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Superfamily TENTHREDINOIDEA
Family TENTHREDINIDAE

This is the largest sawfly family with over 5,000 world species. Representatives of the
Tenthredinidae are found in all regions of the world, but it is a dominant family only in the
Northern Hemisphere where their predominance increases northwards. Most members of this
family are cylindrical, waspHke insects ranging in size from 2.5 mm. to about 15 mm. in length.
Most have nine-segmented antennae though some have fewer and some more. Habits and host
plants are diverse. Most larvae feed externally on the foliage of the host plant, but some are
leafminers, gallformers, or shootborers. Most larvae are caterpillarlike.

The division of the family into subfamilies has always been controversial, and there are nearly
as many different arrangements as authors. There are current differences between North
American and European workers. Eight subfamilies are utilized in this catalog, most of the units
being the same as those previously recognized in the North American literature. Most are readi-
ly separated on the basis of wing venational differences.

For general references to this family, see those listed under Suborder Symphyta.

Subfamily SELANDRIINAE

Members of this subfamily are associated with ferns, grasses, and sedges. The Selandriinae
are poorly represented in the northern regions of the world, but it is one of the dominant groups
in the tropics; for example, nearly 200 species have been described from the Americas south of
the United States.

Revision: Smith, 1969. U. S. Dept. Agr. Tech. Bui. 1398, 48 pp. (Nearctic species).

Taxonomy: Malaise, 1963. Ent. Tidskr. 84: 159-215 (key to world genera).

Tribe STRONGYLOGASTRINI

Genus ERIOCAMPIDEA Ashmead

Eriocampidea Ashmead, 1898. Canad. Ent. 30: 256.

Type-species: Eriocampidea arizonensis Ashmead. Orig. desig.
Cockerellonis MacGillivray, 1908. Canad. Ent. 40: 365.

Type-species: Cockerellonis occidentalis MacGillivray. Orig. desig.

A small genus found from southwestern United States into Mexico.

Eriocampidea arizonensis Ashmead
N. Mex., Ariz.; Mexico.

Eriocampidea arizonensis Ashmead, 1898. Canad. Ent. 30: 256. ♀.
Cockerellonis occidentalis MacGillivray, 1908. Canad. Ent. 40: 365. 9 (?).

Genus HEMITAXONUS Ashmead

Hemitaxonus Ashmead, 1898. Canad. Ent. 30: 311.

Type-species: Taxonus dubitatus Norton. Orig. desig.
Epitaxonus MacGillivray, 1908. Canad. Ent. 40: 365.

Type-species: Taxonus albidopictits Norton. Orig. desig.
Saklbergia Forsius, 1910. Meddel. Soc. Fauna et Flora Fenn. 36: 49.

Type-species: Saklbergia struthiopteridis Forsius. Monotypic.

Larvae of this holarctic genus feed on ferns. Mature larvae may bore into soft wood or other
substances where they pupate or overwinter.

Revision: Smith, 1966. Ent. Soc. Wash., Proc. 68: 113-120.

Hemitaxonus albidopictus (Norton)
N. S., N. B., Que. to Ala. w. to Minn., Iowa, Kans. Host: Onoclea
sensibilis.
Taxonus albido-pictus Norton, 1868. Amer. Ent. Soc, Trans. 2: 213. ♀, ♂.
Taxonus amicus Norton, 1868. Amer. Ent. Soc, Trans. 2: 213. ♀, ♂.
Hemitaxonus rnfopectus Rohwer, 1910. U. S. Natl. Mus., Proc. 41: 204. ♀.

Hemitaxonus dubitatus (Norton)
N. S., N. B., Que. to N. C. w. to Minn., Ill, Tex. Host: Onoclea sensibilis.
Taxonus dubitatus Norton, 1862. Boston Soc. Nat. Hist, Proc. 9: 119. ♀, ♂.




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Biology: Dyar, 1897. N. Y. Ent. Soc, Jour. 5: 20. — Krombein, 1960. Ent. News 71: 30 (in
sumach pith).

Hemitaxonus multicinctus Hall
Maine, Md., Ont., Ohio. Host: Athyrium thelypteroides.

Hemitaxonus viidticinchis Hall, 1917. Ent. Soc. Wash., Proc. 19: 28. Egg, larva.

Biology: Hall, 1917. Ent. Soc. Wash., Proc. 19: 28.

Hemitaxonus primarius Smith
B. C, Oreg., Calif.

Hemitaxonus primarius Smith, 1966. Ent. Soc. Wash., Proc. 68: 119. ♀, ♂.

Genus STRONGYLOGASTER Dahlbom

Tenthredo subg. Strongylogaster Dahlbom, 1835. Conspectus Tenthred., Siricid., Oryssin.
Scand., p. 13.

Type-species: Tenthredo cingulata Fabricius. Desig. by MacGillivray, 1908.
Tkrinax Konow, 1885. Wien. Ent. Ztg. 4: 19.

Type-species: Thrinax contigua Konow. Desig. by MacGillivray, 1908.
Pseudotaxonus Costa, 1894. Prosp. Imen. Ital., p. 157.

Type-species: Tenthredo filicis Klug. Monotypic.
Poiystichophagiis Ashmead, 1898. Canad. Ent. 30: 310.

Type-species: Tenthredo filicis Klug. Orig. desig.
Prototaxonus Rohwer, 1910. Canad. Ent. 42: 49.

Type-species: Prototaxonus typicus Rohwer. Monotypic.

Larvae of this genus feed on the underside of fronds of ferns. Mature larvae may bore into
soft wood, bark of trees, or other substances for overwintering and pupation. Adults are found
around their host plants in the spring months. The genus is holarctic with about 25 world spe-
cies. Some authors consider Thrinax and Pseudotaxonus as distinct genera.

Strongylogaster distans Norton
Colo., Utah w. to B. C, Wash., Oreg., Calif. Host: Pteridium aquilinum,.
Strongylogaster distayis Norton, 1868. Amer. Ent. Soc, Trans. 2: 220. ♀.
Strongylogaster pacificus MacGillivray, 1893. Canad. Ent. 25: 241. ♀, ♂.

Biology: Beer, 1955. Kans. Univ. Sci. Bui. 37: 223-249.

Strongylogaster impressata Provancher
N. S., N. B., Que. to N. C. w. to Minn., Colo., N. Mex. Host: Pteridium
aquilinumin-
Strongylogaster impressata Provancher, 1878. Nat. Canad. 10: 170. ♀.
Strongylogaster luctuosus Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 12. ♀.
Taxonus rufipes Harrington, 1889. Canad. Ent. 21: 97. ♂.
Thrinax pullatus MacGillivray, 1921. Psyche 28: 34. ♂.

Strongylogaster macula (Klug)
Ont., B. C; Europe to Japan. Host: Athyrium and Pteridium. These hosts
were recorded in Europe.
Tenthredo (Allantus) ynacula Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 217. ♀.
Thrinax intermedia Konow, 1885. Wien. Ent. Ztg. 4: 22. ♀.
Thrinax contigua Konow, 1885. Wien. Ent. Ztg. 4: 22. ♀, ♂.

Strongylogaster multicincta Norton
N. S., N. B., Que. to Fla. w. to Minn., 111. Host: Pteridium aquilinum.
Strongylogaster ynulticinctus Norton, 1862. Ent. Soc. Phila., Proc. 1: 143. ♀.
Strongylogaster annulosus Norton, 1868. Amer. Ent. Soc, Trans. 2: 211. ♀.
Strongylogaster longulus Norton, 1868. Amer. Ent. Soc, Trans. 2: 216. ♂.
Strongylogaster alboannulatus Rohwer, 1912. U. S. Natl. Mus., Proc 43: 237. ♀.
Strongylogaster annulosus var. cingulescens Ross, 1932. Canad. Ent. 64: 250. ♀, ♂.

Biology: Hogh, 1966. Kans. Ent. Soc, Jour. 39: 347-354.

Strongylogaster polita Cresson
Conn., Ont, N. Y.

Strongylogaster politus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 20. ♀.

Strongylogaster politus Provancher, 1882. Nat. Canad. 13: 296. ♀. Preocc.

Strongylogaster remota Rohwer
Que., Pa.

Strongylogaster remotus Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 237. ♀.

Strongylogaster rufigastra (Kincaid)
Alaska, B. C.

Paraselandria rufigastra Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 346. ♀.




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Strongylogaster soriculatipes Cresson
N. B., Que. to Md. w. to Minn., 111. Host: Pteridium aquilinum.
Stwngylogaster soriculatipes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 20. ♀.
Strongylogaster soriculatus Provancher, 1882. Nat. Canad. 13: 296. ♀.

Biology: Dyar, 1895. Amer. Ent. Soc, Trans. 22: 310.

Strongylogaster tacita (Norton)
N. B., Que. to Fla. w. to Wis., 111. Ecology: Adults collected from Pteridium
aquilinum.
Tenthredo (Allantus) tacita Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Allantus tacitus Norton, 1860. Boston Jour. Nat. Hist. 7: 246. ♀, ♂.
Strongylogaster unicus Norton, 1862. Boston Soc. Nat. Hist., Proc. 9: 120. ♀.

Strongylogaster tibialis Cresson
Idaho, B. C, Wash., Oreg., Calif. Host: Pteridiuyn aquilininn.
Strongylogaster tibialis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 19. ♀.
Selandria marginata Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 8. ♀.
Strongylogaster primativus MacGillivray, 1893. Canad. Ent. 25: 241. ♀.
Prototaxomis typicus Rohwer, 1910. Canad. Ent. 42: 50. ♀, ♂.

Biology: Beer, 1955. Kans. Univ. Sci. Bui. 37: 223-249.

Strongylogaster tuberculiceps Rohwer
Fla.; Bermuda(?), specimen may be mislabeled.

Strongylogaster tuberculiceps Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 396. ♀.
Strongylogaster melanogaster Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 397. ♂.

Tribe ANEUGMENINI

Genus ANEUGMENUS Hartig

Empbytus subg. Aneugmenus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst
Einleitung Naturgesch. Hym., p. 253.

Type-species: Tenthredo (Emphytus) coronata Klug. Monotypic
Colposelandria Enslin, 1912. Tijdschr. v. Ent. 55: 110.

Type-species: Colposelandria jacobsoni Enslin. Orig. desig.
Selandria subg. Atoposelandria Enslin, 1913. Deut. Ent. Ztschr., Beih., p. 197.

Type-species: Selandria furstenberge^isis Konow. Orig. desig.
Polyselandria MacGillivray, 1914. Canad. Ent. 46: 104.

Type-species: Selayidria floridana MacGillivray. Orig. desig.

Larvae of this holarctic genus feed on ferns. In North America, the genus extends into Mex-
ico.

Revision: Ross, 1930. Canad. Ent. 62: 184-189.

Aneugmenus flavipes (Norton)
Newfoundland to Fla. w. to Man., Minn., Colo. Host: Pteridium aquilinum.
Selandria flavipes Norton, 1862. Boston Soc. Nat. Hist, Proc. 7: 222. ♀, ♂.
Aneugmenus flavitarsis Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 390. ♀.
Aneugmenus flavipes occidentalis Rohwer, 1911. U. S. Natl. Mus., Proc 41: 390. ♀.

Aneugmenus floridella Ross
Ga., Fla.

Selandria floridana MacGillivray, 1895. Canad. Ent. 27: 281. ♂. Preocc. by Cresson, 1880.

Selandria (Aneugmenus) flavipes vslt. floridella Ross, 1930. Canad. Ent. 62: 189. N. name
for floridana MacGillivray.

Aneugmenus padi (Linnaeus)
B. C, Wash., Oreg.; Europe to Siberia, N. Africa, Asia Minor. Probably
introduced. Host: Pteridium aquilinum.

Tenthredo padi Linnaeus, 1761. Fauna Suecica, Ed. 2, p. 390.

Tenthredo (Allantus) stramiyieipes Klug, 1818. Mag. Gesell. Naturf. Freunde Berlin 8: 75. ♀, ♂.

Tenthredo (Eiuphytus) coronata Klug, 1818. Mag. Gesell. Naturf. Freunde Berlin 8: 276. ♀.

Tenthredo albipes Lepeletier, 1823. Monog. Tenthred., p. 105.

Selandria rufitarsis Brulle, 1832. Exped. Moree, Zool., v. 2, p. 394. ♀.

Selandria analis Thomson, 1871. Hym. Scand., v. 1, p. 239. ♀.

Selandria cereipes Vollenhoven, 1873. Tijdschr. v. Ent. 16: 13.

Selandria vollenhoveni Gribodo, 1880. Soc. Ent. Ital., Bui. 12: 7.

Selandria bimaculata Cobelli, 1892. Verh. Ges. Wien. 42: 71. ♂.




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Selandria oglohleni Dovnar-Zapolskii, 1930. Rev. Russe d'Ent. 24: 90. ♀.
Selandria urbis Ross, 1930. Canad. Ent. 62: 186. ♀.

Taxonomy: Benson, 1968. Brit. Mus. (Nat. Hist.) Ent., Bui. 22: 134.

Aneugmenus scutellatus Smith
Ariz.; Mexico. Ecology: Adults collected from ferns.

Aneugmenus scutellatus Smith, 1969. U. S. Dept. Agr. Tech. Bui. 1398, p. 28. ♀, ♂.

Genus BIRKA Malaise

Birka Malaise, 1944. Arkiv for Zool. 35A: 4.

Type-species: Tenthredo (,Alla7itus) cinereipes Klug. Orig. desig.

European species of this genus feed on grasses and sedges.

Birka nordica Smith
Man., Alta., Yukon, Alaska, B. C.

Birka nordica Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1398, p. 29. ♀, ♂.

Genus NESOSELANDRIA Rohwer

Nesoselayidria Rohwer, 1910. U. S. Natl. Mus., Proc. 37: 657.

Type-species: Parasetandria iinitatrix Ashmead. Orig. desig.
Neobusarbia Takeuchi, 1928. Formosa Nat. Hist. Soc, Trans. 18: 40.

Type-species: Neobtisarbia flavipes Takeuchi. Orig. desig.
Melisandra Benson, 1939. Ent. Monthly Mag. 75: 110.

Type-species: Selandria morio (Fabricius) Enslin. Orig. desig.

Most species of this genus are found in Asia.

Nesoselandria morio (Fabricius)
Que., N. H., N. Y., Ont., B. C; Europe to Siberia, Japan.
Tenthredo morio Fabricius, 1781. Species Insectorum, v. 1, p. 414.
Tenthredo ulmi Schrank, 1802. Fauna Boica, p. 237.

Tenthredo {Selandria) tristis Lepeletier, 1823. Monog. Tenthred., p. 106. ♀, ♂.
Emphytus infiiscatus Eversmann, 1847. Soc. Imp. Nat. Moscou, Bui. 20: 29. ♀.
Selandria Fabricii Konow, 1885. Wien. Ent. Ztg. 4: 300.
Aneugmenus brunneus Magretti, 1886. Soc. Ent. Ital., Bol. 18: 25.

Genus EUSTROMBOCEROS Rohwer

Stromboceros subg. Eustromboceros Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 394.

Type-species: Stromboceros (Eustromboceros) melanoptenis Rohwer. Orig. desig.
Stromboceros subg. Prostromboceros Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 235.

Type-species: Stromboceros (Eustromboceros) melanoptenis Rohwer. Orig. desig.

A small genus found from Arizona to Central America.
Taxonomy: Smith, 1971. Amer. Ent. Soc, Trans. 97: 532-533.

Eustromboceros rufocaudatus Smith
S. Ariz.

Eustromboceros rufocaudatus Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1398, p. 30. ♂.

Genus STROMBOCERIDEA Rohwer

Stromboceros subg. Stromboceridea Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 392.

Type-species: Stromboceros (Strain boceridea) pilosuhis Rohwer. Orig. desig.
Petersiana Jorgensen, 1913. Buenos Aires Mus. Nac de Hist. Nat., An. 24: 278.

Type-species: Petersinianad) niveana Jorgensen. Monotypic.
Goniocenis Malaise, 1942. Ent. Tidskr. 63: 93 N. syn.

Type-species: Stromboceros albilabris Konow. Orig. desig.

This is a rather large genus found from southern Arizona to northern Argentina.

Stromboceridea nigricans (Norton)
S. Ariz, to Guatemala.

Strongylogaster nigricans Norton, 1868. Amer. Ent. Soc, Trans. 2: 223. ♀, ♂.

Selandria curialis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 15. ♀.

Stromboceros (Eustromboceros) leucostomus Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 395. ♀.




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Taxonomy: Smith, 1971. Amer. Ent. Soc, Trans. 97: 526 (lectotype for nigricans).

Tribe SELANDRIINI
Genus BRACHYTHOPS Haliday

Coryna Lepeletier and Serville, 1825. Encycl. Meth., Ins., v. 10, p. 567. Preocc. by Bosc,
1802.

Type-species: Tenthredo scapularis Lepeletier and Serville. Desig. by Rohwer, 1911.
Brackythops Haliday, 1839. In Curtis, Brit. Ent., v. 16, p. 764.

Type-species: Brachythops seminigra Haliday. Monotypic.
Paraselandria Ashmead, 1898. Canad. Ent. 30: 255.

Type-species: Selandria flavens Klug. Orig. desig.
Selajidridea Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 388.

Type-species: Selayidridea I'anduzeei Rohwer. Orig. desig.
Pseudoselandria MacGillivray, 1914. Canad. Ent. 46: 103.

Type-species: Pseudoselandria oxalata MacGillivray Orig. desig.

A small holarctic genus closely related to the Palaearctic Selandria Leach. Larvae of this
Genus feed on sedges.

flavens (Klug). Labrador to Alaska s. to N. Y., Wis., Colo.; Europe to Siberia. Host: Carex.

Tenthredo (Allantus) flavens Klug, 1818. Mag. Gesell. Naturf. Freunde Berlin 8: 48. ♀, ♂.

Tenthredo (Coryna) scapularis Lepeletier and Serville, 1825. Encycl. Meth. Ins., v. 10, p.
567. ♀.

Phyllotoma puella Fallen, 1829. Monog. Tenthred. Suec, p. 30. ♀, ♂.

Brachythops seminigra Haliday, 1839. In Curtis, Brit. Ent., v. 16, p. 764.

Selandria flavescens Thomson, 1870. Opusc. Ent, 2nd Fasc, p. 291.

Selandria decolorata Cresson, 1880. Amer. Ent. Soc, Trans. 8: 15. ♂.

Selandridea vandiizeei Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 388. ♀, ♂.

Selandria flavens \ar.flavior Enslin, 1913. Deut. Ent. Ztschr., Beih., p. 144.

Pseudoselandria oxalata MacGillivray, 1914. Canad. Ent. 46: 103. ♀.
wuestneii (Konow). Que., N. W. T., Man., Alta., Yukon, Alaska; Europe. Host: Carex.

Selandria icustneii Konow, 1885. Wien. Ent. Ztg. 4: 122. ♀, ♂.

Tribe ADELESTINI
Genus ADELESTA Ross

Adelesta Ross, 1937. 111. Biol. Monog. 15(2): 68.

Type-species: Selandria nova Norton. Orig. desig.

The single species in this genus is found in North America only.

Adelesta nova (Norton)
N. S., N. B., Que., Maine.

Selandria nova Norton, 1867. Amer. Ent. Soc, Trans. 1: 254. ♀.

Tribe HEPTAMELINI

Genus HEPTAMELUS Haliday

Melicerta Stephens, 1835. Illus. Brit. Ent., v. 7, p. 94. Preocc. by Schrank, 1803.

Type-species: Melicerta ochroleuca Stephens. Monotypic.
Heptamelus Haliday, 1855. Nat. Hist. Rev., v. 2, p. 60.

Type-species: Melicerta ochroleuca Stephens. Monotypic.
Caenoneura Thomson, 1870. Opusc Ent., 2nd Fasc, p. 270.

Type-species: Caenoneura dahlbomi Thomson. Monotypic.

Most species of this genus are found in Asia.

Heptamelus ochroleucus (Stephens)
B. C. (Squamish, Diamond Head Trail); Europe, Japan. Ecology:

Larva bores downward in stems of ferns. Host: Athyrium, Polypodium, and Blechniim.
These hosts were recorded in Europe.
Melicerta ochroleuca Stephens, 1835. Illus. Brit. Ent., v. 7, p. 95.




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Caenoneura dahlhomi Thomson, 1870. Opusc. Ent., 2nd Fasc, p. 271. ♂.

Subfamily DOLERINAE

The two genera in this subfamily are both widespread throughout the subarctic and tem-
perate regions of the Northern Hemisphere; one species is known from Uganda. Dolerus con-
tains about 125 and Lodenis about 16 world species. Larvae feed on horsetails, grasses, sedges,
and rushes. A few species may be of importance when feeding in cultivated crops. Adults fly
early in the spring and are found in their host-plant communities, on nearby flowers such as
those of Crataegus and Prunus, or on willow catkins. Some may feed on the sap of sugar maple.
There is normally a single generation a year, and mature larvae overwinter in an earthen cell in
the ground. Species of this subfamily can be collected in large numbers. The time of emergence
of the adults can be accurately predicted, yet, very little biological work has been done. Larvae
for so few species have been associated that it is impossible to identify them beyond genus.

Revision: Ross, 1931. 111. Biol. Monog. 12(3): 1-116.

Taxonomy: Ross, 1935. Ent. Soc. Wash., Proc. 37: 88-93. —Benson, 1956. Roy. Ent. Soc.
London, Proc, Ser. B: Taxonomy 25: 55-63. —Benson, 1962. Brit. Mus. (Nat. Hist.) Ent.,
Bui. 12: 388-389 (holarctic species).

Genus DOLERUS Panzer

Dolerus Jurine, 1801 (May). Intell. Blatt. Litt.-Ztg. Erlangen, v. 1, p. 163. Suppressed by
Internatl. Comm. Zool. Nomencl. Op. 135, 1939.

Type-species: Tenthredo gonager Fabricius. Desig. by Latreille, 1810.
Dolerus Panzer, 1801 (September). Faunae Ins. German., heft 82, pi. 11.

Type-species: Dolenis pedestris Panzer. Monotypic.
Dosytheus Leach, 1817. Zool. Misc., v. 3, p. 127.

Type-species: Tenthredo eglanteriae Fabricius. Desig. by Brulle, 1846.

Species Group Unicolor

Dolerus asper Zaddach
Mass., Conn., Mich., Ind., S. Dak., Alaska; Europe, Siberia. Host: Cyperaceae
and Gramineae.

Dolerus asper Zaddach, 1859. Programm Friedr. Col. Konigsberg, p. 21. ♂.

Dolerus tectus MacGillivray, 1914. Canad. Ent. 66: 104. ♀, ♂.

Dolerus asper 7negapteroides Muche, 1964. Reichenbachia 4: 31-33. 9 (male misdet.).

Dolerus borealis MacGillivray
Idaho, Wash., Oreg.

Dolenis borealis MacGillivray, 1893. Canad. Ent. 25: 238. ♀.

Dolerus nominatus MacGillivray, 1923. Insecutor Inscitiae Menstruus 11: 34. ♀.

Dolerus illini Ross
Mass., N. Y. w. to Alta., S. Dak., Colo. Ecology: Adults collected in Equisetum,
Eleocharis, and Carex habitats.

Dolerus illini Ross, 1931. 111. Biol. Monog. 12(3): 39. ♀, ♂.

Dolerus illini var. rufilobus Ross, 1931. 111. Biol. Monog. 12(3): 41. ♂.

Dolerus nativus
***authority mismatch
MacGillivray. Colo., Oreg.

Dolerus nativus MacGillivray, 1923. Insecutor Inscitiae Menstruus 11: 32. ♂.

Dolerus nectareus MacGillivray, 1923. Insecutor Inscitiae Menstruus 11: 33. ♂.
neocoilaris narratus MacGillivray. B. C, Idaho, Wash., Oreg.

Dolerus narratus MacGillivray, 1923. Canad. Ent. 55: 65. ♀, ♂.

Dolerus nocuus MacGillivray, 1923. Insecutor Inscitiae Menstruus 11: 34. ♀.
neocoilaris neocoilaris MacGillivray. Que., N. Y. w. to Mont., Nebr., Colo.

Dolerus neocoilaris MacGilHvray, 1908. Canad. Ent. 40: 127. ♀, ♂.

Dolerus refugus MacGillivray, 1908. Canad. Ent. 40: 127. ♀.

Dolerus nitens Zaddach
N. H. to Md., w. to Mich., Ohio, III., Wash., Oreg.; Europe. Ecology: Probably
grasses and sedges are hosts; adults sometimes swept from cultivated grass crops. Host:
Fesiuca sp.

Dolerus coracimis Stephens, 1835. Illus. Brit. Ent., v. 7, p. 87. Preocc. by Klug, 1814.

Dolerus nitens Zaddach, 1859. Programm Friedr. Col. Konigsberg, p. 16. ♀, ♂.

Dolerus anthracinus Thomson, 1871. Hym. Scand., v. 1, p. 291. Preocc. by Klug, 1814.

Dolerus varispinus Konow, 1884. Deut. Ent. Ztschr. 28: 344. Preocc. by Hartig, 1837.




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Dolerus coruscans Konow, 1890. Wien. Ent. Ztg. 9: 10. N. name for anthracinus Thomson.
Dolems wanda Ross, 1935. Ent. Soc. Wash., Proc. 37: 91. ♀.

Biology: Kamm, 1975. Environ. Ent. 4: 312-314.

Dolerus unicolor (Beauvois)
Que. to Ga. w. to Man., S. Dak., Colo. Host: Phleum pratense.
Tenthredo unicolor Beauvois, 1805. Ins. Afr. Amer., p. 97. ♂.
Tenthredo thoracinus Beauvois, 1805. Ins. Afr. Amer., p. 97. ♀.

Dolerus arvensis Say, 1824. In Keating, Narr. Long's 2nd Exped., v. 2 (App.), p. 319. ♀.
Dolerus thoracicus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 230.

Biology: Riley and Marlatt, 1891. U. S. Dept. Agr., Insect Life 4: 168-179. —Forbes, 1885.
14th Rpt. State Ent. Ill, p. 100.

Species Group Sericeus

Dolerus kennedyi Ross
Ohio.

Dolerus kennedyi Ross, 1935. Ent. Soc. Wash., Proc. 37: 90. ♀.

Dolerus sericeus
***authority mismatch
centralis Ross. 111.

Dolerus sericeus centralis Ross, 1931. 111. Biol. Monog. 12(3): 46. ♀, ♂.

Dolerus sericeus
***authority mismatch
neosericeus MacGillivray. N. B. to N. Y. w. to Wis.

Dolerus neosericeus MacGillivray, 1908. Canad. Ent. 40: 125. ♀.

Dolerus necosericeus (!) MacGillivray, 1923. 111. Univ. Bui. 20(50): 13.

Dolerus sericeus
***authority mismatch
parasericeus MacGillivray. N. Y., B. C.

Dolerus parasericeus MacGillivray, 1908. Canad. Ent. 40: 125. ♀.

Dolerus polysericeus MacGillivray, 1908. Canad. Ent. 40: 125. ♀.

Dolerus sericeus
***authority mismatch
sericeus Say. Que. to Del. w. to B. C, Minn., Mo.

Dolerus sericeus Say, 1824. In Keating, Narr. Long's 2nd Exped., v. 2 (App.), p. 320. ♀.

Dolems colosericeiis MacGillivray, 1908. Canad. Ent. 40: 125. ♀.

Dolerus monosericeus MacGillivray, 1908. Canad. Ent. 40: 126. ♀, ♂.

Species Group Collaris

Dolerus abdominalis (Norton)
Mass. to Ga. w. to 111., Kans.

Dosytheus abdominalis Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 153. 9,8.

Dolerus agcistus MacGillivray
Maine to Va. w. to Man., Mont., Wyo., Colo., Calif. Ecology: Adults
collected in Eqnisetutn - Carex habitats.

Dolerus agcistus MacGillivray, 1908. Canad. Ent. 40: 129. ♀.

Dolerus agcistus var. maroa Ross, 1931. 111. Biol. Monog^ 12(3): 61. ♀. N. syn.

Dolerus bicolor (Beauvois)
Que. to Va. w. to Man., Wis., Nebr., Kans.

Tenthredo bicolor Beauvois, 1805. Ins. Afr. Amer., p. 96. ♀.

Dolerus lesticus MacGillivray, 1914. Canad. Ent. 46: 105. ♀. N. syn.

Dolerus graenicheri MacGillivray, 1914. Canad. Ent. 46: 107. ♀. N. syn.

Dolerus bicolor var. nigrita Ross, 1931. 111. Biol. Monog. 12(3): 59. ♀. N. syn.

Dolerus clypealis Ross
Mich., N. W. T., Alta., Mont., Colo., Utah, Wash., Oreg., Calif.

Dolerus clypealis Ross, 1931. 111. Biol. Monog. 12(3): 50. ♀.

Dolerus clypealis var. nigrilabris Ross, 1931. 111. Biol. Monog. 12(3): 51. ♀.

Dolerus collaris Say
Que. to Pa. w. to B. C, Mont., Colo. Host: Eleocharisi?).

Dolerus collaris Say, 1823. West. Quart. Rptr. 2: 720. ♀.

Dosytheus tnaculicollis Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 153. ♀.

Dolerus icterus MacGillivray, 1908. Canad. Ent. 40: 127. ♀.

Dolerus collaris var. erebus Ross, 1931. 111. Biol. Monog. 12(3): 55. ♀.

Dolerus distinctus Norton
Oreg., Calif.

Dolerus distinctus Norton, 1872. Amer. Ent. Soc, Trans. 4: 82. ♀.

Dolerus eurybis Ross
Que., Maine w. to Man., Minn., Colo.

Dolerus eurybis Ross, 1931. 111. Biol. Monog. 12(3): 65. ♀, ♂.

Dolerus idahoensis Ross
Idaho (Moscow).

Dolerus idahoensis Ross, 1931. 111. Biol. Monog. 12(3): 63. ♀.

Dolerus interjectus Ross
Idaho, Wash., Calif.

Dolerus interjectus Ross, 1931. 111. Biol. Monog. 12(3): 63. ♀, ♂.




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Dolerus moramus Ross
N. Y., 111., Minn. Colo.

Dolenis 7noramus Ross, 1931. 111. Biol. Monog. 12(3): 62. ♀, ♂.

Dolerus neoagcistus MacGillivray
Maine, Ont. to Va. w. to Nebr., Kans., Miss.

Dolenis neoagcistus MacGillivray, 1923. Brooklyn Ent. Soc, Bui. 18: 55. ♀.

Dolenis neoshigjius MacGillivray, 1923. Brooklyn Ent. Soc, Bui. 18: 55. ♀.

Dolerus nortoni Ross
Maine, Ont. to Md. w. to Mich., 111.

Dolenis 7iorto7ii Ross, 1931. 111. Biol. Monog. 12(3): 51. ♀, ♂.

Dolenis norioni var. nigritella Ross, 1931. 111. Biol. Monog. 12(3): 53. ♀.

Dolerus osagensis Ross
N. J., Okla.

Dolenis osagensis Ross, 1935. Ent. Soc. Wash., Proc. 37: 89. ♀, ♂.

Dolerus piercei Rohwer
N. J., Nebr., Kans.

Dolenis piercei Rohwer, 1909. Canad. Ent. 41: 10. ♀.

Dolerus salmani Ross
Calif.

Dolenis salmani Ross, 1935. Ent. Soc. Wash., Proc. 37: 88. ♀, ♂.

Dolerus versus Norton
N. Y. to Va. w. to Ohio, Mo., Tex.

Dolenis versa Norton, 1867. Amer. Ent. Soc, Trans. 1: 239. ♀, ♂.

Species Group Similis

Dolerus aprilis (Norton)
Que. to Va. w. to Sask., Colo. Host: Equisetum.

Dosythens aprilis Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 152. ♀.

Dolenis dysporus MacGillivray, 1908. Canad. Ent. 40: 128. ♀.

Dolenis nocivus MacGillivray, 1923. 111. Univ. Bui. 20(50): 12. ♀.

Dolerus apriloides MacGillivray
Que., Conn. w. to Man., Minn., 111.

Dolerus apriloides MacGillivray, 1908. Canad. Ent. 40: 126. ♀.

Dolerus coloradensis Cresson
Mont., Wyo., Colo., N. Mex., Ariz.

Dolenis colorade7isis Cresson. 1880. Amer. Ent. Soc, Trans. 8: 11. ♀, ♂.

Dolerus elderi Kincaid
Newfoundland to Alaska s. to N. J., 111., N. Mex., Utah, Calif.; Europe to
Siberia. Ecology: Widespread. Host: Equisetum.

Dolenis pusillus Jakovlev, 1891. Soc. Ent. Rossica, Horae 26: 32. 9. Preocc by Lepeletier,
1823.

Dolenis elderi Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 259. ♀, ♂.

Dolerus cohaesiis MacGillivray, 1908. Canad. Ent. 40: 128. ♀.

Dolenis nutricius MacGillivray, 1923. Canad. Ent. 55: 159. ♂.

Loderus acricuhis MacGillivray, 1923. 111. Univ. Bui. 20(50): 20. ♀.

Dolenis nyctelius MacGillivray, 1923. N. Y. Ent. Soc, Jour. 30: 163. ♂.

Dolerus nemorosus MacGillivray, 1923. N. Y. Ent. Soc, Jour. 30: 164. ♀.

Dolenis pullulus Zhelochovtsev, 1928. Zool. Anz. 79(3-4): 107. ♀, ♂.

Dolenis elderi var. auraneus Ross, 1931. 111. Biol. Monog. 12(3): 69. ♀, ♂.

Dolenis elderi var. nibicanus Ross, 1931. 111. Biol. Monog. 12(3): 70. ♀, ♂.

Dolenis elderi var. melamis Ross, 1931. 111. Biol. Monog. 12(3): 70. ♀, ♂.

Taxonomy: Benson, 1956. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 25: 55.

Dolerus nasutus
***authority mismatch
MacGillivray. Alta., Mont., Colo., N. Mex., B. C, Idaho, Wash., Oreg., Calif.

Dolenis nasutus MacGillivray, 1923. Canad. Ent. 55: 65. ♀.

Dolenis nugatorius MacGillivray, 1923. Canad. Ent. 55: 66. ♀, ♂.

Dolenis novellus MacGillivray, 1923. Canad. Ent. 55: 67. ♀.

Dolerus similis
***authority mismatch
nescius MacGillivray. B. C, Wash., Nev.

Dolenis nescius MacGillivray, 1923. 111. Univ. Bui. 20(50): 12. ♀.

Dolerus similis
***authority mismatch
similis (Norton). Newfoundland to Alaska s. to Va., Ky., Tex., N. Mex., Calif. Ecology:
Widespread. Host: Equisetum.

Dosytheus similis Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 152. ♀.

Dolenis tibialis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 52. ♂.

Dolenis conjugatus MacGillivray, 1908. Canad. Ent. 40: 128. ♀.

Dolenis plesiiis MacGillivray, 1908. Canad. Ent. 40: 129. ♀.

Dolerus inspectus MacGillivray, 1908. Canad. Ent. 40: 128. ♂.

Dolerus stugnus MacGillivray, 1908. Canad. Ent. 40: 129. ♀.

Dolenis acritus MacGillivray, 1908. Canad. Ent. 40: 130. ♀.




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Dolents simitlaiis Rohwer, 1909. Canad. Ent. 41: 10. ♀.

Dolenis inspiratus MacGillivray, 1914. Canad. Ent. 46: 105. ♀.

Dolerus 7iefastus MacGillivray, 1923. Canad. Ent. 55: 66. ♀.

Dolenis nicaeus MacGillivray, 1923. Canad. Ent. 55: 68. ♀.

Dolerus yiovkius MacGillivray, 1923. Canad. Ent. 55: 67. ♀.

Dolerus nummarius MacGillivray, 1923. Canad. Ent. 55: 159. ♀.

Dolerus nummatus MacGillivray, 1923. Canad. Ent. 55: 159. ♀.

Dolerus yiervosus MacGillivray, 1923. Insecutor Inscitiae Menstruus 11: 31. ♀.

Dolerus imnbosus MacGillivray, 1923. Insecutor Inscitiae Menstruus 11: 33. ♀, ♂.

Dolerus necessarius MacGillivray, 1923. Insecutor Inscitiae Menstruus 11: 35. ♀.

Dolerus sitnilis var. conjediis Ross, 1931. 111. Biol. Monog. 12(3): 80. ♀.

Dolerus similis var. nordanus Ross, 1931. 111. Biol. Monog. 12(3): 80. ♀, ♂.

Dolerus similis war. funiatus Ross, 1931. 111. Biol. Monog. 12(3): 81. ♀, ♂.

Dolerus tejoniensis (Norton)
N. Me.x., Utah, Nev., Oreg., Calif.

Dosytkeus Tejoniensis Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 154. ♀.

Dolerus coccinifera Norton, 1872. Amer. Ent. Soc, Trans. 4: 82. ♀.

Dolerus coccinifer Dalla Torre, 1894. Cat. Hym., v. 1, p. 4. Emend.

Dolerus yukonensis
***authority mismatch
yukonensis Norton. Alta., Alaska; Eurasia. Host: Equisetum. Other subspecies
are Palaearctic.

Dolerus similis var. yukonensis Norton, 1872. Amer. Ent. Soc, Trans. 4: 82. ♂.

Dolerus lateralis Konow, 1895. Wien. Ent. Ztg. 14: 75. ♀.

Dolerus articola Kiaer, 1898. Tromsoe Mus. Aarsh. 19: 58.

Dolerus nuntius MacGillivray, 1923. Canad. Ent. 55: 158. ♀.

Dolerus nundinus MacGillivray, 1923. Canad. Ent. 55: 159. ♂.

Taxonomy: Benson, 1956. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 25: 58-59.

Species Group Gessneri

gessneri konowi MacGillivray. Alaska, B. C, Wash., Oreg., Calif.; Siberia. Host: Equisetum.
D. gessneri gessneri Andre is European.
Dolerus konowi MacGillivray, 1914. Canad. Ent. 46: 106. ♀, ♂.
Dolerus nunterosus MacGillivray, 1923. Canad. Ent. 55: 67. ♀, ♂.
Dolerus yiidulus MacGillivray, 1923. Insecutor Inscitiae Menstruus 11: 31. ♀.
Dolerus kamtchaticus Malaise, 1932. Arkiv for Zool. 23A: 18. ♀, ♂.

Taxonomy: Benson, 1956. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 25: 59.

Dolerus pratensis (Linnaeus)
111., Man., Alta.; Europe to Siberia. Host: Equisetiun.
Tenfhredo pratensis Linnaeus, 1758. Syst. Nat., Ed. 10, p. 556. ♀.

Tenthredo (Dolerus) dubius Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 299. ♀, ♂.
Tenthredo (Dolerus) timida Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 300. ♀, ♂.
Tenthredo (Dolerus) deserta Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 300. ♂.
Dolerus frisoni Ross, 1931. 111. Biol. Monog. 12(3): 91. ♂.

Taxonomy: Malaise and Benson, 1934. Arkiv for Zool. 26A: 5 (D. pratensis of authors, not
L=D. gennanicus (Fabricius)). —Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 389.
subfasciatus neoaprilis MacGillivray. Newfoundland to N. W. T., Alaska s. to N. Y., 111., Nebr.,
Colo. Ecology: Widespread. Host: Equisetuyn. D. subfasciatus subfasciatus Smith is
found in Japan.
Dolerus neoaprilis MacGillivray, 1908. Canad. Ent. 40: 126. ♀.
Dolerus nivatus MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 164. ♀.

Taxonomy: Benson, 1956. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 25: 59.

Genus LODERUS Konow

Loderus Konow, 1890. Deut. Ent. Ztschr. 34: 240.

Type-species: Tenthredo pratorum Fallen. Orig. desig.

Taxonomy: Benson, 1956. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 25: 62 (status of
N. Amer. forms).




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eversmanni acidus MacGillivray. Que., Maine to Pa. w. to Alaska, 111.; East Asia. Host:

Equisetum. L. eversmanni eversniamii (Kirby) is European.
Loderus acidns MacGillivray, 1923. 111. Univ. Bui. 20(50): 20. ♀, ♂.
genucinctus niger Rohwer. Wash., Oreg., Calif. Host: Equisetum. L. genucinctus genucinctus

(Zaddach) is Palaearctic.
Loderus niger Rohwer, 1910. Canad. Ent. 42: 49. ♀, ♂. Preocc. in Dolerus but not in

Loderus.
Dolerus napaeus MacGillivray, 1923. Canad. Ent. 55: 65. ♀.
Dolerus Jiepotulus MacGillivray, 1923. Canad. Ent. 55: 68. ♂.
Dolerus iiauficus MacGillivray, 1923. Insecutor Inscitiae Menstruus 11: 35. ♀.
pratorum albifrons (Norton). Newfoundland to Alaska s. to N. Y., Nebr., Calif.; East Asia.

Host: Equisetum. L. pratorum pratorum (Fallen) is European.
Dosytkeus apricus var. albifrons Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 152. ♀.
Dolerus minusculus MacGillivray, 1908. Canad. Ent. 40: 126. ♀.
Loderus accuratus MacGillivray, 1923. 111. Univ. Bui. 20(50): 19. ♀.
vestigialis apricus (Norton). N. B. to Va. w. to Alaska, Man., Wash., Nebr., Colo. Host:

Equisetum. L. vestigialis vestigialis (Klug) is European.
Tenthredo (Dosytheus) aprica Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Dosytheus apricus Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 152. ♀.
Dolerus hictatus MacGillivray, 1908. Canad. Ent. 40: 127. ♀.
Dolerus femur-ruhnim Rohwer, 1909. Canad. Ent. 41: 9. ♀.
Loderus acerbus MacGillivray, 1923. 111. Univ. Bui. 20(50): 19. ♀.
Loderus alticinctus MacGilHvray, 1923. 111. Univ. Bui. 20(50): 20. ♀.
Loderus ancisus MacGillivray, 1923. 111. Univ. Bui. 20(50): 21. ♀.

Subfamily SUSANINAE

Revision: Smith, 1969. Ent. Soc. Wash., Proc. 71: 13-23.

Genus SUSANA Rohwer and Middleton

Siisana Rohwer and Middleton, 1932. Ent. Soc. Wash., Proc. 34: 93.

Type-species: Susaua mpressi Rohwer and Middleton. Orig. desig.

Members of this exclusively North American genus feed on plants of the family Cupressaceae.

Susana annulata Smith
Calif. (San Mateo Co., Marin Co.). Ecology: Adults collected from Cupressus
macrocarpa.

Susana annulata Smith, 1969. Ent. Soc. Wash., Proc. 71: 22. ♀, ♂.

Susana cupressi Rohwer and Middleton
S. Calif. Host: Cupressus.

Susana cupressi Rohwer and Middleton, 1932. Ent. Soc. Wash., Proc. 34: 94. ♀, ♂.

Susana fuscala Wong and Milliron
B. C. Host: Juniperus scopulomm.

Susana fuscala Wong and Milliron, 1972. Canad. Ent. 104: 1025-1026. 9, larva.

Susana juniperi (Rohwer)
N. Mex. (Las Vegas Hot Springs). Host: Juniperus.

Platycampus juniperi Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 386. ♀.

Susana oregonensis Smith
Idaho, Nev., Oreg. Host: Juniperus.

Susana oregonensis Smith, 1969. Ent. Soc. Wash., Proc. 71: 21. ♀.

Susana punctata Smith
Calif. (Inyo Co.). Ecology: Adult collected from Juniperus.

Susana punctata Smith, 1969. Ent. Soc. Wash., Proc. 71: 19. ♀.

Susana rufa Smith
N. Mex., Ariz. (Chiricahua Mts.).

Susana rufa Smith, 1969. Ent. Soc. Wash., Proc. 71: 21. ♀.

Subfamily NEMATINAE

The Nematinae are a large and diverse group, especially in habits and host plants. It is the
dominant sawfly group in the arctic and subarctic regions of the world where they comprise
nearly the entire sawfly fauna. Their abundance decreases rapidly southward and only a few
species extend into Mexico. Only two species, both Pristiphora, are known from South America.




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Larval habits range from free leaf feeders to gall formers, others are leaf edge rollers, leaf
miners, and some live and feed inside the fruits of rosaceous plants. The taxonomy of many
groups is imperfectly known and few have been recently studied. Because of their abundance in
the arctic region, many represent holarctic species, and a worker is met with the major obstacle
of having to become familiar with the innumerable forms described from Eurasia. Variation is
another factor since the extreme arctic climate may influence major morphological and color dif-
ferences within species. For example, females of Amauroneviatus abnonnis (Holmgren) are
normally brachypterous, but females with normal wings also exist. Some of the gall forming
groups apparently contain species complexes where there are practically no adequate
morphological characters for species separation and where identity may have to be based on
specific host records and accompanying biological information. Extensive field work and
knowledge of the world fauna will be essential to the study of most groups, and such facilities
are not available to most taxonomists.

Revision: Marlatt, 1896. U. S. Dept. Agr. Div. Ent. Tech. Ser. 3, 135 pp.

Biology: Marlatt, 1895. Ent. Sec. Wash., Proc. 3: 263-267 (notes on some gall forming species).

Tribe CLADIINI

Revision: Rohwer and Middleton, 1922. U. S. Natl. Mus., Proc. 60: 1-46 (also biology).
—Smith, 1974. Amer. Ent. Soc, Trans. 100: 1-28.

Genus CLADIUS Illiger

Cladius Illiger, 1807. Mag. Insektenk. 6: 190.

Type-species: Tenth redo diffonnis Panzer. Desig. by Latreille, 1810.

Only one of several species of this holarctic genus is found in North America.

Cladius difformis (Panzer)
N. S., Que. to B. C. s. to Va., Ky., Mo., CaHf.; Europe to Siberia. Ecology:
Widespread, common on cultivated roses. Host: Rosa. Bristly rose slug.

Tenthredo diffonnis Panzer, 1799. Faunae Ins. German. 6: 62.

Nematus crassicomis Stephens, 1835. Illus. Brit. Ent., v. 7, p. 28.

