MacSwain, 1939. South. Calif. Acad. Sci., Bui. 38: 89, pi. 16, figs. 11-14

(life history).

Dasymutilla saetigera Mickel
Ariz. (Baboquivari Mts.).

Dasymutilla saetigera Mickel, 1928. U. S. Natl. Mus., Bui. 143: 211. ♀.

Dasymutilla satanas Mickel
Ariz., Nev., Calif.; Mexico (Baja California).

Dasymutilla satanas Mickel, 1928. U. S. Natl. Mus., Bui. 143: 239. ♀.

Dasymutilla mimula Mickel, 1928. U. S. Natl. Mus., Bui. 143: 255. ♂.

Taxonomy: Barr and Hurd, 1947. Pan-Pacific Ent. 23: 86. ♀, ♂.

Dasymutilla scaevola (Blake)
Mass. south to Ga., west to Mont, and Ariz. Host: Cerceris clypeata Dahlb.?,
C.finitima Cr.?
Mutilla (Sphaeropthalma) obscura Blake, 1871. Amer. Ent. Soc, Trans. 3: 239. ♂. Preocc.
Mutilla (Sphaeropthalma) scaevola Blake, 1871. Amer. Ent. Soc, Trans. 3: 247. ♀.
Mutilla (Sphaeropthalma) admetus Blake, 1872. Amer. Ent. Soc, Trans. 4: 74. ♂.
Sphaerophthalma (!) macer (!) Blake, 1886. Amer. Ent. Soc, Trans. 13: 227. ♂.
Sphaerophthalma (!) macerata Blake, 1886. Amer. Ent. Soc, Trans. 13: 286. Emend.



_n




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Mntnia maceixi (!) Dalla Torre, 1897. Cat. Hym., v. 8, p. 56.

Mutilla {DasyniHtilla) apachea Viereck, 1908. Amer. Ent. Soc, Trans. 33: 386. ♂.

Biology: Shappirio, 1948. Brooklyn Ent. Soc, Bui. 42: 162-163 (host record). — Shappirio,

1949. Brooklyn Ent. Soc, Bui. 43: 157-158 (host records).

Dasymutilla scitula Mickel
Utah, Nev., Calif.

Dasymutilla scitula Mickel, 1928. U. S. Natl. Mus., Bui. 143: 244. ♀, ♂.

Dasymutilla serenitas Mickel
Te.x. (Cotulla, Eastland Co.).

Dasymutilla serenitas Mickel, 1928. U. S. Natl. Mus. , Bui. 143: 289. ♂.

Dasymutilla sicheliana (Saussure)
Ariz.; Mexico (Durango).

Mutilla Sicheliana Saussure, 1868 (1867). Soc. Ent. France, Ann. (4) 7: 360. ♀.

Sphaeroplitlial))>a (!) prunotincta Cockerell, 1895. Ent. News 6: 60. ♀.

Spliaerophtltalvia (!) thera Cameron, 1895. Biol. Cent.-Amer., Hym., v. 2, p. 358. ♀. Preocc.
in Mutilla.

Mutilla gynaecologica Dalla Torre, 1897. Cat. Hym., v. 8, p. 45. N. name.

Taxonomy: Mickel, 1965. Ent. Soc. Wash., Proc. 67: 3 (synonymy).

Dasymutilla snoworum (Cockerell)
Nebr. south to Tex., west to Mont, and Ariz.

Sphaeroplithalnia (!) snoworum Cockerell, 1897. In Cockerell and Fox, Acad. Nat. Sci.
Phila., Proc 49: 135. ♂.

Mutilla poecilonota Melander, 1903. Amer. Ent. Soc, Trans. 29: 301. ♀.

Dasymutilla sophrona Mickel
Ariz.

Dasymutilla sophrona Mickel, 1928. U. S. Natl. Mus., Bui. 143: 271. ♂.

Dasymutilla stevensi Mickel
Iowa, N. Dak., south to Tex., Colo., Utah, N. Mex., Ariz.

Dasymutilla stevensi Mickel, 1928. U. S. Natl. Mus., Bui. 143: 99. ♀.

Dasymutilla medora Mickel, 1928. U. S. Natl. Mus., Bui. 143: 101. ♂.

