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CATALOGUE OF THE EPIPHLOEINAE (COLEOPTERA: CLERIDAE)



Weston Opitz
Kansas Wesleyan University
785-827-5541 ext. 2342
opitz@kwu.edu


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ACANTHOCOLLIS OPITZ, 2009:XX.


Type species: Acanthocollis Opitz, 2009: XX.

Generic synonyms: None.

Most recent generic revision: Opitz, W. 2009. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VIII. The genera Acanthocollis, new genus, Stegnoclava, new genus, and Ichnea Laporte. The Coleopterist Bulletin XXXX
Post Corporaal (1950) citations: None.
Distribution: Brazil.
Number of Species: 1.

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Acanthocollis melanura (Klug), 1842: 376. Type locality: Brazil. Primary
type: lectotype ? (ZMHB).
Figure (nontype specimen).
Distribution: Brazil: Mato Grosso, Rio de Janeiro

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AMBOAKIS OPITZ, 2006: 118.


Type species: Teutonia nova Opitz, 1997: 63.

Generic synonyms: None.

Most recent generic revision: Opitz, W. 2006. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part III. The genera Parvochaetus, n. gen., Amboakis, n. gen., and Ellipotoma Spinola. Insecta Mundi 20 (3-4): 97-164.
Post Corporaal (1950) citations: Opitz, 2006: 118; 2008a: 4.
Distribution: Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Cuba, Honduras, M‚xico, Panam , Per£, and Venezuela,
Number of Species: 24.

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Amboakis anapsis Opitz, 2006: 120 locality: Venezuela: Apure: Sarare River. Primary
type: holotype ? (MNHN).
Figure (holotype).
Distribution: Venezuela: Apure.

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Amboakis atra Opitz, 2006: 134 locality: Argentina: Salta: Campo Quijano. Primary
type: holotype ? (IMLA).
Figure (paratype).
Distribution: Argentina: Salta.

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Amboakis barinas Opitz, 2006: 122 locality: Venezuela: Barinas: Rio Socopo. Primary
type: holotype ? (IZAV).
Figure (holotype).
Distribution: Venezuela: Barinas.

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Amboakis binotonis Opitz, 2006: 124 locality: Cuba: Cienfuegos: Soledad. Primary type:
holotype ? (MCZC).
Figure (holotype).
Distribution: Cuba: Cienfuegos.

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Amboakis capitata (Gorham), 1877: 248 locality: Brazil: Amazonas: Ega. Primary
type: lectotype ? (BMNH).
Figure (paralectotype).
Distribution: Brazil: Amazonas: Mato Grosso.

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Amboakis cauca Opitz, 2006: 140 locality: Colombia: Valle del Cauca: PNN Farallones de Cali, Anchcaya, 03ø26'N 76ø48'. Primary type: holotype ? (IAVH).
Figure (holotype).
Distribution: Colombia: Valle del Cauca.

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Amboakis charis Opitz, 2006: 126 locality: Honduras: Olancho: Parque Nacional La
Muralla. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Honduras: Olancho. Panam : Chiriqu¡.

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Amboakis epiomidia Opitz, 2006: 141 locality: Panam : Chiriqu¡: Tol‚. Primary type:
holotype ? (BMNH).
Figure (paratype).
Distribution: Panam : Chiriqu¡.

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Amboakis erythrohapsis Opitz, 2006: 129 locality: M‚xico: Guerrero: Highway 134, 6.9
km NE junction 200. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: M‚xico: Guerrero.

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Amboakis flavicollis (Zayas), 1988: 61 locality: Cuba: Oriente: Cabo Cruz. Primary type:
syntype, gender not known (FDZC).
Figure (nontype specimen)
Distribution: Cuba: Cienfuegos: Granma: Oriente: Las Villas.

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Amboakis funebris Opitz, 2006: 124 locality: Brazil: Amazonas: Reserva Ducke. Primary
type: holotype ? (MZSP).
Figure (holotype).
Distribution: Brazil: Amazonas.