Tenthredo {Cladius) isomera Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bet. Zool.
Mass., p. 583. Nom. nud.

Cladius isoynerus Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 223. ♀.

Cladius gracilicomis Konow, 1884. Deut. Ent. Ztschr. 28: 314. ♀, ♂.

Cladius major Cobelli, 1892. Zool.-Bot. Gesell. Wien, Verb. 42: 70. ♀.

Biology: Murtfeldt, 1890. U. S. Dept. Agr., Ent. Bui. 22: 78. —Chittenden, 1908. U. S. Dept.
Agr., Bur. Ent. Circ. 105, 6 pp. —Middleton, 1922. In Rohwer and Middleton, U. S. Natl.
Mus., Proc. 60: 13. —Middleton, 1922. U. S. Dept. Agr., Farmers' Bui. 1252, 4 pp.

Genus PRIOPHORUS Dahlbom

Nematus subg. Priophorus Dahlbom, 1835. Conspectus Tenthred., Siricid., Oryssin., Scand.,
p. 4.

Type-species: Priophorus pilicomis Dahlbom. Desig. by Rohwer, 1911.
Stevensia BruUe, 1846. In Lepeletier, Hist. Nat. Ins., Hym., v. 4, p. 667. No species
included.

Type-species: Priophorus varinesV.) Lepeletier. Desig. by Rohwer, 1922. Rohwer
must have meant Pristiphora varipes Lepeletier.

Priophorus betulae Rohwer
Newfoundland to N. Y. w. to Alta., S. Dak. Host: Betula.

Priophorus betulae Rohwer, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 27. ♀, ♂.

Priophorus infuscatus (MacGillivray)
Ont., N. H., N. Y. w. to Mich., 111. Host: Salix, Populus{?).

Craterocercus infuscatus MacGillivray, 1916. Conn. State Geol. and Nat. Hist. Survey Bui. 22: 106. ♀.
Trichiocampus pacatus MacGillivray, 1921. Ent. News 32: 48. ♀.
Trichiocaynpus paetnlus MacGillivray, 1921. Ent. News 32: 48. ♀.
Trichiocampus palliolatus MacGillivray, 1921. Ent. News 32: 49. ♀.




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Priophonis. salicivorus Rohwer, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc.

60: 29. ♀, ♂.

Priophorus morio (Lepeletier)
N. B. to B. C. s. to Pa., Mich., Mont., CaHf.; Europe to Siberia. Ecology:

Widespread. Host: Rubus. Introduced into New Zealand and Tasmania; attempted

introduction into Hawaii from Oreg. and Calif, to aid in biological control of Rubus.
Cladiiis vtorio Lepeletier, 1823. Monog. Tenthred., p. 58. ♀, ♂.
Nemahis bnillei Dahlbom, 1835. Conspectus Tenthred., Siricid., Oryssin. Scand., p. 7.
Cladins tener Zaddach, 1859. Programm Friedr. Col. Konigsberg, p. 11. ♀.
Cladius tristis Zaddach, 1859. Programm Friedr. Col. Konigsberg, p. 11. ♀, ♂.
Priophonis rubivorus Rohwer, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 28. ♀.
Priophorus montanus Rohwer, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 31.9.
Priophonis nibi Rohwer, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 32. ♀.

Biology: Essig and Hoskins, 1944. Calif. Dept. Agr., Ext. Serv. Cir. 87: 37. —Smith and Kido,
1949. Hilgardia 19: 43-54. -Nakao, 1967. Hawaii. Ent. Soc, Proc. 19: 339.

Priophorus pallipes (Lepeletier)
Newfoundland to Alaska s. to Va., Tenn., 111., Colo., Oreg.; Eurasia.

Ecology: Widespread. Host: Pninus, Crataegus, Alnus.
Tenthredo albipes Fallen, 1808. Svenska Ventensk. Akad., Handl. 29: 110. Preocc. by

Geoffroy, 1785.
Cladius pallipes Lepeletier, 1823. Monog. Tenthred., p. 59. ♀, ♂.
Pristiphora varipes Lepeletier, 1823. Monog. Tenthred., p. 61. ♂.
Cladius !wm?(?us Stephens, 1835. Illus. Brit. Ent., v. 7, p. 23.
Neynatus vielaiiostigma Stephens, 1835. Illus. Brit. Ent., v. 7, p. 35.
Cladius aequalis Norton, 1872. Amer. Ent. Soc, Trans. 4: 78. ♂.
Priophonis discors Konow, 1894. Wien. Ent. Ztg. 13: 88.
Cladius solifaris Dyar, 1895. Canad. Ent. 27: 192. 9, larva.
Priophonis modestius MacGillivray, 1921. Ent. News 32: 49. ♀.
Priophonis muuditus MacGillivray, 1921. Ent. News 32: 50. ♀.
Priophonis crataegi Rohwer, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 26. ♀.
Priophonis pnini Rohwer, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 32. ♀, ♂.
Priophonis virginiayius Rohwer, 1922. hi Rohwer and Middleton, U. S. Natl. Mus., Proc. 60:34. ♀.
Priophonis plesius Rohwer, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 35. ♀.

Biology: Middleton, 1922. hi Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 1-46 (notes
following descriptions of species listed above).

Genus TRICHIOCAMPUS Hartig

Cladius subg. Trichiocanipns Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst
Einleitung Naturgesch. Hym., p. 176.

Type-species: Neniaius grandis Lepeletier. Desig. by Rohwer, 1911.

Trichiocampus gregarius (Dyar)
Maine to Pa. w. to Ont, Mich. Host: Populiis.

Cladius (Trichiocanipus) gregarius Dyar, 1895. Canad. Ent. 27: 191. 9, 3, larva.

Biology: Middleton, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 11.

Trichiocampus simplicicornis (Norton)
N. S. to N. Y. w. to Sask.; N. Mex. and Calif. Host: Salix, sometimes
Populus.
Cladius simplicicornis Norton, 1869. Amer. Ent. Soc, Trans. 2: 367. ♂.
Priophorus irregularis Dyar, 1900. N. Y. Ent. Soc, Jour. 8: 28. 6 . Larva.
Trichiocampus patchiae MacGillivray, 1921. Ent. News 32: 48. ♀.
Priophonis rnoratus MacGillivray, 1921. Ent. News 32: 50. ♀.

Biology: Middleton, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 9.




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vitninalis (Fallen). Newfoundland to N. J. w. to Mich.; B. C; Greenland; Europe and north

Asia. Independently introduced into both east and west N. Amer.; first found in N. Y. in
1880's. Host: Populus, sometimes Salix.

Tenthredo viminalis Fallen, 1808. Svenska Vetensk. Akad., Handl. 29: 177.

Nematus grandis Lepeletier, 1823. Monog. Tenthred., p. 61. ♀.

Cladius eucera Bouche, 1834. Naturgesch. des Ins., p. 140. ♀, ♂.

Cladius liiteicortiis Stephens, 1835. Illus. Brit. Ent., v. 7, p. 24.

Cladius hiteiventris Dahlbom, 1836. Prodrom. Hym. Scand., p. 102.

Aulacoments Ititesceris Lintner, 1888. Ins. N. Y. Rpt. 4: 44. ♀, ♂.

Platycantpus victoria MacGillivray, 1920. Canad. Ent. 52: 61. ♀.

Biology: Middleton, 1922. In Rohwer and Middleton, U. S. Natl. Mus., Proc. 60: 7. — Viereck,
1925. Canad. Ent. 57: 43. -Beique, 1961. Canad. Ent. 93: 1085-1097. -Smirnoff and
Beique, 1959. Canad. Ent. 91: 379-381 (polyhedral disease).

Tribe NEMATINI

Genus HOPLOCAMPA Hartig

Hoplocampa Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung
Naturgesch. Hym., p. 276.

Type-species: Tenthredo (Allantus) brevis Klug. Desig. by Rohwer, 1911.
Macgillivraya Ashmead, 1898. Canad. Ent. 30: 257. Preocc. by Forbes, 1852.

Type-species: Macgillivraya oregonensis Ashmead. Orig. desig.
Macgillivrayella Ashmead, 1899. In Smith, Ins. of N. J., p. 606. N. name for Macgillivraya.

Larvae of this holarctic genus live and feed inside the fruits of Rosaceae. Species that attack
cultivated crops such as cherries, pears, and apples may be serious pests. Adults emerge in the
spring when their host plant is in bloom and the females oviposit in the ovaries. Sometimes
adults can be collected in large numbers from their host blossoms. The larvae feed in the
developing fruit of the host, the fruit usually falling off when the larvae mature. The larvae
enter the ground where they overwinter. All species are single brooded.

Revision: Rohwer, 1911. U. S. Dept. Agr. Bur. Ent. Tech. Ser. 20(4): 139-148. —Ross, 1943.
Amer. Ent. Soc, Trans. 69: 61-92 (as Holocampa (!)).

Taxonomy: Benson, 1968. Brit. Mus. (Nat. Hist.) Ent., Bui. 22: 199 (key to three species
groups).

Biology: Bird, 1927. Ent. Soc. Amer., Ann. 20: 481-487.

Hoplocampa alpestris Rohwer
Alta., Mont., Colo., B. C, Wash. Ecology: Adults collected from Amelanchier
flowers.
Hoplocampa {Hoplocampa) alpestris Rohwer, 1911. U. S. Dept. Agr. Bur. Ent. Tech. Ser.
20(4): 142. ♀, ♂.

Hoplocampa bioculata Rohwer
Colo., Utah, Idaho, B. C, Wash., Oreg., Calif. Ecology: Adults collected
from Amelanchier flowers.
Hoplocampa bioculata Rohwer, 1908. Canad. Ent. 40: 179. ♀.
Hoplocampa (Hoplocampa) koebelei Rohwer, 1911. U. S. Dept. Agr. Bur. Ent. Tech. Ser.

20(4): 142. ♀.
Hoplocampa padiisa MacGillivray, 1923. 111. Univ. Bui. 20(50): 17. ♂.

Hoplocampa brevis (Klug)
R. I., Conn., Ont., N. Y., Pa., Md.; Europe. Introduced, first found in Ontario in
1964. Host: Pynis. An economic pest of pear.
Tenthredo (Allantus) brevis Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 53. ♀.

Taxonomy: Dustan, 1966. Canad. Ent. 98: 267 (occurrence in Canada). —Smith, 1966. U. S.
Dept. Agr., Coop. Econ. Ins. Rpt. 16(12): 228-230 (separation of brevis and testudinea
adults and larvae).

Biology: Thomas, 1936. Ann. Appl. Biol. 23: 633-639.




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Hoplocampa cookei (Clarke)
B. C, Idaho, Wash., Oreg., Calif. Host: Prunus. A pest of cultivated cherries.
Cherry fruit sawfly.
Dolenis cookei Clarke, 1906. Canad. Ent. 38: 351. ♀, ♂.

Hoplocampa (Hoplocampa) califomica Rohwer, 1911. U. S. Dept. Agr., Bur. Ent. Tech.
Ser. 20(4): 123. ♀, ♂.

Biology: Foster, 1913. U. S. Dept. Agr., Bur. Ent. Bui. 116, pp. 73-79. — Essig, 1914. Calif.
State Comm. Hort., Monthly Bui., p. 31. — Duruz, 1922. Calif. Dept. Agr., Monthly Bui.
9(4): 393-399. -Essig and Hoskins, 1944. Calif. Dept. Agr., Ext. Serv. Cir. 87: 47.

Hoplocampa halcyon (Norton)
Maine to N. C. w. to Alta., 111. Host: Amelanchier.

Tenthredo (Selandria) halcyon Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Selandria halcyon Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 222. ♀, ♂.
Selandria flavicomis Provancher, 1878. Nat. Canad. 10: 100. ♂.
Hoplocampa (Hoplocampa) xantha Rohwer, 1911. U. S. Dept. Agr., Bur. Ent. Tech. Ser.

20(4): 144. ♀.

Morphology: Bird, 1926. Ent. Soc. Amer., Ann. 19: 268-279 (external anatomy of adult).
—Bird, 1927. Ent. Soc. Amer., Ann. 20: 481-486 (external anatomy of larva).
Idaho Ross. Idaho, Wash., Oreg.

Hoplocampa idaho Ross, 1943. Amer. Ent. Soc, Trans. 69: 81. ♀, ♂.

Hoplocampa lacteipennis Rohwer
Maine to N. Y. w. to Man., Alta., Mont., Colo. Host: Prunus virginiana
and probably other species of Pninus.
Hoplocampa {Macgillivrayella) lacteipennis Rohwer, 1910. Canad. Ent. 42: 244. ♀.
Hoplocampa {Macgillivrayella) xanthura Rohwer, 1910. Canad. Ent. 42: 244. ♂.
Hoplocampa (Macgillivrayella) pallida Rohwer, 1911. U. S. Dept. Agr., Bur. Ent. Tech.
Ser. 20(4): 141. ♀.

Biology: Bird, 1928. Sci. Agr. 8: 479-501.
makila Ross. Oreg.

Hoplocampa makila Ross, 1943. Amer. Ent. Soc, Trans. 69: 68. ♀.

Hoplocampa marlatti Rohwer
Maine to Ga. w. to Alta., Iowa, Colo. Ecology: Adults collected from flowers
of wild cherry, Prunus sp.
Hoplocampa (Hoplocampa) marlatti Rohwer, 1911. U. S. Dept. Agr., Bur. Ent. Tech. Ser.
20(4): 143. ♀, ♂.

Hoplocampa montanicola Rohwer
Maine, Ont. to B. C. s. to N. Y., Mich., S. Dak., Colo., Utah, Calif.

Ecology: Widespread. Host: Pninus virginiana and probably other species of Pninus.
Hoplocampa (Hoplocampa) montanicola Rohwer, 1911. U. S. Dept. Agr., Bur. Ent. Tech.

Ser. 20(4): 145. ♀, ♂.
Hoplocampa (Hoplocampa) orbitalis Rohwer, 1911. U. S. Dept. Agr., Biu*. Ent. Tech. Ser.

20(4): 141. ♀.
Hoplocampa (Hoplocampa) occidentalis Rohwer, 1911. U. S. Dept. Agr., Bur. Ent. Tech.
Ser. 20(4): 144. ♀, ♂.

Biology: Bird, 1927. Ent. Soc. Amer., Ann. 20: 481-486 (xantha, not Rohwer).

Hoplocampa nalema Ross
Wash. (Mt. Rainier).

Hoplocampa nalema Ross, 1943. Amer. Ent. Soc, Trans. 69: 82. ♀, ♂.

Hoplocampa neneti Ross
Maine, N. H., Ont. Ecology: Adults collected from Amelanchier.

Hoplocampa yieneti Ross, 1943. Amer. Ent. Soc, Trans. 69: 77. ♂.

Hoplocampa oregonensis
***authority mismatch
(Ashmead). Mont., Idaho, Oreg.

Macgillivraya oregonensis Ashmead, 1898. Canad. Ent. 30: 257. ♀, ♂.

Hoplocampa oskina Ross
Maine, N. Y. w. to Mich., Iowa, Kans. Ecology: Adults collected from Crataegus
flowers.
Hoplocampa oskina Ross, 1943. Amer. Ent. Soc, Trans. 69: 71. ♀, ♂.

Hoplocampa pallipes MacGillivray
N. H., N. Y. to Mich.; Alta. w. to B. C, Oreg., Calif. Ecology: Adults
collected from Amelanchier flowers.
Hoplocampa pallipes MacGillivray, 1893. Canad. Ent. 25: 239. ♀.

Hoplocampa (Hoplocampa) nevadensis Rohwer, 1911. U. S. Dept. Agr., Bur. Ent. Tech.
Ser. 20(4): 143. ♀.




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Hoplocampa ritcheri Ross
N. C, 111., Wash. Ecology: Adults collected from Crataegus flowers.

Hoplocampa ritcheri Ross, 1943. Amer. Ent. Soc, Trans. 69: 72. ♀, ♂.

Hoplocampa sialica Ross
Maine to Md. w. to Mich. Ecology: Adults collected from Amelanchier flowers.

Hoplocampa sialica Ross, 1943. Amer. Ent. Soc, Trans. 69: 67. ♀, ♂.

Hoplocampa spala Ross
B. C, Oreg.

Hoplocampa spala Ross, 1943. Amer. Ent. Soc, Trans. 69: 83. ♂.

Hoplocampa stricklandi Ross
Maine, N. Y., Mich., Alta.

Hoplocampa stricklandi Ross, 1943. Amer. Ent. Soc, Trans. 69: 73. ♀, ♂.

Hoplocampa testudinea (Klug)
N. H., Vt., Mass., R. I., Conn., N. Y., N. J.; B. C; Europe. Accidentally

introduced into both east and west N. Amer., first found in N. Y. in 1939 and in B. C. in
1940. Host: Mains malus. A pest of apples.
Tenthredo (Allantus) testudinea Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 60. ♀.

Biology: Miles, 1932. Ann. Appl. Biol. 14: 420-431. — Downes and Andison, 1942. Ent. Soc.
Brit. Columbia, Proc 39: 13-16. — Pyenson, 1943. Jour. Econ. Ent. 36: 218-221.

Hoplocampa texas Ross
Tex. Ecology: Adults collected from Crataegus flowers.
Hoplocampa texas Ross, 1943. Amer. Ent. Soc, Trans. 69: 68. ♀, ♂.

Genus CAULOCAMPUS Rohwer

Caidocampus Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 239.

Type-species: Priophorus acericaulis MacGillivray. Orig. desig.

Revision: Smith, 1968. Ent. Soc Wash., Proc. 70: 126-129.

Caulocampus acericaulis (MacGillivray)
Conn., Ont., N. Y., Md., Ala., Mich., 111. Ecology: Larva mines in leaf
petioles. Host: Acer.
Priophorus acericaulis MacGillivray, 1906. Canad. Ent. 38: 306. ♀.

Biology: Britton, 1906. Ent. News 17: 313-320.

Caulocampus matthewsi Smith
N. Y.

Caulocampus matthewsi Smith, 1968. Ent. Soc. Wash., Proc. 70: 128. ♀.

Genus HEMICHROA Stephens

Genus HEMICHROA Subgenus HEMICHROA Stephens

Hemichroa Stephens, 1835. Illus. Brit. Ent., v. 7, p. 55.

Type-species: Tenthredo alni Linnaeus. Desig. by Westwood, 1840.
Leptocerca Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung Naturgesch.

Hym., p. 228.

Type-species: Tenthredo alni Linnaeus. Desig. by Rohwer, 1911.
Engages Gistel, 1848. Naturgesch. Thierr. f. hoh SchuL, p. 9. N. name for Leptocerca

Hartig.
Leptocercus Thomson, 1871. Hym. Scand., v. 1, p. 76. Emend.

Hemichroa crocea
***authority mismatch
(Geoffrey). Newfoundland to Alaska s. to Conn., N. Y., N. Mex., Idaho, Greg.; Europe

to Siberia. Ecology: Widespread. Host: Ahius. Striped alder sawfly.
Tenthredo crocea Geoffrey, 1785. In Fourcroy, Ent. Paris, p. 364.
Tenthredo rufa Panzer, 1799. Faunae Ins. German., v. 6, t. 72, fig. 2.
Hemichroa stigma Stephens, 1835. Illus. Brit. Ent., v. 7, p. 55.
Dineura unicolor Rudow, 1872. Stettin Ent. Ztg. 30: 218.

Neynatus ardens Zaddach, 1882. Phys. Okonom. Gesell. Konigsberg., Schr. 23: 133.
Dineura americana Provancher, 1882. Nat. Canad. 13: 292. ♀.
Hemichroa pallida Ashmead, 1890. Colo. Biol. Assoc, Bui. 1: 15. ♀.
Hemichroa dyari Rohwer, 1918. Ent. Soc Wash., Proc. 20: 171. ♀, ♂.
Hemichroa washingtonia Rohwer and Middleton, 1931. Ent. Soc. Wash., Proc. 34: 97. ♀, ♂.

Biology: Ross, 1932. Canad. Ent. 64: 247-248. -Hopping, 1937. Canad. Ent. 69: 243-249.
—Reeks and Smith, 1945. Acad. Nat. 2: 11.




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Genus HEMICHROA Subgenus VARNA Ross

Hemichroa subg. Varna Ross, 1937. 111. Biol. Monog. 15(2): 79.
Type-species: Nematus militaris Cresson. Orig. desig.

Hemichroa amelanchieridis (Rohwer)
Maine, Que. to N. Y.; Alta. to Colo. w. to B. C, Oreg. Host:
Amelanchier.
Pteronidea amelanchieridis Rohwer, 1920. U. S. Natl. Mus., Proc. 57: 215. ♀.

Hemichroa militaris (Cresson)
Ont. to Alaska s. to R. I., Pa., 111., Mont., Nev. Ecology: Widespread. Host:
Pnmus, Amelanchier.
Nematus militaris Cresson, 1880. Amer. Ent. Soc, Trans. 8: 7. ♀.
Nematus thoracicus Harrington, 1893. Canad. Ent. 21: 58. ♀.

Genus CRATEROCERCUS Rohwer

Craterocercus Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 385.

Type-species: Hemichroa phytophagica Dyar. Orig. desig.

Larvae of this exclusively North American genus feed on Quercus.

Revision: Rohwer, 1918. Ent. Soc. Wash., Proc. 20: 162-165. —Smith, 1969. Ent. Soc. Wash.,
Proc. 71: 153-161.

Craterocercus albipes (Cresson)
Oreg., Nev., Calif. Ecology: Adults collected from Quercus.
Mesoneura albipes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 11. ♀.
Craterocercus califoniicus Rohwer, 1917. U. S. Natl. Mus., Proc. 53: 234. ♀.

cordleyi MacGillivray. Oreg., Calif.

Craterocercus cordleyi MacGillivray, 1923. 111. Univ. Bui. 20(50): 9. ♀.

Craterocercus fraternalis (Norton)
N. H. to Pa. w. to Wis., Ill, Kans., Tex. Host: Quercus alba.

Hemichroa fratenialis Norton, 1872. Amer. Ent. Soc, Trans. 4: 81. ♂.

Craterocercus quercivorus Rohwer, 1918. Ent. Soc. Wash., Proc. 20: 164. ♀, ♂.

Craterocercus cervinus MacGillivray, 1923. 111. Univ. Bui. 20(50): 9. ♀.

Craterocercus circulus MacGillivray, 1923. 111. Univ. Bui. 20(50): 9. ♀.

Craterocercus furcatus Smith
Conn., N. J., 111.

Craterocercus furcatus Smith, 1969. Ent. Soc. Wash., Proc. 71: 160. ♀.

Craterocercus obtusus (Klug)
Conn, to Fla. w. to III., Kans., Ark., Tex. Host: Quercus alba, Q. coccinea.

Tenthredo {Allaiitus) obtusa Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 55. ♀.

Hemichroa albidovariata Norton, 1872. Amer. Ent. Soc, Trans. 4: 81. ♀.

Hemichroa phytophagica Dyar, 1898. N. Y. Ent. Soc, Jour. 6: 125. 9, larva.

Craterocercus floridanus Rohwer, 1912. U. S. Natl. Mus., Proc 43: 238. ♀, ♂.

Genus PLATYCAMPUS Schiodte

Leptopus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung Naturgesch.
Hym., p. 184. Preocc. by Mayer, 1835.

Type-species: Nematus (Leptopus) hypogastricus Hartig. Monotypic
Platycamptis Schiodte, 1839. Mag. Zool. (ser. 2) 1: 20, footnote; pi. 6-10. N. name for

Leptopus Hartig.
Erasminus Gistel, 1848. Naturgesch. Thierr. f. hoh SchuL, p. 9. N. name for Leptopus

Hartig.
Camponiscus Newman, 1869. Entomologist 4: 215.

Type-species: Camponiscus healaei Newman. Monotypic.

A small holarctic genus with two North American representatives.

Platycampus albostigmus (Rohwer)
N. S., Maine to Alaska s. to N. Y., Mich., Minn., Colo., Nev., Wash.
Ecology: Widespread.
Camponiscus albostigmus Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 105. ♀.

Platycampus americanus (Marlatt)
Que., Maine to Alaska s. to Pa., N. Mex., Idaho. Ecology: Widespread.
Host: Populus.
Camponiscus americanus Marlatt, 1896. Canad. Ent. 28: 251. ♀, ♂.
Camponiscus smithi Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 105. ♀.
Platycampus vierecki MacGillivray, 1920. Canad. Ent. 52: 60. ♀.




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Genus ANOPLONYX Marlatt

Anoplonyx Marlatt, 1896. U. S. Dept. Agr. Div. Ent. Tech. Ser. 3: 18.

Type-species: Nematiis pectoralis Lepeletier. Desig. by Rohwer, 1911.
Marlaftia Ashmead, 1898. Canad. Ent. 30: 287.

Type-species: Hemichroa laricis Marlatt. Orig. desig.

Larvae of this holarctic genus feed on Larix.

Taxonomy: Wong, 1955. Canad. Ent. 87: 224-227 (larvae). —Woods, 1962. Ent. Soc. Brit.
Columbia, Proc. 59: 17 (species in B. C).

Anoplonyx canadensis Harrington
Newfoundland to Maine w. to Yukon, n. B. C, Sask., Minn. Host:
Larix laricina.
Anoplonyx canadeyisis Harrington, 1902. Canad. Ent. 34: 94. ♀.

Platycampus (.Anoplonyx) itascus Rohwer and Middleton, 1932. Ent. Soc. Wash., Proc. 34: 96. ♂.

Anoplonyx laricivorus (Rohwer and Middleton)
Mont., B. C, Idaho. Host: Larix occideyitalis.

Platycampus {Anoplonyx) laricivorus Rohwer and Middleton, 1932. Ent. Soc. Wash., Proc. 32:95. ♀, ♂.

Anoplonyx luteipes (Cresson)
Newfoundland to Maine, N. H. w. to Yukon, n. B. C, Minn., 111. Host: Larix
laricina.
Dineura luteipes Cresson, 1880. Amer. Ent. Soc, Trans. 8:11. ♂.
Hemichroa laricis Marlatt, 1896. Canad. Ent. 28: 257. ♀.

Biology: Reeks and Smith, 1945. Acad. Nat. 2:11.
Morphology: Bracken, 1961. Canad. Ent. 93: 573-593.

Anoplonyx occidens Ross
Mont., B. C, Idaho, Oreg. Host: Larix occidentalis.

Platycampus (Anoplonyx) laricis Rohwer and Middleton, 1932. Ent. Soc. Wash., Proc. 34: 95. ♀, ♂. Preocc. by Marlatt, 1896.
Anoplonyx occidens Ross, 1937. 111. Biol. Monog. 15(2): 82. N. name.

Genus ADELOMOS Ross

Adelomos Ross, 1935. Canad. Ent. 67: 201.

Type-species: Adelomos cleone Ross. Orig. desig.

Adelomos cleone Ross
Ky., 111.

Adelomos cleone Ross, 1935. Canad. Ent. 67: 202. ♀.

Genus NEOPAREOPHORA MacGillivray

Neopareophora MacGillivray, 1908. Canad. Ent. 40: 289.

Type-species: Neopareophora ynartini MacGillivray. Orig. desig.
Dineuridea Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 240.

Type-species: Marlattia erythrothorax Rohwer. Orig. desig.

A small Nearctic genus with several unidentified species in addition to those listed below.

Revision: Ross, 1935. Canad. Ent. 67: 204-205.

Neopareophora erythrothorax (Rohwer)
Fla. (Jacksonville).

Marlattia erythrothorax Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 385. ♀.

Neopareophora litura (Klug)
N. B., Que. to Va. w. to 111. Ecology: First instar larvae feed inside the buds,

subsequent instars feed on the leaves. Host: Vacciniujn, especially low-bush blueberry.
Dineura litura Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 83. ♀.
Dineura linitus Norton, 1867. Amer. Ent. Soc, Trans. 1: 240. ♀.
Dineura lateralis Norton, 1867. Amer. Ent. Soc, Trans. 1: 240. ♀.
Neopareophora martini MacGillivray, 1908. Canad. Ent. 40: 289. ♀.

Biology: Maxwell, 1953. Canad. Ins. Pest Rev. 31(1): 100. — Neilson, 1955. Canad. Ent. 87:
356-357. -Neilson, 1958. Canad. Ent. 90: 229-234.



56 Hymenoptiera in America North of Mexico

Genus NEMATINUS Rohwer

Nemafimis Rohwer, 1911. U. S. Dept. Agr., Bur. Ent. Tech. Ser. 20: 99.
Type-species: Tenthredo abdominalis Panzer. Orig. desig.

A holarctic genus which is found in North America from N. S. to Alaska south to N. Y., Mich.,
S. Dak., Colo., and Calif. This genus has not been studied, and the following is merely a list of the
described species with their type localities.

Nematinus ochreatus (Rohwer)
N. B. (St. John).

Pteronus ochreatus Rohwer, 1910. U. S. Natl. Mus., Proc. 38: 198. ♀.

Nematinus oronus (Kincaid)
Alaska (Yukatat Bay).

Pachynematus oronus Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 347. ♀.

Nematinus pontanioides (Marlatt)
Oreg. (Mt. Hood).

Neinatus pontanioides Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 89. ♀.

Nematinus unicolor (Dyar)
N. Y. (Keene Valley). Host: Betula alba.

Neynatus unicolor Dyar, 1895. Amer. Ent. Soc, Trans. 22: 308. Larva. Dyar's description
precedes that of Marlatt, 1896; previous listings give Marlatt as author.

Genus PACHYNEMATUS Konow

Pachynemattis Konow, 1890. Deut. Ent. Ztschr., pp. 233, 238, 246.

Type-species: Nematiis capreae Panzer. Desig. by Rohwer, 1911.

Known hosts for this genus in North America are grasses, sedges, and dock. Some species in
Europe also feed on Salicaceae and Coniferae. Several species reared from wheat and other
grain crops may be of economic importance. So far as is known, the grass feeding species are
single brooded with the flight and feeding activities occurring early in the spring. Some of the
species on sedges apparently have several generations a year. Very little biological work has
been done. The taxonomy of this genus is almost entirely based on males. Females of a number
of species cannot be separated; consequently, the lists of unplaced taxa below. Larvae are com-
monly collected, but so few have been associated with the adults that it is impossible to identify
them except to genus.

Taxonomy: Ross, 1945. Ent. Soc. Wash., Proc. 47: 105-120 (key to N. Amer. species).
—Benson, 1948. Ent. Monthly Mag. 84: 58-65 (British species). —Benson, 1962. Brit. Mus.
(Nat. Hist.) Ent., Bui. 12: 405-406 (holarctic species). —Benson, 1963. Roy. Ent. Soc.
London, Proc, Ser. B: Taxonomy 32: 162-164 (Rumicis group).

Species Group Clitellatus

Pachynematus anolita Ross
Kans., Tex., Mont., Colo. Host: Wheat.

Pachynematus apicalis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 107. ♂.

Preocc. in Pachynematus by apicalis Hartig.
Pachynematus anolita Ross, 1951. In Muesebeck et. al., U. S. Dept. Agr., Agr. Monog. 2:
39. N. name for apicalis Marlatt.

Pachynematus auratus Marlatt
N. Y., Ohio, Ind., 111., Iowa, Kans., Mont.

Pachynemattis auratus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 99. ♀, ♂.

Pachynematus clibrichellus (Cameron)
N. H. to Alaska; arctic Eurasia. Host: Carex.
Nematus clibrichellus Cameron, 1878. Fauna Scotland, v. 1, p. 32. ♂.
Nematus Thomsoni Cameron, 1882. Ent. Soc. London, Trans., p. 540. ♀, ♂.
Nematus clibrichensis Cameron, 1885. Monog. Brit. Phytophagous Hym., v. 2, p. 88. ♂.
Pachynematus pubescens Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 100. ♀.
Pachynematus gotarius Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 348. ♂.
Pachynematus venustus MacGillivray, 1921. Calif. Acad. Sci., Proc. (ser. 4) 11(14): 190. ♀, ♂.
Pachyyiematus vermis MacGillivray, 1921. Calif. Acad. Sci., Proc. (ser. 4) 11(14): 191. ♂.

Pachynematus clitellatus (Lepeletier)
Labrador, Que.; Europe. Introduced ?. Host: Gramineae.

Tenthredo capreae Panzer, 1799. Faunae Ins. German, v. 6, p. 65, t. 8. Preocc. by Linnaeus, 1758.
Nematus clitellatus Lepeletier, 1823. Monog. Tenthred., p. 62. ♀.
Nematus trisignatus Foerster, 1854. Naturh. Ver. Rheinlande, Verb. 11: 292. ♀.



Superfamily TENTHREDINOIDEA 57

Pachynematus clypeatus Marlatt
Man., Minn., Alta., Mont.

Pachynematus clypeatus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 102. ♀.

Pachynematus excisus (Thomson)
Greenland; arctic Europe.

Nematus excisus Thomson, 1862. Svenska Vetensk. Akad., Ofvers. Forh. 19: 628. ♀.

Pachynematus extensicornis (Norton)
Maine, Ont. to Md. w. to Alta., Mont., Wyo., Colo.; Europe. Host:
Wheat, probably other Gramineae.
Nematus extensicornis Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 159. ♂.
Nematus marylandicus Norton, 1864. Ent. Soc. Phila., Proc. 3: 7. ♀.
Nematus aureopectus Norton, 1867. Amer. Ent. Soc, Trans. 1: 219. ♀.
Pachynematus affinis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 97. ♀, ♂.
Pachynematus tritici Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 106. ♂.
Pachynematus foveolatus Konow, 1903. Ztschr. System. Hym. Dipt. 3: 379. ♀, ♂.
Pachynematus truncatus Benson, 1948. Ent. Monthly Mag. 84: 62. ♂.

Pachynematus jamesi Ross
Colo., Utah.

Pachynematus jamesi Ross, 1945. Ent. Soc. Wash., Proc. 47: 112. ♂.

Pachynematus kirbyi (Dahlbom)
Que. to Alaska s. to Maine, Pa., Ill, Iowa, S. Dak., Mont.; Europe to Siberia.
Ecology: Widespread. Host: Carex.
Nematus kirbyi Dahlbom, 1835. Clavis Hym. Syst., p. 40. ♂.
Nematus flaviventris Hartig, 1840. Stettin Ent. Ztg. 1: 24.
Nematus diaphanus Eversmann, 1847. Soc. Nat. Moscou, Bui. 20: 19.
Nematus lanbripennis Eversmann, 1847. Soc. Nat. Moscou, Bui. 20: 20.
Neynatus turgidus Zaddach, 1875. Phys. Okonom. Gesell. Konigsberg, Schr. 16: 82.
Nematus suadus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 10. ♀.
Pachynematus zaddachi Konow, 1903. Ztschr. System. Hym. Dipt. 3: 377. ♀, ♂.
Pachynematus iiifumatus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 107. ♂.
Pachyneynatus refractarius MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 31. ♀.

Pachynematus miscus Ross
Oreg., Calif.

Pachynematus yniscus Ross, 1945. Ent. Soc. Wash., Proc. 47: 114. ♂.

Taxonomy: Middlekauff, 1955. Pan-Pacific Ent. 31: 196.

Pachynematus moerens (Foerster)
Alta., Alaska; Europe. Host: Gramineae.

Nematus moerens Foerster, 1854. Naturh. Ver. Rheinlande, Verb. 11: 294. ♀.
Nematus pteuralis Thomson, 1862. Ofvers. Svensk. Vet. Akad., Forh. 19: 628.
Pachynematus falonus Ross, 1945. Ent. Soc. Wash., Proc. 47: 113. ♂.

Pachynematus nevadensis Marlatt
Nev.

Pachynematus nevadensis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 110. ♂.
nigrritibialis Rohwer. N. Y., Mich., Ohio, 111., Nebr., Kans.

Pachynematus nigritibialis Rohwer, 1909. Canad. Ent. 41: 17. ♀.

Pachynematus painteri Ross
Kans., Idaho. Host: Wheat.

Pachyneynatus painteri Ross, 1945. Ent. Soc. Wash., Proc. 47: 115. ♀, ♂.

Pachynematus parvilabris (Thomson)
N. W. T., Greenland; arctic Europe. Specimen collected on Ward Hunt
Is., 83° 05' N. May be the most northerly sawfly found.

Nematris pannlabris Thomson, 1862. Ofvers. Svensk. Vet. Akad., Forh. 19: 638. ♀, ♂.

Pachyneynatus iyiopinatus Lindqvist, 1949. Notulae Ent. 28: 82.

Pachynematus abstnisus Lindqvist, 1949. Notulae Ent. 28: 83.

Pachynematus setator Ross
N. Y., Ind., 111., Oreg. Host: Festuca.

Pachynematus setator Ross, 1945. Ent. Soc. Wash., Proc 47: 113. ♂.

Biology: Kamm, 1975. Environ. Ent. 4: 312-314 (in Oreg.).

Pachynematus smithae Ross
N. H.; Europe.

Pachyyieynatns smithae Ross, 1945. Ent. Soc. Wash., Proc. 47: 116. ♂.

Pachynematus sporax Ross
Sask., Idaho, Wash., Oreg., Calif. Host: Wheat.

Pachyyieynatns sporax Ross, 1945. Ent. Soc. Wash., Proc. 47: 115. ♂.

Taxonomy: Middlekauff, 1955. Pan-Pacific Ent. 31: 196-198 (female).
Biology: Armitage, 1954. Calif. Dept. Agr., Bui. 54: 73.




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Pachynematus thoracicus Marlatt
Mont.

Pachynemaius thoracicus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 108. ♂.

Pachynematus uvator Ross
Utah, B. C, Wash.

Pachynematus uvator Ross, 1945. Ent. Soc. Wash., Proc. 47: 114. ♂.

Unplaced Taxa of Group Clitellatus

The following species were described from females. Because species separation of Pachyyie-
maius is based mostly on males, they can not be placed as yet.

absyrtus MacGilHvray. Oreg.

Pachynematus absyrtus MacGillivray, 1923. 111. Univ. Bui. 20(50): 27. 2.

Pachynematus academus MacGillivray
Oreg.

Pachynematus academus MacGiUivray, 1923. 111. Univ. Bui. 20(50): 27. ♀.

Pachynematus ater (MacGillivray)
Wash.

Messa ater MacGillivray, 1893. Canad. Ent. 25: 238. ♀.

Pachynematus coloradensis Marlatt
Colo.

Pachynematus coloradensis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 101. ♀.

Pachynematus ebenus (Cresson)
Colo.

Aulacomerus^ ebenus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 10. ♀.

Pachynematus gamus Smith
n. name. Nev.

Pachynematus graminis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 101. ♀.
Preocc. in Pachynematus by Cameron, 1874.

Pachynematus hoodi Marlatt
Oreg. (Mt. Hood).

Pachynematus hoodi Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 104. ♀.

Pachynematus luteipes (Cresson)
n. comb. Nev.

Nematus luteipes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 6. ♀.

Pachynematus pleuricus (Norton)
Colo.

Nematus pleuricus Norton, 1867. Amer. Ent. Soc, Trans. 1: 208. ♀.

Pachynematus robinsonae (Rohwer)
Colo.

Lygaeonematus robinsonae Rohwer, 1915. U. S. Natl. Mus., Proc 49: 212. ♀.

Pachynematus stigmatus (Norton)
Mass.

Tenthredo (Nematus) stigmatus Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Nematus stigmatus Norton, 1861. Boston Soc. Nat. Hist, Proc. 8: 161. ♀.

Species Group Vagus

aurantiacus Marlatt. Que. to N. Y. w. to Alta., Mont., 111. Host: Carex.

Pachynematus aurantiacus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 95. ♀.

6.
Pachynematus pjinctulatns Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 103. ♀.
Pachynematus allegatus MacGillivray, 1923. Canad. Ent. 55: 162. ♀.

Pachynematus corniger (Norton)
Newfoundland to Ga. w. to Minn., Kans. Host: Carex.
Nematus coniiger Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 159. ♂.
Nematus snbalbatus Norton, 1864. Ent. Soc. Phila., Proc. 3: 7. ♀.
Pachynematus repertus MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 31. ♀.
Pachynematus remissus MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 32. 9. Synonymy of

Pachynematus remissus
***authority mismatch
may be questionable.

Pachynematus montivagus Marlatt
N. H.

Pachynematus montivagus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 101. ♀.

Pachynematus obductus (Hartig)
Mass. to B. C. s. to N. Y., Colo., Nev.; Greenland; Iceland, Europe to

Siberia. Ecology: Widespread. Host: Gramineae.
Nematus obductus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung

Naturgesch. Hym., p. 201. ♀, ♂.
Neynatus conductus Ruthe, 1859. Stettin Ent. Ztg. 20: 306. ♀.
Nematus grayninis Cameron, 1874. Ent. Monthly Mag. 10: 221. ♀.
Nematus palliventris Cresson, 1880. Amer. Ent. Soc, Trans. 8: 5. ♀.
Pachynematus boulderensis Rohwer, 1920. U. S. Natl. Mus., Proc 57: 218. ♀.




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Pachynematus rufocinctus MacGillivray
Conn.

PachynematHs rufocinctus MacGillivray, 1916. Conn. State Geol. and Nat. Hist. Survey
Bui. 22: 117. ♀, ♂.

Pachynematus vagus (Fabricius)
Ont. to Alta., Idaho s. to N. Y., 111., Minn., Mont.; Europe to Siberia. Ecology:
Widespread. Host: Carex, Salix.

Tenthredo vaga Fabricius, 1781. Species Insectorum, v. 1, p. 415.

Nematus leucogaster Hartig, 1840. Stettin Ent. Ztg. 1: 23.

Nemahis inconspicuus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 141. ♀.

Pachynematus corticosus MacGillivray, 1901. N. Y. State Mus., Bui. 47: 584. ♀.

Pachynematus roscidus MacGillivray, 1901. N. Y. Ent. Soc, Jour. 29: 31. ♀.

Unplaced Taxa of Group Vagas

Pachynematus abdominalis Marlatt
Wash.

Pachynematus abdoinmalis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 104. ♀.

Pachynematus carolinensis Marlatt
N. C.

Pachynematus carolinensis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 109. ♂.

Pachynematus koebelei Marlatt
Oreg.

Pachynematus koebelei Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 108. ♂.

Pachynematus robustiformis Rohwer
Colo.

Pachynematus robustiforrnis Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 108. ♀.

Pachynematus robustus Marlatt
Mont.

Pachyyiematus robustus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 102. ♀.

Species Group Rumicis

Pachynematus freyi Lindqvist
N. W. T., Yukon; Europe to Siberia.

Pachynematus freyi Lindqvist, 1949. Notulae Ent. 28: 81. ♀.

Pachynematus rumicis (Linnaeus)
Man., Alaska; Iceland, Europe to Siberia. Host: Rumex.

Tenthredo rumicis Linnaeus, 1758. Syst. Nat., Ed. 10, v. 1, p. 559.

Ne7natus flavipennis Cameron, 1876. Ent. Monthly Mag. 12: 189. ♀.

Genus PIKONEMA Ross

Pikonema Ross, 1937. 111. Biol. Monog. 15(2): 86.

Type-species: Nematus dimmockii Cresson. Orig. desig.

Larvae of this Nearctic genus feed on species of Picea.
Revision: Ross, 1938. Ent. Soc. Wash., Proc. 40: 17-20 (adults and larvae).
Taxonomy: Wong, 1951. Ont. Ent. Soc, Ann. Rpt. 1951, p. 65 (cocoons).

Pikonema alaskensis (Rohwer)
Newfoundland to Alaska s. to Mass., Mich., Minn., Wyo., Idaho, B. C.

Ecology: Widespread. Host: Picea canadensis, P. engelmannii, P. excelsa, P. glauca, P.
mariana, P. rubens, P. pungens, P. sitchensis, P. abies. Yellowheaded spruce sawfly.
Pachynematus alaskensis Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 387. ♀, ♂.
Pachynematus piceae Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 387. ♀, ♂.

Biology: Brown, 1938. Ont. Ent. Soc, Ann. Rpt. 1937, p. 14. —Nash, 1939. Jour. Econ. Ent.
32: 330-334. —Brown, 1941. Canad. Dept. Agr., Pub. 712, p. 6. — Schaffner, 1943. Jour.
Forestry 41: 585. —Reeks and Smith, 1945. Acad. Nat. 2: 11. —Bradley, 1951. Canad. Ent.
83: 130. (parasites). —Wilson, 1962. U. S. Dept. Agr., Forest Pest Leaflet 69, 4 pp.

Pikonema dimmockii (Cresson)
Newfoundland to N. W. T., B. C. s. to N. B., N. H., Man., Colo., Wyo.,
Idaho. Ecology: Widespread. Host: Picea canadeiisis, P. engelmannii, P. glauca, P.
mariajia, P. rubens, P. sitchensis, P. abies. Greenheaded spruce sawfly.
Nematus dimmockii Cresson, 1880. Amer. Ent. Soc, Trans. 8: 6. ♀.
Nematus ochreatus Harrington, 1889. Canad. Ent. 21: 95. ♀.

Biology: Brown, 1938. Ont. Ent. Soc, Ann. Rpt. 1937, p. 14. —Brown, 1941. Canad. Dept.
Agr., Pub. 712, p. 6. —Reeks and Smith, 1945. Acad. Nat. 2: 11.

Pikonema ruralis (Cresson)
Oreg., Calif., Nev.

Nematus ruralis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 5. ♀.




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Genus EITELIUS Kontuniemi

Eitelius Kontuniemi, 1966. Ann. Ent. Fenn. 31: 46.

Type-species: Pachynematus dentatus Lindqvist. Orig. desig.

Prior to 1967, the species below was placed in the genus M icronematus Konow which is now a
synonym of Pristiphora.

Taxonomy: Wong, 1967. Canad. Ent. 99: 1101-1104.

Eitelius gregarius (Marlatt)
N. S., Mass., Conn., N. J. Host: Salix.

Pachynematus gregarius Marlatt, 1896. Canad. Ent. 28: 256. ♀, ♂.
Pachynematus varus MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 30. ♀.
Pontania decrepita MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 33. ♀.