Taxonomy: Mickel, 1936. Ent. Soc. Amer., Ann. 29: 44 (synonymy).

Dasymutilla subhyalina Mickel
Calif. (Andrade).

Dasymutilla subhyalina Mickel, 1928. U. S. Natl. Mus., Bui. 143: 281. ♂.

Dasymutilla sulcatulla Mickel
Tex. (Rio Grande in Brewster Co.).

Dasymutilla sulcatulla Mickel, 1928. U. S. Natl. Mus., Bui. 143: 155. ♀.

Dasymutilla testaceiventris (Fox)
Oreg., Calif.

Mutilla testaceiventris Fox, 1899. Amer. Ent. Soc, Trans. 25: 242. ♂.

Dasymutilla texanella Mickel
Kans., Tex., Colo.

Mutilla texana Blake, 1879. Amer. Ent. Soc, Trans. 7: 250. 9. Preocc

Dasymutilla texanella Mickel, 1928. U. S. Natl. Mus., Bui. 143: 96. 9. N. name.

Dasymutilla thetis (Blake)
Ariz.

Sphaerophthalma (!) thetis Blake, 1886. Amer. Ent. Soc, Trans. 13: 214. ♀.

Dasymutilla vandala Mickel
Tex.

Dasymutilla vandala Mickel, 1928. U. S. Natl. Mus., Bui. 143: 74. ♂.

Dasymutilla vesta
***authority mismatch
errans Rohwer. S. Dak. south to Tex., Colo.

Dasymutilla errans Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 457. ♀.

Dasymutilla bosquensis Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 457. ♀.

Dasymutilla texensis Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 460. ♀.

Dasymutilla vesta
***authority mismatch
sappho (Fox). N. C. south to Fla., Miss. Host: Bembix cinerea Handl.

Mutilla sappho Fox, 1899. Amer. Ent. Soc, Trans. 25: 239. ♀.

Biology: Krombein, 1958. U. S. Dept. Agr., Monog. 2, Sup. 1: 104 (host).

Dasymutilla vesta
***authority mismatch
vesta (Cresson). East of Rocky Mts. in Canada and U. S. Host: Trypargilum politum

(Say); Nomia m. melanderi Ckll.
Mutilla Vesta Cresson, 1865. Ent. Soc Phila., Proc 4: 436. ♀.
Scolia unicincta Provancher, 1882. Nat. Canad. 13: 6. cJ (9 misdet.).
Mutilla monozona Dalla Torre, 1897. Cat. Hym., v. 8, p. 64. N. name for Scolia unicincta

Provancher, not Mutilla unicincta Lucas.
Mutilla agenor Fox, 1899. Amer. Ent. Soc, Trans. 25: 245. ♂.
Mutilla zella Rohwer, 1910. Ent. Soc. Wash., Proc. 12: 50. ♀.
Pycnomntilla harmoniiformis Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 455. ♂.




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Dasij)initilla ferrugatella Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 458. ♀.
Dasymntilla'coloradella Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 458. ♀.
Dasymutilla coloraclella virginica Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 459. ♀.
Dasymutilla coloroclella kaviloopseusis Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 459. ♀.
Dasymufilla wenillae Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 461. ♀.
Daxyinutilla Carolina Rohwer, 1912. U. S. Natl. Mus., Proc. 41 462. ♀.

Biology: Fattig, 1943. Emory Univ. Mus. Bui. 1: 4 (host record). — Krombein, 1958. U. S.
Dept. Agr., Monog. 2, Sup. 1: 104 (host record).

Dasymutilla vestita (Lepeletier)
N. Dak. south to Tex., west to Alta. and Calif., south in Me.xico to Oaxaca.

Host: Anthophora occidentalis (Cr.), Diadasia enavata (Cr.); Megachile periliirta Ckll.
Mutilla vestita Lepeletier, 1845. Hist. Nat. Ins. Hym., v. 3, p. 634. ♀, ♂.
Mutilla Montezumae Lepeletier, 1845. Hist. Nat. Ins. Hym., v. 3, p. 634. ♀.
Mutilla fulvohirta Cresson, 1865. Ent. Soc. Phila., Proc. 4: 433. ♂.
Sphaerophthalma (!) townsendi Cockerell, 1894. Ent. News 5: 199. ♂.
Sphaerophthalma (!) aspasia Cameron, 1895. Biol. Cent.-Amer., Hym., v. 2, p. 370. ♂.