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Amboakis incondita Opitz, 2006: 134 locality: Bolivia: Cochabamba: Cochabamba.
Primary type: holotype ? (MNHN).
Figure (holotype).
Distribution: Bolivia: Cochabamba.

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Amboakis katatonis Opitz, 2006: 130 locality: M‚xico: Michoac n. Uruap n. Primary
type: holotype ? (FSCA).
Figure (paratype).
Distribution: M‚xico: Jalisco: Michoac n.

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Amboakis linitis Opitz, 2006: 132 locality: Dominican Republic: Mout Diego de Ocampo.
Primary type: holotype ? (MCZC).
Figure (holotype).
Distribution: Dominican Republic.

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Amboakis mica Opitz, 2006: 135 locality: Brazil: Par : Jacareacanga. Primary type:
holotype ? (DZUP).
Figure (paratype).
Distribution: Brazil: Par .

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Amboakis micula Opitz, 2006: 135 locality: Brazil: Mato Grosso: Diamantino, Facendo
Rio Arenos. Primary type: holotype ? (AMNH).
Figure (holotype).
Distribution: Per£: Huanuco. Bolivia: Cochabamba: Santa Cruz. Brazil: Mato Grosso.

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Amboakis nitida (Gorham), 1877: 248 locality: Brazil: Amazonas: Ega. Primary type:
lectotype ? (BMNH).
Figure (paralectotype).
Distribution: Brazil: Amazonas.

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Amboakis nova (Opitz), 1997: 63 locality: Brazil: Nova Teutonia: Santa Catharina.
Primary type: holotype ? (MZSP).
Figure (homotype).
Distribution: Brazil: Goias: Paran : Pernambuco: Rio de Janeiro: Nova Teutonia: Sao Paulo.

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Amboakis prolata Opitz, 2006: 136 localiy: Brazil: Paran : Rondon, 24ø38'S 54ø07'W.
Primary type: holotype ? (IZAV).
Figure (holotype).
Distribution: Brazil: Paran .

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Amboakis rudis Opitz, 2006: 142 locality: Bolivia: Cochabamba: Cochabamba. Primary
type: holotype ? (MNHN).
Figure (holotype).
Distribution: Bolivia: Cochabamba.

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Amboakis selva Opitz, 2006: 143 locality: Costa Rica: Heredia: Estaci¢n Biologica La
Selva, 10ø26'N 84ø01'W. Primary type: holotype ? (INBC).
Figure (paratype).
Distribution: Costa Rica: Heredia.

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Amboakis stenosis Opitz, 2006: 145 locality: M‚xico: Oaxaca: Sierra de Zempoaltepetl.,
15.2 km S Benito Juarez Juarez. Primary type: holotype ? (LACM).
Figure (holotype).
Distribution: M‚xico: Oaxaca.

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Amboakis taruma Opitz, 2006: 138 locality: Brazil: Amazonas: 1 km W Taruma Falls.
Primary type: holotype ? (MCNZ).
Figure (holotype).
Distribution: Brazil: Amazonas.

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Amboakis vesca Opitz, 2006: 143 locality: Bolivia: Cochabamba: Cochabamba. Primary
type: holotype ? (MNHN).
Figure (holotype).
Distribution: Bolivia: Cochabamba.


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DECOROSA OPITZ, 2008a: 4.


Type species: Decorosa iviei Opitz, 2008a: 8.

Generic synonyms: None.

Most recent generic revision: Opitz, 2008a. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part V. Decorosa Opitz, a new genus of checkered beetles from Hispaniola with description of its four new species. American Museum Novitates 3628: 1-19.
Post Corporaal (1950) citations: None.

Distribution: Dominican Republic.

Number of species: 4.

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Decorosa aladecoris Opitz, 2008a: 6. Type locality: Dominican Republic: La Vega:
Constanza. Primary type: holotype ? (USNM).
Figure (holotype).