Genus PRISTIPHORA Latreille

Pristiphora Latreille, 1810. Consid. Generales Crust. Arachn. Ins., p. 294.

Type-species: Pieronus testaceus Jurine. Orig. desig.
Diphadnus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung
Naturgesch. Hym., p. 225.

Type-species: N ematus fuscicomis Hartig. Desig. by Rohwer, 1911.
Lygaeonematus Konow, 1890. Deut. Ent. Ztschr., pp. 233, 238, 247.

Type-species: Nematus pini Retzius. Desig. by Rohwer, 1911.
Micronematus Konow, 1890. Deut. Ent. Ztschr., pp. 233, 239, 248.

Type-species: Neynatus pullus Foerster. Desig. by Rohwer, 1911.
Gymnonychus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 122.

Type-species: Gymnonychus califomicus Marlatt. Orig. desig.
Neotomostethus MacGillivray, 1908. Canad. Ent. 40: 290.

Type-species: Neotomostethus hyalinus MacGillivray. Orig. desig.
Pristiphora subg. Sala Ross, 1937. 111. Biol. Monog. 15(2): 85. Preocc. by Walker, 1867.

Type-species: Nematus chloreus Norton. Orig. desig.
Lygaeonematus subg. Lygaeotus Lindqvist, 1952. Notulae Ent. 32: 82. Invalid, type-species

not designated.
Lygaeonematus subg. Lygaeophora Lindqvist, 1952. Notulae Ent. 32: 82. Invalid,
type-species not designated.

A large, nearly cosmopolitan genus with diverse host plants. Two species from southern Brazil
are the only known South American Nematinae. Several phyletic lines are evident in this genus
which will eventually allow separation into more convenient species groupings.

Revision: Marlatt, 1896. U. S. Dept. Agr. Div. Ent. Tech. Ser. 3: 113-125.
Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 394-396 (holarctic species).

Pristiphora abbreviata (Hartig)
Ont., Conn, to Va. w. to Wis., Kans., Colo., Utah; B. C, Wash., Oreg.,

Calif.; Europe. Ecology: A pest of pear trees. Larva feeds on foliage. Introduced. Host:
Pyrus communis. California pearslug.
Nematus abbreviatus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung

Naturgesch. Hym., p. 205. ♀.
Gymnonychus califomicus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 122. ♀.

Taxonomy: Smith, 1967. Ent. Soc. Wash., Proc. 69: 92 (distribution). —Wong, 1967. Canad.
Ent. 99: 1101 (compared with species of Eitelius).

Biology: Nougaret, Davidson, and Newcomer, 1916. U. S. Dept. Agr., Bui. 438, 24 pp.

Morphology: Zocchi, 1948. Bol. Ent. Bologna 17: 225-232 (external structure of adult and
larva).

Pristiphora acidovalva Wong
N. B., Que. w. to B. C, Calif. Host: Salix.

Pristiphora acidovalva Wong, 1969. Canad. Ent. 101: 970. ♀, ♂.

Biology: Lindquist and Miller, 1969. Canad. Ent. 101: 973-975 (also description of larva).

Pristiphora aphanta Wong and Ross
Que., Ont., N. Y.

Pristiphora aphanta Wong and Ross, 1960. Canad. Ent. 92: 196. ♀.




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Pristiphora aquilegiae (Vollenhoven)
Ont.; Europe. Introduced. Host: Aquilegia.
Nematus aquilegiae Vollenhoven, 1866. Tidskr. v. Ent. 11: 202. ♀.

Biology: McNay, 1963. Canad. Ins. Pest Rev. 41: 103. — McNay, 1964. Phytoprotection 45: 91.

Pristiphora banksi Marlatt
N. Y. to N. C; Wyo. Host: Vacciniiim.

Pristiphora bayiksi Marlatt, 1896. U. S. Dept. Agr. Div. Ent. Tech. Ser. 3: 117. 9 (male
misdet.).

Pristiphora bivittata (Norton)
Prince Edward Is., N. B. w. to B. C. s. to N. C, III, Iowa, Oreg. Ecology:
Widespread. Host: Spiraea.
Nematus bivittatus Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 158. ♀.
Neviatus luteolus Norton, 1867. Amer. Ent. Soc, Trans. 1: 200. ♂.

Pachynematus occidentalis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 109. ♂.
Pristiphora dyari Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 118. ♀. N. syn.
Pristiphora koebelei Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 119. ♀.
Pristiphora occidentalis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 121. ♂.

Preocc. by occidentalis Marlatt on p. 109 of same publication.
Periclista patchi MacGillivray, 1923. 111. Univ. Bui. 20(50): 29. ♀.

Biology: Neilson, 1958. Canad. Ent. 90: 229-234.

Pristiphora borea (Konow)
Labrador, Que. w. to Alaska, B. C; Europe. Host: Betula, Salix.
Lygaeonematus boreus Konow, 1904. Ztschr. System. Hym. Dipt. 4: 253. ♀, ♂.
Nematus astutus Cameron, 1885. Monog. Brit. Phytophagous Hym., v. 2, p. 77. ♀.

Synonymy may be questionable.
Pachynematus lappo^iicus Enslin, 1916. Deut. Ent. Ztschr., Beih., p. 463. ♀, ♂.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 395.

Pristiphora breadalbanensis (Cameron)
Que., N. W. T., Man., Colo., Alta., Yukon, Alaska, B. C; Europe.

Nematus breadalbanensis Cameron, 1882. London Ent. Soc, Trans., p. 531.

Nematus Whitei Cameron, 1878. Fauna Scotland, v. 1, p. 35. 6. Synonymy may be
questionable.

Lygaeonematus tromsoensis Kiaer, 1898. Tromsoe Mus. Aarsh. 19: 48.

Lygaeonematus corpulentus Konow, 1904. Ztschr. System. Hym. Dipt. 4: 252. ♀, ♂.

Lygaeonematus arcticola Enslin, 1916. Deut. Ent. Ztschr., Beih., p. 514. ♀, ♂.

Pristiphora cadma Wong and Ross
Ont., Sask., Alaska, B. C. Host: Betula.

Pristiphora cadma Wong and Ross, 1960. Canad. Ent. 92: 198. ♀.

Pristiphora carolinensis Marlatt
N. C.

Pristiphora carolinensis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 120. ♂.

Possibly belongs in Pachynematus.

Pristiphora chlorea (Norton)
N. B. to Fla. w. to Man., Oreg., Tex. Host: Quercus.
Nematus chloreus Norton, 1867. Amer. Ent. Soc, Trans. 1: 221. ♀.
Nematus pallifrons Cresson, 1880. Amer. Ent. Soc, Trans. 8: 6. ♂.
Nematus decoratus Provancher, 1888. Addit. Corr. Faune Ent. Canada Hym., p. 349. ♂.
Pteronus quercus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 67. ♀. Preocc. in

Pristiphora by Hartig, 1837.
Nematus pergandei Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 90. ♀.
Amauronematus vacalus MacGillivray, 1923. 111. Univ. Bui. 20(50): 6. ♀. N. syn.

Pristiphora cincta Newman
Newfoundland w. to N. W. T., Alaska s. to Ala., Tenn., Mich., Minn., Colo., B.

C; Europe. Ecology: Widespread. Host: Betula, Vaccinium, Salix.
Pristiphora ciyicta Newman, 1837 (January). Ent. Mag. 4: 259.
Nemattis quercus Hartig, 1837 (March). Fam. Blattwespen und Holzwespen, nebst

Einleitung Naturgesch. Hym., p. 188.
Tenthredo borealis Zetterstedt, 1838. Ins. Lapponica 1: 353. ♀.
Nemattis idiota Norton, 1867. Amer. Ent. Soc, Trans. 1: 177. ♂.
Nematus identidem Norton, 1867. Amer. Ent. Soc, Trans. 1: 177. ♀.
Pristiphora hoodi Marlatt, 1896. U. S. Dept. Agr. Div. Ent. Tech. Ser. 3: 119. 9. Synonymy

questionable.
Pristiphora coloradensis Marlatt, 1896. U. S. Dept. Agr. Div. Ent. Tech. Ser. 3: 121. ♂.

Synonymy questionable.
Pristiphora idiotiformis Rohwer, 1910. U. S. Natl. Mus., Proc 38: 199. ♀.




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Pristiphora cincta ah. nigriventris Hellen, 1943. Notulae Ent. 23: 71

Biology: Neilson, 1958. Canad. Ent. 90: 229-234 (idiota on Vaccinium).

Pristiphora coactula (Ruthe)
Labrador w. to N. W. T., Alaska; Colo.; Europe. Host: Salix, Vacciniumi?).
Nematus coactulus Ruthe, 1859. Stettin Ent. Ztg. 20: 307. ♀.
Nematus Winnipeg Norton, 1867. Amer. Ent. Soc, Trans. 1: 198. ♀.
Nematus alpinus Thomson, 1871. Hym. Scand., v. 1, p. 98. ♀, ♂.
Lygaeonematus pachyvalvis Konow, 1904. Ztschr. System. Hym. Dipt. 4: 253. ♀, ♂.
Lygaeonematus winnipegensis Konow, 1905. In Wytsman, Gen. Ins., v. 29, p. 68. Emend, of

Winnipeg.
Pristiphora dawsoni Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 109. ♀.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 395.

Pristiphora elaphita Wong and Ross
Sask., N. Mex., Alta., B. C, Calif.

Pristiphora elaphita Wong and Ross, 1960. Canad. Ent. 92: 194. ♀.

Pristiphora erichsonii (Hartig)
Newfoundland to Alaska s. to Md., W. Va., Mich., Ill, Minn., Mont, Idaho,

Oreg.; Europe to Japan. Ecology: Widespread. Possibly introduced. Host: Larix laricina,

L. occidentalis, L. lyallii, and European and Asian larch species and hybrids. Larch

sawfly. Numerous papers have appeared on this destructive species; therefore, it is

feasible to give only selected references below.
Nematus Leachei Dahlbom, 1835. Conspectus Tenthred., Siricid., Oryssin. Scand., p. 10.

Nom. nud.
Nematus Leachii Dahlbom, 1835. Clavis Hym., p. 27. Larva. Suppressed by Internatl.

Comm. Zool. Nomencl., Opinion 906, 1970, Bui. Zool. Nomencl. 27: 8.
Nematus erichsonii Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung

Naturgesch. Hym., p. 187. ♀.
Nematus notabilis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 7. ♀.

Taxonomy: Coppel and Leius, 1958. 10th Internatl. Cong. Ent., Proc 1: 231-238 (variations in
populations from Canada, (ilreat Britain, and Japan). —Wong, 1974. Canad. Ent. 106:
1121-1131 (identification and origin of strains in N. Amer.).

Biology: Ives, 1955. Canad. Ent. 87: 301-311 (moisture and selection of cocoon sites). —Ives,
1955. Canad. Jour. Zool. 33: 370-388 (estimation of egg populations). — Drooz, 1956. U. S.
Dept. Agr., Forest Pest Leaflet 8, 4 pp. (rev. 1967). -Graham, 1956. Forest Sci. 2: 132-160
(Lake states). — Ruppel, 1958. Ent. Soc. Brit. Columbia, Proc. 55: 32-35 (history in B. C).
— Buckner, 1958. 10th Internatl. Cong. Ent., Proc. 4: 353-361 (mammal predators). —Ives
and Prentice, 1958. Canad. Ent. 90: 331-338 (sampling techniques). —Ives and Prentice,
1959. Canad. Ent. 91: 496-500 (estimates of parasitism). —Graham and Satterlund, 1959.
Forest Sci. 5: 28-36 (weather and water levels on cocoons). —Ives and Turnock, 1959.
Canad. Ent. 91: 650-661 (estimates of cocoon populations). —Buckner, 1959. Canad. Ent.
91: 275-282 (cocoon predation by mammals). —Turnock, 1960. Canad. Ent. 92: 500-516.
—Drooz, 1960. U. S. Dept. Agr., Tech. Bui. 1212, 52 pp. (biology and control; references to
most of literature up to 1956). —Turnock, 1960. Canad. Ent. 92: 659-662 (estimation of
adult populations). —Maw, 1960. Ent. Gaz. 11: 43-49 (Great Britain). —Ives, 1960. Canad.
Ent. 92: 668-674. —Heron, 1961. Canad. Ent. 93: 431-433 (temperature and post-diapause
development). —Nairn, et ai, 1962. Canad. Ent. 94: 242-255 (outbreaks and effects on
forest stands). —Turnock and Ives, 1962. Canad. Ent. 94: 897-902 (cocoon mortality).
—Ives, 1962. Canad. Ent. 94: 256-268 (mortality of eggs and larvae). —Ives, 1963. Canad.
Ent. 95: 887-892 (defoliation effect on larval survival). — Pschorn-Walcher and Eichhorn,
1963. Comm. Inst. Biol. Control, Tech. Bui. 3: 51-82 (ecology and natural control in central
Europe). — Frilli, 1964. Milan Univ. Cattolica Sacro Cuore Fac Agr., Ann. 4: 528-586
(morpho-biological studies). —Buckner and Turnock, 1965. Ecology 46: 223-226 (avian
predators). —Ives and Nairn, 1966. Canad. Ent. 98: 768-777 (water levels and
overwintering survival). — Schmiege, 1966. Canad. Ent. 98: 671-672 (Alaska). —Smirnoff,
1966. Jour. Invert. Path. 8: 360-364 (a microsporidian parasite). —Turnock and McLeod,
1966. Ent. Soc Manitoba, Proc 22: 55-60 (n. Man.). —Heron, 1966. Canad. Ent. 98: 561-578
(reproductive capacity). —Ives, 1967. Canad. Ent. 99: 1121-1131 (mortality causes).
—Heron, 1967. Canad. Ent. 99: 1150-1155 (heat tolerance of larvae). —Elliott and




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Muldrew, 1967. Canad. Ent. 99: 321-323 (rearing cage). — Muldrew, 1967. Canad. Ent. 99:
312-321 (biology and dispersal of an introduced parasite). —Ives, 1967. Canad. Ent. 99:
607-622 (invertebrate predators and prey). —Ives, 1968. Manitoba Ent. 2: 5-36 (population
dynamics techniques). —Ives, 1968. Canad. Ent. 100: 373-385 (survival and water levels).
—Smirnoff, 1968. Jour. Invert. Path. 10: 417-424 (diseases). —Smirnoff and Chu, 1968.
Jour. Invert. Path. 10: 12 (microsporidian infection). —Heron, 1968. Canad. Ent. 100:
470-475 (dyes as markers for behavioral studies). — Turnock and Muldrew, 1968. Forest
Chronicle 44: 38 (control by introduced parasite). — Harman, 1970. Chesapeake Sci. 11:
183-190 (Md.). —Harman, 1971. Chesapeake Sci. 12: 40-44 (natural mortality of feeding and
cocooning stages). —Heron, 1971. Canad. Ent. 103: 1153-1155 (temperature tolerance of
pronymphs and pupae).

Morphology: Wong, 1956. Canad. Ent. 88: 545 (gynandromorphs and intersexes). —Wong,
1963. Canad. Ent. 95: 897-920 (external morphology of male, female, and larva).

Pristiphora gelida Wong
N. W. T., Yukon, Alaska, Alta., Colo.

Pristiphora gelida Wong, 1968. Jour. Nat. Hist. 2: 185. ♀, ♂.

Pristiphora geniculata (Hartig)
Newfoundland, Ont. s. to W. Va.; Europe to Siberia. Introduced. Host:
Sorbus. Mountain ash sawfly.
Nematus geniculatus Hartig, 1840. Stettin Ent. Ztg. 1: 26. ♀.
Nemahis cheilon Zaddach, 1883. Phys. Okonom. Gesell. Konigsberg, Schr. 24: 145. 9, tJ.

Biology: Schaffner, 1936. Jour. Econ. Ent. 29: 469. —Forbes and Daviault, 1964. Canad. Ent.
96: 1117-1133. —Smirnoff, 1968. Jour. Invert. Path. 10: 436-437 (virus disease).

Pristiphora gronblomi (Lindqvist)
Alaska; N. Europe.

Lygaeotus gronblomi Lindqvist, 1952. Notulae Ent. 32: 101. ♀, ♂.

Pristiphora hucksena Wong and Ross
B. C.

Pristiphora hucksena Wong and Ross, 1960. Canad. Ent. 92: 104. ♀.

Pristiphora hyalina (MacGillivray)
N. Y.

Neotomostethus hyalinus MacGillivray, 1908. Canad. Ent. 40: 290. ♀.

Pristiphora jocularis Cresson
Oreg., Nev.

Pristiphora jocularis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 3. ♀, ♂.

Pristiphora labradoris (Norton)
Labrador, Ont., N. W. T., Man., Alta., Yukon.

Nematus labradoris Norton, 1867. Amer. Ent. Soc, Trans. 1: 196. ♀.

Pristiphora lata (Cresson)
Oreg., Nev.

Ne^natus latus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 4. ♀.

Pristiphora lativentris (Thomson)
N. W. T., Man., Alaska; Europe.

Nematus lativentris Thomson, 1871. Hym. Scand., v. 1, p. 99. ♀, ♂.

Nematus scoticus Cameron, 1881. Ent. Soc. London, Proc, p. 365. ♀.

Nematus extremus Holmgren, 1883. Ent. Tidskr. 4: 148. ♀, ♂.

Pristiphora buccoda Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 350. ♀.

Lygaeonematus apicola Konow, 1904. Ztschr. System. Hym. Dipt. 4: 254. 9,6. Synonymy
of apicola may be questionable.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 395.

Pristiphora leechi Wong and Ross
Mont., Idaho, B. C, Wash. Host: Larix occidentalis.

Pristiphora leechi Wong and Ross, 1960. Canad. Ent. 92: 193. ♀, ♂.

Pristiphora lena Kincaid
Newfoundland to N. W. T., Alaska; Wyo., Colo. Host: Picea.

Pristiphora lena Kincaid, 1900. Wash. Acad. Sci., Proc 2: 351. ♂.

Pristiphora maura Rohwer
Que., N. W. T., Yukon, Alaska, Man., Alta., Colo.

Pristiphora maura Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 109. ♂.

Pristiphora melanocarpa (Hartig)
Que., N. W. T., Yukon, Alaska, Man.; Europe. Host: Betula.

Nematus ynelanocarpus Hartig, 1840. Stettin Ent. Ztg. 1: 27. ♀.

Nematxis wustneii Stein, 1885. Wien. Ent. Ztg. 4: 304. ♀, ♂.

Pristiphora ortinga Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 349. ♀. N. syn.

Pristiphora micronematica
***authority mismatch
Malaise. N. W. T., Man.; Europe to Siberia. Host: Salix.

Pristiphora microneynatica Malaise, 1932. Arkiv for Zool. 23A: 58. ♀.

Lygaeonematus (Lygaeophora) leucostoma Lindqvist 1952. Notulae Ent. 32: 108. ♀, ♂.




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Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 394. — Lindqvist, 1971.
Notulae Ent. 51: 128-129.

Pristiphora mollis (Hartig)
Labrador to N. W. T., Alaska s. to N. C, Colo.; Europe. Host: Vaccinium.
Nematus mollis Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung

Naturgesch. Hym., p. 201.
Pachynematus orarius Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 348. ♀.
Pachyyiematus penegalensis Enslin, 1911. Deut. Ent. Ztschr. 2: 176.9.
Lygaeonematus mollis var. albipes Lindqvist, 1942. Notulae Ent. 22: 107.
Lygaeonematus mollis var. rufonotata Lindqvist, 1942. Notulae Ent. 22: 107.
Pachynematus kontkaneni Lindqvist, 1960. Notulae Ent. 40: 35. ♀.

Pristiphora nigra Marlatt
Wash.

Pristiphora nigra Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 114. ♀.

Pristiphora paedida (Konow)
Alaska; Europe to Siberia.

Lygaeonematus paedidus Konow, 1904. Ztschr. System. Hym. Dipt. 4: 253.

Pristiphora kamtchatica Malaise, 1932. Arkiv for Zool. 23A: 56. ♀, ♂.

Pristiphora pallidiventris (Fallen)
Labrador to Alaska s. to Md., Mich., Iowa, Calif.; Europe to Japan.

Ecology: Widespread. Host: Rubus, Geum, Potentilla, and other Rosaceae. These hosts
were recorded in Europe.

Nematus pallidiventris Fallen, 1808. Svenska Vetensk. Akad., Handl. 29: 120. ♀.

Pristiphora myosotidis Stephens, 1835. Illus. Brit. Ent., v. 7, p. 25.

Nematus ephippiger Hartig, 1840. Stettin Ent. Ztg. 1: 24. ♀.

Nematus flavicomus Tischbein, 1846. Stettin Ent. Ztg. 7: 77. ♀.

Nematus nigricans Eversmann, 1847. Soc. Nat. Moscou, Bui. 20: 16. ♀.

Nematus caudalis Eversmann, 1847. Soc. Nat. Moscou, Bui. 20: 16. ♂.

Nematus breviusculus Eversmann, 1847. Soc. Nat. Moscou, Bui. 20: 17. ♀.

Nematus gemellus Foerster, 1854. Naturh. Ver. Rheinlande, Verh. 11: 425. ♀.

Nematus cirrhostomus Zaddach, 1882. Phys. Okonom. Gesell. Konigsberg., Schr. 23: 195. ♀.

Pristiphora zella Rohwer, 1909. Canad. Ent. 41: 20. ♀. N. syn.

Pristiphora pallicoxa Rohwer, 1910. U. S. Natl. Mus., Proc. 38: 200. ♀. N. syn.

PristophoraV.) xanthotrachela Rohwer, 1913. U. S. Natl. Mus., Proc. 45: 281. ♀. N. syn.

Pristiphora pallidiventris var. haemorrhoidalis Enslin, 1916. Deut. Ent. Ztschr., Beih., p.
526. ♀, ♂.

Pristiphora pallidiventris var. stigmatica Enslin, 1916. Deut. Ent. Ztschr., Beih., p. 526. ♀.

Pristiphora ostiaria MacGillivray, 1920. Canad. Ent. 52: 236. ♀, ♂. N. syn.

Nematus (Pristiphora) pallidiventris var. flaviapex Hellen, 1948. Notulae Ent. 28: 45.

Nematus (Pristiphora) pallidiventris var. nigrofemoratus Hellen, 1948. Notulae Ent. 28:
45.

Pristiphora paloma Wong and Ross
N. Y. to Fla. w. to HI.

Pristiphora paloma Wong and Ross, 1960. Canad. Ent. 92: 196. ♀.

Pristiphora parbeta Wong and Ross
Man., Alaska. Host: Betula papyrifera.

Pristiphora parbeta Wong and Ross, 1960. Canad. Ent. 92: 198. ♂.

Pristiphora pseudocoactula (Lindqvist)
Man. (Churchill); Europe.

Lygaeonematus (Lygaeonematus) pseudocoactula Lindqvist, 1952. Notulae Ent. 32: 115. ♀.

i.

Pristiphora relativa Norton
N. W. T.

Pristiphora relativa Norton, 1867. Amer. Ent. Soc, Trans. 1: 177. ♀.

Pristiphora reuteri (Lindqvist)
Maine, N. H., Man.; Europe. Host: Salix.

Lygaeonematus (Lygaeophora) reuteri Lindqvist, 1960. Notulae Ent. 40: 33. ♀, ♂.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 395. — Vikberg, 1966. Ann.
Ent. Fenn. 32: 252-253 (larva).
ruflpes Lepeletier. Newfoundland to B. C. s. to N. C, Tenn. Mo., Colo., Utah, Calif.; Europe to
Siberia. Ecology: Widespread. Host: Ribes.
Pristiphora pallipes Lepeletier, 1823. Monog. Tenthred., p. 60. ♀. Preocc. in Pristiphora by

Fallen, 1808.
Pristiphora rufipes Lepeletier, 1823. Monog. Tenthred., p. 60. ♀.
Pristiphora fusca Lepeletier, 1823. Monog. Tenthred., p. 60.



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Tenthredo (Neviatus) pallicomis Harris, 1835. hi Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Tenthredo (Neynatus) labrata Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Nematus appendiculatus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst

Einleitung Naturgesch. Hym., p. 202. ♀.
Nematus (Diphadnus) fiiscicomis Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst

Einleitung Naturgesch. Hym., p. 225. ♀.
Nematus cathoraticus Foerster, 1854. Naturh. Ver. Rheinlande, Verh. 11: 325. ♀.
Nematus pallicomis Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 160. ♀.
Nematus pallicomis var. labratus Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 160. ♀.
Nematus vitreipennis Kawall, 1864. Soc. Nat. Moscou, Bui. 37: 295. ♀.
Pristiphora grossulariae Walsh, 1866. Pract. Ent. 1: 123. 9,6. larva.
Nematus peletieri Andre, 1880. Spec. Hym. Eur. Alg. 1: 5. ♀.

Nematus pumilus Zaddach, 1883. Phys. Okonom. Gesell. Konigsberg., Schr. 24: 172. ♂.
Nematus hypobalius Zaddach, 1884. Phys. Okonom. Gesell. Konigsberg., Schr. 25: 154. ♀.
Nematus ghilianii Costa, 1894. Prosp. Imen. Ital. 3: 73.

Biology: Miles, 1932. Bui. Ent. Res. 23: 10-15.

Pristiphora serrula Wong and Ross
Maine, N. H., B. C, Wash., Oreg., Calif.

Pristiphora semda Wong and Ross, 1960. Canad. Ent. 92: 196. ♀.

Pristiphora siskiyouensis Marlatt
N. S. to N. W. T., Yukon, Alaska s. to Va., Wis., Calif. Ecology:
Widespread. Host: Betula, Alnus
Pristiphora siskiyouensis Marlatt, 1896 U. S. Dept. Agr. Div. Ent. Tech. Ser. 3: 116. ♀, ♂.
Pristiphora betulavora Rohwer, 1920. U. S. Natl. Mus., Proc. 57: 219. ♀, ♂.

Taxonomy: Middleton, 1922. U. S. Natl. Mus., Proc. 61: 24 (larva).

Biology: Wong, 1954. Canad. Ent. 86: 154.

Pristiphora staudingeri (Ruthe)
Labrador to Alaska s. to Man., Colo., Alta.; Europe to Siberia. Host:
Salix.
Nematus staudingeri Ruthe, 1859. Stettin Ent. Ztg. 20: 306. ♀, ♂.
Nematus circularis Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 350. ♀. N. syn.
Pristiphora hyperborea Malaise, 1921. Ent. Tidskr. 42: 11. ♀, ♂.
Pristiphora asperlatus Benson, 1935. Roy. Ent. Soc. London, Trans. 83: 36. ♀, ♂.

Pristiphora sycophanta Walsh
Labrador to Alaska s. to Conn., Va., Mo., Colo., Oreg. Ecology: Widespread.
Host: Salix.
Pristiphora sycophanta Walsh, 1866. Ent. Soc. Phila., Proc. 6: 263. ♀.
Pristiphora tibialis Norton, 1867. Amer. Ent. Soc, Trans. 1: 76. ♀.
Nematus trivialis Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 140. Unnecessary n. name for

Pristiphora tibialis Norton

Pristiphora murtfeldtiae Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 117. ♀.
Pristiphora anaka Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 350. ♀. N. syn.

Pristiphora valvangula Wong and Ross
N. Y. (?)

Pristiphora valvangula Wong and Ross, 1960. Canad. Ent. 92: 194. ♂.

Pristiphora venatta Wong and Ross
Idaho.

Pristiphora venatta Wong and Ross, 1960. Canad. Ent. 92: 196. ♀.

Genus CROESUS Leach

Croesus Leach, 1817. Zool. Misc., v. 3, p. 129.

Type-species: Tenthredo septentrionalis Linnaeus. Monotypic.

A small holarctic genus characterized by the enlarged, flattened hind basitarsus. Larvae feed
externally on the foHage of the host plant.

Revision: Benson, 1963. Ent. Tidskr. 84: 25 (world species groups). —Smith, 1972. Ent. Soc.
Wash., Proc. 74: 169-180 (Nearctic species, adults and larvae).

Croesus castaneae Rohwer
N. Y., Pa., Md., Va., Ky. Host: Castanea dentata.

Croesus castaiieae Rohwer, 1915. U. S. Natl. Mus., Proc. 49: 213. ♀, ♂.




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Taxonomy: Middleton, 1922. U. S. Natl. Mus., Proc. 61: 14 (larva).

Croesus curvarius Smith
Conn. Host: Corylus.

Croesus curvarius Smith, 1972. Ent. Soc. Wash., Proc. 74: 175. 9, larva.

Croesus latitarsus Norton
Que. to Fla. w. to Alaska, B. C, Utah. Host: Betula. Dusky birch sawfly.

CraesusV.) latitarsus Norton, 1862. Ent. Soc. Phila., Proc. 1: 199. ♀.

Biology: Jack, 1888. Psyche 5: 41. —Brown, 1940. Canad. Dept. Agr., Forest Ins. Survey 4th
Ann. Rpt., p. 16. —Wong, 1954. Canad. Ent. 86: 154.

Croesus varus (Villaret)
Que. (Montreal); Europe. Introduced (?). Host: Ahius.
Nematus varus Villaret, 1832. Soc. Ent. France, Ann. 1: 306. ♂.

Taxonomy: Buckle, 1930. Canad. Ent. 62: 21 (record of only known specimen taken in N.
Amer.).

Genus NEMATUS Panzer

Nematus Panzer, 1801. Faune Ins. German., heft 82, pi. 10.

Type-species: Tenthredo (Nematus) lucida Panzer. Monotypic.
Hypolaepus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 324.

Type-species: Hypolaepus abbotii Kirby. Monotypic.
Holcocneme Konow, 1890. Deut. Ent. Ztschr., pp. 233, 238, 246.

Type-species: Tenthredo crassa Fallen. Desig. by Rohwer, 1911.
Pteronidea Rohwer, 1911. U. S. Dept. Agr. Bur. Ent. Tech. Ser. 20: 98.

Type-species: Nematus ventralis Say. Orig. desig.

A large holarctic genus associated with various trees and shrubs. The larvae are external leaf
feeders.

Revision: Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 44-75, 87-90.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent, Bui. 12: 404-405 (holarctic species).

Species Group Abbotii

Nematus abbotii (Kirby)
Ont. to Ga. w. to Wis., 111. Host: Robinia.

Hypolaepus abbotii Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 324. ♀.

Nematus crassus (Fallen)
N. W. T. (Mackenzie River, Reindeer Depot); Eurasia. Host: Betula, Populus,
Salix, Rumex. These hosts recorded in Europe.

Tenthredo crassa Fallen, 1808. Svenska Vetensk. Akad., Handl. 29: 106.

Nematus sulcipes Hartig, 1840. Stettin Ent. Ztg. 1: 23. ♀, ♂.

Nematus longispinis Kriechbaumer, 1885. Correspbl. Naturh. Ver. Regensburg 39: 13.

Nematus laticulus (Norton)
N. Y. to N. C. w. to Wis., S. Dak., 111.

Croesus laticulus Norton, 1869. Amer. Ent. Soc, Trans. 2: 368. ♀.

Neinatus procidentius Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 245. ♂.

Amauronematus rufipes Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 78. ♂.

Species Group Ventralis

Nematus chalceus (Marlatt)
Mont, Alaska, B. C, Wash. Host: SalixiO, Alnus(^).

Ammironematus chalceus Marlatt 1896. U. S. Dept. Agr., Div. Ent Tech. Ser. 3: 84. ♀.

Nematus erratus (MacGillivray)
Maine, Man., Alaska. Host: Populus.

Pteronidea erratus MacGillivray, 1923. N. Y. Ent Soc, Jour. 31: 170. ♀.

Nematus exactus (MacGillivray)
Maine.

Pteronidea exactus MacGillivray, 1920. Canad. Ent. 52: 235. ♂.

Amauronematus vanus MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 27. ♀.

Amauronematus venaticus MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 29. ♂.

Nematus fulvicrus Provancher
Que. to N. Y. w. to Alta., Colo. Host: Salix, Populus{i).

Nematus fulvicrus Provancher, 1882. Nat. Canad. 13: 291. ♀.

Messa salicis Ashmead, 1890. Colo. Biol. Assoc, Bui. 1: 15. Preocc. in Nematus by
Linnaeus, 1758.

Nematus salicicola Dalla Torre, 1894. Cat. Hym., v. 1, p. 257. N. name for salicis
Ashmead.

Pteronidea evanida MacGillivray, 1920. Canad. Ent. 52: 233. ♀, ♂.




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Nematus hudsoniimagnus Dyar
Maine to N. J. w. to Alaska, Man., Ill, Tex. Host: Populus.

Neynatus Htidsoiiii magnus Dyar, 1895. Amer. Ent. Sec, Trans. 23: 306. 9, larva. This
name is available under Article 26(A) of the Internatl. Code Zool. Nomencl.

Pteronus populi Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 59. ♀.

Pteronus hudsonii Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 59. 9. Emend.

Pterojius lombardae Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 73. ♂.

Pteronidea edura MacGillivray, 1920. Canad. Ent. 52: 233. ♀.

Pteronidea effusa MacGillivray, 1920. Canad. Ent. 52: 233. ♀.

Pteronidea effeta MacGillivray, 1920. Canad. Ent. 52: 234. ♀.

Pteronidea emerita MacGillivray, 1920. Canad. Ent. 52: 234. ♀.

Pteronidea enidita MacGillivray, 1920. Canad. Ent. 52: 234. ♀.

Nematus iridescens Cresson
Mont., Wye, Utah w. to B. C, Wash., Oreg., Calif.; Mexico. Host: Salix.

Nematiis iridescens Cresson, 1880. Amer. Ent. Soc, Trans. 8: 5. ♂.

Pteronus occidentalis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 48. ♀. N. syn.

Nematus limbatus Cresson
N. B., Que. to Pa. w. to Sask., Ohio. Host: Salix, Populus.

Nematus limbatus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 8. ♀.

Pteronus harringtoni Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 53. ♀, ♂.

Pteronus fylesi Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3:54. ♀, ♂.

Nematus ventralis Say
Que. to N. C. w. to Alta., Mont., Colo. Host: Salix. Willow sawfly.

Nematus ventralis Say, 1824. In Keating, Narr. Long's 2nd Exped., v. 2 (App.), p. 315. ♀.

Nematus suratus Fitch, 1856. N. Y. Agr. Soc, Trans. 3: 315. ♀.

Pteronus latiis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 48. ♀. Preocc. by
Nematus latus Cresson, 1880.

Hypolaepus viereckii Bradley, 1903. Canad. Ent. 35: 47. ♀.

Pteronidea edita MacGillivray, 1920. Canad. Ent. 52: 235. ♂.

Biology: Howard, 1888. U. S. Dept. Agr., Insect Life 1: 33-37. — Rohwer, 1915. Ent. Soc.
Wash., Proc 17: 197 (mating habits).

Morphology: Judd, 1955. Ont. Ent. Soc, 85th Ann. Rpt. 1954, p. 32 (anatomy and histology of
digestive tract of larva).

Species Group Atriceps

Nematus atriceps (Marlatt)
Wis., Colo., Utah w. to B. C, Wash., Oreg., Calif.

Pteronus atriceps Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 52. ♀.

Pteronus califomicus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 60. ♀. N. syn.

Nematus auratus (Marlatt)
Wash., Oreg., Calif.

Pteronus auratus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 60. ♀.

Amauronematus coquilletti Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 84. ♀.
N. syn.

Nematus coloradensis (Marlatt)
Alta., Mich., N. Dak., Ariz. w. to Oreg., Calif.

Pteronus coloradensis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 52. ♀, ♂.

Species Group Erythrogaster

Nematus carpini (Marlatt)
Conn., N. Y., N. J., Fla. Host: Carpinus.

Pteronus carpini Marlatt, 1898. Canad. Ent. 30: 303. ♀.

Nematus corylus Cresson
Mass. to Md. w. to Mich., 111. Host: Corylus.

Nematus corylus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 8. ♀.

Pteronus coryli Konow, 1905. In Wytsman, Gen. Ins., v. 29, p. 55. Emend.
erythrogaster Norton. Que. to Fla. w. to Minn., 111. Host: Ahius.

Nematus erythrogaster Norton, 1865. Ent. Soc. Phila., Proc 4: 8. ♀.

Pteronus dyari Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 58. ♀, ♂.

Pteronidea pulchella Rohwer, 1913. U. S. Natl. Mus., Proc. 45: 279. ♀.

Pteronidea alnivora Rohwer, 1920. U. S. Natl. Mus., Proc. 57: 213. ♀, ♂.

Taxonomy: Middleton, 1922. U. S. Natl. Mus., Proc. 61: 22 (larva).

Nematus latifasciatus Cresson
N. B. to Pa. w. to Ont., Mich. Host: Betula.

Nematus latifasciatus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 7. ♀.
Nematus rufocinctus Harrington, 1893. Canad. Ent. 25: 58. ♀. N. syn.




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Species Group Magnus

Nematus magnus (Marlatt)
Que., Maine. Host: Salix.

Pteronus magnus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 67. ♀.

Species Group Oligospilus

Nematus attus Smith
n. name. Que., Maine to N. Y. and Mich. Host: Populus.

Amauronematus dyari Marlatt, 1896 (October). Canad. Ent. 28: 255. ♀, ♂. Preocc. by
Pteronus dyari Marlatt, 1896(February), U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 58.

Nematus costillensis (Rohwer)
Colo.

Pteronus costillensis Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 106. ♀.

Nematus effrenatus (MacGillivray)
Alaska.

Pteronidea effrenatus MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 171. ♀.

Nematus excavatus (Marlatt)
n. comb. No. Amer., locality unknown.

Amauronematus excavatus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 85. ♀.

Nematus excessus (MacGillivray)
Alaska.

Pteronidea excessus MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 170. ♀.

Nematus currani Ross
B. C, Oreg. Host: Populus.

Pteronidea nigriventris Curran, 1926. Canad. Ent. 58: 233. ♀, ♂. Preocc. in Nematus by
Holmgren, 1883.

Nematus currani Ross, 1951. hi Muesebeck, et al., U. S. Dept. Agr., Agr. Monog. 2: 45. N.
name.

Nematus hypomelas (Rohwer)
Colo.

Pteronus hypomelas Rohwer, 1908. Canad. Ent. 40: 48. ♀.

Nematus kukukiana (Kincaid)
n. comb. Alaska.

Pontania kukukiana Kincaid, 1900. Wash. Acad. Sci., Proc 2: 356. ♀.

Nematus lipovskyi Smith
Maine, N. H., Mass., N. J., Pa., Md., Va., Ala. Host: Azalea.

Nematus lipovskyi Smith, 1974. Ent. Soc. Wash., Proc. 76: 205-206. ♀.

Nematus notatus (Rohwer)
Colo.

Pteronus notattis Rohwer, 1908. Canad. Ent. 40: 49. ♀.

Nematus oligospilus Foerster
Que. to Alaska s. to Va., Ind., Kans., Colo., Calif.; Mexico; Europe to
Himalayas. Ecology: Widespread. Host: Salix.

Nematus oligospihis Foerster, 1854. Naturh. Ver. Rheinlande, Verb. 11: 284. ♀.

Nematus mendicus Walsh, 1866. Ent. Soc. Phila., Proc. 6: 261. ♀.

Nematus trivittatus Norton, 1867. Amer. Ent. Soc, Trans. 1: 218. ♀.

Nematus dorsivittatus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 10. ♀.

Nematus salicivonis Cameron, 1882. Ent. Monthly Mag. 18: 194.

Nematus sylvestris Cameron, 1884. Ent. Monthly Mag. 20: 266.

Pteronus koebelei Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 71. ♀.

Pteronidea vanduzeei Rohwer, 1913. U. S. Natl. Mus., Proc. 45: 280. ♀.

Pteronidea elelea MacGillivray, 1923. Canad. Ent. 55: 162. ♀.

Nematus pinguidorsum Dyar
Maine to N. Y. w. to Sask., N. Dak. Host: Betula.

Nematus pinguidorsum Dyar, 1895. Amer. Ent. Soc, Trans. 22: 303. Larva.

Amauronematus vacivus MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 28. ♂. N. syn.

Biology: Wong, 1954. Canad. Ent. 86: 154.

Nematus reticulatus Holmgren
Labrador to Alaska s. to N. H., Man.; Europe to Siberia. Ecology:
Widespread. Host: Vaccinitim.
Nematus reticulatus Holmgren, 1883. Ent. Tidskr. 4: 143. ♀.
Nemattis marionellus Holmgren, 1883. Ent. Tidskr. 4: 144.
Nematus occipitalis Holmgren, 1883. Ent. Tidskr. 4: 144.
Nematus anceps Holmgren, 1883. Ent. Tidskr. 4: 145.
Nematus yyiysticus Holmgren, 1883. Ent. Tidskr. 4: 145.
Nematus parvulus Holmgren, 1883. Ent. Tidskr. 4: 146.
Nematus picticollis Holmgren, 1883. Ent. Tidskr. 4: 147.
Nematus lieyiterictis Holmgren, 1883. Ent. Tidskr. 4: 146.
Pontania popofiana Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 353. ♀, ♂.
Pontania glinka Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 355. ♀, ♂.




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Pontania teyiuitarsis Konow, 1901. Ztschr. System. Hym. Dipt. 1: 130.
Pontania poppii Konow, 1904. Ztschr. System. Hym. Dipt. 4: 230.
Pontania forsiusi Enslin, 1915. Deut. Ent. Ztschr., Beih., p. 354.
Amauronematus alberich Benson, 1934. Ann. and Mag. Nat. Hist. 14: 208.
Pteronidea nubium Benson, 1935. Ent. See. London, Trans. 83: 30.
Amauronematus alsius Benson, 1935. Ent. Soc. London, Trans. 83: 32.
Pteronidea leptostignia Lindqvist, 1957. Notulae Ent. 37: 111.
Pteronidea nigrita Lindqvist, 1957. Notulae Ent. 37: 113.
Pteronidea wolteri Lindqvist, 1957. Notulae Ent. 37: 115. ♀.
Pteronidea roberti Lindqvist, 1957. Notulae Ent. 37: 117.
Pteronidea lindbergi Lindqvist, 1957. Notulae Ent. 37: 105.
Pteronidea rutilipes Lindqvist, 1959. Notulae Ent. 39: 59. ♀.
Pteronidea stdciceps Lindqvist, 1959. Notulae Ent. 39: 57. ♀.
Pteronidea aspera Lindqvist, 1959. Notulae Ent. 39: 58. ♀.
Pteronidea thunbergi Lindqvist, 1959. Notulae Ent. 39: 58.

Taxonomy: Benson, 1961. Resultats des recherches scientifiques entreprises au Pare
National Suisse, Band VII, pp. 192-193.

Nematus subpallidus (MacGillivray)
n. comb. N. W. T.

Pontania subpallida MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 30, p.

110. ♀.

Nematus tibialis Newman
Ont. , Maine to Fla. w. to Minn., Kans., Colo., Ark., Ariz.; Wash., Greg.;

Europe. Host: Robiyiia pseudacacia, Gleditsia triacanthos. Introduced with host into

Europe.
Nematus tibialis Newman, 1837. Ent. Mag. 4: 260. ♀.
Nematus trilineatus Norton, 1867. Amer. Ent. Soc, Trans. 1: 215. ♀.
Nematus similaris Norton, 1880. U. S. Dept. Agr., Ann. Rpt. for 1879, p. 224. ♀.
Nematus robiniae Forbes, 1885. State Ent. 111. Rpt. 14: 116. ♀.
Pteronus solitariiis Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 107. ♀.

Nematus vancouverensis (Marlatt)
Maine to Alaska s. to Pa., Iowa, Colo., Ariz., Calif. Ecology:

Widespread. Host: Populus.
Pteronus vancouverensis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 70. ♀, ♂.
Atnauro7iematus califomicus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 85. ♀.

Preocc. by Pteronus califortiicus Marlatt, p. 60 of same publication. N. syn.
Pachyneyyiatus vemalis Rohwer, 1909. Canad. Ent. 41: 18. ♀. N. syn.
Pteromis wrighti Rohwer, 1911. Ent. Soc Wash., Proc 13: 31. N. name for

Amauro7iematus califomicus Marlatt.
Aynauronemattcs Valerius MacGillivray, 1923. 111. Univ. Bui. 20(50): 6. 6.N. syn.

Biology: Brown, Eads, and Clark, 1969. Jour. Econ. Ent. 62: 672 (Nematus sp.).

Nematus villosus Thomson
N. W. T.; Europe.

Nematus villosus Thomson, 1862. Ofvers. Svensk. Vetensk. Akad., Forh. 19: 629. ♀.

Nematus viridescens Cameron
Maine, Ont., N. Y., Man., Sask., Alaska; Europe. Host: Betula.

Nematus viridescens Cameron, 1885. Monog. Brit. Phytophagous Hym., v. 2, p. 122.

Pteronus hyalinus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 67. ♀. N. syn.

Biology: Wong, 1954. Canad. Ent. 86: 158 (hyalinus).

Nematus zebratus (Kincaid)
Alaska.

Pteronus zebratus Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 357. ♀.

Species Group Ribesii

Nematus cockerelli (Rohwer)
Colo.

Pteronus cockerelli Rohwer, 1909. Canad. Ent. 41: 88. ♀.

Nematus edwardsii Cresson
Calif.

Neynatiis edwardsii Cresson, 1880. Amer. Ent. Soc, Trans. 8: 7. ♀.

Nematus equina (MacGillivray)
Maine.

Pteronidea equina MacGillivray, 1920. Canad. Ent. 52: 235. ♀.

Nematus foveatus (Marlatt)
Wash., Oreg.

Pteronus foveatus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 55. ♀.




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Pteronidea eledra MacGillivray, 1923. 111. Univ. Bui. 20(50): 30. ♀.

Nematus jugicola Thomson
Man. (Churchill); Europe. Host: Salix.

Nematus jugicola Thomson, 1871. Hym. Scand., v. 1, p. 146. ♀.

Nematus marlattii Dyar
N. H. Host: Alnus.

Nematus Marlattii Dyar, 1895. Amer. Ent. Soc, Trans. 22: 305. 9, larva.