Preocc. in Mutilla.
Mutilla anpasioides Dalla Torre, 1897. Cat. Hym., v. 8, p. 12. N. name.
Ephuta californica var. eucliwa Cockerell, 1897. Ann. and Mag. Nat. Hist. (6) 20: 513. ♀.

Taxonomy: Mann, 1915. Psyche 22: 178, fig. 1 (gynandromorph).

Biology: Mickel, 1928. U. S. Natl. Mus., Bui. 143: 72. —Krombein, 1958. U. S. Dept. Agr.,

Monog. 2, Sup. 1: 104 (host records).

Dasymutilla waco (Blake)
Tex.

Mutilla (Sphaeropthahna) Waco Blake, 1871. Amer. Ent. Soc, Trans. 3: 238. ♀.

Dasymutilla wileyae Mickel
Tex.

Dasymutilla wileyae Mickel, 1928. U. S. Natl. Mus., Bui. 143: 177. ♀.

Dasymutilla zelaya (Blake)
Okla., Tex., N. Mex., Ariz.

Mutilla (Spliaeroptltaima) Zelaya Blake, 1871. Amer. Ent. Soc, Trans. 3: 234. ♂.

Genus LOMACHAETA Mickel

Lomachaeta Mickel, 1936. Ent. Soc Amer., Ann. 29: 289.

Type-species: Lomachaeta liicksi Mickel. Orig. desig.

Revision: Mickel, 1940. Pan-Pacific Ent. 16: 127-131.

Lomachaeta coloradensis Mickel
Colo. (Boulder Co.).

Lomachaeta coloradensis Mickel, 1936. Ent. Soc. Amer., Ann. 29: 292. ♂.

Lomachaeta formosula Mickel
Ga. (Clarke Co.); Mexico (Coahuila).

Lo7nacbaeta fonuosula Mickel, 1940. Pan-Pacific Ent. 16: 130. ♂.

Lomachaeta hicksi Mickel
Calif.

Lomachaeta hicksi Mickel, 1936. Ent. Soc. Amer., Ann. 29: 289. ♀, ♂.

Lomachaeta minutula Mickel
Okla., Tex. (Type from Tenn., Mo. or Ark.).

Lomachaeta minutula Mickel, 1936. Ent. Soc. Amer., Ann. 29: 294. ♀.

Lomachaeta punctinota Mickel
111. (Carbondale).

Lomachaeta punctinota Mickel, 1936. Ent. Soc. Amer., Ann. 29: 293. ♂.

Lomachaeta variegata Mickel
Wash., Calif., Nev., Ariz., N. Mex. Host: Solierella plenoculoides similis
(Brid.), S. blaisdelli (Brid.), nesting in stems of Eriogonum elatum.

Lomachaeta variegata Mickel, 1940. Pan-Pacific Ent. 16: 128. ♀, ♂.

Biology: Parker, 1962. Pan-Pacific Ent. 38: 116 (host records). —Parker and Bohart, 1966.
Pan-Pacific Ent. 42: 95 (host records).

Genus SMICROMUTILLA Mickel

Sviicrovnifilla Mickel, 1964. Pan-Pacific Ent. 40: 108.

Type-species: Smicromutilla powelli Mickel. Orig. desig.

Smicromutilla powelli Mickel
Calif. (San Luis Obispo Co.). Host: Diodontus occidentalis Fox?
Smicromutilla powelli Mickel, 1964. Pan-Pacific Ent. 40: 108, 1 fig. c5, 2.




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Unplaced Taxa of Mutillidae

Mutilla contractu Say, 1836. Boston Jour. Nat. Hist. 1: 295. S. Type destroyed.

Mutilla exulans Fabricius, 1775. Systema Ent., p. 397. 9. Described from Drury coll.; type

probably lost.
Mutilla versicolor Fabricius, 1775. Systema Ent., p. 398. 9. Described from Drury coll.; type

probably lost.
Mutilla vigilans Say, 1836. Boston Jour. Nat. Hist. 1: 296. S. Type destroyed.


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