Distribution: Dominican Republic: La Vega,

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Decorosa iviei Opitz, 2008a: 8. Type locality: Dominican Republic: San Juan: Pico
Duarte weather station. Primary type: holotype ? (USNM).
Figure (holotype).

Distribution: Dominican Republic: San Juan, La Vega,

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Decorosa limatula Opitz, 2008a: 8. Type locality: Dominican Republic: La Vega: Loma
Vieja. Primary type: holotype ? (MCZC).
Figure (holotype).

Distribution: Dominican Republic: La Vega,

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Decorosa neiba Opitz, 2008a: 11. Type locality: Dominican Republic: San Juan: Cierra
de Neiba, 9.1 km WSW Hondo Valle. Primary type: holotype ?.
Figure (holotype).

Distribution: Dominican Republic: San Juan.



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DIAPROMECES OPITZ, 1997: 58.


Type species: Diapromeces aclydis Opitz, 1997: 59

Generic synonyms: None.

Most recent generic revision: Opitz, 2007. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part IV. The genera Pyticeroides Kuwert and Diapromeces Opitz. Entomologica Basiliensia et Collectionis Frey 29: 77-166.
Post Corporaal (1950) citations: Opitz, 1997: 58. Opitz, 2007: 138.

Distribution: Brazil.

Number of species: 1.

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Diapromeces aclydis Opitz, 1997: 59. Opitz, 2007: 138. Primary type: holotype ?.

Figure (holotype).

Distribution: Brazil: Santa Catharina: Nova Teutonia.


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ELLIPOTOMA SPINOLA, 1844: 36.


Type species: Ellipotoma tenuiformis Spinola, 1844: 38.
Generic synonyms: None.
Most recent generic revision: Opitz, W. 2006. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part III. The genera Parvochaetus, n. gen., Amboakis, n. gen., and Ellipotoma Spinola. Insecta Mundi 20(3-4): 97-164.
Post Corporaal (1950) citations: Opitz, 1997: 60. Opitz, 2006: 145.
Distribution: Honduras. Costa Rica. Panam . Guyana. Ecuador. Per£. Brazil.
Number of Species: 2.

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Ellipotoma tenuiformis Spinola, 1844: 38. Type locality: Colombia. Primary type: holotype ? (MNHN). Opitz, 2006: 150.
Figure

Distribution: Honduras: Olancho. Costa Rica: Heredia, Puntarenas. Panam :

Chiriqu¡, Col¢n, Panam . Guyana: Mazaruni-Potaro. Ecuador: Tungurahua. Per£: Amazonas, Hu nuco. Brazil: Amazonas, Par , Santa Catarina.

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Ellipotoma turmalis, Opitz, 2006: 152. Type locality: Bolivia: Cochabamba. Primary
type: holotype ? (MNHN).
Figure (holotype).

Distribution: Bolivia: Cochabamba. Argentina: Salta.



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EPIPHLAEUS SPINOLA, 1841: 75.


Type species: Enoplium duodecimmaculatus Klug, 1842: 370.

Generic synonyms: None.

Most recent generic revision: Opitz, W. 2008b. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VI. The genera Epiphlaeus Spinola and Opitzius Barr. Annales Zoologoci (Warszawa) 58(1): 1-34.
Post Corporaal (1950) citations: None.

Distribution: Colombia to Brazil.

Number of species: 7.

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Epiphlaeus adonis Opitz, 2008b: 21 locality: French Guiana: Guyane. Primary type:
holotype ? (MNHN).
Figure (holotype).
Distribution: French Guiana: Guyane..

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Epiphlaeus duodecimmaculatus Klug, 1842: 370 locality: Brazil: Par . Primary type:

holotype ? (ZMHB). Opitz, 200b: 11.
Figure (nontype specimen).
Distribution: Panam : Panam . Colombia: Amazonas, Caquet . Venezuela: Bolivar. Guyana: East Demerara-West Coast Berbice. Per£: Jun¡n. Brazil: Amazonas, Mato Grosso, Par , Rond“nia.