Nematus nigropectus Cresson
Wash., Oreg., Nev.

Nematus nigropectus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 6. ♀.

Pteronus bicolor Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 56. ♀.
Questionable synonym.

Nematus ostryae (Marlatt)
N. J. to Va. Host: Ostrya.

Pteronus ostryae Marlatt, 1896. Canad. Ent. 28: 252. ♀.

Nematus ribesii (Scopoli)
Que., Ont. to Wash. s. to D. C, Ohio, 111., Mo., Kans., Colo., Oreg.; Europe to
Caucasus. Ecology: Widespread. Introduced. Host: Ribes. Imported currantworm.

Tenthredo ribesii Scopoli, 1763. Ent. Cam., p. 280.

Tenthredo ventricosa Latreille, 1805. Hist. Nat. Crust. Ins., v. 13, p. 131.

Tenthredo salicis Fallen, 1808. Svenska Vetensk. Akad., Handl. 29: 118. ♀.

Nematus dimidiata Lepeletier, 1823. Monog. Tenthred., p. 68. ♀.

Nematus trimaculata Lepeletier, 1823. Monog. Tenthred., p. 69. ♀.

Nematus grossulariae Moore, 1831. Lit. and Phil. Soc. Manchester, Mem. 2: 114-134.

Nematus depressa Hartig, 1840. Stettin Ent. Ztg. 1: 24. ♀.

Nematus macrocera Hartig, 1840. Stettin Ent. Ztg. 1: 25.
Nematus ribis Dufour, 1847. Soc. Ent. France, Ann. 2: 571-581. ♀, ♂.

Pteronus longicomis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 72. ♂. Preocc.
by Nematus longicomis Eschscholtz, 1822.

Pteronus arapahonum Cockerell, 1906. Ent. News 17: 220. ♂.

Pteronidea ribesii \ds. feminina Enslin, 1916. Deut. Ent. Ztschr., Beih., p. 419.

Pteronidea ribesii var. konowi Enslin, 1916. Deut. Ent. Ztschr., Beih., p. 420.

Pteronidea aceris Rohwer, 1915. U. S. Natl. Mus., Proc. 49: 211. ♀.

Pteronidea equatia MacGillivray, 1923. 111. Univ. Bui. 20(50): 30. ♂.

Biology: Caesar and Garlick, 1920. Canad. Ent. 52: 106-108. — Kemner, 1924. Centralanst. for
Forsoksv. pa Jordbruksomradet Meddel. 43: 1-33 (in Sweden). —Miles, 1932. Bui. Ent. Res.
23: 1-6. — Walden, 1933. Conn. Agr. Expt. Sta., Bui. 346: 416-427. — Rautapaa, 1970. Ann.
Ent. Fenn. 36: 118-120 (feeding deterrents, host preference).

Morphology: Middleton, 1921. Ent. Soc. Wash., Proc. 23: 173 (terminal abdominal structures).
— Snodgrass, 1928. Smith. Misc. Coll. 81: 121. — Shafiq, 1954. Jour. Microscopical Sci. 95: 93-114 (embryonic development).

Nematus salicisodoratus Dyar
Maine to Va. w. to Wis., 111., Ark. Host: Salix, Populus.
Nematus salicis-odoratus Dyar, 1894. Canad. Ent. 26: 187. 9,6, larva.
Pteronus odoratus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 65. Emend.
Pteronus comelli Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 65. ♀, ♂.
Pteronidea winnanae Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 386. ♂.
Pteronidea mendicana Rohwer, 1920. U. S. Natl. Mus., Proc. 57: 214. ♀.
Pteronidea plesia Rohwer, 1920. U. S. Natl. Mus., Proc. 57: 216. ♀.
Pontania dotata MacGillivray, 1921. N. Y. Ent. Soc, Jour. 21: 34. ♂.

Taxonomy: Middleton, 1922. U. S. Natl. Mus., Proc 51: 21 (larva of plesia).

Biology: Warren, 1957. Kans. Ent. Soc, Jour. 30: 11.

Nematus superbus (Provancher)
Que., N. H.

Eriocampa superba Provancher, 1885. Addit. Corr. Faune Ent. Canada, Hym., p. 351. ♀.

Pteronus antennatus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 61. ♀.

Nematus tricolor (Marlatt)
N. H.

Pteronus tricolor Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 56. ♀.

Nematus umbratus Thomson
Ont., Que., Wash. (?), Oreg. (?).; Europe. Host: Alnus, Betula.

Nematus umbratus Thomson, 1871. Hym. Scand., v. 1, p. 142. ♀.

Nematus collinus Cameron, 1882. Ent. Soc. London, Trans., p. 534. ♀.




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Nematus unicolor (Marlatt)
Calif.

Pteronus unicolor Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 72. ♀.

Unplaced Taxa of Nematus Panzer

Nematus integer Say
Ind.

Tenthredo (Nematus) Integra Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 583. Nom. nud.

Nematus integer Say, 1836. Boston Jour. Nat. Hist. 1: 218. ♀.

Nematus minutus (Marlatt)
Wash. (Olympia).

Pachynematus minutus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 110. ♂.

Nematus nigristigmus (Provancher)
Que.

Emphyt2is nigristigma Provancher, 1888. Addit. Corr. Faune Ent. Canada, Hym., p. 348. ♀.

Nematus obscurus Norton
Mass.

Nematus obscurus Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 160. ♀.

Nematus pacificus (Marlatt)
Wash. (Olympia).

Pteronus pacificus Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 49. ♀.

Nematus semirufus Kirby
Mass.

Tenthredo (Neviatus) fulvipes Harris, 1835. hi Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 583. Nom. nud.

Nematiis fulvipes Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 159. ♀. Preocc. by Fallen,
1808.

Nematus semirufus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 148. N. name for fulvipes
Norton.

Nematus flavipesil) Smith, 1966. Ent. Soc. Wash., Proc. 68: 248.

Nematus tetraopsis Provancher
Calif.

Nematus tetraopsis Provancher, 1895. Nat. Canad. 22: 79. ♀.

Nematus wrangeli (Marlatt)
Alaska.

Pachynematus wrangeli Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 109. ♂.

Pteronus rivularis Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 358. ♂.

Genus PONTANIA Costa

Pontania Costa, 1859. Fauna del Regno di Napoli, Imen., pt. 3, p. 20.

Type-species: Nonatus gallicola Stephens. Desig. by Marlatt, 1896.

Larvae of this large, holarctic genus form closed, pea-shaped or bean-shaped galls on the
leaves of Salix or Populus. This genus is in need of revision, and the following is primarily a list
of the described species and their type localities except for the work by Benson (1960). Though
Benson (1960) established species groups, many of the North American forms have not been stu-
died adequately to permit such an arrangement.

Revision: Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 20-43.

Taxonomy: Benson, 1960. Brit. Mus. (Nat. Hist.) Ent, Bui. 8: 369-384. —Benson, 1962. Brit.
Mus. (Nat. Hist.) Ent., Bui. 12: 403-404 (holarctic species).

Biology: Smith, 1970. Ent. Soc. Amer., Ann. 63: 36-51.

Morphology: Smith, 1968. Ent. Soc. Amer., Ann. 61: 1-389-1407.

Pontania agilis (Cresson)
Nev.

Nematus agilis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 9. ♂.

Pontania arctica (MacGillivray)
N. W. T., Yukon, Alaska, Man. Ecology: Forms leaf galls. Host: Salix
reticulata.
Euura arctica MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G, p. 5G. 9.
Euura arbortiva MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G, p. 4G. 9.
Pontania delicatula MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G, p. 8G. ♂.
Pontania deminuta MacGilHvray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G, p. 9G. 9.




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Taxonomy: Benson, 1960. Brit. Mus. (Nat. Hist.) Ent., Bui. 8: 377. -Vikberg, 1970. Ann. Ent.

Fenn. 36: 19 (not holarctic).

Pontania arctophilae Benson
Man. (Churchill). Ecology: Ovipositing on Salix arctophila.

Pontania arctophilae Benson, 1960. Brit. Mus. (Nat. Hist.) Ent, Bui. 8: 373. ♀.

Pontania atriventris Marlatt
Oreg. (Mt. Hood).

Pontania atriventris Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 38. ♀.

Pontania beckettae Benson
Man. (Churchill). Ecology: Produces pea-shaped galls attached to midvein
on underside of leaves. Host: Salix planifolia, S. discolor, S. glaiica, S. reticulata.

Pontania beckettae Benson, 1960. Brit. Mus. (Nat. Hist.) Ent., Bui. 8: 378. 9, gall.

Pontania borealis Marlatt
N. Y. Ecology: Produces galls. Host: Salix sericea.

Pontania borealis Marlatt, 1898. Canad. Ent. 30: 302. ♀.

Pontania brachycarpae Rohwer
Colo. Ecology: Adult collected from Salix brachycarpae.

Pontania brachycarpae Rohwer, 1908. Canad. Ent. 40: 46. ♀.

Pontania bruneri Marlatt
Wyo. (Robinson's Ranch). Ecology: Produces leaf galls. Host: Salix longifolia.

Ponta7iia bruneri Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 35. 9, gall.

Pontania californica Marlatt
Calif. Ecology: Produces a leaf gall. Host: Salix.

Po7itania californica Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 38. 9, 6, gall.

Pontania consors Marlatt
N. Y. Ecology: Produces leaf galls. Host: Salix sericea.

Pontania coji.wrs Marlatt, 1898. Canad. Ent. 30: 302. ♀, ♂.

Pontania crassicornis Rohwer
Ont. Ecology: Produces galls. Host: Salix humilis.

Pontania crassicornis Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 241. ♂.

Pontania crassipes (Thomson)
N. W. T., Yukon, Man., Alaska; n. Europe, Switzerland. Ecology: In
Europe, makes pea-shaped galls on main vein usually on underside of leaves. Host:
Salix herbacea, S. polaris, S. arctica, S. retusa, S. arbuscula, S. inyrsiiiites, S.
lapponum, S. reticulata.

Neynatus crassipes Thomson, 1871. Hym. Scand., v. 1, p. 162. ♀.

Nematiis herbaceae Cameron, 1875. Glasgow Nat. Hist. Soc, Proc. 2: 304.

Pontania ora Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 354. ♀.

Pontania atrata MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G, p. 6G. ♂.

Pontania lorata MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G, p. 8G. ♂.

Pontania polaris Malaise, 1920. Ent. Tidskr. 40: 112.

Pontania enslini Zirngiebl, 1937. Festschr. zum 60 Geburtst. von Prof. Dr. Embrik Strand
3: 337.

Pontania arbusculae Benson, 1941. Ent. Soc. London, Proc. 10: 134.

Pontania algida Benson, 1941. Ent. Soc. London, Proc. 10: 134.

Pontania aquilonis Benson, 1941. Ent. Soc. London, Proc. 10: 135.

Taxonomy: Benson, 1960. Brit. Mus. (Nat. Hist.) Ent., Bui. 8: 379.

Biology: Vikberg, 1970. Ann. Ent. Fenn. 36: 18 (in Finland).

Pontania daedalus MacGillivray
N. Y.

Pontania daedalus MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 33. ♀.

Pontania dedecora MacGillivray
N. Y.

Po7itayiia dedecora MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 32. ♀.

Pontania demissa MacGillivray
N. Y.

Pontania dentissa MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 33. ♀.

Pontania derosa MacGillivray
N. Y.

Pontania derosa MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 34. ♀.

Pontania devincta MacGillivray
Maine.

Pojitania devincta MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 34. ♀, ♂.

Pontania dolichura (Thomson)
Man. (Churchill); n. Europe, Austria, Switzerland. Ecology: Produces

Galls. Host: Salix planifolia, S. purpurea, S. phylicifolia, S. yiigricans, S. arbuscula, S.
lapponum, S. myrsinites, S. lanata, S. glabra, S. incana, S. retusa, S. sachalinensis.
These hosts were recorded in Europe, except for S. planifolia which was recorded in
Canada.
Nematus dolichurus Thomson, 1871. Hym. Scand., v. 1, p. 164. ♀, ♂.




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N €7)1 at us fern oralis Cameron, 1875. Glasgow Nat. Hist. Soc, Proc. 2: 295.
Pontania robbinsi Benson, 1935. Ent. Soc. London, Trans. 83: 26.

Taxonomy: Benson, 1960. Brit. Mus. (Nat. Hist.) Ent, Bui. 8: 380.

Biology: Vikberg, 1970. Ann. Ent. Fenn. 36: 13 (in Finland).

Pontania dubius (Marlatt)
Mass.

Pteronus dubius Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 74. ♂.

Pontania gracilis Marlatt
Va. Ecology: Produces a leaf gall. Host: Salix.

Pontania gracilis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 39. 9, gall.

hopping! (Ross). B. C, Wash. Ecology: Produces a leaf gall. Host: Salix.

Pteronus kincaidi Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 55. ♀, ♂. Preocc.
by Pontania kincaidi Marlatt, p. 33.

Eiiura hoppingi Ross, 1937. 111. Biol. Monog. 15(2): 88. N. name.

Pontania kincaidi Marlatt
Wash. (Olympia).

Pontania kincaidi Marlatt. 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 33. ♀.

Pontania lucidae Rohwer
Ont. Ecology: Produces galls. Host: Salix lucida.

Pontania lucidae Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 242. ♀, ♂.

Pontania monile Marlatt
Utah (American Fork Canyon). Ecology: Produces leaf galls. Host: Salix.

Pontania monUe Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 43. gall, larva.

Pontania myrtillifoliae Benson
Man. (Churchill). Host: Salix tnyrtillifolia.

Pontania myrtillifoliae Benson. 1960. Brit. Mus. (Nat. Hist.) Ent. Bui. 8: 372. ♀.
paciflca Marlatt. Calif. Ecology: Produces a leaf gall. Host: Salix lasiolepis.

Pontania pacifica Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 35. ♀.

Biology: Caltagirone, 1964. Ent. Soc. Amer., Ann. 57: 279-291.

Pontania peninsularis Kincaid
Alaska.

Pontania peninsularis Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 354. ♀.

Pontania petiolaridis Rohwer
Ont Ecology: Produces a leaf gall. Host: Salix petiolaris.

Pontania petiolaridis Rohwer, 1917. In Cosens, Ont Ent. Soc, Ann. Rpt. 47: 18-19. ♀, ♂.

Po7itania petiolaris Felt. 1917. N. Y. State Mus., Bui. 200: 34.

Pontania placenta (Norton)
Labrador.

Nematus placentus Norton, 1867. Amer. Ent. Soc, Trans. 1: 213. ♀, ♂.

Pontania proxima (Lepeletier)
Maine, Ont., N. Y.; Europe. Ecology: Produces reddish leaf galls in rows
on each side of midrib. Probably introduced with host Host: Salix fragilis, S. alba.
Willow redgall sawfly.

Neynatus proximus Lepeletier, 1823. Monog. Tenthred., p. 67. ♀.

Nematus gallicola Stephens. 1835. Illus. Brit. Ent., v. 7, p. 36.

Messa hyalina Norton, 1864. Ent. Soc. Phila., Proc 4: 8. ♀.

Hoplocampa gallicola Cameron, 1877. Ent. Monthly Mag. 14: 156.

Euura flavipes Cameron, 1885. Monog. Brit. Phytophagous Hym., v. 2, p. 211.

Biology: Carleton, 1939 Linn. Soc. London. Jour. 40: 575-624.

Pontania pumila Rohwer
N. B.

Pontania pumila Rohwer, 1910. U. S. Natl. Mus., Proc 38: 198. ♀, ♂.

Pontania pyriformis Marlatt
Calif. (Placer Co.). Ecology: Produces a leaf gall. Host: Salix califomica

(?).
Pontania pyriformis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 43. gall, larva.

Pontania resinicola Marlatt
Calif. (Los Angeles). Ecology: Produces leaf galls. Host: Salix califomica.

Po7itania resinicola Marlatt, 1896. U. S. Dept. Agr., Div. Ent Tech. Ser. 3: 30. 9, 6, gall

Pontania rugulosa Marlatt
Mich. Ecology: "reared (?) from willow gall."

Pontania rugulosa Marlatt, 1896. U. S. Dept Agr., Div. Ent. Tech. Ser. 3: 41. ♂.

Pontania s-desmodioides (Walsh)
111. Ecology: Produces a leaf gall. Host: Salix hum His.

Neynatus s. desmodioides Walsh, 1866. Ent Soc. Phila., Proc. 6: 257. 9, 6, gall, larva.

Nematus salicis-des7nodioides Dalla Torre, 1894. Cat. Hym., v. 1. p. 259. Emend.

Pontania s-pisum (Walsh)
111. Ecology: Produces a leaf gall. Host: Salix discolor.

Nematus s. pisum Walsh, 1866. Ent. Soc Phila., Proc 6: 258. 9,6, gall, larva.

Nematus salicis-pisuyn Thomas. 1881. State Ent. 111. Rpt. 10: 68. gall, larva.




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Pontania s-pomum (Walsh)
111. Ecology: Produces a leaf gall. Host: Salix.

Neynatus s. pomum Walsh, 1866. Ent. Soc. Phila., Proc. 6: 255. 9,6, gall, larva-

Neinatus hospes Walsh, 1866. Ent. Soc. Phila., Proc. 6: 261. ♀, ♂.

Nematus salicis-pomuyn Thomas, 1881. State Ent. 111. Rpt. 10: 68. gall, larva.

Poniania pomum Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 36. Eme^nd.

Pontania stigmatalis Marlatt
Oreg. (Mt. Hood).

Pontania stigmatalis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 39. ♀.

Pontania subatrata MacGillivray
Alaska.

Pontania subatrata MacGillivray, 1921. Calif. Acad. Sci., Proc. (ser. 4) 11: 189. ♂.

Pontania sublorata MacGillivray
Alaska.

Pontania sublorata MacGillivray, 1921. Calif. Acad. Sci., Proc. (ser. 4) 11: 190. ♂.

Pontania sulphurea Marlatt
Mont., Nev.

Pontania sulphurea Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 41. ♀, ♂.

Pontania truncata Marlatt
Calif.

Pontania truncata Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 38. ♀.

Genus PHYLLOCOLPA Benson

Phyllocolpa Benson, 1960. Ent. Monthly Mag. 96: 60.

Type-species: Nematus leucapsis Tischbein. Orig. desig.

Larvae of this holarctic genus live in rolled leaves or rolled leaf edges of Salix or Populus.
This genus is in need of study. The following is mostly a list of described species with their type
localities without regard to possible synonymy. Benson (1960) established two species groups.

Revision: Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 24-30 (Pontania, Group I).
— Ross, 1929. Ent. Soc. Wash., Proc. 31: 91-97 (Marlatt's Group I of Pontania).

Taxonomy: Benson, 1960. Brit. Mus. (Nat. Hist.) Ent., Bui. 8: 380-381 (section A of
Pontania). —Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 403 (holarctic species).

Biology: Smith, 1970. Ent. Soc. Amer., Ann. 63: 36-51.

Morphology: Smith, 1968. Ent. Soc. Amer., Ann. 61: 1389-1407.

Phyllocolpa acuminata (Marlatt)
Mich.

Pontania aciuninata Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 32. ♀.

Phyllocolpa acutiserra (Lindqvist)
Man. (Churchill); Lapland, Scotland. Host: Salix.

Pontania acutiserra Lindqvist, 1949. Notulae Ent. 28: 66. ♀, ♂.

Biology: Vikberg, 1970. Ann. Ent. Fenn. 36: 11 (in Finland).

Phyllocolpa agama (Rohwer)
CaHf.

Pontania agama Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 242. ♀.

Pontania foveata Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 243. ♀.

Phyllocolpa bozemani (Cooley)
Sask., Alta., Mont. Host: Populus. Poplar leaffolding sawfly.

Pontania bozemani Cooley, 1903. Canad. Ent. 35: 197. ♀, ♂.

Biology: Cooley, 1903. Mont. Agr. Expt. Sta. Bui. 46: 113-118.

Phyllocolpa coriacea (Benson)
Alta. (Lake Louise); Scandinavia, Switzerland. Host: Salix.
Nematus (Pontania) coriacea Benson, 1953. Ent. Monthly Mag. 89: 150. ♀, ♂.

Biology: Vikberg, 1970. Ann. Ent. Fenn. 36: 12 (in Finland).

Phyllocolpa cressoni (Marlatt)
Wash.

Pontania cressoni Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 26. ♀.

Phyllocolpa excavata (Marlatt)
Colo., Alaska, Calif.; Europe to Siberia. Ecology: Larva Hves in rolled leaf
edge. Host: Salix petandra.

Pontania excavata Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 30. ♀, ♂.

Pontania megacephala Rohwer, 1908. Canad. Ent. 40: 47. ♂.

Pontania destricta MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 168. ♀.

Pontania apicifrons Malaise, 1932. Arkiv for Zool. 23A: 33. ♀, ♂.

Pontania carinifrons Benson, 1940. Ent. Monthly Mag. 76: 210. ♀, ♂.

Taxonomy: Benson, 1960. Ent. Monthly Mag. 96: 60.




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Biology: Vikberg, 1970. Ann. Ent. Fenn. 36: 11 (in Finland).

Phyllocolpa leavitti (Rohwer)
N. B.

Po7ita7iia leavitti Rohwer, 1910. U. S. Natl. Mus., Proc. 38: 199. ♀.

Phyllocolpa leucostoma (Rohwer)
Colo.

Pontania leucostoma Rohwer, 1908. Canad. Ent. 40: 45. ♂.

Phyllocolpa mariana (Ross)
B. C. (Vancouver). Ecology: Larva makes a leaf fold. Host: Populus
balsaniifera.

Pontania mariana Ross, 1929. Ent. Soc. Wash., Proc. 31: 91. 9, S, larva.

Phyllocolpa marlatti (Ross)
Oreg.

Pojitania ynarlatti Ross, 1929. Ent. Soc. Wash., Proc. 31: 93. ♀.

Phyllocolpa melanosoma (Rohwer)
Colo.

Pontania melanosoyna Rohwer, 1908. Canad. Ent. 40: 48.

Phyllocolpa mellina (Cresson)
Nev.

Nematus melliyms Cresson, 1880. Amer. Ent. Soc, Trans. 8: 10. ♀.

Phyllocolpa nevadensis (Cresson)
Nev.

Nematus nevadensis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 9. ♀, ♂.
ni^ipecta (Rohwer), n. comb. Calif.

Pontania nevadensis var. nigripecta Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 243. ♀, ♂.

Phyllocolpa nigrita (Marlatt)
n. comb. Mich.

Pontania nigrita Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 27. ♀.

Phyllocolpa parva (Cresson)
Nev.

Nematus parvus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 5. ♀.

Phyllocolpa pectoralis (Marlatt)
111. (Algonquin).

Pontania pectoralis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 31. ♀.

Phyllocolpa pepii (Ross)
Mont. (Florence).

Pontania pepii Ross, 1929. Ent. Soc. Wash., Proc 31: 96. ♀.

Phyllocolpa popuella (Ross)
Man., S. Dak. Ecology: Larva makes a leaf fold. Host: Populus deltoides.

Pontania popuella Ross, 1929. Ent. Soc. Wash., Proc. 21: 93. ♀, ♂.

Phyllocolpa populi (Marlatt)
Mass., Conn., Ont., N. J. Host: Populus.

Pontania populi Marlatt, 1896. Canad. Ent. 28: 253. ♀.

Phyllocolpa robusta (Marlatt)
D. C. (?), Mich.

Pontania robusta Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 32. ♀, ♂.

Phyllocolpa stipata (MacGillivray)
n. comb. Alaska.

Pontania stipata MacGillivray, 1921. Calif. Acad. Sci., Proc. (ser. 4) 9: 188.

Phyllocolpa terminalis (Marlatt)
N. Y. Host: Salix.

Pontania terminalis Marlatt, 1896. Canad. Ent. 28: 253. ♀, ♂.

Phyllocolpa tundra (Kincaid)
Alaska.

Pontania tundra Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 352. ♀, ♂.

Phyllocolpa utensis (Rohwer)
n. comb. Colo.

Pontania utensis Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 107. ♀.

Genus EUURA Newman

Two subgenera have been proposed, Euura for those species whose larvae form stem, twig, or
petiole galls on Salix, and Gemmura for those species whose larvae form bud galls on Salix.
This genus is in need of study. The following is primarily a list of described species with their
type localities without regard to possible synonymy. According to Smith (1968) conventional use
of morphological characters for species separation is impossible in this group and identification
may have to be based on specific host data and biological information; therefore, the exact
identity of many of the described species may never be known.

Revision: Rohwer, 1909. N. Y. Ent. Soc, Jour. 17: 7-25.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 402-403 (holarctic species).
—Smith, 1968. Ent. Soc Amer., Ann. 61: 1389-1407 (Calif species).




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Biology: Smith, 1968. Ent. Soc. Amer, Ann. 61: 1389-1407. —Smith, 1970. Ent. Soc. Amer.,
Ann. 63: 36-51.

Morphology: Smith, 1968. Ent. Soc. Amer., Ann. 61: 1389-1407. —Smith, 1972. Int. Jour. Ins.
Morph. and Embryol. v. 1, 1: 321-365 (external genitalia of Euura).

Genus EUURA Subgenus EUURA Newman

Euura Newman, 1837(January). Ent. Mag. 4: 259.

Type-species: Euura gallae Newman. Desig. by Rohwer, 1911.
Nematus subg. Cryptocampus Hartig, 1837(March). Fam. Blattwespen und Holzwespen,
nebst Einleitung Naturgesch. Hym., p. 221.

Type-species: Nematus (Cryptocampus) medullaris Hartig. Desig. by Rohwer, 1911.

Euura albiricta Cresson
Nev.

Euura albiricta Cresson, 1880. Amer. Ent. Soc, Trans. 8: 4. ♀.

Euura atra (Jurine)
N. B., Que., Ont., Alta.; Europe. Ecology: Attacks many of the narrow-leaved

willows. Host: Salix repens, S. viminalis, S.fragilis, S. purpurea, S. babylonica, S. alba
var. vitelliua.
Pteronus a^fr Jurine, 1807. Nouv. Meth. Class. Hym., p. 6, fig. 6b. 9.
Cryptocampus angustus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst

Einleitung Naturgesch. Hym., p. 222.
Euura nigra Provancher, 1888. Addit. Corr. Faune Ent. Canada Hym., p. 346. ♀. Preocc.
by Norton, 1867.

Biology: MacCall and Titus, 1972. Canada Dept. Environment, Forestry Service, Bi-Monthly
Research Notes 28: 8.

Euura bakeri (Rohwer)
Calif. Ecology: Produces a stem gall. Host: Salix.
Cryptocampus bakeri Rohwer, 1910. Canad. Ent. 42: 51. ♀, ♂.

Euura bebbianae (Rohwer)
Colo. Ecology: Produces a stem gall. Host: Salix / bebbiana.

Cryptocampus bebbianae Rohwer, 1909. N. Y. Ent. Soc, Jour. 17: 18. ♀, ♂.
brewcriae Smith. Calif, (inner Coast Ranges, Yolo Co., Napa Co.). Ecology: Produces a stem
gall. Host: Salix breweri.
Enura (Euura) breweriae Smith, 1968. Ent. Soc. Amer., Ann. 61: 1405. 9, 6, gall.

Euura cooperae Cockerell
N. Mex. Ecology: Produces a stem gall on narrow-leaved willow. Host:
Salix.
Euura cooperae Cockerell, 1901. Ann. and Mag. Nat. Hist. (ser. 7) 7: 337. ♀.

Euura cosensii Rohwer
Ont. Ecology: Produces a petiole gall. Host: Salix humilis.

Euura cosensii Rohwer, 1915. U. S. Natl. Mus., Proc 49: 213. ♀, ♂.

Euura exiguae Smith
Calif, (north of Transverse Ranges up to 750m.). Ecology: Produces a stem
gall. Host: Salix exiguae.
Euura {Euura) exiguae Smith, 1968. Ent. Soc. Amer., Ann. 61: 1402. 9,6, gall.

Euura geyerianae Smith
Oreg. (Klamath Co.), and probably s. in Calif. Sierras. Ecology: Produces a
stem gall. Host: Salix geyeriana.
Euura (Euura) geyerianae Smith, 1968. Ent. Soc. Amer., Ann. 61: 1405. 9, 6, gall.

Euura lasiolepis Smith
Calif. (Monterey Co. to Mendocino Co. west to Sierra Nevada). Ecology:
Makes a stem gall. Host: Salix lasiolepis.
Euura (Euura) lasiolepis Smith, 1968. Ent. Soc. Amer., Ann. 61: 1406. 9, 6, gall.

Euura lemmoniae Smith
Calif. (Sierra Nevada and adjacent ranges from Mt. Lassen and s.).
Ecology: Produces a stem gall. Host: Salix lemmonii.
Euura (Euura) lemmoniae Smith, 1968. Ent. Soc. Amer., Ann. 61: 1406. 9, S, gall.

Euura macgillivrayi (Rohwer)
Colo. Ecology: Produces a stem gall. Host: Salix luteosericea.

Cryptocampus macgillivrayi Rohwer, 1909. N. Y. Ent. Soc, Jour. 17: 16. ♀, ♂.

Euura maura (Rohwer)
Colo. Ecology: Adult collected from Salix.

Cryptocampus maura Rohwer, 1909. N. Y. Ent. Soc, Jour. 17: 23. ♀, ♂.

Euura minuta MacGillivray
Iowa.

Euura minuta MacGillivray, 1914. Canad. Ent. 46: 366. ♀.

Euura moenia MacGillivray
Oreg.

Euura moenia MacGillivray, 1923. 111. Univ. Bui. 20(50): 17. ♀, ♂.




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Euura nigrella Rohwer
Ont.

Euura nigrella Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 241. ♀.

Euura nigra
***authority mismatch
Norton. Labrador.

Euura orbitalis var. nigra Norton, 1867. Amer. Ent. See, Trans. 1: 71. ♀.

Euura orbitalis Norton
Conn., N. Y. Ecology: Adults collected from Salix.

Euura orbitalis Norton, 1862. Ent. Soc. Phila., Proc. 1: 144. ♀, ♂.

Euura pallistigmus (Rohwer)
N. B.

Cryptocampus pallistigrnus Rohwer, 1910. U. S. Natl. Mus., Proc. 38: 200. ♀, ♂.

Euura parva Rohwer
Colo.

Euura parva Rohwer, 1908. Canad. Ent. 40: 176. ♀.

Euura perdita Rohwer
Colo. Ecology: Produces galls. Host: Salix.

Euura perdita Rohwer, 1909. Canad. Ent. 41: 20. ♀, ♂.

Euura propinqua (Rohwer)
N. Mex. Ecology: Produces a stem gall. Host: Salix.

Cryptocampus propinqua Rohwer, 1909. N. Y. Ent. Soc, Jour. 17:24. ♂.

Euura salicicola Smith
111. Ecology: Produces a stem gall. Host: Salix alba.

Euura salicicola Smith, 1879. North Amer. Ent. 1: 41. ♀, ♂.
scouierianae Smith. Calif. (Sierra Nevada and adjacent ranges from Mt. Shasta southward).
Ecology: Produces a stem gall. Host: Salix scouleriana.

Euura {Euura) scouierianae Smith, 1968. Ent. Soc. Amer., Ann. 61: 1406. 9,6, gall.

Euura serissimae Rohwer
Ont. Ecology: Produces a petiole gall. Host: Salix serissi7na.

Euura serissiynae Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 240. ♀.

Euura s-gemma Walsh
111. Ecology: Produces twig galls. Host: Salix humilis.

Euura s. gemma Walsh, 1866. Ent. Soc. Phila., Proc. 6: 250. 9, d, gall, larva.

Euura genuina Norton, 1867. Amer. Ent. Soc, Trans. 1: 79. 9, 6, gall.

Euura s-nodus Walsh
Ont., Mich., Ohio., 111. Ecology: Produces a stem gall. Host: Salix interior.

Euura s. nodus Walsh, 1866. Ent. Soc. Phila., Proc. 6: 253. ♂.

Cryptocampus salicis-nodus Dalla Torre, 1894. Cat. Hym., v. 1, p. 278. Emend.

Cryptocampus nodus Konow, 1905. In Wytsman, Gen. Ins., v. 29, p. 51. Emend.

Biology: Judd, 1954. Canad. Ent. 86: 542-546.

Euura s-ovum Walsh
N. Y., 111. Ecology: Produces a twig gall. Host: Salix.

Euura s. ovuyn Walsh, 1866. Ent. Soc. Phila., Proc. 6: 252. 9,6, gall.

Euura perturbans Walsh, 1866. Ent. Soc. Phila., Proc. 6: 254. 9,6, gall.

Euura salicis-ovimi Thomas, 1881. State Ent. 111. Rpt. 10: 69. gall, larva.

Euura s-ovulum Walsh
III. Ecology: Produces a stem gall. Host: Salix humilis.

Euura s. ovulum Walsh, 1866. Ent. Soc. Phila., Proc. 6: 253. gall, larva.

Cryptocampus salicis-ovulum Dalla Torre, 1894. Cat. Hym., v. 1, p. 278. Emend.

Genus EUURA Subgenus GEMMURA Smith

Euura subg. Gemmura Smith, 1968. Ent. Soc. Amer., Ann. 61: 1401.
Type-species: Cryptocampus mucronatus Hartig. Orig. desig.

Euura brachycarpae Rohwer
Colo.

Euura brachycarpae Rohwer, 1908. Canad. Ent. 40: 176. ♀, ♂.

Euura lanatae Malaise
Man. (Churchill), Alta. (Lake Louise); n. Europe. Ecology: Produces bud galls.
Host: Salix la^iata, S. calcicola.
Euura lanatae Malaise, 1920. Ent. Tidskr. 40: 105. ♀, ♂.

Euura mucronata (Hartig)
Man. (Churchill), Alaska; Europe. Ecology: Larva feeds inside buds of
many species of willow. Host: Salix.
Neynatus (.Cryptocampus) mucronatus Hartig, 1837. Fam. Blattwespen und Holzwespen,

nebst Einleitung Naturgesch. Hym., p. 223. ♀, ♂.
Euura nigritarsis Cameron, 1885. Monog. Brit. Phytophagous Hym., v. 2, p. 214.
Eiiura insularis Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 352. ♀, ♂.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 402.




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Genus AMAURONEMATUS Konow

Amauroneviatus Konow, 1890. Deut. Ent. Ztschr., pp. 233, 237, 246.

Type-species: N ematus fallax Lepeletier. Desig. by Rohwer, 1911.
Pontoprista Malaise, 1921. Ent. Tidskr. 42: 12.

Type-species: Pontoprista siiavis var. fusca Malaise. Desig. by Ross, 1937.

A large holarctic genus common in the arctic and subarctic regions. This genus is in need of
study. Several phyletic lines are evident which will eventually allow more convenient species
groupings.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 396-402 (holarctic species).

Amauronematus abnormis (Holmgren)
Baffin Is., N. W. T., Alaska; n. Europe, Switzerland, Siberia. Females
are commonly brachypterous.
Neynatus abnormis Holmgren, 1883. Ent. Tidskr. 4: 148. ♀.
Amauronematus tolli Konow, 1907. E. Zoologie 1: 20.

Aniauronematus aulatiis MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G,
p. 16G.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent, Bui. 12: 401.

Amauronematus amentorum (Foerster)
N. H. to Oreg. s. to Va., Mich., Wis., Colo.; Greenland; Iceland; Europe
to Siberia. Ecology: Widespread. Larva feeds on catkins. Host: Salix.

Nematus amentoruvi Foerster, 1854. Naturh. Ver. Rheinlande, Verb. 11: 337. ♀.

Nematus s^iavis Ruthe, 1859. Stettin Ent. Ztg. 20: 308.

Pontania atra Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 37. ♀. N. syn.

Pontania maura Rohwer, 1908. Canad. Ent. 40: 47. ♀. N. syn.

Pontania amentivora Rohwer, 1915. U. S. Natl. Mus., Proc. 49: 209. ♀.

Pontoprista kamtchatica Malaise, 1931. Arkiv for Zool. 23A: 53. ♀.

Amauronematus arcticola
***authority mismatch
(Dalla Torre). N. W. T., Alaska; arctic Europe and Siberia. Host: Salix.

Nematus arcticus Thomson, 1871. Hym. Scand., v. 1, p. 134. ♀. Preocc. by Holmgren, 1869.

Nematus arcticola Dalla Torre, 1894. Cat. Hym., v. 1, p. 207. N. name.

Aynauronematus arcticola Enslin, 1915. Deut. Ent. Ztschr., Beih., p. 389. N. name.

Amauronematus mcluckiei Benson, 1935. Roy. Ent. Soc. London, Trans. 83: 31. ♀, ♂.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 398.

Amauronematus azaleae Marlatt
N. H. Host: Azalea. Recorded from other eastern states but records are
based on larvae and their identity is not certain.
Amauronematus azaleae Marlatt, 1896. Canad. Ent. 28: 254. ♀, ♂.

Biology: Smith, 1974. Ent. Soc. Wash., Proc. 76: 204.

Amauronematus borealis (Marlatt)
Greenland (Disco Is.).

Nematus borealis Marlatt, 1892. In Fox, Acad. Nat. Sci. Phila., Proc. 44: 133. ♂.

Amauronematus brunneus (Norton)
Md., Wis., Sask., Kans., Colo., Utah.

Nematus brunneus Norton, 1864. Ent. Soc. Phila., Proc. 43: 244. ♀.

Amauronematus knabi Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 244. ♀. N. syn.

Amauronematus completus MacGillivray
Alaska.

Amauronematus completus MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v.
3G, p. 13G.

Amauronematus comstocki Marlatt
N. Y.

Amauronematus comstocki Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 77. ♀.

Amauronematus cooki Marlatt
Mich.

Amauronematus cooki Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 79. ♂.

Amauronematus digestus MacGillivray
Alaska.

Amauronematus digestus MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G,
p. 14G.

Amauronematus egnatia (MacGillivray)
Alta.

Pteronidea egnatia MacGillivray, 1923. Canad. Ent. 55: 162. ♀.

Amauronematus enavata (MacGillivray)
Maine.

Pteronidea enavata MacGillivray, 1920. Canad. Ent. 52: 236. ♀.




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Amauronematus fallax (Lepeletier)
N. S., Ont. to N. W. T., Yukon, Alaska s. to N. Y., Colo.. Oreg.; Europe.

Ecology: Widespread. Host: Salix.
Nematiis fallax Lepeletier, 1823. Monog. Tenthred., p. 64. ♀, ♂.
Nematus trifiircatus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 148.
Amauronematus similis Marlatt, 1896. Canad. Ent. 28: 255. ♀.
Pontavia qtiadrifasciatiis MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G,

p. lOG. 9.
A7naurone7iiatus vescus MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 30. ♀. N. syn.
A7najironematus veneficus MacGillivray, 1923. N. Y. Ent. Soc, Jour. 81: 169. ♀. N. syn.
Amauronematus nuorbinjargi Saarinen, 1949. Ann. Ent. Fenn. 15: 57.
Aviauronematus glacialis Saarinen, 1950. Ann. Ent. Fenn. 16: 45.
Amaiiroyieynatus aniicula Saarinen, 1950. Ann. Ent. Fenn. 16: 47.
Amauro7iematus subiiitens Saarinen, 1950. Ann. Ent. Fenn. 16: 49.
Amauroiiejnatiis septentrionalis Saarinen, 1950. Ann. Ent. Fenn. 16: 54.
Amauronematus festivus Saarinen, 1950. Ann. Ent. Fenn. 16: 58.
Amauronematus propinquus Saarinen, 1950. Ann. Ent. Fenn. 16: 60.
Nematus trifurcus Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 399. Misspelling of

trifurcatus Kirby.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., BuL 12: 899.

Amauronematus gracilis Marlatt
Mass., N. Y.

Amauronematus gracilis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 78. ♀.
gToenlandicus Malaise. N. W. T., Alaska; Greenland; n. Europe. Host: Salix (?).

Amauronematus groenlandicus Malaise, 1933. Skr. om Svalb. og Ischavet 53: 3. ♀.

Amauronematus carboyiarius Hellen, 1951. Acta Univ. Lund (n. s.) 46: 71.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 396-397.

Amauronematus hebes Konow
N. W. T.; Europe.

Amauro7ie7tiatus hebes Konow, 1907. Akad. Nauk. Fiz.-Mat. Otd. Zap. 18(3): 17. ♀.
A7nauro7iematus pristipkori7ius Malaise, 1931. Arkiv for Zool. 23A: 2.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 897.

Amauronematus helleni Lindqvist
Baffin Is., Que., N. W. T., Man.; Scandinavia.

Amaiiro7iematus helleni Lindqvist, 1941. Notulae Ent. 21: 65. ♀, ♂.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 397.

Amauronematus histrio (Lepeletier)
Maine to Alaska s. to 111., Colo.; Eurasia. Ecology: Widespread. Host:
Salix.

Ne7natus histiio Lepeletier, 1823. Monog. Tenthred., p. 63. ♀.

Ne7natus luteotergum Norton, 1861. Boston Soc. Nat. Hist, Proc. 8: 161. ♀.

Nematus fur Walsh, 1866. Ent. Soc. Phila., Proc. 6: 263. ♂.

N€77iatus concolor Norton, 1867. Amer. Ent. Soc, Trans. 1: 196. ♂.

Nematus violaceipeyinis Norton, 1867. Amer. Ent. Soc, Trans. 1: 201. ♂.

Ne7nat7is discolor Cresson, 1880. Amer. Ent. Soc, Trans. 8: 8. ♀.

Netnatus gle7ielge7isis Cameron, 1882. Ent. Soc. London, Trans., p. 535.

A7na7iro7ie7natus viscendus MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 28. ♀, ♂.

Amauronematus hulteni Malaise
Man., N. W. T., Yukon; Siberia.

Amauroneynatus variabilis var. hulteni Malaise, 1931. Arkiv for Zool. 23A: 43. ♀, ♂.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 397.

Amauronematus indicatus MacGillivray
Alaska (Camden Bay).

A7na7iro7iematiis i7idicatus MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1918-1918, v. 8G,
p. 14G.

Amauronematus isolatus Kincaid
N. W. T., Alaska.

Amauro7ie7natus isolatus Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 858. ♀.
Amauroue7tiatus 7nagnus MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G,

p. 17G. N. syn.
A77iauro7ie7natus veridicus MacGillivray, 1923. N. Y. Ent. Soc, Jour. 81: 170. ♀.

Amauronematus itelmena (Malaise)
Man., N. W. T.; Siberia. Ecology: Adult collected from Salix alexensis.
Po7itoprista itelme7ia Malaise, 1931. Arkiv for Zool. 23A: 54. ♀.




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Amauronematus leptocephalus (Thomson)
Man.; arctic Eurasia.

Nematus leptocephalus Thomson, 1862. Ofvers. Svensk. Vet. Akad., Forh. 19: 632. ♀.

Amauronematus leucolaenus (Zaddach)
Ont, Man., N. W. T., Sask., Alta., Yukon, Alaska; Eurasia.

Nematus leucolaenus Zaddach, 1882. Phys. Okonom. Gesell. Konigsberg., Schr. 23: 196. ♀.

Pontania imga Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 354. ♀.

Pteronidea egeria MacGillivray, 1923. Canad. Ent. 55: 161. ♀. N. syn.

Amauronematus saarineiii Lindqvist, 1945. Notulae Ent. 25: 106.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 402.

Amauronematus lineatus (Harrington)
Maine, Ont., N. Y. w. to III, Nebr., Kans. Host: Salix.
Nematus lineatus Harrington, 1893. Canad. Ent. 25: 59. ♀.
Ajnauronematus li7icolnensis Rohwer, 1909. Canad. Ent. 41: 19. ♀.

Biology: Ross, 1935. HI. State Acad. Sci., Trans. 28: 261-262 (life history; female, male, larva

described).

Amauronematus longicaudus (Hellen)
Man., B. C; Lapland.

Nematus (Amauronematus) longicauda Hellen, 1948. Notulae Ent. 27: 113. ♀.

Amauronematus microphyes (Foerster)
Ont. (Ottawa); Europe. Host: Salix.

Nematus microphyes Foerster, 1854. Naturh. Ver. Rheinlande, Verb. 11: 344. ♀.

Pontoprista romani Malaise, 1921. Ent. Tidskr. 42: 14.

Pontoprista nana Lindqvist, 1949. Notulae Ent. 28: 68.

Amauronematus neglectus (Kirby)
Que., Ont., HI., N. W. T., Man., Colo., Yukon, B. C; Lapland, Siberia. Host:
Salix.

Nematus neglectus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 147. ♀.

Amauronematus intennedius Malaise, 1931. Arkiv for Zool. 23A: 39.

Amauroneynatus spiniserra Malaise, 1921. Ent. Tidskr. 42: 119.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent, Bui. 12: 400-401. —Lindqvist, 1972.

Notulae Ent. 52: 69.

Amauronematus nigriventris (Holmgren)
Alaska; n. Europe.

Nematus nigriventris Holmgren, 1883. Ent. Tidskr. 4: 146. ♀.

A7nauronematus varianus MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G,
p. 16G. 9 (male misdet.). N. syn.

Pteronidea melanostoma Rohwer, 1920. U. S. Natl. Mus., Proc. 57: 212. ♀. N. syn.

Amauronematus nigrofemoratus (Cresson)
Nev.

Nematus nigrofemoratus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 4. ♀.

Amauronematus nitidipleuris Malaise
Man. (Churchill); Greenland; Europe, Siberia.

Amauronematus nitidipleuris Malaise, 1931. Arkiv for Zool. 23A: 45. ♀.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent, Bui. 12: 402.

Amauronematus orbitalis Marlatt
Colo., Oreg.

Aynauronematus orbitalis Marlatt, 1896. U. S. Dept Agr., Div. Ent. Tech. Ser. 3: 80. ♀.

Amauronematus oregonensis Marlatt
Oreg. (Mt. Hood).

Amauronematus oregonensis Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 80. ♀.

Amauronematus peralus Smith
n. name. Colo., Nev.

Nematiis pectoralis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 9. 9 Preocc. by Lepeletier,
1823.

Amauronematus polaris (Holmgren)
N. W. T., Alta., Alaska; n. Europe. Host: Salix.

Neynatus polaris Holmgren, 1883. Ent. Tidskr. 4: 143. ♀.

Pontania islandica Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 355. ♀.

Pteronus shumagensis Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 357. ♀.

Ptero7iidea parriserrata Lindqvist, 1944. Notulae Ent. 24: 23.

Aynauroneynatyis leucopyga Lindqvist, 1948. Notulae Ent. 28: 69.

Amauronematus subpolaris Lindqvist, 1948. Notulae Ent. 28: 71.

Pteronidea bipicta Lindqvist, 1959. Notulae Ent. 39: 56.

Pteronidea sordidiapex Lindqvist, 1959. Notulae Ent. 39: 57.

Taxonomy: Benson, 1961. Ergeb. Wiss. Unters. Schweiz. Nat. Parks 7(N. F.)(44): 190.




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Amauronematus pravus Konow
Alaska; n. Eurasia. Host: Salix.

A)nauro7iematus pravus Konow, 1896. Termes. Fuzetek 18: 176. ♀.

Amauronematus rapax (Cresson)
Nev.

Nematus rapax Cresson, 1880. Amer. Ent. Soc, Trans. 8: 4. ♂.

Amauronematus sagmarius Konow
Ont., Man., N. W. T., Alta., Alaska; Eurasia. Host: Salix.
Aniauronematus sagDiarius Konow, 1896. Termes. Fuzetek 18: 177. ♀, ♂.
Ayuauroneyyiatiis viduatiims Malaise, 1931. Arkiv for Zool. 23A: 40.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 399.

Amauronematus semilacteus (Zaddach)
B. C (Vernon); n. and cent. Europe. Host: Salix.

Neiuatus semilacteus Zaddach, 1883. Phys. Okonom. Gesell. Konigsberg., Schr. 24: 148. ♀.

Taxonomy: Vikberg and Kontuniemi, 1971. Ann. Ent. Fenn. 37: 89-95 (male, female, larva).

Amauronematus stordalensis (Strand)
Baffin Is., Ellesmere Is., N. W. T., Alaska; Siberia.

Nematus stordalensis Strand, 1905. Second Norw. Arctic Exped. "Fram" Rpt., no. 3, p. 7. ♀, ♂.
Nematus marginifer Strand, 1905. Second Norw. Arctic Exped. "Fram" Rpt., no. 3, p. 8. ♂.
Amauronematus cogitatus MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G, p. 150.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 400. —Vikberg, 1970. Ann.
Ent. Fenn. 36: 84-91 (description, distribution, separation from related species).

Amauronematus tillbergi Malaise
N. W. T., Alaska; arctic Europe and Siberia. Host: Salix.
Amauronematus tillbergi Malaise, 1920. Ent. Tidskr. 40: 125. ♀.
Amauronematus rex Benson, 1948. Ent. Monthly Mag. 84: 32. ♀.

Taxonomy: Lindqvist, 1972. Notulae Ent. 52: 71.

Amauronematus trifasciatus
***authority mismatch
(MacGilHvray). N. W. T.

Pontania trifasciata MacGillivray, 1919. Rpt. Canad. Arctic Exped. 1913-1918, v. 3G, p.
IIG. 9.

Amauronematus variator (Ruthe)
Man., N. W. T., Yukon, Alaska; Eurasia. Host: Salix.
Nematus variator Ruthe, 1859. Stettin Ent. Ztg. 20: 308. ♀.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hst.) Ent., Bui. 12: 398-399.

Amauronematus venerandus MacGillivray
Maine.

Amauronematus venerandus MacGilHvray, 1921. N. Y. Ent. Soc, Jour. 29: 30. ♂.

Amauronematus ventosus MacGillivray
Alaska.

Amauronematus ventosus MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 169. ♀.

Amauronematus verbosus MacGillivray
Maine.

Amauronematus verbosus MacGillivray, 1921. N. Y. Ent. Soc, Jour. 29: 29. ♀.

Amauronematus vicinalis (Cresson)
CaHf.

Neynatus vicinalis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 4. ♀.

Amauronematus viduatus (Zetterstedt)
Ont., Sask.; Europe.

Tenthredo viduata Zetterstedt, 1838. Ins. Lap., v. 1, p. 351. ♀.

Amauroneinatus viduatoides Lindqvist, 1959. Notulae Ent. 39: 127.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 402.

whitncyi Rohwer. Alaska.

Amauronematus whitneyi Rohwer, 1920. U. S. Natl. Mus., Proc 57: 216. ♀.

Amauronematus xanthus Rohwer
Nebr.

Amauronematiis xanthus Rohwer, 1909. Canad. Ent. 41: 18. ♀.

Genus DECANEMATUS Malaise

Decanematxis Malaise, 1931. Arkiv for Zool. 23A: 31.

Type-species: Decanematus longiserra Malaise. Orig. desig.

A small genus found in northern Canada and northern Europe.

Revision: Wong, 1968. Canad. Ent. 100: 84-86.




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Decanematus dulichus Wong
N. W. T. (Reindeer Depot, MacKenzie Delta).
Decaneviahis dulichus Wong, 1968. Canad. Ent. 100: 84. ♀.

Tribe PSEUDODINEURINI

Genus PSEUDODINEURA Konow

Dolerus subg. Pehnatopus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst
Einleitung Naturgesch. Hym., p. 244. Preocc. by Fischer de Waldheim, 1824.

Type-species: Dolenis (.Pehnatopus) nimutus Hartig. Monotypic.
Pseudodmeura Konow, 1885. Wien. Ent. Ztg. 4: 297.

Type-species: Tenthredo (AUaiitus) parvula Klug. Desig. by Rohwer, 1911.
Pkyllopais Hering, 1934. Internatl. Ent. Ztschr. 28: 353. N. name for Pelmatopus Hartig.

Several unidentified species are known from western United States.

Pseudodineura parva (Norton)
Maine to Pa. w. to 111. Ecology: Leafminer. Host: Hepatica.
Dineura parvus Norton, 1867. Amer. Ent. Soc, Trans. 1: 241. ♂.

Genus KERITA Ross

Kenta Ross, 1937. 111. Biol. Monog. 15 (2): 80.

Type-species: Kerita fidala Ross. Orig. desig.

An unidentified species is also known from the western United States.

Kerita fidala Ross
111. Ecology: A leafminer. Host: Mertensia.
Kerita fidala Ross, 1937. 111. Biol. Monog. 15 (2): 80. ♀.

Tribe PRISTOLINI

Revision: Wong, 1968. Canad. Ent. 100: 1049-1057.

Genus PRISTOLA Ross

Pristola Ross, 1945. Pan-Pacific Ent. 21: 153.

Type-species: Pristola macnabi Ross. Orig. desig.

Pristola clarki Wong
Colo. (Echo Lake, Mt. Evans, 10,600 ft.).

Pristola clarki Wong, 1968. Canad. Ent. 100: 1050. ♀, ♂.
macnabi Ross. Newfoundland, N. H., Man., Alta., B. C, Idaho, Oreg. Ecology: Larva lives in
berries. Host: Vacciniuin membranaceum.
Pristola macnabi Ross, 1945. Pan-Pacific Ent. 21: 154. ♀, ♂.

Genus MELASTOLA Wong

Melastola Wong, 1968. Canad. Ent. 100: 1053.

Type-species: Gymnonychus resinicolor Marlatt. Orig. desig.

Melastola ferruginosa Wong
Alaska, B. C, Wash.

Melastola ferruginosa Wong, 1968. Canad. Ent. 100: 1054. ♀, ♂.

Melastola resinicolor (Marlatt)
Alaska.

Gymnonychus resinicolor Marlatt, 1896. U. S. Dept. Agr., Div. Ent. Tech. Ser. 3: 125. ♀.
Pachynematus sahlbergi Konow, 1908. Ztschr. System. Hym. Dipt. 8: 83. ♀.

Unplaced Taxa of Nematinae

The following species are of questionable generic assignment due to inadequate descriptions
and inaccessibility, loss, or poor condition of type material. Three species described by Cameron
from northern Mexico are included; though not strictly within the scope of this Catalog, they un-
doubtedly represent southern extensions of northern species.

Hoplocampa? atriceps Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 168. Ga.

Nematjis calais Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 144. 9. Arctic America, MacKenzie

River.
Nematus castaneus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 147. 9. Hudson Bay, Canada, St.

Martin's Falls, Albany River.




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Neniatus crass7is Eschscholtz, 1822. Entomographien, p. 213. Isle of Unalaska, Alaska. Preocc.

by Fallen, 1808; see obtusus Kirby.
Nematus extraneus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 142. 9. Hudson Bay, Canada, St.

Martin's Falls, Albany River.
Neviatus fallax Norton, 1867. Amer. Ent. See, Trans. 1: 198. 6. Labrador. Preocc. by

Lepeletier, 1823; see nortonii Dalla Torre.
Nematus hudsonkus Norton, 1867. Amer. Ent. Sec, Trans. 1: 207. 9. Fort Good Hope,

MacKenzie River, N. W. T., Canada.
Nematus inquilinus Walsh, 1866. Ent. Soc. Phila., Proc. 6: 260. 9, cJ. Rock Island, 111. Ecology:

"Bred from Cecidomyidous gall S. rhodoides Walsh."
Nematus lateralis Norton, 1867. Amer. Ent. Soc, Trans. 1: 211. 9. Maine, N. Y.
Nematus longicomis Eschscholtz, 1822. Entomographien, p. 98. Unalaska Isle, Alaska.
Nematus longicortiis Say, 1835. Boston Jour. Nat. Hist. 1: 219. Iowa. Preocc. by Eschscholtz,

1822; see longulicomis Norton.
Nematus longulicomis Norton, 1867. Amer. Ent. Soc, Trans. 1: 214. N. name for longicomis

Say.
Nematus 77ialacus Norton, 1867. Amer. Ent. Soc, Trans. 1: 196. 9. Labrador.
Euura viexicana Cameron, 1884. Ent. Soc. London, Trans., p. 482. N. Sonora, Mexico.
Nematus mexicanus Cameron, 1884. Ent. Soc. London, Trans., p. 481. N. Sonora, Mexico.
Nematus mouela Norton, 1867. Amer. Ent. Soc, Trans. 1: 198. 6. Labrador.
Tenthredo (Nematus) monochroma Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Nematus monochroma Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 161. 9. Mass.
Hemichroa yiigricans Cameron, 1884. Ent. Soc. London, Trans., p. 482. N. Sonora, Mexico.
Nematus nigritus Norton, 1861. Boston Soc. Nat. Hist, Proc. 8: 159. 6. Conn.
Nematus nortonii Dalla Torre, 1894. Cat. Hym., v. 1, p. 246. N. name for fallax Norton.
Nematus obtusus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 148. N. name for crassus

Eschscholtz.
Nematus proximatus Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 160. 9. Maine, Mass., Conn.
Nematus quercicola Walsh, 1866. Ent. Soc. Phila., Proc 6: 260. Nom. nud.
Nematus rufo-fasciatiis Norton, 1867. Amer. Ent. Soc, Trans. 1: 205. 9. MacKenzie River, N.

W. T., Canada.
Pontania salicum Cockerell, 1905. In Aldrich, Smiths. Misc. Coll., v. 46, p. 163. Nom. nud.
Nematus satkatchewan Norton, 1867. Amer. Ent. Soc, Trans. 1: 200. 9. "Lake

Sathkatchewan."
Nematus sumptus Norton, 1867. Amer. Ent. Soc, Trans. 1: 207. 6. Maine.
Nematus vertebratus Say, 1836. Boston Jour. Nat. Hist. 1: 218. 9. Ind.

Subfamily HETERARTHRINAE

Most of these sawflies are small black insects, among the smallest of the Tenthredinidae. Lar-
vae of the Caliroini feed externally, skeletonizing the foliage of the host, and larvae of the
Heterarthrini and Fenusini are all leafminers.

Revision: Smith, 1971. U. S. Dept. Agr., Tech. Bui. 1420, 84 pp. (adults and larvae of Nearctic
species). —Smith, 1973. Ent. Soc Wash., Proc. 75: 337-345 (Neotropical species).

Taxonomy: Smith, 1967. Ent. Soc. Wash., Proc. 69: 277-284 (review of subfamily in N. Amer.).

Tribe CALIROINI

Genus ENDELOMYIA Ashmead

Endelomyia Ashmead, 1898. Canad. Ent. 30: 256.

Type-species: Selandria rosae Harris. Orig. desig.

Endelomyia aethiops (Fabricius)
Que., Conn, to Va. w. to B. C, Oreg.; Europe. Ecology: Widespread; a
common pest of cultivated roses. Host: Rosa. Rose-slug.
Tenthredo aethiops Fabricius, 1781. Species Insectorum, v. 1, p. 416.
Tenthredo (Selandria) atratula Dahlbom, 1835. Conspectus Tenthred., Sircid., Oryssin.

Scand., p. 10.
Selandria rosae Harris, 1841. Rpt. Ins. Mass. Injurious to Veg., p. 380.




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Eriocampa Livonensis Gimmerthal, 1844. Stettin Ent. Ztg. 5: 38. ♀.

Eriocampa Jiitida Tischbein, 1846. Stettin Ent. Ztg. 7: 113.

Tenthredo (Sela7idria) dolosa Eversmann, 1847. See. Nat. Moscou, Bui. 20: 33. ♀.

Sela7idria sorror Vollenhoven, 1869. Tijdschr. v. Ent. 12: 123.

Eriocampa testaceipes Cameron, 1874. Ent. Monthly Mag. 11: 128. ♂.

Poecilosoma nigricolle Cameron, 1882. Monog. Brit. Phytophagous Hym., v. 1, p. 218.

Biology: Riley, 1892. U. S. Dept. Agr., Insect Life 5: 6-11. —Chittenden, 1908. U. S. Dept.
Agr., Cir. 105, 12 pp. -Middleton, 1922. U. S. Dept. Agr., Farmers' Bui. 1252, 14 pp.
—Miles, 1935. Ann. Appl. Biol. 22: 116-133.

Genus CALIROA Costa

Caliroa Costa, 1859. Fauna del Regno di Napoli, Imen., pt. 3, p. 59.

Type-species: Caliroa sebetia Costa. Monotypic.
Eriocampoides Konow, 1890. Deut. Ent. Ztschr., p. 239.

Type-species: Tenthredo limacina Retzius. Desig. by MacGillivray, 1909.
Periclistoptera Ashmead, 1898. Canad. Ent. 30: 255.

Type-species: Monostegia alba Norton. Orig. desig.

Larvae of this genus are sluglike in appearance and feed on the underside of the foliage of the
host. Host records not listed below are Acer, Ceanothus, and Cotoneaster, all from larvae that
cannot be identified. Larvae have not been associated with adults for many species and are all
very similar; therefore, adults are needed in most cases for species determination.

Caliroa annulipes (Klug)
Ont.; Europe. Host: Betula papyrifera.

Tenthredo {Allantus) annulipes Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 70.

Caliroa cerasi (Linnaeus)
Que., Ont. to Fla. w. to B. C, Wash., Oreg., Calif., N. Mex.; Eurasia; Africa;
Argentina, Chile, Uruguay; Australia, Tasmania, New Zealand; China. Ecology:
Widespread; most commonly a pest of pear trees. Probably native to Europe and
introduced to other parts of the world, including N. Amer., by commerce. Host: Pyrus,
Pninus, Crataegus, Sorbus, and probably many other Rosaceae. Pear-slug.

Tenthredo cerasi Linnaeus, 1758. Syst. Nat., Ed. 10, p. 557.

Tenthredo li7nacina Retzius, 1783. Genera et Species Insectorum, p. 73.

Tenthredo cerasi Peck, 1799. Mass. Agr. Rpt., p. 9.

Tenthredo (Allantus) adumbrata Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 64. ♀.

Caliroa laudata MacGillivray, 1909. Canad. Ent. 41: 356. ♀.

Caliroa lacinata MacGillivray, 1909. Canad. Ent. 41: 357. ♀.

Biology: Peck, 1799. Mass. Agr. Rpt., p. 9. —Harris, 1841. Rpt. Ins. Mass. Injurious to Veg.,

p. 383. — Winchell, 1865. Boston Soc. Nat. Hist., Proc. 9: 321-325. —Riley, 1870. Amer. Ent.

and Bot. 2: 296. -Marlatt, 1897. U. S. Dept. Agr., Cir. 26 (ser. 2), 7 pp. —Webster, 1912. N.

Y. Ent. Soc, Jour. 20: 125-130 (number of moults). —Webster, 1912. Iowa Agr. Expt. Sta.,

Bui. 130: 167-193. —Wilson, 1913. Oreg. Agr. Expt. Sta., Rpt. for 1911-12, pp. 217.

— Ewing, 1917. Ent. Soc. Amer., Ann. 10: 330-336 (parthenogenesis). — Britton, 1921. Conn.

(State) Agr. Expt. Sta., Bui. 226: 199-201. —Miles, 1935. Ann. Appl. Biol. 22: 116-133.

—Carl, 1972. Ztschr. Angew. Ent. 71: 58-83.

Caliroa distincta Smith
B. C, Oreg., Nev., Calif.

Caliroa distincta Smith, 1971. U. S. Dept. Agr., Tech. Bui. 1420, p. 17. ♀.

Caliroa fasciata (Norton)
Ont., Maine to Va. w. to Iowa, 111. Host: Quercus velutina, Q. palustris, Q.
rubra, and probably other oaks.

Selandria fasciata Norton, 1864. Ent. Soc. Phila., Proc. 3: 9. ♀.

Caliroa floridana Smith
Fla.

Caliroa floridana Smith, 1971. U. S. Dept. Agr., Tech. Bui. 1420, p. 19. ♀.

Caliroa hyalina Smith
Oreg. Ecology: Adults collected from cherry.

Caliroa hyalina Smith, 1971. U. S. Dept. Agr. Tech. Bui. 1420, p. 19. ♀, ♂.

Caliroa labrata MacGillivray
Alta., B. C, Oreg., Nev., Calif. Host: Salix.

Caliroa labrata MacGillivray, 1909. Canad. Ent. 41: 360. ♂.

Caliroa liturata MacGillivray
Conn, to Fla. w. to Mont., Colo., La. Host: Prunus, peach, and plum.

Caliroa liturata MacGillivray, 1909. Canad. Ent. 41: 349. ♀.




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Caliroa lineata MacGillivray, 1909. Canad. Ent. 41: 350. ♀.

Caliroa loricata MacGillivray, 1909. Canad. Ent. 41: 351. ♀.

Caliroa (Eriocampoides) aviygdalina Rohwer, 1911. Ent. News 22: 263. ♀, ♂.

Biology: Cushman, 1911. U. S. Dept. Agr., Bur. Ent. Bui. 97: 91-102.

Caliroa lobata MacGillivray
Vt. to Va. w. to Iowa, Tex. Host: Quercus palustris, Q. velutina, and
probably other oaks.

Caliroa lobata MacGillivray, 1909. Canad. Ent. 41: 355. ♀.

Caliroa lorata MacGillivray
Maine to Ala. w. to Wis. Host: Castanea dentata.

Caliroa lorata MacGillivray, 1909. Canad. Ent. 41: 355. ♀.

Eriocampoides castaneae Rohwer, 1917. U. S. Natl. Mus., Proc. 53: 152. ♀.

Caliroa lunata MacGillivray
Newfoundland to Va. w. to Mich., 111., Tex.

Caliroa lunata MacGillivray, 1909. Canad. Ent. 41: 353. ♀.

Caliroa lata MacGillivray, 1909. Canad. Ent. 41: 361. ♀.

Caliroa nyssae Smith
Pa. to Ga. w. to Miss. Host: Nyssa sylvatica.

Caliroa Jiyssae Smith, 1971. U. S. Dept. Agr., Tech. Bui. 1420, p. 25. ♀, ♂.

Caliroa obsoleta (Norton)
Ont., Mass. to Ga. w. to Minn., Iowa, 111. Host: Quercus alba, Q. stellata, Q.
prinus, and probably other oaks.

Selayidria obsoleta Norton, 1867. Amer. Ent. Soc, Trans. 1: 254. ♀.

Selandria quercus alba Norton, 1867. Amer. Ent. Soc, Trans. 1: 258. ♀, ♂.

Selandria quercus-alba Cresson, 1880. Amer. Ent. Soc, Trans. 8: 60. Emend.

Eriocampoides quercus Konow, 1905. hi Wytsman, Gen. Ins., v. 29, p. 74. Emend.

Caliroa petiolata Smith
Pa., Md., Va. Host: Quercus palustris.

Caliroa petiolata Smith, 1971. U. S. Dept. Agr., Tech. Bui. 1420, p. 28. ♀, ♂.

Caliroa quercuscoccineae (Dyar)
Maine to N. C. w. to Minn., 111., Mo., La. Host: Quercus coccinea, Q.
velutina, Q. palustris, Q. alba, and probably other oaks.

Monostegia quercus-coccineae Dyar, 1894. Canad. Ent. 26: 42. 9 , larva.

Tribe HETERARTHRINI
Genus HETERARTHRUS Stephens

Phyllotoma Fallen, 1829. Monog. Tenthred. Suec, p. 25. Preocc. by Leach, 1819.

Type-species: Phyllotoma vagans Fallen. Desig. by Rohwer, 1911.
Heterarthrus Stephens, 1835. Illus. Brit. Ent., v. 7, p. 94.

Type-species: Tenthredo [Emphytus) ochropoda Klug. Monotypic.
Decatna Stephens, 1835. Illus. Brit. Ent., v. 7, p. 94.

Type-species: Decatria fuscipennis Stephens. Monotypic.
Druida Newman, 1838. Ent. Mag. 5: 484.

Type-species: Druida parviceps Newman. Monotypic.
Heterarthus Cameron, 1882. Monog. Brit. Phytophagous Hym., v. 1, p. 282. Emend.
Phlebatrophia MacGillivray, 1909. Canad. Ent. 41: 345.

Type-species: Phlebatrophia mathesoni MacGillivray. Monotypic.

This genus is Palaearctic with one introduced species in North America. All larvae are leaf-
miners.

Heterarthrus nemoratus (Fallen)
Newfoundland, N. S., N. B., Que., Maine, N. H., Mass., N. Y., Ont.; Europe.
Ecology: A leafminer. Introduced, first found at Pictou, N.S. in 1908. Host: Betula.
Hylotoma nemorata Fallen, 1808. Svenska Vetensk. Akad., Handl. 29: 47.
Phyllotoma nemoralis Fallen, 1829. Monog. Tenthred. Suec, p. 35.
Fenusa parviceps Newman 1837. Ent. Mag. 4: 261.

Phyllotoma tenella Zaddach, 1859. Programm Friedr. Col. Konigsberg, p. 28. ♀.
Phlebatrophia mathesoni MacGillivray, 1909. Canad. Ent. 41: 345. ♀.

Biology: Pierson, 1929. Jour. Econ. Ent. 22: 588-594. — Pierson, Taylor, and Wilkins, 1930.
Maine Forest Service, Cir. 1, 8 pp. —Glasgow, 1932. Jour. Econ. Ent. 25: 693-695.
— Pierson and Brower, 1936. Maine Forest Service and Hardwood Assoc, Bui. 11, 37 pp.
— Gobeil, 1937. Prov. de Quebec Min. des Terre et Forets, Bui. 1, 7 pp. — Lindquist, 1959.
Canad. Ent. 91: 625-627.




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Tribe FENUSINI

Genus METALLUS Forbes

Metallus Forbes, 1885. State Ent. 111. Rpt. 14: 87.

Type-species: Metallus rubi Forbes. Monotypic.
Entodecta Konow, 1886. Wien. Ent. Ztg. 5: 184.

Type-species: Tenthredo (Allantiis) pumila Klug. Desig. by MacGillivray, 1909.
Polybates MacGillivray, 1909. Ent. Soc. Amer., Ann. 2: 264.

Type-species: Polybates slossonae MacGillivray. Monotypic.

Metallus bensoni Smith
N. Y., B. C.

Metallus bensoni Smith, 1971. U. S. Dept. Agr., Tech. Bui. 1420, p. 34. ♀.

Metallus capitalis (Norton)
Newfoundland to N. Y. w. to 111.; Alaska, B. C, Oreg. Ecology: A leafminer.
Host: Rtibus.
Selandria capitalis Norton, 1867. Amer. Ent. Soc, Trans. 1: 247. ♀.
Entodecta humilis Konow, 1908. Ztschr. System. Hym. Dipt. 8: 84. ♀.
Polybates slossonae MacGillivray, 1909. Ent. Soc. Amer., Ann. 2: 265. ♀.
Polybates secundus Rohwer, 1910. U. S. Natl. Mus., Proc. 38: 202. ♀.

Metallus rohweri MacGillivray
N. S., Maine to Fla. w. to Ont., Wis., Ill, Mo. Ecology: A leafminer.
Host: Rubus.
Metallus rubi Forbes, 1885. State Ent. 111. Rpt. 14: 87. ♂. Preocc. by Boie, 1848.
Metallus rohweri MacGillivray, 1909. Ent. Soc. Amer., Ann. 2: 267. ♀.
Metallus bethunei MacGillivray, 1914. Canad. Ent. 46: 366. ♀, ♂.

Biology: Houghton, 1908. Ent. News 19: 212-216 {capitalis, not Norton). —Houghton, 1910.
Del. Col. Agr. Expt. Sta., Bui. 87: 3-15 (capitalis, not Norton). —Daniel, 1928. N. Y. State
Agr. Expt. Sta., Tech. Bui. 133: 3-38.

Genus MESSA Leach

Messa Leach, 1817. Zool. Misc., v. 3, p. 126.

Type-species: Tenthredo hortiilana Klug. Monotypic.
Fenusella Enslin, 1914. Deut. Ent. Ztschr., Beih., p. 270.

Type-species: Fenusa Wustneii Konow. Orig. desig.
Melanobates MacGillivray, 1916. Conn. State Geol. and Nat. Hist. Survey, Bui. 22: 158.
Type-species: Parabates leucostomus Rohwer. Monotypic.

Messa hortulana (Klug)
Mass.; Europe. Ecology: A leafminer. Introduced (?). Host: Populus.

Tenthredo (Emphytus) hortulana Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 276. ♀.
Phaenusa doederleinii Destefani, 1883. Nat. Sicil. 3: 12. ♀, ♂.
Phaenusa ticinensis Magretti, 1886. Soc. Ent. Ital., Bui. 18: 25. ♀.

Messa leucostoma (Rohwer)
N. B., Minn., Iowa, Colo., Alta., Wash., Oreg., Calif. Ecology: Adults
collected from Populus.
Parabates leucostoma Rohwer, 1910. U. S. Natl. Mus., Proc. 38: 202. ♀.
Blennocampa amara MacGillivray, 1923. Canad. Ent. 55: 161. ♀.

Messa nana (Klug)
Maine, Mass., N. Y., Ont.; Europe. Ecology: A leafminer. Introduced, first
recorded from Maine in 1967. Host: Betula.
Tenthredo (Allantus) yiana Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 72.
Tenthredo intercus Zetterstedt, 1838. Ins. Lapp., v. 1, p. 340.
FenuSa quercus Cameron, 1885. Monog. Brit. Phytophagous Hym., v. 2, p. 222.

Biology: Lindquist and Thomson, 1970. Canad. Ent. 102: 108-111.

Messa populifoliella (Townsend)
N. B. to Conn. w. to Man., S. Dak., Colo., N. Mex., Ariz., Calif.
Ecology: A leafminer. Host: Populus.
Blennocampa populifoliella Townsend, 1893. Canad. Ent. 25: 304.
Scolioneura populi Marlatt, 1895. Ent. Soc. Wash., Proc. 3: 235. ♀, ♂.
Scolioneura luteopicta Rohwer, 1911. U. S. Natl. Mus., Proc. 3: 235. ♀, ♂.




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Biology: Townsend, 1892. U. S. Dept. Agr., Insect Life 4: 26-27 (Cottonwood leafminer).
— Townsend, 1892. Zoe 3: 234-236 (Tineidae ? larvae). —Underwood and Titus, 1968.
Canad. Ent. 100: 407-411. -Brown and Eads, 1969. Jour. Econ. Ent. 62: 672-674.

Messa wuestneii (Konow)
Que. to Alaska s. to Colo., Calif.; Europe. Ecology: A leafminer. Host:
Salix.
Fenusa wustneii Konow, 1894. Wien. Ent. Ztg. 13: 91. ♀, ♂.
Fenusa alaskana Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 345. ♂.

Genus SETABARA Ross

Parabafes MacGillivray, 1909. Ent. Soc. Amer., Ann. 2: 262. Preocc.

Type-species: Parabates histrionicus MacGillivray. Orig. desig.
Setabara Ross, 1951. In Muesebeck, et al., U. S. Dept. Agr., Agr. Monog. 2: 31. N. name.

A monotypic genus found only in western North America.

Setabara histrionica (MacGillivray)
Colo., Idaho, Nev., Wash., Oreg., Calif. Ecology: Adults collected
from Primus.
Parabates histrionicus MacGillivray, 1909. Ent. Soc. Amer., Ann. 2: 263. ♀.

Genus BIDIGITUS Smith

Bidigitus Smith, 1967. Ent. Soc. Wash., Proc. 69: 280.

Type-species: Profenusa platani Burks. Orig. desig.

Bidigitus platani (Burks)
Calif. Ecology: A leafminer. Host: Platanus racemosa.
Profenusa platanaei}.) Burks, 1957. Ent. News 68: 207. 9, tJ.

Biology: Brown and Eads, 1965. Calif. Agr. Expt. Sta., Bui. 818: 14, 22-32.

Genus PROFENUSA MacGillivray

Profenusa MacGillivray, 1914. Canad. Ent. 46: 364.

Type-species: Profenusa collaris MacGillivray. Monotypic.
Gimnea Malaise, 1964. Ent. Tidskr. 85: 33.

Type-species: Tenthredo pygmaea Klug. Orig. desig.

Profenusa alumna (MacGillivray)
Maine to Va. w. to Wis., 111. Ecology: A leafminer. Host: Quercus rubra
and probably other oaks. References to alumna as a leafminer of Betula are actually

Profenusa thomsoni (Konow)

Messa alumna MacGillivray, 1923. 111. Univ. Bui. 20(50): 23. ♀.
Messa arnica MacGillivray, 1923. 111. Univ. Bui. 20(50): 23. ♂.
Messa appota MacGillivray, 1923. 111. Univ. Bui. 20(50): 24. ♂.
Profenusa mainensis Smith, 1966. Ent. Soc. Amer., Ann. 59: 720. 9, larva.

Profenusa canadensis (Marlatt)
Que., Ont. to D. C. w. to Iowa, Mo., Ark. Ecology: A leafminer. Host:
Crataegus, to a lesser extent Primus.
Scolioneura canadensis Marlatt, 1895. Ent. Soc. Wash., Proc. 3: 235. ♀.
Profenusa collaris MacGillivray, 1914. Canad. Ent. 46: 364. ♀, ♂.

Biology: Parrott and Fulton, 1915. N. Y. Agr. Expt. Sta., Bui. 411: 551-580. — Parrott and

Fulton, 1915. Jour. Agr. Res. 5: 519-528. -Hamilton, 1943. N. Y. Agr. Expt. Sta., Bui. 703:

59-61. -Hamilton, 1950. Jour. Econ. Ent. 43: 694-696.

Profenusa inspirata (MacGillivray)
Oreg., Nev., Calif. Ecology: A leafminer. Host: Quercus.

Parabates inspiratus MacGillivray, 1909. Ent. Soc. Amer., Ann. 2: 264. ♀.

Profenusa lucifex (Ross)
Maine, Ont, N. Y., 111. Ecology: Leafminer. Host: Quercus alba, Q. macrocarpa.

Fenusa lucifex Ross, 1936. 111. State Acad. Sci., Trans. 29: 266. ♀.

Biology: Lindquist and Jackson, 1965. Canad. Dept. Forestry, Bimonthly Prog. Rpt. 21(4): 1.

Profenusa thomsoni (Konow)
Que., Maine, Vt., Conn., Ont., Man.; Ecology: A leafminer. Introduced.
Host: Betula.
Fenusa thomsoni Konow, 1886. Wien. Ent. Ztg. 5: 270. ♀.




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Biology: Lindquist, 1959. Canad. Ent. 91: 625-627 (ahimna, not MacGillivray). —Watson,
1959. Canad. Ent. 91: 618-625 {ahimna not MacGillivray). —Martin, 1960. Canad. Ent. 92:
376-384.

Genus NEFUSA Ross

Nefusa Ross, 1951. In Muesebeck et al., U. S. Dept. Agr., Agr. Monog. 2, p. 31.
Type-species: Nefusa anita MacGillivray. Orig. desig.

The single species in this genus is known only from North America.

Nefusa ambigua (Norton)
Que., Maine to Tenn. w. to Wis. Ecology: A leafminer. Host: Viola.
Fenusa ambiguus Norton, 1867. Amer. Ent. Soc, Trans. 1: 225. ♂.
Messa anita MacGillivray, 1923. 111. Univ. Bui. 20(50): 23. ♀.

Biology: Shaw, 1940. Jour. Econ. Ent. 33: 951 {Fenusa sp.).

Genus PROLATUS Smith

Prolatus Smith, 1967. Ent. Soc. Wash., Proc. 69: 282.
Type-species: Prolatus artus Smith. Orig. desig.

A monotypic genus known only from western North America.

Prolatus artus Smith
Oreg.

Prolatus artus Smith, 1967. Ent. Soc. Wash., Proc. 69: 283. 9, 6 (?).

Genus FENUSA Leach

Fenusa Leach, 1817. Zool. Misc., v. 3, p. 126.

Type-species: Tenthredo (Emphytus) puynila Klug. Monotypic.
Kaliosysphinga Tischbein, 1846. Stettin Ent. Ztg. 7: 79.

Type-species: Kaliosysphinga dohrnii Tischbein. Monotypic.
Aphadnurus Costa, 1859. Fauna del Regno di NapoH, Imen., pt. 3, p. 40.

Type-species: Aphadnurus tantillus Costa. Monotypic.
Caliosysphinga Konow, 1905. In Wytsmann, Gen. Ins., v. 29, p. 89. Emend.
Kaliofenusa Viereck, 1910. N. J. State Mus., Ann. Rpt. 1909, p. 591.

Type-species: Fenusa ulmi Sundevall. Monotypic.

A genus of several species most of which are holarctic though two of them were probably in-
troduced into this continent.

Fenusa dohrnii (Tischbein)
Newfoundland to Pa. w. to B. C, Oreg., Utah., N. Mex.; Europe; Japan; S.
Africa. Ecology: A widespread leafminer. Host: Alnus. European alder leafminer.
Kaliosysphinga dohrnii Tischbein, 1846. Stettin Ent. Ztg. 7: 80.
Fenusa curtis Norton, 1861. Ent. Soc. Phila., Proc. 1: 199. ♀.
Phaenusa melanopoda Cameron, 1876. Glasgow Nat. Hist. Soc, Proc. 3: 6.
Fenusa nigricans Thomson, 1870. Opusc. Ent., 2nd Fasc, p. 271.
Fenella westwoodi Cameron, 1882. Monog. Brit. Phytophagous Hym., v. 1, p. 289.

Biology: Slingerland, 1905. N. Y. (Cornell) Agr. Expt. Sta. Bui. 233: 49-62. — Frankenhuyzen,
1970. Ent. Ber. 30: 49-52 (in Netherlands).

Fenusa pusilla (Lepeletier)
Newfoundland to Md. w. to Ont., Minn., Iowa; Wash., Oreg.; Europe.

Ecology: A leafminer. Introduced, first reported in Conn, in 1925. Host: Betula. Birch

leafminer.
Tenthredo (Eynphytus) pumila Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 277.

Preocc.
Dolerus (Fenusai) pusillus Lepeletier, 1823. Monog. Tenthred., p. 120.
Tenthredo pygmaea Zetterstedt, 1838. Ins. Lapp., v. 1, p. 340. ♀, ♂.
Aphadnurus tantillus Costa, 1859. Fauna del Regno di Napoli, Imen., pt. 3, p. 41. ♂.
Fenusa fuliginosa Healy, 1869. Entomologist 4: 225-227.
Fenusa minima Brischke, 1883. Schrift. Naturf. Gesell. Danzig 4: 264. ♀.




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Biology: Britton, 1925. Conn. (State) Agr. Expt. Sta. Bui. 265: 340. —Friend, 1931. Jour.
Econ. Ent. 24: 171-177. —Friend, 1933. Conn. (State) Agr. Expt. Sta. Bui. 348: 291-364.
—Cheng and LeRoux, 1965. Ent. Soc. Quebec, Ann. 10: 173-188. — Schread, 1971. Conn.
(State) Agr. Expt. Sta. Bui. 693: 5-19. —Cheng and LeRoux, 1970. Canad. Ent. 102:
995-1002 (factors in survival of immature stages).

Fenusa ulmi Sundevall
Que., Ont., Mass., N. Y., Mich.; Europe. Ecology: A leafminer. Introduced.
Host: Ulmus. Elm leafminer.
Femisa ulmi Sundevall, 1844. Skand. Naturf. Forhandl., p. 249.
Feiiusa intennedia Thomson, 1871. Hym. Scand., v. 1, p. 186. ♀.
Messa alsia MacGilhvray, 1923. 111. Univ. Bui. 20(50): 22. ♀.

Biology: Felt, 1898. N. Y. State Mus., Bui. 5(23): 237 (an elm leafminer). — Slingerland, 1905.
N. Y. (Cornell) Agr. Expt. Sta. Bui. 233: 49-62.

Genus FENELLA Westwood

Fenella Westwood, 1840. Introduct. Modern Class. Ins., v. II, p. 54.

Type-species: Fenella yiigrita Westwood. Monotypic.
Melinia Costa, 1859. Fauna del Regno di Napoli, Imen., pt. 3, p. 41.

Type-species: Melinia minutissima Costa. Desig. by Rohwer, 1911.
Paraphyllotoma Forsius, 1930. Notulae Ent. 10: 103.

Type-species: Paraphyllotoma judaica Forsius Orig. desig.

Taxonomy: Benson, 1953. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 22: 136-138 (key to
European species).

Fenella nigrita Westwood
Conn., Ont., Mich.; Europe. Ecology: A leafminer. Introduced (?). Host:
Potentilla, Agrimonia. Agriynonia was recorded as a host-plant in Europe.
Fenella nigrita Westwood, 1840. Introduct. Modern Class. Ins., v. II, p. 54.

Subfamily BLENNOCAMPINAE

A diverse subfamily with various hosts. Most of these sawflies are small to medium, stout in-
sects. Most larvae are external leaf feeders, and adults fall and feign death when disturbed.

Revision: Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, 179 pp. (adults and larvae of
Nearctic species).

Tribe LYCAOTINI

Revision: Smith, 1973. Pan-Pacific Ent. 49: 93-101 (S. Amer.).

Taxonomy: Benson, 1966. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 35: 75-77 (key to
world genera).

Genus BLENNOGENERIS MacGillivray

Blennogeneris MacGillivray, 1923. 111. Univ. Bui. 20(50): 8.

Type-species: Blennocampa typicella MacGillivray. Monotypic.
Lycaotella Ross, 1932. Canad. Ent. 64: 41.

Type-species: Selandria (Hoplocampa) spissipes Cresson. Orig. desig.

Blennogeneris coloradensis (Rohwer)
Man., Colo., Idaho, Utah, Wash.

Lycaota coloradensis Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 384.

Blennogeneris gittinsi Smith
Idaho, Wash.

Blemtogeneris gittinsi Smith, 1969. U. S. Dept. Agr. Tech. Bui. 1397, p. 24. ♀.

Blennogeneris spissipes (Cresson)
Ont., Minn., Colo. w. to B. C, Wash., Oreg., Calif. Ecology: Larva lives in
terminal bud gall. Host: Symphoricarpos.

Selandria (Hoplocampa) spissipes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 14. ♀.

Selandria [Hoplocampa) lenis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 14. ♂.

Lycaota bnineri Rohwer, 1908. N. Y. Ent. Soc, Jour. 14: 104. ♀. N. syn.

Lycaota spissipes bninneus Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 384. ♀.

Blennocampa typicella MacGillivray, 1923. 111. Univ. Bui. 20(50): 8. ♂.

Biology: Ross, 1932. Canad. Ent. 64: 42-44.




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Genus LYCAOTA Konow

Lycaota Konow, 1903. Ztschr. System. Hym. Dipt. 3: 147.

Type-species: S€la7idria sodalis Cresson. Desig. by Rohwer, 1911.

Lycaota bouquetensis Smith
S. Calif.

Lycaota bouquetensis Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 29. ♀.

Lycaota janetae Smith
Oreg., Calif. Ecology: Adult collected from Symphoricarpos.

Lycaota janetae Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 29. ♀, ♂.

Lycaota sodalis (Cresson)
Sask., Colo., Utah w. to B. C, Wash., Oreg.

Selandria sodalis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 14. ♀.

Lycaota fusca Rohwer, 1908. Canad. Ent. 40: 180. ♂.

Tribe TOMOSTETHINl
Genus TOMOSTETHUS Konow

Tomostethus Konow, 1886. Wien. Ent. Ztg. 5: 184, 214.

Type-species: Tenthredo nigrita Fabricius. Desig. by Rohwer, 1911.

A small holarctic genus.

Tomostethus multicinctus (Rohwer)
Mass., Ont. to N. C. w. to S. Dak., Kans., Okla., Tex.; Oreg., Calif. Host:
Fraxiyiiis aiuericana, F. oregona. Brown-headed ash sawfly.
Monophadnus multicinctus Rohwer, 1909. Canad. Ent. 41: 90. ♂.

Biology: Sasscer, 1911. Ent. Soc. Wash., Proc. 13: 107-110. — Langford and McConnell, 1935.
Jour. Econ. Ent. 28: 208-210. — MacNay, 1957. Canad. Ins. Pest Rev. 35: 184. — Armitage,
1950. Calif. Dept. Agr., Bui. 39: 183.

Genus TETHIDA Ross

Tetliida Ross, 1937. 111. Biol. Monog. 15(2): 96.

Type-species: Tenthredo cordigera Beauvois. Orig. desig.

This genus contains a single North American species.

Tethida cordigera (Beauvois)
N. B., Que. to Fla. w. to Sask., Mont, S. Dak., Nebr., Kans., Tex. Host:
Fraxiniis americana, F. nigra, F. pennsylvanica. Black-headed ash sawfly.

Tenthredo cordigera Beauvois, 1809. Ins. Afr. Amer., p. 97. ♀.

Tenthredo (Selandria) tarda Harris, 1835. hi Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 583. Nom. nud.

Tenthredo tardus Say, 1836. Boston Jour. Nat. Hist. 1: 218. ♀.

Selayidria dubia Cresson, 1865. Ent. Soc. Phila., Proc. 4: 244. ♀.

Biology: Osborne, 1884. Canad. Ent. 16: 148-152. —Osborne, 1884. Ont. Ent. Soc, Ann. Rpt.
15: 32-34. -Packard, 1890. U. S. Ent. Comm. Rpt. 4 (Bui. 7, rev.), p. 544.

Tribe PHYMATOCERINI

Genus PHYMATOCERA Dahlbom

Tenthredo subg. Pkymatocera Dahlbom, 1835. Conspectus Tenthred., Siricid., Oryssin.
Scand., p. 11.

Type-species: Tenthredo (Allantus) aterrima Klug. Monotypic.
Pectinia Brulie, 1846. In Lepeletier, Hist. Nat. Ins., Hym., p. 664.

Type-species: Tenthredo (.Allantus) aterrima Klug. Orig. desig.
Melanoselandria Ashmead, 1900. In Smith, N. J. State Bd. Agr. Ann. Rpt. 27: 606. Nom.

nud.
Hypargyricus MacGillivray, 1908. Canad. Ent. 40: 290.

Type-species: Hypargyricus infuscatus MacGillivray. Orig. desig.
Melanoselandria MacGillivray, 1909. Canad. Ent. 41: 404.

Type-species: Melanoselandria zabriskiei MacGillivray. Monotypic.

Species of this holarctic genus are associated with Liliaceae.




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Phymatocera fumipennis (Norton)
Que. to Ala. w. to Wis., 111., Mo. Host: Smilacma.

Selandria fiimipennis Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 222. ♀, ♂.

Phymatocera offensa (MacGillivray)
Que., Maine to B. C. s. to Ont., 111., Colo., Utah, Calif. Host: Smilacina.

Paracharactus offensus MacGillivray, 1923. 111. Univ. Bui. 20(50): 28. ♀.

Phymatocera racemosae Smith
Que. to N. C. w. to Wis., Ill; B. C. Ecology: Adults collected from
Smilacinai?), Polygonatum(^).

Phymatocera racemosae Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 49. ♀, ♂.

Phymatocera similata (MacGillivray)
Ont., N. Y. to B. C. s. to Mich., Ill, Iowa, Nebr., Colo., Utah, Calif.
Host: Smilacina.

Rhadinoceraea similata MacGillivray, 1908. Canad. Ent. 40: 290. ♀.

Hypargyricus infuscatus MacGillivray, 1908. Canad. Ent. 40: 290. ♀.

Rhadinoceraea lucida Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 229. ♀.

Phymatocera smilacinae Smith
Que. to Va. w. to Wis., 111. Ecology: Adults collected from Smilacina and
Polygonatum.

Phymatocera smilacinae Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 55. ♀, ♂.

Unplaced Taxa of Phymatocera Dahlbom

Phymatocera rusculla (MacGillivray)
Oreg. (Marys Peak).

Monophadmis ruscullus MacGillivray, 1923. Psyche 30: 80. ♂.

Phymatocera zabriskiei (MacGillivray)
No locality.

Melanoselandria zabriskiei MacGillivray, 1909. Canad. Ent. 41: 404.

Genus PARACHARACTUS MacGillivray

Paracharactus MacGillivray, 1908. Canad. Ent. 40: 292.