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Epiphlaeus fundurufus Opitz, 2008b: 20 locality: Costa Rica: Heredia: Biological Station
La Selva, 10ø26'N 84ø'01'W. Primary type: holotype ? (INBC).
Figure (paratype).
Distribution: Costa Rica: Alajuela, Guanacaste, Heredia. Panam : Panam . Colombia: Amazonas, Choc¢.

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Epiphlaeus princeps Gorham, 1886: 340 locality: Panam : Chiriqu¡: Bugaba. Primary type: lectotype ? (MNHN). Opitz, 2008b: 20.
Figure (paratype).
Distribution: Panam : Chiriqu¡.

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Epiphlaeus pulcherrimus Gorham, 1877: 246 locality: Brazil: Amazonas: Ega. Primary type: lectotype ? (INBC). Opitz 2008b: 21.
Figure (lectotype).
Distribution: Brazil: Amazonas.

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Epiphlaeus quattuordecimmaculatus Chevrolat, 1876: 27 locality: Brazil. Primary type:
lectotype ? (MNHN). Opitz 2008b: 13.
Figure (homotype).
Distribution: Brazil: Goi s, Guanabara, Santa Catarina. Argentina: Misiones.

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Epiphlaeus tigrinus Opitz, 2008b: 15 locality: Panam : Col¢n: Fort Sherman. Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: Nicaragua: Matagalpa. Costa Rica: Alajuela. Panam : Col¢n: Panam .


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HAPSIDOPTERIS OPITZ, 1997: 62.


Type species: Hapsidopteris diastenus Opitz, 1997: 62.

Generic synonyms: None.

Most recent generic revision: Opitz, W. 2008. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nem‚sio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus, new genus. Zootaxa 1754: 1-40.
Post Corporaal (1950) citations: Opitz, 1997: 62. Opitz, 2008: 5.
Distribution: M‚xico: Veracruz.
Number of species: 1.

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Hapsidopteris diastenus Opitz, 1997: 62. Type locality: M‚xico: Veracruz: Jalapa. Opitz, 2008: 5. Primary type: holotype ? (MNHN). Opitz, 2008c: 6.
Figure (holotype)
Distribution: M‚xico: Veracruz.

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ICHNEA LAPORTE, 1863: 55.


Type species: Ichnea lycoides Laporte, 1836: 55.

Generic synonyms: None.

Most recent generic revision: Opitz, W. 2009. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VIII. The genera Acanthocollis, new genus, Stegnoclava, new genus, and Ichnea Laporte. The Coleopterist Bulletin XXXX
Post Corporaal (1950) citations: Knull, 1951: 303 (Neichnea). Ekis, 1975: 49. Opitz, 1997: 57; 2002: 278.
Distribution: Argentina, Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Honduras, M‚xico, Panam , Paraguay, Per£, United States of America, Venezuela.
Number of species: 19.

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Ichnea acanthomelina Opitz, 2009: XX. Type locality: Brazil: Rond“nia, 62 km SW
Ariquemes, near Fazenda Rancho Grande. Opitz. Primary type: holotype ?.
(MZSP).
Figure (holotype)
Distribution: Brazil: Mato Grosso. Par , Rond“nia.

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Ichnea aequinoctialis Spinola, 1844: 22. Type locality: Panam : Col¢n: Madden forest.
Primary type: neotype ? (MIUP). Ekis, 1975: 49.
Figure (neotype).
Distribution: Honduras: Atl ntida, Cort‚s, Olancho, Yoro. Panam : Boca del
Toro, Chiriqu¡, Col¢n, Panam , Darien. Colombia: Boyac , Choc¢,
Meta, Magdalena, Valle del Cauca,.Venezuela: Amazonas, Aragua,
Bolivar, Tachira,.Falc¢n. Guyana: Bartica. French Guiana: Cayenne.
Ecuador: Napo. Per£: Huanuco, Junin, Loreto. Bolivia: Cochabamba, La Paz, Santa Cruz. Brazil: Alagoas, Amazonas, Bahia, Espiritu Santo, Go s, Par , Rio de Janeiro, Rio grande do Sul, Rond“nia,
Santa Catarina, Tocantins, Paraguay: Caazap , Itap£a. Argentina:
Misiones.

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Ichnea aterrima (Klug). Type locality: M‚xico. Primary type: holotype ?
(ZMHB).
Figure (homotype)
Distribution: United States of America: Arizona. New Mexico. M‚xico: Chiapas,
Jalisco, Morelos, Nayarit, Oaxaca, Tamaulipas, Veracruz.
Guatemala: Baja Verapaz, Chiquimula, El Progresso,
Huehuetenango, Izabal, Sacatep‚quez. Zacapa. Belize: Orange Walk.
Honduras: Atl ntida, Cort‚s, El Paraiso, Francisco Moraz n,
Olancho. Nicaragua: Granada, Matagalpa. Costa Rica: Cartago,
Guanacaste, San Jos‚. Panam : Chiriqu¡, Panam . Colombia: Cauca.
Venezuela: Falc¢n. Ecuador: Carch¡.

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Ichnea atra Opitz, 2009: XX. Type locality: Brazil: Amazonas: 1 km W Taruma Falls
Primary type: holotype ? (MZSP).
Figure (holotype)
Distribution: Guyana: Rupunnuni. Brazil: Amazonas. Suriname: Commewijne.

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Ichnea callanga Opitz, 2009: XX. Type locality: Per£: Callanga. Primary type: holotype ? (ZMHB).
Figure (paratype)
Distribution: Colombia: Huila. Bolivia: La Paz. Per£: Cuzco. Brazil: Mato
Grosso.

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Ichnea digna Wolcott, 1927: 93. Type locality: Costa Rica: Alajuela: San Carlos (=
Ciudad Quesada) Primary type: holotype, gender not known (USNM).
Figure (nontype specimen).
Distribution: Costa Rica: Alajuela, Guanacaste, Heredia, Lim¢n. Panam : Bocas
del Toro, Chiriqu¡, Panam . Colombia: Cauca.

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Ichnea dimidiatepennis Spinola, 1844: 162. Type locality: Colombia.
Primary type: lectotype ? (MIZT).
Figure (notype specimen)
Distribution: Colombia.

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Ichnea divisa (Chevrolat), 1843: 37. Type locality: Brazil.
Primary type: lectotype ? (MNHN).
Figure (nontype specimen).
Distribution: Panam : Col¢n, Panam . Colombia: Magdalena.Venezuela: Bolivar.
French Guiana: Cayenne. Ecuador: Napo. Per£: Loreto. Bolivia:
Santa Cruz. Brazil: Goi s, Mato Grosso, Rond“nia, Santa Catarina.

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Ichnea frenata (Erichson), 1847: 86. Type locality: Per£. .
Primary type: Lectotype ? (ZMHB).
Figure (nontype specimen).
Distribution: Nicaragua: Rio San Juan. Costa Rica: Heredia, Puntarenas. Panam :
Chiriqu¡, Col¢n, Darien, Panam . Colombia: Amazonas, Cauca,
Meta. Venezuela: Amazonas, Bolivar, Tachira. Per£: Hu nuco, Jun¡n,
Loreto, Madre de Dios, San Mart¡n, Tambopata. Ecuador: Esmeraldas, Napo, Sucumbios. .Bolivia: Cochabamba, La Paz, Santa
Cruz. Brazil: Amazonas, Goi s, Mato Grosso, Par .

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Ichnea gregata Opitz, 2009: XX. Type locality: Per£: Ancash: Utcuyacu, Tarma Junin
Primary type: holotype ? (AMNH).
Figure (holotype)
Distribution: Per£: Ancash.

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Ichnea gremia Opitz, 2009: XX. Type locality: Ecuador: Esmeraldas, vicinity Bisly.
Primary type: holotype ? (FSCA).
Figure (FSCA).
Distribution: Ecuador: Esmeraldas.