Type-species: Paracharactus obscuratus MacGillivray. Orig. desig.
Dicrostema Benson, 1952. Handb. Ident. Brit. Ins., v. 6, pt. 2(b), p. 101.

Type-species: Selandria gracilicomis Zaddach. Orig. desig.

A small holarctic genus. Adults are most commonly collected by sweeping grasses and sedges.

Paracharactus montivagus (Cresson)
Mont., Idaho, Wash., Oreg., Nev., Calif.

Selandria montivaga Cresson, 1880. Amer. Ent. Soc, Trans. 8: 13. ♀.

Selaiidria scelesta Cresson, 1880. Amer. Ent. Soc, Trans. 8: 14. ♂.

Paracharactus niger (Harrington)
Que., Maine to Md. w. to Minn., Iowa, Kans. Host: Carex.

Phymatocera nigra Harrington, 1889. Canad. Ent. 21: 96. ♀.

Monophadnus distinctus MacGillivray, 1908. Canad. Ent. 40: 291. ♀.

Tomostethus nortonii MacGillivray, 1908. Canad. Ent. 40: 291. ♀.

Paracharactus rudis (Norton)
Que., Maine to Ga. w. to Sask., S. Dak., Colo.

Selandria rudis Norton, 1861. Boston Soc. Nat. Hist., Proc 8: 221. ♀.

Selandria nifula Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 221. ♂.

Paracharactus obscuratus MacGillivray, 1908. Canad. Ent. 40: 293. ♀.

Genus RHADINOCERAEA Konow

Species of this holarctic genus are associated with Liliaceae and Iridaceae.

Genus RHADINOCERAEA Subgenus RHADINOCERAEA Konow

Rhadinoceraea Konow, 1886. Wien. Ent. Ztg. 5: 184, 211.

Type-species: Tenthredo (.Allantus) micans Klug. Desig. by Rohwer, 1911.

Rhadinoceraea brysonensis Smith
Calif. (Bryson).

Rhadinoceraea (Rhadinoceraea) brysonensis Smith, 1969. U. S. Dept. Agr., Tech. Bui.
1397, p. 67. ♀.

Rhadinoceraea ctenidium Smith
Calif. (Mt. Pinos, Kern Co., 8900 ft.).

Rhadinoceraea (Rhadinoceraea) ctenidiuyn Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397,
p. 67. ♀.

Rhadinoceraea nigra (Rohwer)
S. Calif. Host: Calockortus venustus.

Paracharactus niger Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 231. ♀.




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Rhadinoceraea utahensis Smith
N. Utah.

Rhadinoceraea (Rhadinoceraea) utahensis Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397,
p. 70. ♀, ♂.

Genus RHADINOCERAEA Subgenus VERATRA Smith

Rhadinoceraea subg. Veratra Smith, 1969. U. S. Dept. Agr. Tech. Bui. 1397, p. 70.
Type-species: Selandria nubilipennis Norton. Orig. desig.

Rhadinoceraea aldrichi (MacGillivray)
Alta., Idaho, B. C, Wash., Oreg., Nev., Calif. Host: Veratrum.
Pareophora aldrichi MacGillivray, 1923. 111. Univ. Bui. 20(50): 28. ♀, ♂.

Rhadinoceraea insularis (Kincaid)
Alaska, B. C, Wash., Oreg., Calif. Host: Veratrum.

Monophadnus insularis Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 346. ♂.

Rhadinoceraea jacintensis Smith
S. Calif.

Rhadinoceraea (Veratra) jacintensis Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 75. ♀, ♂.

Rhadinoceraea nubilipennis (Norton)
N. B., Que. s. to N. C. Host: Veratnim.

Selandria nubilipennis Norton, 1867. Amer. Ent. Soc, Trans. 1: 252. ♀, ♂.

Neopareophora scelesta MacGillivray, 1908. Canad. Ent. 40: 289. ♀.

Monophadnus planus MacGilHvray, 1921. Brooklyn Ent. Soc, Bui. 16: 23. ♂. Preocc. by

Klug, 1814.
Monophadnus vapularis Benson, 1930. Entomologist 63: 107. N. name for planus

MacGillivray.

Genus LAGONIS Ross

Lagonis Ross, 1937. 111. Biol. Monog. 15(2): 99.

Type-species: Selandria nevadensis Cresson. Orig. desig.

A small genus known from western North America and east Asia.

Lagonis nevadensis (Cresson)
Alta., Wyo., Utah w. to B. C, Wash., Oreg., Calif. Host: Sambucus.
Selandria nevadensis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 13. ♀.
Paracharactus leucostomus Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 230. ♀.

Genus MONOPHADNUS Hartig

Tenthredo subg. Monophadnus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst
Einleitung Naturgesch. Hym., p. 271.

Type-species: Tenthredo albipes Gmehn. Desig. by Ashmead, 1898.
Monophadnus subg. Doderia Malaise, 1935. Ent. Tidskr. 56: 167.

Type-species: Tenthredo (Allantus) spinolae Klug. Orig. desig.

Monophadnus aequalis
***authority mismatch
MacGiUivray. Ont., Maine to Md. w. to Alta., Colo. Host: Ranunculus.
Monophadnus aequalis MacGillivray, 1908. Canad. Ent. 40: 292. ♀.
Monophadnus plicatus MacGillivray, 1908. Canad. Ent. 40: 292. ♀, ♂.
Monophadnus transversus MacGillivray, 1908. Canad. Ent. 40: 292. ♀.
Monophadnus tnincatus Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 232. ♀.

Monophadnus assaracus MacGillivray
Oreg.

Monophadnus assaracus MacGillivray, 1923. 111. Univ. Bui. 20(50): 26. ♂.

Monophadnus bakeri Smith
Md., Va., 111., Mont., Kans.

Monophadnus bakeri Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 86. ♀.

Monophadnus californicus (Rohwer)
Oreg., Calif. Host: Ranunculus.

Neocharactus californicus Rohwer, 1909. Canad. Ent. 41: 89. ♀.

Monophadnoides conductus MacGillivray, 1923. 111. Univ. Bui. 20(50): 24. ♀.

Monophadnus contortus
***authority mismatch
(MacGillivray). Mont., Idaho, B. C, Oreg., Calif.

Monophadnoides contorta MacGillivray, 1923. Psyche 30: 78. ♀.

Monophadnus aeratus MacGillivray, 1923. Psyche 30: 79. ♂.

Apha7iisus obsitiis MacGillivray, 1923. 111. Univ. Bui. 20(50): 7. ♀.

Apha7iisus occiduus MacGillivray, 1923. 111. Univ. Bui. 20(50): 7. ♀.

Monophadnus lattini Smith
N. H., Mass., Conn., Wis., Man., Minn.

Monophadnus lattini Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 91. ♀, ♂.




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Monophadnus pallescens (Gmelin)
Newfoundland to N. Y. w. to Mich.; B. C; Europe. Introduced (?). Host:

Ranunculus. The host was recorded in Europe.
Tenthredo albipes Gmelin, 1790. In Linnaeus, Syst. Nat., Ed. 13, p. 2667. Preocc. by

Geoffroy, 1785.
Tenthredo pallescens Gmelin, 1790. In Linnaeus, Syst. Nat., Ed. 13, p. 2668.
Selajidria tiliae Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 221. ♀.
Monophadnus bipurictatus MacGillivray, 1908. Canad. Ent. 40: 292. ♀.
Monophadnus furvus Benson, 1930. Entomologist 63: 107. Unnecessary n. name for

bipunctata MacGillivray, but also preocc. by Konow, 1898.

Genus STETHOMOSTUS Benson

Stethomostus Benson, 1939. Ent. Monthly Mag. 75: 111.

Type-species: Tenthredo fuliginosa Schrank. Orig. desig.

Stethomostus fuliginosus (Schrank)
N. S., Que. to Md. w. to Mich.; Europe; Japan. Introduced (?). Host:
Ranunculus.
Tenthredo fuliginosa Schrank, 1781. Enum. Ins. Austriae, p. 334.
Tenthredo (Selandria) fiiscus Lepeletier, 1823. Monog. Tenthred., p. 106. ♂.
Tenthredo (Selandria) fraxini Lepeletier, 1823. Monog. Tenthred., p. 106. ♀, ♂.
Monophadnus fukaii Rohwer, 1910. U. S. Natl. Mus., Proc. 39: 108. ♀, ♂.

Genus EUTOMOSTETHUS Enslin

Tomostethus subg. Eutomostethus Enslin, 1914. Deut. Ent. Ztschr., Beih., p. 286.

Type-species: Tenthredo luteiventris Klug. Orig. desig.
Tomostethus subg. Atomostethus Enslin, 1914. Deut. Ent. Ztschr., Beih. p. 287.

Type-species: Tenthredo ephippium Panzer. Orig. desig.
Tomostethopsis Sato, 1928. Insecta Matsumurana 2: 178.

Type-species: Tomostethopsis metallicus Sato. Orig. desig.
Forsia Malaise, 1932. Arkiv for Zool. 23: 29.

Type-species: Forsia tomostethi Malaise. Orig. desig.

Most of the species of this genus are found in Asia.

Eutomostethus ephippium (Panzer)
Newfoundland, N. S., Que., Ont. s. to Md.; Tex.; B. C, Wash.; Europe, N.
Africa, Asia Minor. Introduced (?). Host: Poa and other Gramineae. Hosts recorded in
Europe.
Tenthredo ephippium Panzer, 1798. Faunae Ins. German., heft 52, pi. 5. ♀.
Selandria inhabilis Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 220. ♀.

Eutomostethus luteiventris (Klug)
N. S., Que. to Md. w. to Mich.; Alta., B. C, Wash., Oreg.; Europe. Ecology:
Larva bores in sterile shoots of Juncus until last instar when it emerges and feeds
externally, according to European observations. Introduced (?). Host: Juncus.
Tenthredo luteiventris Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 56.
Tenthredo (Selandria) fuscipennis Lepeletier, 1823. Monog. Tenthred. p. 107. ♀, ♂.

Tribe BLENNOCAMPINI

Genus ARDIS Konow

Ardis Konow, 1886. Wien. Ent. Ztg. 5: 184, 188.

Type-species: Tenthredo (Allantus) bipunctata Klug. Desig. by Rohwer, 1911.
Valco Ross, 1937. 111. Biol. Monog. 15(2): 99.

Type-species: Selandria irrogata Cresson. Orig. desig.

A small holarctic genus associated with roses.

Ardis atrata (Harrington)
B. C., Wash., Oreg., CaHf. Ecology: Adults collected from Rosa.
Monophadnus at rat us Harrington, 1894. Canad. Ent. 26: 193. ♂.
Monophadnoides circina MacGillivray, 1923. 111. Univ. Bui. 20(50): 24. ♂.
Paracharactus obtentus MacGillivray, 1923. 111. Univ. Bui. 20(50): 28. ♀.




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Ardis brunniventris (Hartig)
Maine to N. W. T. s. to N. C, 111., N. Mex, Calif.; Europe to Siberia,
Japan. Ecology: Widespread. Larva bores and feeds in shoots of wild and cultivated
roses. Host: Rosa. Previous reports of Ardis sulcata (Cameron) in N. Amer. were
based on a misidentification and pertain to this species.

Tenthredo (Allantus) hipunctata Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 215. ♀. Preocc. by MuUer, 1776.

Tenthredo (Monophadnns) brunniventris Hartig, 1837. Fam. Blattwespen und Holzwespen,
nebst Einleitung Naturgesch. Hym., p. 274. ♀.

Selandria irrogata Cresson, 1880. Amer. Ent. Soc, Trans. 8: 13. ♀.

Aphanisus odoratiis MacGillivray, 1908. Canad. Ent. 40: 296. ♀.

Aphanisus parallelus MacGillivray, 1923. 111. Univ. Bui. 20(50): 7. ♀.

Taxonomy: Smith, 1965. Pan-Pacific Ent. 41: 9-10 (synonymy). —Smith, 1966. Ent. Soc.
Amer., Ann. 59: 1292 (larva separated from related species).

Biology: Eide, 1948. Jour. Econ. Ent. 41: 819-821. — Middlekauff, 1958. Pan-Pacific Ent. 34:
202.

Genus MONARDIS Enslin

Monardis Enslin, 1914. Deut. Ent. Ztschr., Beih., p. 284.
Type-species: Tenthredo plana Klug. Monotypic.

Monardis pulla Smith
Sask. to Colo., Utah and w. to B. C.

Mo^iardis pulla Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 114. ♀.

Genus APAREOPHORA Sato

Apareophora Sato, 1928. Insecta Matsumurana 2: 185.

Type-species: Apareophora forsythiae Sato Orig. desig.

This small genus is found in east Asia and North America.

Apareophora dyari (Benson)
N. B. to N. Y. and Ont. Host: Spiraea salicifolia.

Bleymocampa spiraeae Dyar, 1895. Canad. Ent. 27: 194. 9, d, larva. Preocc. by Brischke

and Zaddach, 1883.
Bleymocampa dyari Benson, 1930. Entomologist 63: 107. N. name.

Biology: Dyar, 1895. Canad. Ent. 27: 194.

Apareophora rossi Smith
Maine w. to Wis., Minn.

Apareophora rossi Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 118. ♀, ♂.

Genus EUPAREOPHORA Enslin

Eupareophora Enslin, 1914. Deut. Ent. Ztschr., Beih., p. 283.

Type-species: Blennocampa exarmata Thomson. Monotypic.

Only two species are known, the type-species from Europe and the species below.

Eupareophora parca (Cresson)
N. B. to Md. w. to Sask., N. Dak., Iowa, Kans., Tex.; Oreg., Calif. Host.

Fraxinus pennsylvanica, F. nigra, F. americana, F. oregona, Carya illinoensis,

Chionanthus(^).
Selandria parca Cresson, 1880. Amer. Ent. Soc, Trans. 8: 13. ♂.
Periclista chionanthi Dyar, 1898. N. Y. Ent. Soc, Jour. 6: 132. 9, larva.
Monophadnns 7Hinutiis MacGillivray, 1908. Canad. Ent. 40: 291. ♀.
Pareophora guana MacGillivray, 1923. 111. Univ. Bui. 20(50): 28. ♂.
Pareophora guara MacGillivray, 1923. Brooklyn Ent. Soc, Bui. 18: 54. ♀.

Biology: Packard, 1890. U. S. Ent. Comm. Rpt. 5 (Bui. 7, rev.), p. 545 (Selandria sp.).

Genus PERICLISTA Konow

Larvae of this genus feed on the foliage of Qnercus and Carya. The genus is holarctic as well
as being found in Chile and southern Argentina.

Revision: Stannard, 1949. Amer. Ent. Soc, Trans. 75: 7-42.




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Genus PERICLISTA Subgenus PERICLISTA Konow

Perklista Konow, 1886. Wien. Ent. Ztg. 5: 184, 186.

Type-species: Tenthredo (Allantus) lineolata Klug. Desig. by Rohwer, 1911.
Mogerus MacGillivray, 1895. Canad. Ent. 27: 281. Unnecessary n. name for Periclista

Konow.
Isodyctium Ashmead, 1898. hi Dyar, N. Y. Ent. Soc, Jour. 6: 127.

Type-species: Mogerus caryicoius Dyar. Orig. desig.
Apericlista Enslin, 1914. Deut. Ent. Ztschr., Beih., p. 265.

Type-species: Tenthredo albipennis Zaddach. Orig. desig.
Neodisia Malaise, 1964. Ent. Tidskr. 85: 22.

Type-species: Periclista andrei Konow. Orig. desig.
Comaria Malaise, 1964. Ent. Tidskr. 85: 22.

Type-species: Comaria fumipennis Malaise. Monotypic.

Taxonomy: Benson, 1968. Brit. Mus. (Nat. Hist.) Ent., Bui. 22: 152-153 (key to European
species).

Species Group Lineolata

Periclista albicollis (Norton)
Maine, Ont. to Va. w. to Man., Iowa, Kans., Okla., Tex. Host: Quercus alba,
Q. ilicifolia, Q. macrocarpa, Q. rubra, Q. velutina.

Selandria albicollis Norton, 1872. Amer. Ent. Soc, Trans. 4: 85. ♂.

Periclista mutabilis Konow, 1904. Ztschr. System. Hym. Dipt. 4: 241. ♀, ♂.

Periclista quercus Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 232. ♀, ♂.

Periclista linea Stannard
Oreg., Calif. Host: Quercus agrifolia.

Periclista linea Stannard, 1949. Amer. Ent. Soc, Trans. 75: 19. ♀, ♂.

Biology: Beer, 1955. Kans. Ent. Soc, Jour. 28: 19-26 (also description of larva).

Periclista media (Norton)
N. H., Mass. to Fla. w. to Man., Wis., Iowa. Host: Quercus alba.
Selandria media Norton, 1864. Ent. Soc Phila., Proc Z:9. ♀, ♂.
Selayidria (Hoplocanipa) floridana Cresson, 1880. Amer. Ent. Soc, Trans. 8: 12. ♂.
Periclista piirpuridorsum Dyar, 1898. N. Y. Ent. Soc, Jour. 6: 129. ♀, ♂.
Periclista cotifiisa MacGillivray, 1908. Canad. Ent. 40: 291. ♀.
Periclista similaris Rohwer, 1917. U. S. Natl. Mus., Proc. 53: 155. ♀.

Taxonomy: Middleton, 1922. U. S. Natl. Mus., Proc 61: 4-6 (larva, similaris).

Periclista naranga Stannard
Calif.

Periclista naranga Stannard, 1949. Amer. Ent. Soc, Trans. 75: 20. ♀.

Periclista spicula Stannard
Calif.

Periclista spicula Stannard, 1949. Amer. Ent. Soc, Trans. 75: 21. ♂.

Species Group Melanocephala

Periclista bipartita (Cresson)
Mass. to Fla. w. to Mo., Tex. Host: Quercus coccinea.

Selandria (Blennocampa) bipartita Cresson, 1880. Amer. Ent. Soc, Trans. 8: 12. ♂.

Mogerus emarginatus MacGillivray, 1895. Canad. Ent. 25: 281. ♂.

Isodyctium murtfeldtiae Dyar, 1898. N. Y. Ent. Soc, Jour. 6: 135. ♀.

Periclista diluta (Cresson)
Maine, Ont. to Fla. w. to Wis., Kans., Tex. Host: Quercus alba, Q. prinus.

Selandria (Monophadniis) diluta Cresson, 1880. Amer. Ent. Soc, Trans. 8: 12. ♀.

Isodyctium subgregarium Dyar, 1898. N. Y. Ent. Soc, Jour. 6: 134. ♀, ♂.

Taxonomy: Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 127 (male).

Periclista entella MacGillivray
Oreg., Calif. Ecology: Adults collected from Quercus.
Periclista entella MacGillivray, 1923. 111. Univ. Bui. 20(50): 29. ♂.

Taxonomy: Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 128 (female).

Periclista marginicollis (Norton)
Conn., Ont. to Fla. w. to Iowa, Kans., Okla., Tex. Host: Carya, Quercus

alba(n.
Tenthredo (Allantus) ynarginicollis Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot.

Zool. Mass., p. 583. Nom. nud.
Selandria marginicollis Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 220. ♀.
Mogerus caryicoius Dyar, 1897. N. Y. Ent. Soc, Jour. 5: 193. ♀, ♂.




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IsidyctiumV.) atratum MacGillivray, 1908. Canad. Ent. 40: 290. ♀.
Periclista hicoriae Rohwer, 1917. U. S. Natl. Mus., Proc. 53: 154. ♀.
Periclista xanthognatha Rohwer, 1917. U. S. Natl. Mus., Proc. 53: 156. ♀.
Periclista plesia Rohwer, 1920. U. S. Natl. Mus., Proc. 57: 211. ♀.
Periclista pecanivora Rohwer, 1920. U. S. Natl. Mus., Proc. 57: 212. ♀.

Taxonomy: Middleton, 1922. U. S. Natl. Mus., Proc. 61: 2-4 (larva). —Smith, 1969. U. S. Dept.
Agr., Tech. Bui. 1397, p. 129 (larva).

Periclista rileyi (Cresson)
Fla., 111., Mo., Tex.

Selandria (Monophadnus) rileyi Cresson, 1880. Amer. Ent. Soc, Trans. 8: 13. ♀.
Isodyctium floridense Dyar, 1898. N. Y. Ent. Soc, Jour. 6: 134. ♀.

Periclista stannardi Smith
Iowa, Tex.

Periclista stannardi Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 130. ♀.

Periclista sulfurana Stannard
N. Y., Mich., 111., Iowa. Host: Quercus.

Periclista sulfurana Stannard, 1949. Amer. Ent. Soc, Trans. 75: 28. ♀, ♂.

Species Group Californica

Periclista californica Rohwer
Calif.

Periclista californica Rohwer, 1917. U. S. Natl. Mus., Proc 53: 154. ♀.

Periclista electa MacGillivray
Oreg., CaHf. Ecology: Adults collected from Quercus.

Periclista electa MacGillivray, 1923. Psyche 30: 80. ♂.

Periclista vergorba Stannard
CaHf.

Periclista vergorba Stannard, 1949. Amer. Ent. Soc, Trans. 75: 32. ♀.

Genus PERICLISTA Subgenus NEOCHARACTUS MacGillivray

Neocharactus MacGillivray, 1908. Canad. Ent. 40: 293.

Type-species: Neocharactus bakeri MacGilhvray. Monotypic

Aphaiiisus MacGillivray, 1908. Canad. Ent. 40: 295.

Type-species: Aphayiisus lobatus MacGillivray. Orig. desig.

Periclista bakeri (MacGillivray)
Calif. Ecology: Adults collected from Quercus agrifolia.

Neocharactus bakeri MacGillivray, 1908. Canad. Ent. 40: 293. ♂.

Periclista leucostoma Rohwer, 1909. Canad. Ent. 41: 397. ♀, ♂.

Periclista inaequidens (Norton)
N. H. to Md. w. to Wis., Ill, Tex. Host: Quercus alba.

Selandria inaequidens Norton, 1872. Amer. Ent. Soc, Trans. 4: 84. ♀.

Isodyctium infrequens Dyar, 1898. N. Y. Ent. Soc, Jour. 6: 134. ♀.

Periclista pallipes (Provancher)
Calif. Ecology: Adults collected from Quercus agrifolia, Q. wislizenii.

Monophadmis pallipes Provancher, 1895. Nat. Canad. 22: 80. ♀, ♂.

Periclista occidentalis Rohwer, 1909. Canad. Ent. 41: 398. ♀, ♂.

Periclista subtruncata Dyar
N. Y. to Fla. w. to III, Tex. Host: Quercus coccinea.

Tenthredo (Allantus) labiata Klug, 1818. Mag. Gesell Naturf. Freunde Berhn 8: 73. ♀.
Preocc. by Geoff roy, 1785.

Periclista subtnmcata Dyar, 1898. N. Y. Ent. Soc, Jour. 6: 131. ♀.

Aphanisus lobatus MacGillivray, 1908. Canad. Ent. 40: 295. ♀.

Aphanisus muricatus MacGillivray, 1908. Canad. Ent. 40: 296. ♀.

Genus MONOPHADNOIDES Ashmead

Monophadnoides Ashmead, 1898. Canad. Ent. 30: 253.

Type-species: Monophadnus rubi Harris. Orig. desig.
Claremontia Rohwer, 1909. Canad. Ent. 41: 397.

Type-species: Claremontia typica Rohwer. Orig. desig.
Monophadnus subg. Pseudomonophadnus Malaise, 1935. Ent. Tidskr. 56: 167.

Type-species: Tenthredo geniculata Hartig. Orig. desig.
Monophadnus subg. Pseudoblennocampa Malaise, 1935. Ent. Tidskr. 56: 167.

Type-species: Tenthredo {Allantus) tenuicorais Klug. Orig. desig.

A holarctic genus mostly associated with Rosaceae though the host for only one North Amer-
ican species is known.




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Monophadnoides atratus (MacGillivray)
N. W. T. (Norman Wells), Alaska s. to Oreg.
Blennocampa atrata MacGillivray, 1893. Canad. Ent. 25: 239. ♀.
Eryfhraspides ashmeadi Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 345. ♀, ♂.

Monophadnoides conspiculatus MacGillivray
N. S., Ont. s. to N. C, Tenn.

Monophadnoides conspiculata MacGillivray, 1908. Canad. Ent. 40: 293. ♀.

Monophadnoides geniculatus (Hartig)
Labrador to N. W. T. s. to Ga., Ark., Tex., Colo., Calif.; Europe and Asia.

Ecology: Widespread. Host: Rubus. An extremely variable species in N. Amer.

Raspberry sawfly.
Tenthredo geniculatus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung

Naturgesch. Hym., p. 274.
Selandria {Hoplocampa) nibi Harris, 1845. In Darling, New Haven Hort. Soc, Trans., p.

13.
Selayidria nigella Cresson, 1880. Amer. Ent. Soc, Trans. 8: 12. ♀.
Monophadnus hudscmicus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 176. ♂.
Monophadnus atracomis MacGillivray, 1893. Canad. Ent. 25: 239. ♀.
Blennocampa Gillettei Weldon, 1907. Canad. Ent. 39: 304. ♀.
Monophadnoides conspicuus MacGillivray, 1908. Canad. Ent. 40: 293. ♀.
Monophadnoides consobrinus MacGillivray, 1908. Canad. Ent. 40: 294. ♀.
Monophadnoides concessus MacGillivray, 1908. Canad. Ent. 40: 294. ♀.
Monophadnoides crassus MacGillivray, 1908. Canad. Ent. 40: 294. ♀.
Monophadnoides conspersus MacGillivray, 1908. Canad. Ent. 40: 294. ♀.
Monophadnoides costalis MacGillivray, 1908. Canad. Ent. 40: 295. ♀.
Monophadnoides coracinus MacGillivray, 1908. Canad. Ent. 40: 295. ♂.
Monophadnoides collaris MacGillivray, 1908. Canad. Ent. 40: 295. ♀.
Aphanisiis nigritus MacGillivray, 1908. Canad. Ent. 40: 296. ♀.
Aphaniusd) lenis Rohwer, 1909. Canad. Ent. 41: 309. ♀, ♂.
Monophadnoides corytiis MacGillivray, 1923. Psyche 30: 79. ♂.
Monophadnoides consoiius MacGillivray, 1923. 111. Univ. Bui. 20(50): 25. ♀.
Monophadnoides constitutus MacGillivray, 1923. 111. Univ. Bui. 20(50): 25. ♀.
Monophadnoides curiosus MacGillivray, 1923. 111. Univ. Bui. 20(50): 25. ♀.
Monophadnoides kincaidi MacGillivray, 1923. 111. Univ. Bui. 20(50): 26. ♀.
Monophadnoides shawi MacGillivray, 1923. 111. Univ. Bui. 20(50): 26. ♀, ♂.
Paracharactus obversiis MacGillivray, 1923. 111. Univ. Bui. 20(50): 28. ♀.

Biology: Harris, 1850. New England Farmer 2: 33. —Saunders, 1873. Canad. Ent. 5: 101-103.
— Riley, 1877. 111. State Ent. Rpt. 6: 61. —Saunders, 1885. Canad. Ent. 17: 1-2. —Petit,
1900. Mich. Agr. Expt. Sta., Bui. 180: 137-138. -Lowe, 1898. N. Y. State Agr. Expt. Sta.,
Bui. 150, pp. 251-262.

Monophadnoides osgoodi Smith
Maine, N. H., Ont.

Monophadnoides osgoodi Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 146. ♀.

Monophadnoides pauper (Provancher)
Labrador to Md. w. to Alta., Colo.

Selandria paupera Provancher, 1882. Nat. Canad. 13: 293. ♀.
Monophadnoides cordatus MacGillivray, 1908. Canad. Ent. 40: 294. ♀.
Blennocampa abnortna MacGillivray, 1908. Canad. Ent. 40: 296. ♂.
Blennocampa antennata MacGillivray, 1908. Canad. Ent. 40: 296. ♀.
Blennocampa acuminata MacGilHvray, 1908. Canad. Ent. 40: 297. ♀.
Blennocampa adiista MacGillivray, 1908. Canad. Ent. 40: 297. ♀.
Blennocampa aiigulata MacGillivray, 1908. Canad. Ent. 40: 297. ♀.
Blennocampa aperta MacGilHvray, 1908. Canad. Ent. 40: 297. ♀.
Erythraspides tuckeri Rohwer, 1909. Canad. Ent. 41: 145. ♀.
Paracharactus nigrisomus Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 231. ♂.

Monophadnoides quebecensis Smith
Que.

Monophadnoides quebecensis Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 150. ♀.

Monophadnoides typicus (Rohwer)
B. C. to Calif.

Claremontia typica Rohwer, 1909. Canad. Ent. 41: 397. ♀.




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Tribe CERATULINI

Genus CERATULUS MacGillivray

Ceratulus MacGillivray, 1908. Canad. Ent. 40: 454.

Type-species: Ceratulus spectabilis MacGillivray. Orig. desig.

Only a single species is known for this genus.

Ceratulus spectabilis MacGillivray
Tex. Host: Cissus incisa.

Ceratulus spectabilis MacGillivray, 1908. Canad. Ent. 40: 454. ♀, ♂.

Tribe WALDHEIMIINI

Genus WALDHEIMIA Brullc

Waldhei^nia Brulle, 1846. In Lepeletier, Hist. Nat. Ins. Hym., p. 665.

Type-species: Tenthredo brasiliensis Lepeletier. Orig. desig.
Zarca Cameron, 1878. Ent. Soc. London, Trans, p. 142.

Type-species: Zarca apicatis Cameron. Monotypic.
Parazarca Ashmead, 1898. In Dyar, N. Y. Ent. Soc, Jour. 6: 128. N. syn.

Type-species: Parazarca fumipennis Ashmead. Orig. desig.
Calozarca Ashmead, 1898. In Dyar, N. Y. Ent. Soc, Jour. 6: 129. N. syn.

Type-species: Selatidria fascipe7i7iis Norton. Orig. desig.

A large genus found from southwestern United States to Argentina. Of the more than 70
described species, only one reaches north of Mexico.

Revision: Malaise, 1949. Arkiv for Zool. 42A: 33-58 (key to species).

Waldheimia bedeae Smith
S. Ariz.; Mexico.

Waldheimia bedeae Smith, 1969. U. S. Dept. Agr., Tech. Bui. 1397, p. 156. ♀.

Genus ERYTHRASPIDES Ashmead

Erythraspides Ashmead, 1898. In Dyar, N. Y. Ent. Soc, Jour. 6: 128.
Type-species: Tenthredo pygmaea Say. Orig. desig.

A small genus found also in Mexico and Central America.

Erythraspides carbonarius (Cresson)
Maine, Ont. to Ga. w. to Sask., Colo., Tex. Host: Oenothera.
Selayidria carbonaria Cresson, 1880. Amer. Ent. Soc, Trans. 8: 12. ♀.
Selandria parva Cresson, 1880. Amer. Ent. Soc, Trans. 8: 12. ♂.
Blennocampa abjecta MacGillivray, 1921. Brooklyn Ent. Soc, Bui. 16: 22. ♀.
Blennocampa absona MacGillivray, 1921. Brooklyn Ent. Soc, Bui. 16: 22. ♂.

Erythraspides vitis (Harris)
N. H. to Fla. w. to Iowa, Ark., La. Host: Vitis. Grape sawfly.

Tenthredo pygmaea Say, 1824. In Keating, Narr. Long's 2nd Exped. (App.) p. 318. ♀, ♂.

Preocc. by Klug, 1814.
Tenthredo (Selandria) vitis Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 586. Nom. nud.
Selandria vitis Harris, 1841. Rpt. Ins. Mass. Injurious to Veg., Ed. 1, p. 378.
Selandria caryae Norton, 1869. In Packard, Guide Study Ins., p. 224. Adults only, larvae
are Eriocavipa juglandis (Fitch).

Biology: Horsfall, 1929. Ent. News 40: 174-177.

Genus HALIDAMIA Benson

Halidamia Benson, 1939. Ent. Monthly Mag. 75: 111.

Type-species: Blennocampa affinis (Fallen). Orig. desig.

The single species in this genus was probably introduced into North America.

Halidamia affinis (Fallen)
Conn, to Md. w. to Ont, Wis.; Europe. Probably introduced. Host: Galium.
Host recorded in Europe.
Hylotoma affinis Fallen, 1807. Svenska Vetensk. Akad., Handl. 28: 207.
Hylotoma assimilis Fallen, 1807. Svenska Vetensk. Akad., Handl. 28: 204. ♀.




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Tenthredo {Allantus) kyalina Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 58. ♀.
Blennocampa formosella Costa, 1882. Rend. Accad. Sci. fis Napoli, p. 198.

Subfamily ALLANTINAE

Members of this subfamily are associated with various plants. Some hosts are strawberries,
raspberries, roses, violets, and dogwood. After feeding is completed, larvae of many species bore
into fruits, wood, or other suitable substances for pupating or overwintering. The occurrence of
larvae in these secondary hosts is sometimes more significant if in cultivated fruits or structural
wood, but it is purely incidental depending on their proximity to the true host plant. Adults of
this subfamily are medium to large sawflies and may be either rather stout or elongated.

Tribe ERIOCAMPINI

Genus PSEUDOSIOBLA Ashmead

Pseudosiobla Ashmead, 1898. Canad. Ent. 30: 308.

Type-species: Allaritus excavatus Norton. Orig. desig.

Pseudosiobla cephalanthi Rohwer
Mass., N. Y., Mo. Host: Cephalanthus occidentalis.

Pseudosiobla cephalanthi Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 404. ♂.

Pseudosiobla excavata (Norton)
Maine to Fla. w. to 111., Tex. Host: Cephalanthus.

Allantus excavatus Norton, 1862. Ent. Soc. Phila., Proc. 1: 143. ♀.

Siobla robiista Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 253. ♀.

Taxonus floridanus Provancher, 1888. Addit. Corr. Faune Ent. Canada Hym., p. 352. ♂.

Genus ERIOCAMPA Hartig

Eriocampa Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung
Naturgesch. Hym., p. 279.

Type-species: Tenthredo ovata Linnaeus. Desig. by Rohwer, 1911.
Eriocampa subg. Brachyocampa Zirngiebl, 1956. Munchen Ent. Gesell. Mitt. 46: 326.
Type-species: Eriocampa dorpatica Konow. Orig. desig.

Eriocampa juglandis (Fitch)
N. B., Que., Ont. to N. C. w. to Minn., Nebr. Host: Juglans.

Tenthredo (Allantus) obesa Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Selandriai juglandis Fitch, 1857. N. Y. State Agr. Soc, Trans. 16: 467. Larva.
Allantus obesus Norton, 1860. Boston Jour. Nat. Hist. 7: 260. ♂.
Sciapteryx rotundus Norton, 1867. Amer. Ent. Soc, Trans. 1: 242. ♀.
Eriocampa rotundiformis Rohwer, 1909. Canad. Ent. 41: 16. ♂.

Eriocampa ovata (Linnaeus)
Que., Mass., N. Y., Ont, B. C, Wash.; Europe. Probably introduced
independently into east and west N. Amer. Host: Alnus rubra.
Tenthredo ovata Linnaeus, 1761. Fauna Suecica, Ed. 2, p. 392.
Tenthredo gossypina Retzius, 1783. Genera et Species Insectorum, p. 72.
Tenthredo leucogona Schrank, 1802. Fauna Boica, v. 2, p. 251.

Biology: Bouchard, 1961. Ent. Soc. Quebec, Ann. 6: 69-80. —Borden and Dean, 1971. Ent. Soc.
Brit. Columbia, Proc. 68: 26-28 (in B. C).

Morphology: Hsin, 1935. Ztschr. Angew. Ent. 22: 253-294 (epidermal glands of larva).

Genus DIMORPHOPTERYX Ashmead

Dimorphopteryx Ashmead, 1898. Canad. Ent. 30: 308.

Type-species: Allantus pinguis Norton. Orig. desig.

Revision: Rohwer, 1915. U. S. Natl. Mus., Proc. 48: 445-448.

Taxonomy: Middleton, 1915. U. S. Natl. Mus., Proc. 48: 497-501 (larvae).

Dimorphopteryx abnormis Rohwer
Ont., Maine to N. C. w. to Sask., Mont., 111. Host: Pninus, Pynis,
Amelanchier, Crataegus.
Dimorphopteryx ahiormis Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 406. ♀, ♂.
Dimorphopteryx desidiosiis MacGillivray, 1923. 111. Univ. Bui. 20(50): 10. ♀.




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Dimorphopteryx ithacus MacGilHvray, 1923. 111. Univ. Bui. 20(50): 10. ♀.

Dimorphopteryx salinus MacGillivray, 1923. 111. Univ. Bui. 20(50): 11. ♀.

Dimorphopteryx autumnalis Rohwer
Va., Wis. Host: Querciis.

Dimorphopteryx autumnalis Rohwer, 1915. U. S. Natl. Mus., Proc. 48: 447. ♀.

Dimorphopteryx quercivora Rohwer, 1915. U. S. Natl. Mus., Proc. 48: 448. ♂.

Dimorphopteryx melanognathus Rohwer
Newfoundland to Va. w. to Ont., Mich. Host: Betula papyrifera.

Dimorphopteryx melanognathus Rohwer, 1910. U. S. Natl. Mus., Proc. 38: 205. ♀.

Diynorphopteryx enucleatus MacGillivray, 1923. 111. Univ. Bui. 20(50): 10. ♀.

Dimorphopteryx scopulosus MacGillivray, 1923. 111. Univ. Bui. 20(50): 12. ♀.

Biology: Dyar, 1895. Amer. Ent. Soc, Trans. 22: 311 {pinguis, not Norton). — Dyar, 1897. N.
Y. Ent. Soc, Jour. 8: 199. (pinguis, not Norton).

Dimorphopteryx pinguis (Norton)
N. S., P. E. I., N. B., Que. to Va. w. to Sask., Colo. Host: Alnus, Betula.
Allantzts pinguis Norton, 1860. Boston Jour. Nat. Hist. 7: 244. ♀.
Sciapteryx punctum Provancher, 1878. Nat. Canad. 10: 72. ♂.

Dimorphopteryx pinguis errans Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 406. ♀, ♂.
Dimorphopteryx coloradensis Rohwer, 1915. U. S. Natl. Mus., Proc. 49: 206. ♀.
Dimorphopteryx ormiis MacGillivray, 1923. 111. Univ. Bui. 20(50): 11. ♀.
Dimorphopteryx morsei MacGillivray, 1923. 111. Univ. Bui. 20(50): 11. ♀.

Biology: Wong, 1954. Canad. Ent. 86: 154.

Dimorphopteryx virginicus Rohwer
Pa., Ohio s. to Ga. Host: Casta7iea dentata.

Dimorphopteryx pinguis virginica Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 406. ♀, ♂.
Dimorphopteryx castaneae Rohwer, 1915. U. S. Natl. Mus., Proc. 48: 447. ♀, ♂.

Tribe EMPRIINI

Genus EMPRIA Lepeletier

Empria Lepeletier, 1828. Encycl. Meth., Diet. Ins., p. 571.

Type-species: Dolenis (Empria) palliynacula Lepeletier. Desig. by Brulle, 1846.
Poecilostoma Dahlbom, 1835. Conspectus Tenthred., Siricid., Oryssin. Scand., p. 13.

Type-species: Tenthredo guttatum Fallen. Desig. by Rohwer, 1911.
Prosecris Gistel, 1848. Naturgesch. Thierr. f. hoh. Schul., p. 10. Unnecessary n. name for

Poecilostoyna.
Poecilosoma Thomson, 1870. Opusc. Ent. 2: 265. Emend, of Poecilostoma. Preocc. by

Huebner, 1819, and four others.
Poecilostomidea Ashmead, 1898. Canad. Ent. 30: 256.

Type-species: Emphytiis maculatus Norton. Orig. desig.
Tetratneura Ashmead, 1898. Canad. Ent. 30: 256.

Type-species: Selandria ignotus Norton. Orig. desig.
Tetraneura Konow, 1905. In Wytsman, Gen. Ins., v. 29, pp. 102, 171. Emend. Preocc. by

Hartig, 1814.
Parataxonus MacGillivray, 1908. Canad. Ent. 40: 367.

Type-species: Taxonus multicolor Norton. Orig. desig.
Leucempria Enslin, 1913. Deut. Ent. Ztschr., Beih., p. 187.

Type-species: Tenthredo candidata Fallen. Orig. desig.
Empria subg. Trieynpria Enslin, 1914. Deut. Ent. Ztschr., Beih., p. 213.

Type-species: Empria tridens Konow. Orig. desig.

Species of this holarctic genus show considerable color variation; consequently the long list of
synonymies under some species. This genus also extends into Mexico.

Revision: Ross, 1936. Pan-Pacific Ent. 12: 172-178.

Empria alpina Benson
N. W. T., B. C; Europe. Host: Salix spp.

Empria alpina Benson, 1938. Soc. Brit. Ent., Trans. 5: 190, 195. ♀, ♂.

Empria candidata (Fallen)
Que., Maine, N. H., N. W. T., Man., Alta., Yukon, B. C. Alaska; Europe to
Siberia. Host: Betula.
Tenthredo candidata Fallen, 1807. Svenska Vetensk. Akad., Handl. 28: 105.
Tenthredo (Allantus) repanda Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 77. ♀.




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Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 390 (distribution).

Empria coryli (Dyar)
N. H., N. Y. w. to Wis., Iowa, Mo. Host: Corylus.

Harpiphonis maciilatus var. coryli Dyar, 1897. N. Y. Ent. Soc, Jour. 5: 194. 9, larva.
Empria mellipes Rohwer, 1910. Canad. Ent. 42: 175. ♀.
Empria caetrata MacGillivray, 1911. Canad. Ent. 43: 305. ♀.

Biology: Dyar, 1897. N. Y. Ent. Soc, Jour. 5: 194.
ignota (Norton). Maine to Pa. w. to Alaska, B. C, Wash., Oreg., Calif. Ecology: Widespread.
Adult captured from Salix.

Selandria ignotus Norton, 1867. Amer. Ent. Soc, Trans. 1: 257. ♀.

Monostegia kincaidi MacGillivray, 1893. Canad. Ent. 25: 239. ♀.

Empria calda MacGillivray, 1911. Canad. Ent. 43: 307. ♀.

Empria cata MacGillivray, 1911. Canad. Ent. 43: 307. ♂.

Empria castigata MacGillivray, 1911. Canad. Ent. 43: 309. ♀.

Empria casca MacGillivray, 1911. Canad. Ent. 43: 310. ♀.

Empria evecta MacGillivray, 1911. Canad. Ent. 43: 310. ♀.

Empria confirmata MacGillivray, 1911. Canad. Ent. 43: 341. ♀.

Empria coiicitata MacGillivray, 1911. Canad. Ent. 43: 342. ♂.

Empria culpata MacGillivray, 1911. Canad. Ent. 43: 343. ♀.

Empria cerina MacGillivray, 1921. Psyche 28: 34. ♀, ♂.

Empria cirrha MacGillivray, 1923. 111. Univ. Bui. 20(50): 16. ♀.

Empria cithara MacGillivray, 1923. 111. Univ. Bui. 20(50): 17. ♀.

Empria improba (Cresson)
Maine, Ont. w. to N. W. T., Alaska, B. C, Oreg., Colo., Calif. Ecology:
Widespread. Adults collected from Salix.

Emphytus improbus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 11. ♀.

Empria salicis Rohwer, 1910. Canad. Ent. 42: 174. ♀, ♂.

Empria contexta MacGillivray, 1911. Canad. Ent. 43: 345. ♀.

Empria maculata (Norton)
Newfoundland to N. C. w. to Alaska, B. C, Oreg., Calif. Ecology:
Widespread; recorded from nearly every state in its range. Sometimes a pest of
cultivated strawberries. Host: Fragaria, Potentilla, Rubus.

Emphytus maculatus Norton, 1861. Boston Soc. Nat. Hist., Proc 8: 157. 9, tJ.

Poecilostoma convexa MacGillivray, 1909. Canad. Ent. 41: 402. ♀.

Empria distincta Rohwer, 1910. Canad. Ent. 42: 173. ♀.

Empria submaculata Rohwer, 1910. Canad. Ent. 42: 174. ♀, ♂.

Empria melanostoma Rohwer, 1910. Canad. Ent. 42: 175. ♀, ♂.

Monosoma maura Rohwer, 1910. U. S. Natl. Mus., Proc. 38: 204. ♀.

Empria callosa MacGillivray, 1911. Canad. Ent. 43: 305. ♀.

Empria celsa MacGillivray, 1911. Canad. Ent. 43: 306. ♀.

Empria callida MacGillivray, 1911. Canad. Ent. 43: 306. ♀.

Empria caprina MacGillivray, 1911. Canad. Ent. 43: 307. ♀, ♂.

Eynpria casta MacGillivray, 1911. Canad. Ent. 43: 308. ♀, ♂.

Empria celebrata MacGillivray, 1911. Canad. Ent. 43: 308. ♀.

Empria captiosa MacGillivray, 1911. Canad. Ent. 43: 308. ♀.

Empria caeca MacGillivray, 1911. Canad. Ent. 43: 308. ♀.

Empria cariosa MacGillivray, 1911. Canad. Ent. 43: 309. ♀.

Empria candidula MacGillivray, 1911. Canad. Ent. 43: 310. ♀.

Empria canora MacGillivray, 1911. Canad. Ent. 43: 310. ♀.

Empria caiita MacGillivray, 1911. Canad. Ent. 43: 311. ♀.

Empria condensa MacGillivray, 1911. Canad. Ent. 43: 342. ♀.

Empria cmnidata MacGillivray, 1911. Canad. Ent. 43: 343. ♀.

Empria ciirata MacGillivray, 1911. Canad. Ent. 43: 345. ♀.

Empria cuveata MacGillivray, 1911. Canad. Ent. 43: 345. ♀.

Empria cupida MacGillivray, 1911. Canad. Ent. 43: 346. ♀.

Empria schwarzi Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 398. ♀.

Empria cadurca MacGillivray, 1923. Canad. Ent. 55: 158. ♀, ♂.