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Ichnea marginella (Klug), 1842: 376. Type locality: Brazil: Par .
Primary type: holotype ? (ZMHB). Ekis, 1975: 49.
Figure (homotype).
Distribution: M‚xico: Sinaloa, Oaxaca, Veracruz. Guatemala: Baja Verapaz. Belize: cayo. Honduras: Atl ntida, Olancho, Santa B rbara. Costa
Rica: Cartago, Puntarenas. Panam : Chiriqu¡, Col¢n, Panam ,
Darien, Veraguas. Choco, Colombia: Amazonas, Cauca, Magdalena,
Meta, Norte de Sandaner, Putumayo.Venezuela: Amazonas, Aragua,
Bolivar, Monagas, Sucre, Tachira. Trinidad: Arima Borough, Saint
Andrew. Guyana: Masaruni-Potaro. Surinam: Surinam. French
Guiana: Cayenne, Guyane. Ecuador: Napo, Sucumbios. Per£:
Huanuco. Bolivia: Cochabamba, Santa Cruz. Brazil: Amazonas,
Esp¡rito Santo, Paran . Paraguay: Alto Paran .

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Ichnea lycoides laporte, 1836: 55. Type locality: Brazil: Bahia: Encruzilhada
Primary type: Neotype ? (CMNH). Ekis, 1975: 49. Menier, 1985: 1071.
Figure (neotype).
Distribution: Bolivia: La Paz. Brazil: Bahia, Espiritu Santo, Sao Paulo.

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Ichnea mimica Gorham, 1877: 412. Type locality: Brazil: Amazonas: Par : Par .
Primary type: lectotype ? (BMNH).
Figure (homotype).
Distribution: Brazil: Par .

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Ichnea opaca (Klug), 1842: 377. Type locality: French Guiana: Cayenne.
Primary type: lectotype ? (ZMHB).
Figure (homotype).
Distribution: Guyana: Masaruni-Potaro, Rupununi. Bartica. French Guiana:
Cayenne, R‚gina. Brazil: Amap .

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Ichnea plumbea Gorham, 1877: 413. Type locality: Brazil: Amazonas.
Primary type: lectotype ? (BMNH).
Figure (nontype specimen).
Distribution: Panam : Panam . Trinidad: Arima Borpugh. Colombia: Amazonas,
Caquet , Putumayo. Ecuador: Napo. Per£: Loreto. Bolivia: La Paz.
Brazil: Amazonas, Mato Grosso, Par , Rond“nia.

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Ichnea praeusta (Klug), 1842: 376. Type locality: Brazil. Primary type: holotype ?
(ZMHB).
Figure (nontype specimen).
Distribution: Brazil: Amazonas, Mato Grosso, Bahia, Esp¡ritu Santo, Minas
Gerais, Rio Grande do Sul, Santa Catarina. Argentina: Misiones.

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Ichnea procera (Schenkling), 1900: 399. Type locality: Per£: Cuzco: Callanga. Primary
type: lectotype ? (DEIG).
Figure (paralectotype).
Distribution: Per£: Cuzco. Bolivia: Cochabamba, Santa Cruz.

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Ichnea tarumaensis Opitz, 2009: XX. Type locality: Brazil: Amazonas: Manaus, 1 km W
Taruma falls
Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Brazil: Amazonas.

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IONTOCLERUS OPITZ, 1997: 56.

Type species: Enoplium humerale Klug, 1842: 373. Opitz, 1997: 56. Opitz, 2008:

8.
Generic synonyms: None.
Most recent generic revision: Opitz, W. 2008. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nem‚sio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus, new genus. Zootaxa 1754: 1-40.
Post Corporaal (1950) citations: Opitz, 1997: 56. Opitz, 2008: 8.

Distribution: Colombia to Argentina.
Number of species: 2.