Biology: Riley, 1867. Prairie Farmer, May, p. 348. —Petit, 1899. Mich. Agr. Expt. Sta. Bui.
175: 365. —Webster, 1915. Iowa Agr. Expt. Sta. Bui. 162, 20 pp. —Webster, 1916. Iowa
Acad. Sci., Proc. 23: 291-297. — Neiswander, 1944. Ohio Agr. Expt. Sta. Bui. 651: 35.




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Empria mexicana (Cameron)
N. Mex., Ariz.; Mexico.

Poecilosoma mexicanum Cameron, 1883. Biol.-Cent. Amer., v. 1, p. 34.

Empria arizonensis Rohwer, 1910. Canad. Ent. 42: 174. ♀.

Empria multicolor (Norton)
N. B., Que. to Ga. w. to Alta., Wash., Oreg., Calif. Ecology: Widespread.
Host: Alnus, Betnla papyrifera, B. lutea.

Strong ylogaster multicolor Norton, 1862. Boston Soc. Nat. Hist., Proc. 9: 120. ♀, ♂.

Emphytus hullensis Provancher, 1886. Addit. Corr. Faune Ent. Canada Hym., p. 25. ♂.

Empria carbasea MacGillivray, 1911. Canad. Ent. 43: 341. ♀.

Aphilodyctium maculatum Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 408. ♀.

Aphilodyctium multicolor erythrogastrum Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 408. ♀, ♂.

Biology: Wong, 1954. Canad. Ent. 86: 154.

Empria nordica Ross
N. W. T., Man.

Empria nordica Ross, 1936. Pan-Pacific Ent. 12: 175. ♀.

Empria obscurata (Cresson)
Maine to Pa. w. to N. W. T., Alaska, B. C, Wash., Oreg., Calif. Ecology:
Widespread; sometimes a pest of cultivated strawberries. Host: Fragaria. Either a
variable species or a species complex.

Selandria obscurata Cresson, 1880. Amer. Ent. Soc, Trans. S: 15. ♀, ♂.

Poecilosoma punctulata Weldon, 1907. Canad. Ent. 39: 304. ♀, ♂.

Empria affinis Rohwer, 1910. Canad. Ent. 42: 173. ♀.

Empria caudelli Rohwer, 1910. Canad. Ent. 42: 173. ♀.

Empria cava MacGillivray, 1911. Canad. Ent. 43: 306. ♀.

Empria cauduca MacGillivray, 1911. Canad. Ent. 43: 309. ♀.

Empria capillata MacGillivray, 1911. Canad. Ent. 43: 341. ♀.

Eynpria condita MacGillivray, 1911. Canad. Ent. 43: 342. ♀.

Empria contorta MacGillivray, 1911. Canad. Ent. 43: 343. ♀.

Empria conferta MacGillivray, 1911. Canad. Ent. 43: 344. ♀.

Empria cwicreta MacGillivray, 1911. Canad. Ent. 43: 344. ♀.

Empria conciliata MacGillivray, 1911. Canad. Ent. 43: 344. ♀.

Empria concisa MacGillivray, 1911. Canad. Ent. 43: 346. ♀.

Empria costata MacGillivray, 1914. Canad. Ent. 46: 103. ♀.

Empria fragariae Rohwer, 1914. Jour. Econ. Ent. 7: 479. ♀.

Empria cista MacGillivray, 1923. 111. Univ. Bui. 20(50): 16. ♀.

Empria cistula MacGillivray, 1923. 111. Univ. Bui. 20(50): 16. ♀.

Biology: Malley, 1889. U. S. Dept. Agr., Insect Life 2: 137-140 (ignota, not Norton). — Malley,
1890. U. S. Dept. Agr., Insect Life 3: 9-12 (ignota, not Norton). — Osborn and Gossard,
1892. Iowa Agr. Expt. Sta. Bui. 18: 512-514 {ignota, not Norton). —Webster, 1915. Iowa
Agr. Expt. Sta., Bui. 162, 20 pp. (fragariae). —Webster, 1916. Iowa Acad. Sci., Proc. 23:
291-297 (fragariae).

Genus PHRONTOSOMA MacGillivray

Phrontosoma MacGillivray, 1908. Canad. Ent. 40: 366.

Type-species: Phrontosoma atnim MacGillivray. Orig. desig.

A small, exclusively North American genus. One unidentified species extends the distribution
of this genus to western United States.

Phrontosoma belfragei (Cresson)
Mass. to Pa. w. to Ont., Wis., Kans., Tex. Ecology: Adults collected from
Comus.
Selandria belfragei Cresson, 1880. Amer. Ent. Soc, Trans. 8: 15. ♀.
Caliroa nortonia MacGillivray, 1894. Canad. Ent. 24: 324. ♂.
Phrontosoma atrum MacGillivray, 1908. Canad. Ent. 40: 367. ♂.
Phrontosoma daeckei MacGillivray, 1908. Canad. Ent. 40: 367. ♀.
Phrontosoma collaris MacGillivray, 1908. Canad. Ent. 40: 367. ♀.

Genus MONOSOMA MacGillivray

Monsoma(l) MacGillivray, 1908. Canad. Ent. 40: 368.




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Type-species: Taxovus inferentia Norton. Orig. desig.
Monosoma Viereck, 1910. N. J. State Mus., Ann. Rpt. 1909, p. 583. Emend.

A small genus with two species, one Nearctic the other Palaearctic.

Monosoma inferentia (Norton)
Newfoundland to N. C. w. to Minn., 111. Host: Alniis.
Poecilosfo7na inferentia Norton, 1868. Amer. Ent. Soc, Trans. 2: 224. ♂.
Strongylogaster albosectus Provancher, 1878. Nat. Canad. 10: 168. ♀.
Monophadnus lineatus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 177. 9. A questionable

synonym.
Monosoma inferentia var. andronosa Ross, 1932. Canad. Ent. 64: 249. ♀.

Genus MONOSTEGIA Costa

Monostegia Costa, 1859. Fauna del Regno di Napoli, Imen., pt. 3, p. 60.

Type-species: Tenthredo (Allantus) luteola Klug. Desig. by Ross, 1951.
Poecilosoma subg. Nematoceros Konow, 1896. Wien. Ent. Ztg. 15: 52, 54.
Type-species: Tenthredo (Allaiitus) luteola Klug. Monotypic.

Monostegia abdominalis (Fabricius)
Que., Ont. to N. J., Pa. w. to Ohio; Europe to Siberia. Probably
introduced. Host: Lysiniacliia numyyiularia, L. vulgaris, L. terrestris, Anagallis.
Tenthredo abdominalis Fabricius, 1798. Sup. Ent. System., p. 216. ♀.
Tenthredo (Allantus) luteola Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 48.
Monostegia martini MacGillivray, 1908. Canad. Ent. 40: 366. ♀.
Monostegia nearctica Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 209. ♀, ♂.

Biology: Price, 1970. Canad. Ent. 102: 491-495.

Genus AMETASTEGIA Costa

Ametastegia Costa, 1882. Accad. delle Sci. Fis. e Mat. Napoli, Rend. 21(10): 198.

Type-species: Ametastegia fulvipes Costa. Monotypic.
Aomodyctium Ashmead, 1898. Canad. Ent. 30: 309. No included species.

Type-species: Strongylogaster abyiortnis Provancher. Desig. by Rohwer, 1911.
Protemphytus Rohwer, 1909. Canad. Ent. 41: 92.

Type-species: Emphytus coloradensis Weldon. Monotypic.
Emphytina Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 399.

Type-species: Emphytina pnlchella Rohwer. Orig. desig.
Simple7nphytus MacGillivray, 1914. Canad. Ent. 46: 363.

Type-species: Simple7nphytus pacificus MacGillivray. Monotypic.
Unitaxonus MacGillivray, 1921. Psyche 28: 32.

Type-species: Uyiitaxonus repentinus MacGillivray. Orig. desig.

Species of this holarctic genus feed on herbaceous plants. On maturing, larvae bore into soft
substances to form a pupal cell. This secondary host may be apples, other fruits, soft wood, or
pith of other plants. Such records as hosts for species of this genus do not constitute the true
host on which the larvae feed.

Revision: Ross, 1937. Arb. uber Morph. u. Taxonom. Ent. 4: 84-91.

Ametastegia aperta (Norton)
Newfoundland to N. C. w. to Mich., Colo.

Tenthredo (Emphytus) aperta Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Emphytus apertus Norton, 1861. Boston Nat. Hist. Soc, Proc. 8: 155. ♀.
Emphytus leucostoina Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 110. ♀. Preocc. by Costa,

1890.
Emphytina plesia Rohwer, 1911. U. S. Natl. Mus., Proc 41: 402. N. name.

Ametastegia articulata (Klug)
Que., Maine to Fla. w. to Minn., Iowa, Ark., Tex. Host: Rumex, Polygonum.
Tenthredo (Emphytus) articulatus Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 284.

6.
Dolenis inomatus Say, 1824. In Keating, Narr. Long's 2nd Exped., v. 2, (App.) p. 319. ♀.
Emphytus halitus MacGillivray, 1923. 111. Univ. Bui. 20(50): 14. ♀.

Taxonomy: Smith, 1973. Ent. Soc. Wash., Proc. 75: 29.




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Ametastegia coloradensis (Weldon)
Labrador, Colo., Alta., B. C, Oreg., Calif.
Emphytus coloradensis Weldon, 1907. Canad. Ent. 39: 304. ♂.
Emphytus hiulcus MacGillivray, 1923. 111. Univ. Bui. 20(50): 15. ♀.

Ametastegia equiseti (Fallen)
Que., Maine to N. C. w. to Alaska, Oreg.; Europe to Siberia. Ecology:

Widespread. Host: Rumex, Polygonum.
Tenthredo equiseti Fallen, 1808. Svenska Vetensk. Akad., Handl. 29: 60.
Tenthredo (Allantus) bicolor Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 204. ♀, ♂.
Tenthredo hizonata Zetterstedt, 1838. Ins. Lapp., v. 1, p. 344. ♀.
Taxonus iyinoniinatiis MacGillivray, 1901. N. Y. State Mus. Bui. 47: 585. ♀.
Stro7igylogastroidea depressata MacGillivray, 1921. Psyche 28: 31. ♀.
Unitaxonus repentinus MacGillivray, 1921. Psyche 28: 31. ♀, ♂.
Unitaxonus nwiicis MacGillivray, 1921. Psyche 28: 31. ♀.
Hemitaxmms dedititius MacGillivray, 1923. Psyche 30: 77. ♂.
Stro7igylogastroidea rufocinctella MacGillivray, 1923. 111. Univ. Bui. 20(50): 32. ♀.

Ametastegia glabrata (Fallen)
N. S., Que. to Md. w. to B. C, Wash., Oreg.; Europe to Siberia. Ecology:

Widespread. If adjacent to host, larva commonly bores into apples or other fruits to

pupate. Host: Runiex. Dock sawfly.
Tenthredo glabrata Fallen, 1808. Svenska Vetensk. Akad., Handl. 29: 108.
Tenthredo (Allantus) agilis Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 208. ♀, ♂.
Tenthredo (Allantus) rufipes Lepeletier, 1823. Monog. Tenthred., p. 239. ♀.
Taxonus nigriso7na Norton, 1862. Boston Soc. Nat. Hist., Proc. 9: 119. ♀.
Strongylogaster abnonnis Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 10. ♀, ♂.
Strong ylogastroidea potulenta MacGillivray, 1923. 111. Univ. Bui. 20(50): 31. ♀.

Biology: Chittenden and Titus, 1905. U. S. Dept. Agr. Bui. 54: 40-43. —Webster, 1908. Jour.
Econ. Ent. 1: 310-311 (in apples). — Dustan and Gilliatt, 1916. Ent. Soc. Nova Scotia, Proc,
pp. 45-48. —Newcomer, 1916. U. S. Dept. Agr., Bui. 265, 40 pp. — Blackman and Stage,
1924. N. Y. State Col. Forestry, Syracuse Univ., Tech. Pub.'l?, v. 24, p. 153 (in dead and
decaying hickory). — Zuk, 1961. Ent. Soc. Brit. Columbia, Proc. 58: 21 (boring holes in
cedar siding).

Ametastegia pallipes (Spinola)
Newfoundland to Md. w. to B. C, Oreg., Calif.; Europe to Siberia; Iceland.

Ecology: Widespread. Host: Viola. Violet sawfly.
Tenthredo pallipes Spinola, 1808. Insectorum Liguriae, v. 2, p. 19.
Tenthredo (Emphytus) grossulariae Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8:

283. ♀.
Dolems leucopodus Lepeletier, 1823. Monog. Tenthred., p. 119. ♀.
Tenthredo pallipes Provancher, 1878. Nat. Canad. 10: 66. 9, 6. Homonym and Synonym.
Emphytus canadense Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 204. N. name for pallipes

Provancher.
Emphytus pallidipes Dalla Torre, 1894. Cat. Hym., v. 1, p. 119. Emend.
E^npria cavata MacGillivray, 1911. Canad. Ent. 43: 305. ♀.
Empria cetaria MacGillivray, 1921. Psyche 28: 33. ♀.
Emphytus halesus MacGillivray, 1923. 111. Univ. Bui. 20(50): 13. ♀.
Emphytus heroicus MacGillivray, 1923. 111. Univ. Bui. 20(50): 14. ♀.
Emphytus hiatics MacGillivray, 1923. 111. Univ. Bui. 20(50): 14. ♀.
E7nphytus hospitus MacGillivray, 1923. 111. Univ. Bui. 20(50): 15. ♀.
E7nphytus hyaci7ithus MacGillivray, 1923. 111. Univ. Bui. 20(50): 16. ♀.

Biology: Dyar, 1894. Canad. Ent. 26: 185 (canade7isis).

Ametastegia recens (Say)
Newfoundland to Ala. w. to Alaska, B. C, Oreg. Ecology: Widespread. Host:

Polygo7ium. Several species are currently going under this name.
Emphytus rece7is Say, 1836. Boston Jour. Nat. Hist. 1: 221. ♂.
Emphytus stra7ni7ieipes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 52. 9. A questionable

synonym.
E7nphytus a7igustus Kincaid, 1900. Wash. Acad. Sci., Proc 2: 360. 6 . A questionable

synonym.
E7nphyti7ia pulchella Rohwer, 1911. U. S. Natl. Mus., Proc 41: 400. ♀.
E7nphyti7ia pallidiscapa Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 401. ♀, ♂.




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Eviphytina virginica Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 401. ♀.
Emphytiis ynasi MacGillivray, 1921. Psyche 28: 31. ♀.

Ametastegia tener (Fallen)
Que., Ont. to Va. w. to B. C, Wash., Oreg.; Europe to Siberia. Ecology:
Widespread. Host: Rumex.
Tenthredo tener Fallen, 1808. Svenska Vetensk. Akad., Handl. 29: 109.
Tenthredo (Empkytus) patellatus Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 283. ♀, ♂.
Dolerus luctuosus Lepeletier, 1823. Monog. Tenthred., p. 119. ♀, ♂.
Dolenis nigritiis Lepeletier, 1823. Monog. Tenthred., p. 120. ♂.
Simplemphytus pacificus MacGillivray, 1914. Canad. Ent. 46: 363. ♀, ♂.
Emphyiina vanduzeei Rohwer, 1915. U. S. Natl. Mus., Proc. 49: 205. ♀.
Empria columna MacGillivray, 1923. Brooklyn Ent. Soc, Bui. 18: 54. ♀.
Emphytus haliartus MacGillivray, 1923. 111. Univ. Bui. 20(50): 14. ♀.
Emphytus haustus MacGillivray, 1923. 111. Univ. Bui. 20(50): 14. ♂.

Genus APHILODYCTIUM Ashmead

Aphilodyctium Ashmead, 1898. Canad. Ent. 30: 310.

Type-species: Strotigylogaster rubripes Cresson. Orig. desig.
Polytaxonus MacGillivray, 1908. Canad. Ent. 40: 368.

Type-species: Taxonus robustus Provancher. Orig. desig.

The single species in this genus is found only in North America.

Taxonomy: Ross, 1937. Arb. uber Morph. u. Taxonom. Ent. 4: 90.

Aphilodyctium fldum (Cresson)
Que., Maine to Md. w. to B. C, Wash., Oreg., Calif. Ecology: Widespread.
Host: Rosa.
Strongylogaster fidus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 19. ♀.
Strong ylogaster rubripes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 20. ♀.
Taxonus robustus Provancher, 1882. Nat. Canad. 13: 294. ♀.
Taxonus parens Provancher, 1889. Addit. Corr. Faune Ent. Canada Hym., p. 9.
Taxo7ius lenis Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 110. ♀, ♂.
Aphilodyctium rubripes nigritarsis Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 408. ♀, ♂.
Taxonus inclinatus MacGillivray, 1923. Psyche 30: 78. ♂.

Tribe ALLANTINI

Genus ALLANTUS Panzer

Allantus Panzer, 1801. Faunae Ins. German., heft 82, pi. 12.

Type-species: Tenthredo (Allantus) togatus Panzer. Monotypic
Emphytus Klug, 1813. Mag. Gesell. Naturf. Freunde Berlin 7: 273.

Type-species: Tenthredo cincta Linnaeus. Desig. by Curtis, 1833.

This is a much larger genus in the Palaearctic region. Three of the seven North American
forms were introduced.

Revision: Ross, 1937. Arb. uber Morph. u. Taxonom. Ent. 4: 91.
Taxonomy: Wong, 1966. Canad. Ent. 98: 852-854.

Allantus albolabris (Rohwer)
Sask., Alaska, B. C, Wash., Oreg.

Emphytus inellipes var. albolabris Rohwer, 1917. U. S. Natl. Mus., Proc. 53: 152. ♀.
Emphytus gemitus MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 163. ♀.

Allantus basalis (Klug)
Newfoundland, N. Y., Tenn.; Europe to Siberia, China. Introduced. Host: Rosa.
Tenthredo {Emphytus) basalis Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 282. ♂.

Allantus cinctus (Linnaeus)
Newfoundland to Va. w. to B. C, Wash.; Europe to Siberia. Ecology:
Widespread; most commonly a pest of cultivated roses. Introduced (?). Host: Rosa,
Fragaria. Curled rose saw fly.
Tenthredo cinctus Linnaeus, 1758. Syst. Nat., Ed. 10, v. 1, p. 557.
Tenthredo togatus Zetterstedt, 1838. Ins. Lapp., v. 1, p. 342. ♀, ♂.
Dolerus cingulatus Blanchard, 1840. Hist. Nat. Ins., v. 3, p. 240.
Emphytus neglectus Zaddach, 1859. Programm Friedr. Col. Konigsberg, p. 27. ♂.




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Emphytus cinctipes Norton, 1867. Amer. Ent. Soc, Trans. 1: 229. ♀, ♂.

Biology: Jack, 1889. Psyche 5: 279. — Britton, 1916. Conn. (State) Agr. Expt. Sta., Ann. Rpt.
1915, pt. 2, pp. 185-186. — Middleton, 1922. U. S. Dept. Agr., Farmers' Bui. 1252, 4 pp.
— McNay, 1958. Canad. Ins. Pest Rev. 36: 141 (on strawberry).

Morphology: Will, 1933. Pa. Acad. Sci., Proc. 7: 30-32 (wing venation variations).

Allantus mellipes (Norton)
N. B., Que. to Fla. w. to Alta., Colo. Host: Fragaria.

Tenthredo (Emphytus) mellipes Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Emphytus mellipes Norton, 1861. Boston Soc. Nat. Hist, Proc. 8: 155. ♀.
Emphytus gillettei MacGillivray, 1902. In Johnson, Colo. Expt. Sta., Ann. Rpt. 15: 113. ♀.

Biology: Johnson, 1902. Colo. Expt. Sta., Ann. Rpt. 15: 113-114.

Allantus nigritibialis Rohwer
N. J., Va; Siberia, China. Possibly introduced. Host: RosaiO- May be
only a color form of cinctus (L.).
Allantus cinctus nigritibialis Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 407. 9. Described
from specimens intercepted in New Jersey from Hong Kong.

Allantus umbonatus Wong
Ont., Man., Sask., Alta. Host: Betula.

Allantus umbonatus Wong, 1966. Canad. Ent. 98: 852. ♀.

Allantus viennensis (Schrank)
N. Y. (Tompkins Co.); Europe. Introduced. Host: Rosa, Rubus.
Tenthredo Vieyuiensis Schrank, 1781. Enum. Ins. Austr., p. 331.

Taxonomy: Smith. 1975. Coop. Econ. Ins. Rpt. 25 (10): 163-165 (first U. S. record).

Biology: Kartasheva, 1964. Zasch. Rast. ot Vred. i Boleznei 9 (10): 41-42. — Scheibelreiter,
1973. Ztschr. Angewandte Ent. 72: 243-244.

Genus MACREMPHYTUS MacGillivray

Macremphytus MacGillivray, 1908. Canad. Ent. 40: 368.

Type-species: Harpiphorus varianus Norton. Orig. desig.

Larvae of this North American genus feed on Comus species of the osier dogwood group.
After feeding, larvae sometimes bore into wood to form pupal cells.

Macremphytus lovetti MacGillivray
Alta., Mont., B. C, Idaho, Oreg. Ecology: Adults collected from Cornus.

Macremphytus lovetti MacGillivray, 1923. Psyche 30: 77. ♀.

Macremphytus semicornis (Say)
Newfoundland to N. Y. w. to B. C. (Terrace), Man., Minn., 111. Host: Comus.

Emphytus semicornis Say, 1836. Boston Jour. Nat. Hist. 1: 220. ♀.

Macremphytus bicornis MacGillivray, 1923. 111. Univ. Bui. 20(50): 21. ♀.
tarsatus (Say). N. B., Que. to Fla. w. to Minn., Ill, Kans., Tex. Host: Comus sp., C. altemifolia.

Tenthredo (Emphytus) tarsata Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 583. Nom. nud.

Emphytus tarsatus Say, 1836. Boston Jour. Nat. Hist. 1: 220. ♀.

Emphytus bollii Norton, 1872. Amer. Ent. Soc, Trans. 4: 80. ♀.

Harpiphorus intemiedius Dyar, 1900. N. Y. Ent. Soc, Jour. 8: 30. 9, larva.

Macremphytus testaceus (Norton)
Maine, Ont. to Ga. w. to Man., Minn., Colo. Host: Comus sp., C. stolonifera
C. candidissima.

Emphytus (Harpiphorus) testaceus Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 156. ♀.

Emphyttis (Harpiphorus) varianus Norton, 1861. Boston Soc. Nat. Hist., Proc. 8: 156. ♀.
6.

Emphytus (Harpiphoms) versicolor Norton, 1867. Amer. Ent. Soc, Trans. 1: 230. ♀.

Biology: Jack, 1889. Garden and Forest 2: 520. —Riley and Howard, 1890. U. S. Dept. Agr.,
Insect Life 2: 239-243. —Will, 1944. Pa. Acad. Sci., Proc 18: 48. —Will, 1959. Pa. Acad. Sci.,
Proc 33: 214.

Morphology: Kreiger, Gilvert, and Wilkinson, 1970. Jour. Econ. Ent. 63: 1322-1323
(microsomal mixed-function oxidase activity).




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Genus TAXONUS Hartig

Taxonus Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung Naturgesch.
Hym., p. 297.

Type-species: Tenthredo (Allantiis) nitida Klug. Desig. by Rohwer, 1911.
Ermilia Costa, 1859. Fauna del Regno di Napoli, Imen., pt. 3, p. 106.

Type-species: Ermilia pulchella Costa. Monotypic.
Strongylogastroidea Ashmead, 1898. Canad. Ent. 30: 308.

Type-species: Alla7itus apicalis Say. Orig. desig.
Parasiobla Ashmead, 1898. Canad. Ent. 30: 308.

Type-species: Allayitus rufociyictus Norton. Orig. desig.
Hypotaxonus Ashmead, 1898. Canad. Ent. 30: 311.

Type-species: Allanhis pallipes Say. Orig. desig.

This genus is better represented in east Asia and eastern North America than in other parts
of the world. Judging from the few records, plants of the Rosaceae serve as hosts.

Taxonus borealis MacGillivray
Newfoundland to N. C, Tenn. w. to Mich., Wis.

Taxo7ius borealis MacGillivray, 1895. Canad. Ent. 27: 78. ♀.

Taxonus epicera (Say)
N. H., Ont. to N. C. w. to Wis., 111. Kans., Tex.

Allantus epicera Say, 1836. Boston Jour. Nat. Hist. 1: 216. ♂.

Taxonus pallicoxus (Provancher)
N. S., N. B., Que. to Va. w. to Alta., 111. Host: Fragaria.

Tenthredo (Taxonus) unicinctus Norton, 1862. Boston Soc. Nat. Hist., Proc. 9: 119. ♀, ♂.
Preocc. by Brulle, 1832.

Strongylogaster pallicoxus Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 11. ♀, ♂.

Strongylogaster pallidicoxis Dalla Torre, 1894. Cat. Hym., v. 1, p. 137. Emend.

Strongylogastroidea nifocinctana MacGillivray, 1923. 111. Univ. Bui. 20(50): 31. ♀.

Taxonus pallidicornis (Norton)
Que., Maine, Ont. to Fla. w. to Wis., Ill, La. Host: Rubiis.

Strongylogaster pallidicomis Norton, 1868. Amer. Ent. Soc, Trans. 2: 216. ♀.

Taxonus pallipes (Say)
N. H. to Ga., Ala. w. to Mich., 111.

Allantus pallipes Say, 1823. West. Quart. Rptr. 2: 72.

Strongylogaster pallidipes Dalla Torre, 1894. Cat. Hym., v. 1, p. 137. Emend.

Strongylogastroidea rufula MacGillivray, 1923. 111. Univ. Bui. 20(50): 32. ♀.

Taxonus proximus (Provancher)
Que., Maine, Ont. s. to Ga.

Strongylogaster proximus Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 12. ♀.

Strongylogastroidea rufinerva MacGillivray, 1923. 111. Univ. Bui. 20(50): 31. ♀.

Taxonus rufocinctus (Norton)
Que., Maine, Ont. to N. C. w. to Minn., 111., Ark.

Allantus rufocinctus Norton, 1860. Boston Jour. Nat. Hist. 7: 248. ♀, ♂.

Taxonus (Parasiobla) rufocinctus virginicus Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 405. ♀.

Strongylogastroidea unicinctella MacGillivray, 1923. 111. Univ. Bui. 20(50): 33. ♀.

Taxonus spiculatus (MacGillivray)
Maine to N. C, Tenn. w. to Ohio, W. Va.

Strongylogastroidea spiculata MacGillivray, 1908. Canad. Ent. 40: 369. ♀.

Taxonus terminalis (Say)
N. S., N. B., Que., Ont. to Fla. w. to Minn., 111., Tex.

Tenthredo tertninalis Say, 1824. In Keating, Narr. Long's 2nd Exped., v. 2 (App.), p. 318. ♀, ♂.

Tenthredo (Allantus) melisoma Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 583. Nom. nud.

Allantus apicalis Say, 1836. Boston Jour. Nat. Hist. 1: 216.

Allantus abdojninalis Norton, 1860. Boston Jour. Nat. Hist. 7: 238. ♀, ♂.

Allayitus rnellosus Norton, 1860. Boston Jour. Nat. Hist. 7: 287. ♀, ♂.

Strongylogaster rufoculus MacGillivray, 1894. Canad. Ent. 26: 327. ♀.

Strongylogastroidea confusa MacGillivray, 1908. Canad. Ent. 40: 369. ♀.

Strongylogastroidea shermani MacGillivray, 1923. 111. Univ. Bui. 20(50): 32. ♀.




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Subfamily TENTHREDININAE

This subfamily contains most of the larger and more colorful members of the Tenthredinidae.
Though commonly collected, hosts and life histories are known for very few species.

Taxonomy: Benson, 1959. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 28: 121-127
(tribes).

Tribe PERINEURINI

Genus LEUCOPELMONUS MacGillivray

Leucopelmonus MacGillivray, 1916. Conn. State Geol. and Nat. Hist. Survey Bui. 22: 83.
Type-species: Leucopelmonus annulatus MacGillivray. Monotypic.

Leucopelmonus annulicornis (Harrington)
Newfoundland, Que., Ont. s. to N. C.

Tenthredo confusus Norton, 1869. Amer. Ent. Soc, Trans. 2: 241. ♂. Preocc. by Lepeletier,

1823.
Tenthredopsisf annulicornis Harrington, 1893. Canad. Ent. 25: 61. ♀, ♂.
Perineura turbata Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 408. ♀, ♂.
Leucopelmonus annulatus MacGillivray, 1916. Conn. State Geol. and Nat. Hist. Survey,
Bui. 22: 83. ♀.

Tribe SCIAPTERYGINI

Genus ZASCHIZONYX Ashmead

Zaschizonyx Ashmead, 1898. Canad. Ent. 30: 257.

Type-species: Hoplocampa montana Cresson. Orig. desig.
Opisthoneura Ashmead, 1898. Canad. Ent. 30: 287.

Type-species: Opisthoneura Crevecoeuri Ashmead. Orig. desig.

Revision: Ross, 1929. Canad. Ent. 61: 272-273.

Taxonomy: Benson, 1959. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 28: 127.

Zaschizonyx montana (Cresson)
Ont., Minn., 111., Iowa, Mo. w. to B. C, Wash., Oreg., Calif. Host:
Syynphoricarpos.

Selandria montana Cresson, 1865. Ent. Soc. Phila., Proc. 4: 244. ♀.

Selandria (Hoplocampa) gentilis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 14. ♂.

Opisthoneura Crevecoeuri Ashmead, 1898. Canad. Ent. 30: 287.

Zaschisonyxd) montana var. drabula Ross, 1929. Canad. Ent. 61: 272. ♀.

ZaschisonyxV-) montana subsp. occidentalis Ross, 1929. Canad. Ent. 61: 272. ♀.

Zaschizonyx pluricincta (Norton)
Ariz., Calif.

Macrophya pluricinctus Norton, 1862. Boston Soc Nat. Hist., Proc. 9: 118. ♀, ♂.

Macrophya albipes Provancher, 1895. Nat. Canad. 22: 95. 9, 6. Preocc by Geoffroy, 1785.

Labidia doanei Rohwer, 1909. Canad. Ent. 41: 91. ♀.

Macrophya provancheri Rohwer, 1909. Canad. Ent. 41: 328. N. name for albipes
Provancher.

Macrophya nigricomis Rohwer, 1909. Canad. Ent. 41: 329. ♀, ♂.

Macrophya occidentalis Rohwer, 1909. Canad. Ent. 41: 330. ♂.

Macrophya bakeri Rohwer, 1909. Canad. Ent. 41: 331. ♂.

Macrophya truncata Rohwer, 1909. Canad. Ent. 41: 331. ♀, ♂.

Macrophya pluricinctella Rohwer, 1909. Canad. Ent. 41: 332. ♀.

Macrophya inultici^icta Rohwer, 1909. Canad. Ent. 41: 333. ♀.

Macrophya melanostoma Rohwer, 1909. Canad. Ent. 41: 333. ♀.

Macrophya napeiisis Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 410. ♀.

Morphology: Matsuda, 1957. Kans. Ent. Soc, Jour. 30: 99-108 (head).

Tribe TENTHREDOPSINI
Genus AGLAOSTIGMA Kirby
Aglaostigma Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 325.




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Type-species: Aglaostigma ehurtieiguttahim Kirby. Monotypic.
Laurentia Costa, 1890. Accad. delle Sci. Fis. e Mat. Napoli, Rend. (ser. 2) 4: 173. Preocc. by
Ragonot, 1888.

Type-species: Laurentia craverii Costa. Monotypic.
Bivena MacGillivray, 1894. Canad. Ent. 24: 328.

Type-species: Bivena maria MacGillivray. Orig. desig.
Homoeoneura Ashmead, 1898. Canad. Ent. 30: 313.

Type-species: Pachyprotasis delta Provancher. Orig. desig.
Neopus Viereck, 1910. In Smith, N. J. State Mus., Ann. Rpt. 1909, p. 585.

Type-species: Tenthredo quattuordecimpunctatus Norton. Orig. desig.
Macrophyopsis Enslin, 1913. Deut. Ent. Ztschr., Beih., p. 128.

Type-species: Macropkya nebulosa Andre. Orig. desig.
Astochus MacGillivray, 1914. Canad. Ent. 46: 107.

Type-species: Astochus fletcheri MacGillivray. Orig. desig.
Kincaidia MacGillivray, 1914. Canad. Ent. 46: 107.

Type-species: Tenthredopsis ruficoma MacGillivray. Orig. desig.
Parallomma Malaise, 1933. Ent. Tidskr. 54: 53.

Type-species: Rhogogaster lichtwardti Konow. Orig. desig.
Neurosiobla Conde, 1935. Notulae Ent. 14: 79.

Type-species: Neurosiobla malaisei Conde. Orig. desig.
Laurentina Malaise, 1937. Rev. d'Ent. Franc. 4: 44.

Type-species: Laurentina ruficomis Malaise. Orig. desig.

Revision: Ross, 1943. Ent. Soc. Wash., Proc. 45: 79-84.

Aglaostigma dilutum (Cresson)
Calif.

Tenthredo diiuta Cresson, 1880. Amer. Ent. Soc, Trans. 8: 24. ♀.

Aglaostigma quattuordecimpunctatum (Norton)
Que., Maine, Ont. s. to N. C. Host: Podophyllum
peltatum.

Tejithredo H-punctatus Norton, 1862. Ent. Soc. Phila., Proc. 1: 143. ♀, ♂.

Aglaostigma rubens (Cresson)
Idaho, Nev. B. C, Wash., Oreg., Calif.

Tenthredo rubens Cresson, 1880. Amer. Ent. Soc, Trans. 8: 24. ♂.

Tenthredo Edwardsii Cresson, 1880. Amer. Ent. Soc, Trans. 8: 24. ♀.

Tenthredo atraveyius MacGiUivray, 1895. Canad. Ent. 27: 283. ♂.

Astochus fletcheri MacGillivray, 1914. Canad. Ent. 46: 108. ♀.

Astochus aldrichi MacGillivray, 1914. Canad. Ent. 46: 137. ♀.

Tenthredo racilia MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 112. ♂.

Tenthredo refractaria MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 113. ♀.

ruflcornum (MacGillivray). Wash., Oreg.

Tenthredopsis ruficoma MacGillivray, 1893. Canad. Ent. 25: 242. ♀.

Aglaostigma semiluteum (Norton)
Que., Maine to S. C. w. to Mont., Iowa, Kans. Host: Impatiens biflora.

Tenthredo semihiteus Norton, 1862. Boston Soc Nat. Hist., Proc 9: 121. ♀, ♂.

Pachyprotasis delta Provancher, 1878. Nat. Canad. 10: 108. ♀.

Bivena maria MacGillivray, 1894. Canad. Ent. 26: 328. ♀.

Aglaostigma jocosa (Provancher)
Que., N. H., Mass., Man.

Tenthredo jocosa Provancher, 1882. Nat. Canad. 13: 298. ♀.

Harpiphorus varipictus Harrington, 1889. Canad. Ent. 21: 96. ♀.

Aglaostigma dentatum Ross, 1943. Ent. Soc. Wash., Proc. 45: 82. ♀, ♂.

Aglaostigma veedee Ross
Calif.

Aglaostigma veedee Ross, 1943. Ent. Soc Wash., Proc. 45: 81. ♀, ♂.

Tribe TENTHREDININI
Genus LAGIUM Konow

Lagium Konow, 1904. Ztschr. System. Hym. Dipt. 4: 246.

Type-species: Tenthredo atroviolaceum Norton. Orig. desig.

Revision: Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 213.



1 10 Hymenoptera in America North of Mexico

Lagium atroviolaceum (Norton)
Que. to Va. w. to Wis., Ill, Kans. Host: Saynbiicus.

Tenthredo (Allantus) atroviolacea Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot.

Zool. Mass., p. 583. Nom. nud.
Allantus atroviolaceus Norton, 1860. Boston Jour. Nat. Hist. 7: 255. ♀, ♂.
Tenthredo atroviolacea var. peratra Dyar, 1897. N. Y. Ent. Soc, Jour. 5: 192. 6, larva.
Lagium angulabre Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 215. ♂.
Lagium planifrons Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 216. ♀, ♂.

Lagium tardum (Norton)
Mass. to Ga. w. to Wis., Ill, Kans.

Tenthredo (Allantus) tarda Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Allantus tardus Norton, 1860. Boston Jour. Nat. Hist. 7: 246. ♀, ♂.
Lagium erythrogastrum Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 214. ♀.

Genus RHOGOGASTER Konow

Rhogogaster Konow, 1884. Deut. Ent. Ztschr. 28: 257.

Type-species: Teyithredo viridis Linnaeus. Desig. by Rohwer, 1911.
Rhogogastera Konow, 1885. Wien. Ent. Ztg. 4: 123. Emend.

Primarily a Palaearctic genus with only four species in North America. Three of the six spe-
cies groups separated by Benson (1965) are represented on this continent.

Revision: Ross, 1943. Pan-Pacific Ent. 19: 129-136.

Taxonomy: Benson, 1965. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 34: 105-112 (world
species and species groups).

Species Group Lateraria

Rhogogaster addenda (Cresson)
Mont, and Colo. w. to B. C, Wash., Oreg., Calif.

Tenthredo addenda Cresson, 1880. Amer. Ent. Soc, Trans. 8: 23. ♀.

Tenthredo vittatipes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 24. ♂.

Rhogogaster tnincatus Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 211. ♀.

Rhogogaster pitohatus Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 211. ♀.

Rhogogaster lateraria (Cresson)
Alta., Mont., Nev. w. to B. C, Wash., Oreg., Calif.

Tenthredo lateraria Cresson, 1880. Amer. Ent. Soc, Trans. 8: 23. 2.

Tenthredo retosta MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 109. ♂.

Species Group Californica

Rhogogaster californica (Norton)
Que. to Yukon s. to N. Y., Man., N. Mex., Utah, Calif.; Norway. Host:
Populus balsamifera, P. tremuloides.
Tenthredo califomicus Norton, 1862. Ent. Soc. Phila., Proc. 1: 198. ♀.
Tenthredopsis Evansii Harrington, 1889. Canad. Ent. 21: 98. ♀.
Tenthredo ripula MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 109. ♂.
Rhogogaster polaris Lindqvist, 1964. Notulae Ent. 44: 121. ♂.

Taxonomy: Benson, 1965. Roy. Ent. Soc London, Proc. Ser. B: Taxonomy 34: 108.

Species Group Viridis

Rhogogaster viridis (Linnaeus)
Yukon and Alaska s. to Man., Mont, Calif.; Eurasia, Japan. Host: Salix,

Populus, Alnus, Ranunculus, Filipendula, Stellaria, Circaea. These hosts recorded in
Europe.

Tenthredo viridis Linnaeus, 1758. Syst. Nat., Ed. 10, v. 1, p. 557. ♀.

Rhogogastera ruga MacGillivray, 1923. Canad. Ent. 55: 160. ♀.

Rhogogastera respectus MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 165. ♀.

Rhogogaster similis Lindqvist, 1958. Notulae Ent. 38: 68.

Taxonomy: Benson, 1963. Notulae Ent. 43: 137-138 (wear and damage of saws).

Morphology: Zacwilichowska, 1953. Polski Pismo Ent. 23: 114-128 (innervation and sense
organs in wings).



Superfamily TENTHREDINOIDEA 111

Genus ISCHYROCERAEA Kiaer

Ischyroceraea Kiaer, 1898. Tromsoe Mus. Aarsh. 19: 68.

Type-species: Ischyroceraea hyperborea Kiaer. Monotypic.

The species below is sometimes given in the genus Eniscia Thomson, 1870. However, the only
valid type-species designation of Eniscia is Tenthredo consobrina Klug as designated by
Rohwer, 1911. Because of this Eniscia is a synonym of Sciapteryx Stephens, 1835, a Palaearctic
genus.

Ta.\onomy: Smith, 1974. Coop. Econ. Ins. Rpt. 24 (35): 707-710 (first N. Amer. record).

Ischyroceraea arctica (Thomson)
Alaska (Umiat); Siberia, Scandinavia.
Eniscia arctica Thomson, 1870. Opusc. Ent. 2: 300.
Ischyroceraea hyperborea Kiaer, 1898. Tromsoe Miis. Aarsh. 19: 68.

Genus TENTHREDO Linnaeus

Tenthredo Linnaeus, 1758. Syst. Nat., Ed. 10, v. 1, p. 343.

Type-species: Tenthredo scrophulariae Linneaus. Desig. by Latreille, 1810.
Labidia Provancher, 1886. Addit. Corr. Faune Ent. Canada Hym., p. 21.

Type-species: Labidia colutnbiana Provancher. Monotypic.
Parastatis Kirby, 1881. Ent. Monthly Mag. 18: 107.

Type-species: Parastatis indica Kirby. Monotypic.
Rethrax Cameron, 1899. Manchester Lit. Phil. Soc, Mem. and Proc. 43: 33.

Type-species: Rethrax carinata Cameron. Monotypic.
Fethalia Cameron, 1902. Bombay Nat. Hist. Soc, Trans. 14: 439.

Type-species: Fethalia nigra Cameron. Monotypic.
Clydostomus Konow, 1908. Ztschr. System. Hym. Dipt. 8: 19.

Type-species: Clydostomus cestatns Konow. Desig. by Rohwer, 1911.
Tenthredella Rohwer, 1910. U. S. Natl. Mus., Proc. 39: 117.

Type-species: Tenthredo afra Linnaeus. Orig. desig.
Zamacrophya Rohwer, 1912. U. S. Natl. Mus., Proc. 39: 221.

Type-species: Zamacrophya nigrilabris Rohwer. Orig. desig.

Probably the largest sawfly genus, with over 700 described world species. Adults are large
and many are brightly colored. They are commonly found on flowers of various plants where
they feed on nectar and pollen and may also be factors in pollination. Adults are also predaceous
on other arthropods. Hosts and biologies are known for very few species. European authors
have devised their own species groups within the genus; however, until the North American
forms are studied in relation to the world fauna, the species groups below are retained from
previous catalogs.

Taxonomy: Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 225-226 (key to species of Labidia
[originalis group]). — Smulyan, 1923. Boston Soc. Nat. Hist., Proc. 36: 385-465 (New
England species).

Species Group Originalis

Tenthredo anomocera Rohwer
Alta., Wyo., Colo. w. to B. C, Oreg., Calif.

Tenthredo (Labidia) anomocenis Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 223. ♀.
Tenthredo (Labidia) alienatus Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 224. ♀.
Tenthredo (Labidia) anomus Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 225. ♀.

Tenthredo devia (Konow)
Alaska (Aleutian Islands to Popof Is.); Japan, N. Siberia to N. Europe.
Allantus devius Konow, 1900. Ent. Nachr. 26: 124. ♀.
Allantus heraclei Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 364. 9,8. Preocc. by Rudow,

1872.
Allantiis alaskanus Enslin, 1910. Rev. Russe d'Ent. 10: 371. N. name for heraclei Kincaid.

Taxonomy: Benson, 1959. Roy. Ent. Soc. London, Proc, Ser. B: Taxonomy 28: 98.

Tenthredo maxima (Norton)
Alta., Mont., Wyo., Colo., N. Mex. w. to Wash., Oreg., Calif.

Allantus unicinctus Norton, 1864. Ent. Soc. Phila., Proc. 3: 9. ♀, ♂. Preocc. by Brulle,

1832.
Allantus 7naximus Norton, 1867. Amer. Ent. Soc, Trans. 1: 263. ♀.




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Allantus unicinctulus Dalla Torre, 1894. Cat. Hym., v. 1, p. 80. N. name for imicinctus
Norton.

Tenthredo ocampa Ross
Ariz.; Mexico.

Zamacrophya nigrilabris Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 222. ♀, ♂. Preocc. by

Friwaldozky, 1876.
Tenthredo ocampa Ross, 1951. In Muesebeck, et al., U. S. Dept. Agr., Agr. Monog. 2: 71. N.
name for nigrilabris Rohwer.

Tenthredo opima (Cresson)
Colo. w. to B. C, Wash., Oreg., Calif.

Allantus opimus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 15. ♀, ♂.
Labidia Columbiana Provancher, 1886. Addit. Corr. Faune Ent. Canada Hym., p. 21. ♂.
Tenthredo (Labidia) opimus coloradensis Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 224. ♀.

Tenthredo originalis (Norton)
Labrador s. to N. Y.

Allantus originalis Norton, 1867. Amer. Ent. Soc, Trans. 1: 261. ♀.

Tenthredo retura Ross
Nev., Calif.

Allaiitus nigriceps Cresson, 1880. Amer. Ent. Soc, Trans. 8: 16. ♀, ♂. Preocc. by Spinola,

1851.
Tenthredo retura Ross, 1951. In Muesebeck, et al., U. S. Dept. Agr., Agr. Monog. 2: 71. N.
name for nigriceps Cresson.

Tenthredo subnigriceps (Rohwer)
N. Mex., Utah w. to Wash., Oreg., Calif.
Allantus subnigriceps Rohwer, 1909. Canad. Ent. 41: 148. ♀.

Species Group Piceocincta

Tenthredo piceocincta (Norton)
Ont. to Alaska s. to N. H., N. Y., B. C, Wash.

AUayitus piceocinctus Norton, 1860. Boston Jour. Nat. Hist. 7: 249. ♀.

Allantus cogitans Provancher, 1878. Nat. Canad. 10: 163. ♀.

Tenthredo basilaris Provancher, 1878. Nat. Canad. 10: 196. 9. Preocc by Say, 1824.

Tenthredo hudsoni Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 318. 9. A questionable

synonym.
Allantus robustus Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 8. ♀.
Tenthredo ruficoxa Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 14. ♀.
Tenthredo ventricus MacGillivray, 1895. Canad. Ent. 27: 284. ♂.
Tenthredo sicatus MacGillivray, 1900. Canad. Ent. 32: 179. ♂.
Tenthredo iunatus MacGillivray, 1900. Canad. Ent. 32: 180. ♀.
Tenthredo harrimani Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 363. ♀.
Tenthredo cynthia Enslin, 1912. Arch. f. Naturgesch. 6: 105. N. name for basilaris

Provancher.
Tenthredo neoslossoni MacGillivray, 1914. Canad. Ent. 46: 138. ♀.
Tenthredo yuasi MacGillivray, 1920. Brooklyn Ent. Soc, Bui. 15: 112. ♀.
Tenthredo smetica MacGillivray, 1920. Brooklyn Ent. Soc, Bui. 15: 113. ♀.
Tenthredo reputinella MacGillivray, 1923. 111. Univ. Bui. 20(50): 33. ♂.

Species Group Basilaris

Tenthredo basilaris Say
N. S., N. B., Que. to N. C. w. to Alta., Minn., Colo., N. Mex.

Tenthredo (Allantus) coronatus Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 583. Nom. nud.

Tenthredo basilaris Say, 1824. In Keating, Narr. Long's 2nd Exped. (App.) p. 316. ♀.

Allantus universus MacGillivray, 1923. 111. Univ. Bui. 20(50): 6. ♀.

Tenthredo nimina MacGillivray, 1923. Canad. Ent. 55: 160. ♀.

Tenthredo fernaldii MacGillivray
Maine, Ont. to Pa. w. to Minn., Colo.

Allantus dtibius Norton, 1860. Boston Jour. Nat. Hist. 7: 241. ♀. Preocc. by Strom, 1768.

Tenthredo fenialdii MacGillivray, 1900. Canad. Ent. 32: 177. ♀.

Tenthredo perplexa MacGillivray
Alaska s. to Oreg.

Tenthredo perplexus MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 104. ♀.

Tenthredo commata Konow, 1908. Ztschr. System Hym. Dipt. 8: 90. ♀, ♂.

Tenthredo rabula MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 112. ♂.

Tenthredo rima MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 110. ♂.




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Species Group Olivacea

Tenthredo olivacea Klug
Yukon, Alaska; Europe to Siberia.

Tenthredo (AHantus) olivacea Klug, 1814. Mag. Gesell Naturf. Freunde Berlin 8: 193. ♀.

6.
Tenthredo dissimulans Kincaid, 1900. Wash. Acad. Sci. , Proc. 2: 363. ♀, ♂.
Rhogogastera respersus MacGillivray, 1923. N. Y. Ent. See, Jour. 31: 165. ♂.

Species Group Rufopecta

tnellicoxa Provancher. Ont. to Va. w. to Minn., W. Va.

Tenthredo mellkoxa Provancher, 1878. Nat. Canad. 10: 198. ♀, ♂.

Tenthredo repleta MacGillivray
Wash., Oreg.

Tenthredo repleta MacGillivray, 1923. N. Y. Ent. See, Jour. 31: 111. ♀, ♂.

Tenthredo rufopecta (Norton)
N. B. to N. C. w. to Wis.

Allantus nifopectus Norton, 1860. Boston Jour. Nat. Hist. 7: 255. ♀, ♂.

Tenthredo rurigena MacGillivray
N. B., Que., Ont. to N. Y. w. to Minn., Colo.

Tenthredo lineata Provancher, 1878. Nat. Canad. 10: 198. ♀. Preocc. by Christ, 1791.
Tenthredo nirigena MacGillivray, 1923. 111. Univ. Bui. 20(50): 36. ♀.

Species Group Pectoralis

Tenthredo associata Enslin
Calif.

AUantus liinbatns Cresson, 1880. Amer. Ent. Soc, Trans. 8: 17. ♀. Preocc. by Klug, 1814.
Tenthredo associata Enslin, 1912. Arch. f. Naturgesch. 6: 103. N. name for limbatus

Cresson.
Tenthredo cryptella Ross, 1951. In Muesebeck, et ai, U. S. Dept. Agr., Agr. Monog. 2: 73.

N. name for limbatus Cresson.

Tenthredo pectoralis Norton
Idaho, Colo. w. to Alaska, B. C, Wash., Oreg., Calif.
Tenthredo pectoralis Norton, 1864. Ent. Soc. Phila., Proc. 3: 11. ♂.
Tenthredo variegatus Norton, 1864. Ent. Soc. Phila., Proc. 3: 12. 6. A questionable

synonym.
Allantus afflictus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 17. ♀, ♂.
Alla7itus nevadensis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 17. ♀, ♂.
Allantus occidaneus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 18. ♀.
Tenthredo parvula Cresson, 1880. Amer. Ent. Soc, Trans. 8: 22. 6. Preocc by Lepeletier,

1823
Tenthredo lateralba MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 108. ♀.
Teyithredo aldrichii MacGillivray, 1900. Canad. Ent. 32: 183. ♀.
Tenthredo atracostus MacGillivray, 1900. Canad. Ent. 32: 183. ♀.
Tenthredo nigrifascia MacGillivray, 1900. Canad. Ent. 32: 184. ♀.
Tenthredo savagei MacGillivray, 1900. Canad. Ent. 32: 184. ♀.
Tenthredo coenobita Enslin, 1912. Arch. f. Naturgesch. 6: 104. N. name for parvula

Cresson.
Tenthredo remissa MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 114. ♂.
Tenthredo reperta MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 115. ♀.
Tenthredo replata MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 115. ♀.
Tenthredo reticentia MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 114. ♀.
Tenthredo refuga MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 167. ♀.
Tenthredo rota MacGillivray, 1923. 111. Univ. Bui. 20(50): 34. ♀.
Tenthredo ndnosa MacGillivray, 1923. 111. Univ. Bui. 20(50): 36. ♀.
Teyithredo rata MacGillivray, 1923. 111. Univ. Bui. 20(50): 37. ♀.

Species Group Angulifera

Tenthredo angulifera (Norton)
N. B., Que. to Pa. w. to Mich., Ohio.

Allantus angulifer Norton, 1860. Boston Jour. Nat. Hist. 7: 252. ♀, ♂.

Carolina (Rohwer). N. C, Ohio.

Tenthredella Carolina Rohwer, 1913. U. S. Natl. Mus., Proc 45: 275. ♀.

Tenthredo dissimilis (Norton)
Maine, N. Y.

Allantus dissimilis Norton, 1860. Boston Jour. Nat. Hist. 7: 250. ♀.
Tenthredo bilineatus MacGillivray, 1895. Canad. Ent. 27: 282. ♀.




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Tenthredo fisheri (Rohwer)
N. J., Md., Va., 111.

Tenthredella fisheri Rohwer, 1913. U. S. Natl. Mus., Proc. 45: 276. ♀.

Tenthredo glacialis (Ross)
111.

Tenthredella glacialis Ross, 1931. Ent. Soc. Amer., Ann. 24: 110. ♀, ♂.

Tenthredo lobata (Norton)
N. Y. to N. C. w. to Wis.

Allanius lobatus Norton, 1860. Boston Jour. Nat. Hist. 7: 253. ♀.

Tenthredo mantha Ross
Conn. w. to Wis., 111.

Tenthredella lobata maculosa Smulyan, 1915. Canad. Ent. 47: 324. ♀, ♂. Preocc. by
Lepeletier, 1823.

Tenthredo mayitha Ross, 1951. In Muesebeck, et ai, U. S. Dept. Agr., Agr. Monog. 2: 73. N.
name.

Species Group Leucostoma

Tenthredo cinctitibiis Norton
N. H., N. Y. w. to Alta.

Tenthredo cinctitibiis Norton, 1869. Amer. Ent. Soc, Trans. 2: 239. ♀.

Tenthredo decorata Provancher, 1878. Nat. Canad. 10: 200. 9. A questionable synonym.

Tenthredo erythromera Provancher
Que., S. Dak., Colo. w. to Alaska, B. C, Wash., Oreg., Calif.

Tenthredo erythromera Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 13. ♀.
Tenthredo atrocoerulea Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 13. ♀.
Tenthredo nigricostata Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 14. ♀.
Tenthredo terminalis Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 14. ♂.

Preocc. by Smith, 1878.
Tenthredo nigrisoma Harrington, 1894. Canad. Ent. 26: 195. ♀. Preocc.
Tenthredo chaonica Enslin, 1912. Arch. f. Naturgesch. 6: 105. N. name for terrninalis

Provancher.
Tenthredo regula MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 166. ♀.
Tenthredo requieta MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 167. ♀.

Tenthredo fraternalis (Ross)
Alta., B. C.

Tenthredella fratenialis Ross, 1931. Ent. Soc Amer., Ann. 24: 113. ♀.

Tenthredo fumipennis Norton
Calif.

Tenthredo fumipennis Norton, 1869. Amer. Ent. Soc, Trans. 2: 236. ♂.

Tenthredo leucostoma Kirby
N. S. to Pa. w. to Alta., S. Dak., Colo.

Tenthredo nifipes Say, 1824. hi Keating, Narr. Long's 2nd Exped. (App.) p. 317. ♀. Preocc.

by Linnaeus, 1758.
Tenthredo leucostoma Kirby, 1837. Fauna Bor.-Amer., v. 4, p. 256.
Tenthredo rufopedibus Norton, 1864. Ent. Soc Phila., Proc. 3: 15. ♂.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 392 (Possibly a geographical
race of Tenthredo atra Linnaeus, a widespread species in Eurasia).

Tenthredo mellipes Rohwer
Colo.

Tenthredo mellipes Rohwer, 1909. Canad. Ent. 41: 105. ♀.

Tenthredo nigritibialis MacGillivray
Wash.

Tenthredo nigritibialis MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 107. ♂.

Tenthredo nimbipennis Cresson
Md., Ill, Ark., Tex.

Tenthredo nimbipennis Cresson, 1872. Amer. Ent. Soc, Trans. 4: 155. ♀.

Tenthredo remea MacGillivray
Oreg.

Tenthredo remea MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 107. ♀.

Tenthredo remora MacGillivray, 1923. N. Y. Ent. Soc, Jour. 21: 108. ♂.

Tenthredo ruficollis Harrington
B. C.

Tenthredo ruficollis Harrington, 1894. Canad. Ent. 26: 195. ♀.

Tenthredo ruma MacGillivray
Pa.

Tenthredo nima MacGillivray, 1923. 111. Univ. Bui. 20(50): 36. ♂.

Tenthredo subcoerulea Eschscholtz
Que. to Alaska s. to Fla., Colo., Calif. Ecology: Widespread.

Tenthredo subcoerulea Eschscholtz, 1822. Entomographien, p. 91.

Allantus flavomarginis Norton, 1860. Boston Jour. Nat. Hist. 7: 254. ♀.

Tenthredo melanosoma Harrington, 1894. Canad. Ent. 26: 194. ♀.




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Tenthredo pallipundus MacGillivray, 1895. Canad. Ent. 27: 282. 9. A questionable

synonym.
Tenthredo sahlbergii Konow, 1908. Ztschr. System. Hym. Dipt. 8: 91. ♀, ♂.

Species Group OccmENTALis

Tenthredo bella Cresson
Colo.

Tenthredo bella Cresson, 1880. Amer. Ent. Soc, Trans. 8: 21. ♀.

Tenthredo occidentalis Cresson
Ont. (?), Wyo., Colo.

Tenthredo occidentalis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 23. ♀.

Tenthredo vapida Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 315. 9, synonym.

Tenthredo femigineipes var. pikei Rohwer, 1909. Canad. Ent. 41: 107. ♀.

Tenthredo varifrons Rohwer, 1909. Canad. Ent. 41: 110.

Tenthredo subrufescens Kirby
Ont.

Tenthredo subrufescens Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 311. ♀.

Species Group Grandis

Tenthredo colon Klug
Labrador, N. B., N. H. w. to Alaska; Europe, Asia Minor, Siberia. Host: Viburnum,
Salix, Rosa, Corylus, Sorbtis. These hosts recorded in Europe.
Tenthredo {Allantus) colon Klug, 1814. Mag. Gesell. Naturf. Freunde BerHn 8: 182. ♀, ♂.
Tenthredo ebumeifrons Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 306. ♀.
Tenthredo nigricollis Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 308. ♀.

Taxonomy: Benson, 1962. Brit. Mus. (Nat. Hist.) Ent., Bui. 12: 392 {Tenthredo livida of
American authors, not Linnaeus).

Tenthredo grandis (Norton)
N. B., Que., Ont. s. to N. C.

Allantus grandis Norton, 1860. Boston Jour. Nat. Hist. 7: 239. ♀.
Tenthredo antennata Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 308. ♀.
Tenthredo dubitatus MacGilhvray, 1897. N. Y. Ent. Soc, Jour. 5: 103. ♂.

Species Group Verticalis

Tenthredo borealis
***authority mismatch
Kirby. Ont., N. Y. w. to Alta., Minn.

Tenthredo cingulata Provancher, 1878. Nat. Canad. 10: 196. ♀. Preocc.

Tenthredo borealis Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 318. ♀.

Tenthredo nitila MacGillivray 1923. Canad. Ent. 55: 160. ♀.

Tenthredo verticalis Say
N. B., Que., Ont. to N. C. w. to Minn.

Tenthredo verticalis Say, 1824. In Keating, Narr. Long's Exped. (App.) p. 317. ♀.

Tenthredo causatus MacGilhvray, 1900. Canad. Ent. 32: 178. ♂.

Tenthredo mbripes MacGilhvray, 1900. Canad. Ent. 32: 178. ♂.

Species Group Semirufa

Tenthredo attracta Norton
"English River."

Tenthredo attractus Norton, 1869. Amer. Ent. Soc, Trans. 2: 240. ♀.

Tenthredo concessa Norton
Ont. to Pa. w. to Alta., S. Dak., Mont., Colo. Host: Betula.
Allantus tricolor Norton, 1860. Boston Jour. Nat. Hist. 7: 247. 9. Preocc
Tenthredo concessus Norton, 1869. Amer. Ent. Soc, Trans. 2: 238. ♀.
Tenthredo bamstoni Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 314. 9, 6. A questionable

synonym.
Tenthredo cressonii Kirby, 1882. List. Hym. Brit. Mus., v. 1, p. 315. ♀.
Tenthredo uniformis Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 317. ♀.
Tenthredella rohweri Smulyan, 1915. Canad. Ent. 47: 323. N. name for tricolor Norton.
Tenthredella tricolor var. savanna Ross, 1931. Ent. Soc. Amer., Ann. 24: 116. ♀.

Biology: Dyar, 1895. Amer. Ent. Soc, Trans. 12: 312.

Tenthredo eximia Norton
Maine to N. J. w. to Alta., Colo.

Tenthredo eximius Norton, 1869. Amer. Ent. Soc, Trans. 2: 231. ♀, ♂.

Tenthredo pallicoxa Provancher, 1878. Nat. Canad. 10: 201. 9. A questionable synonym.

Tenthredo nova MacGilhvray, 1897. N. Y. Ent. Soc, Jour. 5: 105. ♀.




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ferrugincipes Cresson. Idaho, Colo., N. Mex. w. to Alaska, B. C, Oreg., Calif.
Tenthredo femigineipes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 22. ♀.
Tenthredo sectilis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 22. ♀.
Tenthredo luteipes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 21. 9. A questionable

synonym.
Tenthredo tenninatns MacGillivray, 1895. Canad. Ent. 27: 283. ♀.
Tenthredo aequalis MacGillivray, 1895. Canad. Ent. 27: 284. ♀.
Tenthredo nibrisonunus MacGillivray, 1900. Canad. Ent. 32: 181. ♀.
Tenthredo retinentia MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 166. ♀.
Tenthredo nibrica MacGillivray, 1923. 111. Univ. Bui. 20(50): 35. ♀.

Tenthredo lacticincta Cresson
Nev., Calif.

Tenthredo lacticincta Cresson, 1880. Amer. Ent. Soc, Trans. 8: 21. ♀, ♂.

Tenthredo olivatipes MacGillivray
Wash., Calif.

Allantus intemiptus Norton, 1867. Amer. Ent. Soc, Trans. 1: 263. ♂. Preocc. by Fabricius,
1804.

Tenthredo olivatipes MacGillivray, 1900. Canad. Ent. 32: 180. ♀.

Tenthredo asella Enslin, 1912. Arch. f. Naturgesch. 6: 103. N. name for interruptus
Norton.

Tenthredo ornaticeps (Cresson)
Nev., Calif.

Allantus ornaticeps Cresson, 1880. Amer. Ent. Soc, Trans. 8: 16. ♀, ♂.

Tenthredo pallicola
***authority mismatch
MacGiUivray. N. H., Alta., N. Mex.

Tenthredo pallicola MacGillivray, 1895. Canad. Ent. 27: 80. ♀.

Tenthredo pallicola var. beulahensis Rohwer, 1909. Canad. Ent. 41: 108. ♀.

Tenthredo pleuralis Cresson
Colo.

Tenthredo pleuralis Cresson, 1865. Ent. Soc. Phila., Proc. 4: 245. ♀, ♂.

Teiifhredo utensis Rohwer, 1909. Canad. Ent. 41: 109. ♀.

Tenthredo rabida MacGillivray
Oreg.

Tenthredo rabida MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 110. ♀.

Tenthredo ralla MacGillivray
Oreg.

Tenthredo ralla MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 111. ♀.

Tenthredo reduvia MacGillivray
Oreg.

Tenthredo reduvia MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 108. ♀.

Tenthredo rabiosa MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 112. ♀.

Tenthredo resima MacGillivray
Oreg.

Tenthredo resima MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 109. ♀.

Tenthredo rubeola Cresson
Nev.

Tenthredo rubeola Cresson, 1880. Amer. Ent. Soc, Trans. 8: 23. ♂.

Tenthredo rubicunda MacGillivray
N. H.

Tenthredo rubicunda MacGillivray, 1923. 111. Univ. Bui. 20(50): 34. ♀.

Tenthredo semirufa Norton
N. H. to N. C. w. to Idaho, Colo. Host: Betula.

Allantus signatus Norton, 1860. Boston Jour. Nat. Hist. 7: 247. ♀. Preocc. by Scopoli, 1763.

Tenthredo semirufus Norton, 1864. Ent. Soc. Phila., Proc. 3: 12. ♀.

Tenthredo seminibra Norton, 1864. Ent. Soc. Phila., Proc. 3: 14. ♀.

Tenthredo thora Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 310. 9, cJ. A questionable

synonym.
Tenthredo reviota MacGillivray, 1895. Canad. Ent. 27: 81. ♀.
Tenthredo junghannsii MacGillivray, 1900. Canad. Ent. 32: 179. ♀.
Tenthredo denotata Enslin, 1912. Arch. f. Naturgesch. 6: 105. Unnecessary n. name for

Tenthredo basilaris
***authority mismatch
MacGillivray, 1897.
Tenthredo ruina MacGillivray, 1923. 111. Univ. Bui. 20(50): 36. ♀.
Tenthredo nisticaiia MacGillivray, 1923. 111. Univ. Bui. 20(50): 37. ♀.
Tenthredo russa MacGillivray, 1923. 111. Univ. Bui. 20(50): 37. ♀.

Taxonomy: Dyar, 1897. N. Y. Ent. Soc, Jour. 5: 19 (larva and host of remota MacGillivray).

Tenthredo variata Norton
Colo., Wash.

Tenthredo variatus Norton, 1864. Ent. Soc. Phila., Proc. 3: 12. ♀, ♂.
Tenthredo capitatus MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 108. ♀.




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Tenthredo varipicta Norton
Alaska s. to Nev., Calif.

Tenthredo varipictus Norton, 1868. Amer. Ent. Soc, Trans. 2: 234. ♀.
Tenthredo scaevola Cresson, 1880. Amer. Ent. Soc, Trans. 8: 20. ♀.
Tenthredo magnifica MacGillivray, 1893. Canad. Ent. 25: 240. ♀.
Tenthredo Imipes MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 104. ♂.

Species Group Secunda

Tenthredo nortoni (Smulyan)
Mass., N. Y., N. J.

Tenthredo angidatus Norton, 1864. Ent. Soc Phila., Proc 3: 13. ♂. Preocc. by Westwood,
1838.

Tenthredella nortoni Smulyan, 1915. Canad. Ent. 47: 321. ♂.

Tenthredo relativa Rohwer
Nebr.

Tenthredo relativus Rohwer, 1909. Canad. Ent. 41: 108. ♀.

Tenthredo rhammisia MacGillivray
B. C, Wash., Oreg.

Tenthredo rhammisia MacGillivray, 1923. 111. Univ. Bui. 20(50): 33. ♀.

Tenthredo rutata MacGillivray
N. J.

Tenthredo rutata MacGillivray, 1923. 111. Univ. Bui. 20(50): 38. ♀.

Tenthredo secunda MacGillivray
N. H., Mass., N. Y., Ont.

Tenthredo secundus MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 105. ♀.

Tenthredo siynulatus MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 105. ♀.
Strickland! (Ross). Alta., Idaho, Alaska, Oreg.

Teyithredella stricklaiidi Ross, 1931. Ent. Soc Amer., Ann. 24: 117. ♀, ♂.

Tenthredo vellosa Ross
Conn.

Tenthredo formosus Norton, 1864. Ent. Soc. Phila., Proc. 3: 14. ♀, ♂. Preocc. by Klug, 1814.

Tenthredo vellosa Ross, 1951. In Muesebeck, et al., U. S. Dept. Agr., Agr. Monog. 2: 77. N.
name.

Species Group Xantha

Tenthredo macgillivrayi (Smulyan)
N. B., Que., Vt. w. to Alta.

Teyithredella macgillivrayi Smulyan, 1915. Canad. Ent. 47: 323. ♀.

Tenthredo mellina (Norton)
Maine.

Allantiis melliniis Norton, 1860. Boston Jour. Nat. Hist. 7: 248. ♀, ♂.

Tenthredo obscuripennis Cresson
Oreg., Nev., Calif.

Tenthredo obscuripennis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 20. ♀.

Tenthredo rubella Cresson, 1880. Amer. Ent. Soc, Trans. 8: 22. ♂.

Tenthredo prosopa Stannard
N. Y. to Ga.

Tenthredo prosopa Stannard, 1947. Ent. Soc. Amer., Ann. 40: 434. ♀, ♂.

Tenthredo redimacula MacGillivray
N. H., Ont.

Tenthredo, vittata Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 319. 9. Preocc by
Kriechbaumer, 1869.

Tenthredo redimacula MacGillivray, 1895. Canad. Ent. 27: 78. ♀.

Tenthredo co7iso7na Enslin, 1912. Arch. f. Naturgesch. 6: 105. N. name for vittata Kirby.

Tenthredo xantha Norton
Maine, Ont. to Alaska s. to Minn., N. Mex., S. Dak., Utah, Calif. Ecology:
Widespread. Host: Rubiisi^.

Tenthredo xanthus Norton, 1864. Ent. Soc. Phila., Proc. 3: 13. ♀.

Tenthredo nificolor Norton, 1868. Amer. Ent. Soc, Trans. 2: 228. ♀.

Tenthredo nupera Cresson, 1880. Amer. Ent. Soc, Trans. 8: 22. cJ. A questionable
synonym.

Tenthredo castanea Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 319. ♀.

Allantus rubricus Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 9. ♀.

Tenthredo reliquia MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 168. ♀.

Tenthredo resticula MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 168. ♀.

Biology: Smith and Kido, 1949. Hilgardia 19: 54 (host).




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Species Group Bigemina

Tenthredo bigemina (Dyar)
B. C. s. to Nev., Calif.

Allantus annularis Norton, 1867. Amer. Ent. Soc, Trans. 1: 262. ♀, ♂. Preocc. by

Schrank, 1776.
Allantus elegantulus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 17. ♀, ♂. Preocc. by Fallen,

1808.
Labidia opimus var. bigeminus Dyar, 1893. Canad. Ent. 25: 195. ♀.
Tenthredo obliquatus MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 105. ♀, ♂.
Tenthredo elegantula oregona Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 411. ♀.
Tenthredo admeta Enslin, 1912. Arch. f. Naturgesch. 6: 103. N. name for annularis

Norton.
Tenthredo arethtisa Enslin, 1912. Arch. f. Naturgesch. 6: 103. N. name for elegantulus

Cresson.

Unplaced Taxa of Tenthredo Linnaeus

Most of the species in the following list were described from males and belong to groups in
which it is impossible to identify this sex with any degree of certainty.

Tenthredo alpestris Rohwer
Colo.

Tenthredo alpestris Rohwer, 1909. Canad. Ent. 41: 146. ♂.

Tenthredo auraria Konow
Alaska (Ft. Wrangel).

Tenthredo anraria Konow, 1899. Ent. Nachr. 25: 7. ♀.

Tenthredo bivittata Kincaid
Alaska.

Tenthredo bivittata Kincaid, 1900. Wash. Acad. Sci., Proc. 2: 364. ♀.

Tenthredo cinctitibiis
***authority mismatch
abdominalis (Smulyan), n. comb. N.H.

Tenthredella ciiictitibiis abdominalis Smulyan, 1915. Canad. Ent. 47: 324. ♀. Preocc. by
Fabricius, 1798.

Tenthredo discrepans Norton
Ont.

Tenthredo discrepans Norton, 1868. Amer. Ent. Soc, Trans. 2: 235. ♂.

Tenthredo divergens Rohwer
Colo.

Tenthredo divergens Rohwer, 1909. Canad. Ent. 41: 112. ♂.

Tenthredo diversiceps Rohwer
N. B.

Tenthredo diversiceps Rohwer, 1910. U. S. Natl. Mus., Proc. 38: 208. ♂.

Tenthredo frigida MacGillivray
N. H., Wash.

Tenthredo frigida MacGillivray, 1895. Canad. Ent. 27: 80. ♂.

Tenthredo hyalina MacGillivray
N. Y.

Tenthredo hyalina MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 108. ♂. Preocc.

Tenthredo hypoleuca Rohwer
Colo.

Tenthredo hypoleuca Rohwer, 1909. Canad. Ent. 41: 147. ♂.

Tenthredo juga Konow
Alaska.

Tenthredo juga Konow, 1908. Ztschr. System. Hym. Dipt. 8: 89. ♀.

Tenthredo lovetti Benson
Oreg.

Teyithredo rustica MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 113. ♂. Preocc. by

Linnaeus, 1758.
Tenthredo lovetti Benson, 1930. Entomologist 63: 107. N. name.

Tenthredo magnata MacGillivray
Wash.

Tenthredo magnatus MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 107. ♂.

Tenthredo messica MacGillivray
Wash.

Tenthredo rnessica MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 107. ♂.

Tenthredo messicaeformis Rohwer
Colo.

Tenthredo messicaeformis Rohwer, 1909. Canad. Ent. 41: 147. ♂.

Tenthredo mimula Cresson
Colo.

Tenthredo ynimula Cresson, 1880. Amer. Ent. Soc, Trans. 8: 23. ♂.

Tenthredo morosa Cresson
Colo.

Tenthredo morosa Cresson, 1880. Amer. Ent. Soc, Trans. 8: 23. ♂.




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■nutans Norton. Canada.

Tenthredo mutans Norton, 1869. Amer. Ent. Soc, Trans. 2: 236. ♂.

Tenthredo nigricoxi MacGillivray
Wash.

Tenthredo nigricoxi MacGillivray, 1900. Canad. Ent. 32: 181. ♂.

Tenthredo pallipectis MacGillivray
Wash.

Tenthredo pallipectis MacGillivray, 1897. N. Y. Ent. See, Jour. 5: 106. ♂.

Tenthredo reflua MacGillivray
Oreg.

Tenthredo reflua MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 111. ♂.

Tenthredo reposita MacGillivray
Oreg.

Tenthredo reposita MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 116. ♂.

Tenthredo reputina MacGillivray
Oreg.

Tenthredo repntiyia MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 114. ♂.

Tenthredo resegmina MacGillivray
Oreg.

Tenthredo resegmina MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 115. ♂.

Tenthredo restricta MacGillivray
Oreg.

Tenthredo restricta MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 110. ♂.

Tenthredo resupina MacGillivray
Oreg.

Tenthredo resupina MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 113. ♂.
retrovcrsa MacGillivray. Alaska.

Teyithredo retroversa MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 164. ♂.

Tenthredo rotula MacGillivray
N. Y.

Tenthredo rotula MacGillivray, 1923. 111. Univ. Bui. 20(50): 34. ♂.

Tenthredo rudicula MacGillivray
Maine.

Tenthredo rudicula MacGillivray, 1923. 111. Univ. Bui. 20(50): 35. ♀.

Tenthredo rufescens (Norton)
Maine.

Allantus rufescens Norton, 1860. Boston Jour. Nat. Hist. 7: 245. ♀.

Tenthredo rufostigma MacGillivray
Idaho.

Teyithredo nifostigmtis MacGillivray, 1895. Canad. Ent. 27: 283. ♂.

Tenthredo rusticula MacGillivray
Alaska.

Teyithredo rusticula MacGillivray, 1923. N. Y. Ent. Soc, Jour. 31: 166. ♂.

Tenthredo sectiliformis Rohwer
Colo.

Tenthredo sectiliformis Rohwer, 1909. Canad. Ent. 41: 145. ♀.

Tenthredo slossonii MacGillivray
N. H.

Tenthredo slossonii MacGillivray, 1900. Canad. Ent. 32: 179. ♂.

Tenthredo stigmata MacGillivray
Wash.

Tenthredo stigmatus MacGillivray, 1897. N. Y. Ent. Soc, Jour. 5: 108. ♂.

Tenthredo suavis Cresson
Nev.

Teyithredo suavis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 21. ♂.

Tenthredo titusi Rohwer
Colo.

Teyithredo titusi Rohwer, 1909. Canad. Ent. 41: 111. ♂.

Tenthredo varians Norton
Canada.

Teyithredo variayis Norton, 1868. Amer. Ent. Soc, Trans. 2: 235. ♂.

Tenthredo ventralis (Say)
Ark.

Allayitus veyitralis Say, 1823. West. Quart. Rptr. 2: 71.

Tenthredo zelmira Rohwer
Colo.

Tenthredo zelmirus Rohwer, 1909. Canad. Ent. 41; 109. ♂.

Tenthredo zetes Kirby
Ont.

Teyithredo zetes Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 312. ♂.

Tribe MACROPHYINI
Genus MACROPHYA Dahlbom

Teyithredo subg. Macrophya Dahlbom, 1835. Conspectus Tenthred., Siricid., Oryssin.
Scand., p. 11.




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Type-species: Tenthredo rusticus Linnaeus Desig. by Westwood, 1840. T. rusticus in
sense of authors at that time.
Zalagium Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 216.

Type-species: Zalagium clypeatitm Rohwer. Orig. desig.

A large holarctic genus. Little is known of the hosts and biologies for most species.

Taxonomy: Middleton, 1922. U. S. Natl. Mus., Proc. 61: 6-11 (larvae). —Ross, 1931. Ent. Soc.

Amer., Ann. 24: 119-128 (species with white apical antennal segments). —Malaise and

Benson, 1934. Arkiv for Zool. 26A: 4-5 (.Tenthredo nistica L. is Argidae; T. rustica auct.,

nee Linnaeus= Macropliya montana (Scopoli)).

Macrophya alba MacGillivray
Que. to Pa. w. to Wis., Ind.

Macrophya pulchella alba MacGillivray, 1895. Canad. Ent. 27: 285. ♀, ♂.

Macrophya zonata Konow, 1899. Wien. Ent. Ztg. 18: 44. Unnecessary N. name for alba.

Macrophya annulipes Cresson
Nev., Calif.

Macrophya annulipes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 18. ♀.

Macrophya bifasciata (Say)
Conn, to Fla. w. to Okla., Tex.

Tenthredo (Alla^itus) ynedia Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 583. Nom. nud.

Allayitus bifasciatus Say, 1823. West. Quart. Rptr. 2: 72. ♀.

Macrophya medius Norton, 1861. Boston Soc. Nat. Hist., Proc. 9: 118.

Macrophya texa7ia Cresson, 1880. Amer. Ent. Soc, Trans. 8: 52. ♀, ♂.

Macrophya zabriskiei Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 218. ♀, ♂.

Macrophya castaneae Rohwer
Va. Host: Ca.stanea dentata.

Macrophya castaneae Rohwer, 1917. U. S. Natl. Mus., Proc 53: 151. ♂.

Macrophya cinctula (Norton)
Conn, to Ga. w. to Minn., 111.

Tenthredo atroviolacea var. cinctulus Norton, 1869. Amer. Ent. Soc, Trans. 2: 240. ♀.

Allantiis dejectiis Norton, 1860. Boston Jour. Nat. Hist. 7: 249. 6. A questionable
synonym.

Macrophya abbotii Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 269. ♂.

Zalagium clypeatum Rohwer, 1912. U. S. Natl. Mus., Proc 43: 217. ♂.

Macrophya coquilletti (Rohwer)
Calif.

Sciapteryx coquilletti Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 212. ♀, ♂.

Macrophya fascialis Norton
N. Y. to Md. w. to Mich., 111.

Macrophya fascialis Norton, 1867. Amer. Ent. Soc, Trans. 1: 276. ♀.

Macrophya fascialis var. puella Ross, 1931. Ent. Soc. Amer., Ann. 24: 123, 9,6.

Macrophya flavicoxae (Norton)
N. B., Ont. to Ga. w. to Alta., Wis.

Allantus flavicoxae Norton, 1860. Boston Jour. Nat. Hst. 7: 258. ♀.

Allantus incertus Norton, 1860. Boston Jour. Nat. Hist. 7: 258. ♂.

Selandria canadensis Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym., p. 7. ♀.

Macrophya flavicoxis Dalla Torre, 1894. Cat. Hym., v. 1, p. 52. Emend.

Macrophya formosa (Klug)
Mass. to Fla. w. to Ohio.

Tenthredo (Allayitus) formosa Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 115. ♀.

Allantus bicinctus Norton, 1860. Boston Jour. Nat. Hist. 7: 241. ♀, ♂.

Macrophya fumator Norton
Colo., Idaho w. to B. C, Wash., Oreg., Calif.

Macrophya fumator Norton, 1867. Amer. Ent. Soc, Trans. 1: 279. ♀.

Macrophya maura Cresson, 1880. Amer. Ent. Soc, Trans. 8: 18. ♀.

Macrophya jugosa Cresson, 1880. Amer. Ent. Soc, Trans. 8: 18. ♀.

Macrophya bicolorata Cresson, 1880. Amer. Ent. Soc, Trans. 8: 18. ♀.

Macrophya fumatrix Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 275. Misspelling ?.

Synairema pacifica Provancher, 1885. Addit. Corr. Faune Ent. Canada Hym. p. 15. ♀.

Perineura kincaidia MacGillivray, 1895. Canad. Ent. 27: 7. ♀.

Macrophya obnissa MacGillivray, 1923. 111. Univ. Bui. 20(50): 22. ♂.

Macrophya goniphora (Say)
N. Y. to Ga., Ala.

Allantus goniphorus Say, 1836. Boston Jour. Nat. Hist. 1: 216. ♀, ♂.
intermedia (Norton). Conn., N. Y. w. to Wis., Nebr., Mo.

Allantus intermedius Norton, 1860. Boston Jour. Nat. Hist. 7: 242. ♀, ♂.

Macrophya simillima Rohwer, 1917. Ent. News 28: 265. ♀, ♂.




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Macrophya nebraskensis Rohwer
Nebr.

Macrophya sambuci Rohwer, 1909. Canad. Ent. 41: 15. ♀. Preocc. by Latreille, 1805.

Macrophya nebraskensis Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 220. N. name.

Macrophya nigra (Norton)
N. H. to Ala. w. to III, Kans.

AUantus niger Norton, 1860. Boston Jour. Nat. Hist. 7: 239. ♀.

Macrophya nigristigma Rohwer
Conn., Pa., W. Va. Host: Hicoria.

Macrophya nigristigma Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 219. ♀.

Macrophya melanopleura MacGillivray, 1914. Canad. Ent. 46: 139. ♀.

Taxonomy: Middleton, 1922. U. S. Natl. Mus., Proc. 61: 9 (larva).

Macrophya pulchella (Klug)
Ont., Conn, to Va. w. to Wis., Nebr., Colo., Miss. Host: Aster, Solidago,

Rudbeckia.
Tenthredo (AUantus) pulchella Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 8: 121. ♂.
Allantus flavolineatus Norton, 1860. Boston Jour. Nat. Hist. 7: 259. ♀, ♂.
Macrophya lineata Norton, 1867. Amer. Ent. Soc, Trans. 1: 269. ♀.
Macrophya zoe Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 270. ♀.
Macrophya crassicomis Provancher, 1888. Addit. Corr. Faune Ent. Canada Hym., p. 352.

6. A questionable synonym.
Macrophya lineatana Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 220. ♀.
Macrophya confusa MacGillivray, 1914. Canad. Ent. 46: 139. ♀.

Macrophya pulchelliformis Rohwer
Nebr.

Macrophya pulchelliformis Rohwer, 1909. Canad. Ent. 41: 15. ♂.

Macrophya pulcherrima Kirby
Fla.

Macrophya pulcherrima Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 275. ♂.

Macrophya pumila Norton
Alaska s. to CaHf.

Macrophya pu7nilus Norton, 1867. Amer. Ent. Soc, Trans. 1: 272. ♂.

Macrophya oregona Cresson, 1880. Amer. Ent. Soc, Trans. 8: 19. ♀.

Macrophya obaereta MacGillivray, 1923. 111. Univ. Bui. 20(50): 21. ♀.

Macrophya oregona var. dukiae Ross, 1931. Ent. Soc. Amer., Ann. 24: 122. ♀, ♂.

Macrophya punctumalbum (Linnaeus)
Ont.; Europe to Caucasus. Introduced (?). Host: Fraxinus,
Ligustnim. Hosts recorded in Europe.

Tenthredo punctum album Linnaeus, 1767. Syst. Nat., Ed. 12, p. 924.

Tenthredo erythropus Schrank, 1776. Beitr. z. Naturgesch., p. 86.

Macrophya subviolacea Cresson
Calif.

Macrophya subviolacea Cresson, 1880. Amer. Ent. Soc, Trans. 8: 18. ♀.

Macrophya succincta Cresson
Conn., N. Y. to Ga. w. to Sask., Mont., Kans., Tex.

Macrophya succincta Cresson, 1880. Amer. Ent. Soc, Trans. 8: 19. ♀.

Macrophya xanthonota Rohwer, 1912. U. S. Natl. Mus., Proc 43: 218. ♀.

Macrophya trisyllaba (Norton)
N. B., Que. to Md. w. to Alta., Wis., 111., Ala. Host: Sambucus.

Tenthredo (Allantus) trisyllabus Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 583. Nom. nud.

Allantus trisyllabus Norton, 1860. Boston Jour. Nat. Hist. 7: 238. ♀.

Macrophya trisyllabus var. sinannula Ross, 1931. Ent. Soc. Amer., Ann. 24: 127. ♀, ♂.

Taxonomy: Dyar, 1897. N. Y. Ent. Soc, Jour. 5: 192 (larva). —Middleton, 1922. U. S. Natl.

Mus., Proc. 61: 8 (larva).

Macrophya trosula (Norton)
N. H. to Va. w. to Minn., 111.

Tenthredo (Allantus) trosula Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 583. Nom. nud.

Allantus trosula Norton, 1860. Boston Jour. Nat. Hist. 7: 244. ♀.

Macrophya albifacies Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 271. ♀.

Macrophya dyari Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 410. ♀.

Macrophya omata MacGillivray, 1914. Canad. Ent. 46: 139. ♀.

Macrophya varia (Norton)
N. B., Que. to N. Y. w. to Man., Minn., 111.

Allantus varius Norton, 1860. Boston Jour. Nat. Hist. 7: 240. ♀.

Macrophya eurythmia Norton, 1867. Amer. Ent. Soc, Trans. 1: 276. ♀, ♂.

Macrophya iiidonea MacGillivray, 1895. Canad. Ent. 27: 77. ♂.

Macrophya varius \ar.festana Ross, 1931. Ent. Soc. Amer., Ann. 24: 124. ♀.




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Macrophya varius var. nordicola Ross, 1931. Ent. Soc. Amer., Ann. 24: 125. ♀, ♂.

Macrophya zonalis Norton
Que. to Md. w. to Minn.

Macrophya zonalis Norton, 1864. Ent. Soc. Phila., Proc. 3: 11. ♀.

Unplaced Taxa of Macrophya Dahlbom

The names in the following list fall into a difficult complex previously known as the Epinota
complex. The species are merely listed without regard to possible synonymy.

Macrophya albilabris Harrington
Ont. (Ottawa).

Macrophya albilabris Harrington, 1893. Canad. Ent. 25: 60. ♂.

Macrophya albomaculata (Norton)
Maine, Conn. Host: Vibumum(^).

Allantus albomaculata Norton, 1860. Boston Jour. Nat. Hist. 7: 256. ♀, ♂.

Macrophya bellula MacGillivray
N. J.

Macrophya bellula MacGillivray, 1923. Brooklyn Ent. Soc, Bui. 18: 55. ♀.

Macrophya bilineata MacGillivray
Conn., 111. Host: Vibumumi?).

Macrophya bilineata MacGillivray, 1916. Conn. State Geol. and Nat. Hist. Survey, Bui. 16: 96. ♀.

Cassandra Kirby. N. Y.

Macrophya cassandra Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 273. ♀.

Macrophya cesta (Say)
U. S.

Allantus cestus Say, 1836. Boston Jour. Nat. Hist. 1: 216. ♀, ♂.

Macrophya contaminata Provancher
Canada.

Macrophya coyitayimiata Provancher, 1878. Nat. Canad. 10: 105. ♀.

Macrophya fuscoterminata Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 410. ♀.

Macrophya epinota (Say)
N. H. to Va. w. to Kans., Colo. Host: Sambucus.

Tenthredo (Allayitus) sambuci Harris, 1833. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 586. Nom. nud.

Allantus epinotus Say, 1836. Boston Jour. Nat. Hist. 1: 215. ♀, ♂.

Taxonomy: Middleton, 1922. U. S. Natl. Mus., Proc. 61: 6 (larva).

Macrophya errans Rohwer
Pa.

Macrophya errans Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 411. ♀.

Macrophya externa (Say)
Mo., Kans. Host: Carya.

Allantus extemus Say, 1823. West. Quart. Rptr., p. 72. ♀.

Macrophya externiformis Rohwer
Wis.

Macrophya externiformis Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 220. ♀.

Macrophya fistula MacGillivray
N. Y.

Macrophya fistula MacGillivray, 1920. Brooklyn Ent. Soc, Bui. 15: 114. ♀.

Macrophya flaccida MacGillivray
N. Y.

Macrophya flaccida MacGillivray, 1920. Brooklyn Ent. Soc, Bui. 15: 113. ♀.

Macrophya flicta MacGillivray
N. Y.

Macrophya flicta MacGillivray, 1920. Brooklyn Ent. Soc, Bui. 15: 114. ♀.

Macrophya fuliginea Norton
111.

Macrophya fuligineus Norton, 1867. Amer. Ent. Soc, Trans. 1: 273. ♀.
Macrophya fill iginosa Dalla Torre, 1894. Cat. Hym., v. 1, p. 52. Misspelling.

Macrophya melanota Rohwer
111.

Macrophya melanota Rohwer, 1912. U. S. Natl. Mus., Proc 43: 219. ♀.

Macrophya minuta MacGillivray
N. Y.

Macrophya minuta MacGillivray, 1895. Canad. Ent. 27: 286. ♂.

Macrophya mixta MacGillivray
N. H. Host: V iburtmyn(^).

Macrophya mixta MacGillivray, 1895. Canad. Ent. 27: 77. ♀.

Macrophya pannosa (Say)
U. S.

Allantus pannosus Say, 1836. Boston Jour. Nat. Hist. 1: 217. ♀, ♂.

Macrophya propinqua Harrington
Canada.

Macrophya propinqua Harrington, 1889. Canad. Ent. 21: 97. ♀.




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Macrophya proximata Norton
Conn.

Macrophya proximata Norton, 1867. Amer. Ent. Soc, Trans. 1: 270. ♀.

Macrophya punctata MacGillivray
N. Y.

Macrophya punctata MacGillivray, 1895. Canad. Ent. 27: 285. ♀.

Macrophya raui Rohwer
Mo.

Macrophya raui Rohwer, 1917. Ent. News 28: 264. ♀, ♂.

Macrophya tibiator Norton
Ont. to Ga. w. to 111., Kans. Host: Sambucus.

Macrophya tibiator Norton, 1864. Ent. Soc. Phila.. Proc. 3: 10. ♀, ♂.

Genus PACHYPROTASIS Hartig

Pachyprotasis Hartig, 1837. Fam. Blattwespen und Holzwespen, nebst Einleitung
Naturgesch. Hym., p. 295.

Type-species: Tenthredo rapae Linnaeus. Desig. by Westwood, 1840.
Lithracia Cameron, 1902. Bombay Nat. Hist. Soc, Jour. 14: 441.
Type-species: Lithracia flavipes Cameron. Monotypic.

This genus is better represented in Eurasia. Only a single holarctic species is found in North
America.

Pachyprotasis rapae (Linnaeus)
Newfoundland to Alaska s. to N. C, Tenn., Mo., N. Mex., Calif.; Europe to
Siberia. Ecology: Widespread. Host: Solidago, Scrophularia, Betonica, Fraxinus,
Antirrhinum. These hosts recorded in Europe.

Tenthredo rapae Linnaeus, 1767. Syst. Nat., Ed. 12, p. 926.

Tenthredo nigrofasciata Eschscholtz, 1822. Entomographien, p. 96. A questionable
synonym.

Macrophya (Pachyprotasis) omega Norton, 1867. Amer. Ent. Soc, Trans. 1: 280. ♀, ♂.

Synairema Americana Provancher, 1885. Canad. Ent. 17: 50. ♀.

Rhogogaster sayi Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 111. ♀.

Macrophya obyiata MacGillivray, 1923. 111. Univ. Bui. 20(50): 22. ♀.

Unplaced Taxa of Tenthredinidae

Tenthredo (Nematus) melanocephala Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.

Mass., p. 583. Nom. nud.
Emphytus platycerus Say, 1836. Boston Jour. Nat. Hist. 1: 220. 6. Ind.



125




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