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Iontoclerus humeralis (Klug), 1842: 373. Type locality: Brazil: Par . Primary type:
lectotype ? (ZMHB). Opitz, 1997: 56. Opitz, 2008: 10.
Epiphlaeus marginellus Spinola, 1844: 15 (Synonymized by Opitz, 1997: 56).
Epiphlaeus humeralis Spinola, 1844: 15 (Synonymized by Opitz, 1997: 56).
Apolopha vittata Pic, 1936: 127 (Synonymized by Opitz, 1997: 56).
Figure (homotype)
Distribution: Colombia: Putumayo. Guyana: Rupununi. Ecuador: Napo. Brazil:
Amazonas: Mato Grosso: Par : Paran : Rio Grande do Sul:
Rond“nia. Per£: Chambireyac£: Madre de Dios. Bolivia: Santa Cruz. Argentina: Misiones.

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Iontoclerus sericius (Klug), 1842: 373. Type locality: Brazil.. Primary
type: lectotype ? (ZMHB). Opitz, 2008: 12
Figure (homotype).
Distribution: Colombia: Putumayo. French Guiana: Cayenne: Ret De L?Amse.
Brazil: Amazonas: Mato Grosso: Par : Rond“nia. Ecuador: Napo:
Sucumb¡os. Per£: Huallaga: Hu nuco: Madre de Dios. Bolivia: Cochabamba: La Paz.

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KATAMYURUS OPITZ, 1997: 60.

Type species: Katamyurus paxillus Opitz, 1997: 60. Opitz, 2008: 15.
Most recent generic revision: Opitz, W. 2008. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nem‚sio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus, new genus. Zootaxa 1754: 1-40.
Post Corporaal (1950) citations: Opitz, 1997: 60. Opitz, 2008: 15.
Distribution: M‚xico to Nicaragua.
Number of species: 2

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Katamyurus albopaniculus Opitz, 2008: 16. Type locality: M‚xico: Sinaloa: 14 km NE La Cap. De Taxte (= Taste). Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: M‚xico: Durango: M‚xico: Sinaloa.

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Katamyurus paxillus Opitz, 1997: 60. Type locality: Nicaragua: Cerro Cimborazo, 13ø02'N 85ø56'W. Primary type: holotype ? (USNM). Opitz, 2008: 16.
Figure (holotype).
Distribution: Guatemala: Huehuetenango. Honduras: Olancho. Nicaragua:
Matagalpa.

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MADONIELLA PIC, 1935: 10.

Type species: Madoniella minor Pic, 1935: 10.
Most recent generic revision: Opitz, W. 2010. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part X. The genus Madoniella Pic.Bulletin of the American Museum of Natural History XXX
Post Corporaal (1950) citations: Opitz, 1997: 62. Opitz, 2006: 118. Opitz, 2008a: 4.
Distribution: Canada, United States of America, Andros Island, M‚xico,
Guatemala, Cuba, Haiti, Dominican Republic, Jamaica, Puerto Rico,
Dominica, Montserrat, Guana Island, Guadeloupe, Belize,
Honduras, Nicaragua, Panam , Colombia, Venezuela, Guyana,
Costa Rica, Brazil, to Argentina.
Number of species: 74

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Madoniella abacula Opitz, 2010. XX. Type locality: M‚xico: Oaxaca: 14 km NW Diaz
Ordaz. Primary type: Holotype ? (CMNC).
Figure (holotype).
Distribution: M‚xico: Oaxaca.

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Madoniella adona Opitz, 2010. XX. Type locality: Dominican Republic: San Pedro de
Macoris: Mt. Diego de Ocampo. Primary type: holotype ? (MCZC).
Figure (holotype).
Distribution: Dominican Republic: Hato Mayor, San Pedro de Macoris, Santiago.

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Madoniella aktis Opitz, 2010. XX. Type locality: M‚xico: Chiapas: Portugal. 11.2 km SE
Simojovel. Primary type: holotype ? (EMEC).
Figure (paratype).
Distribution: M‚xico: Chiapas.

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Madoniella anapsis Opitz, 2010. XX. Type locality: Dominica: St. Peter: Dublanc.
Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: Dominica: St. Peter, St. Paul. Guadeloupe: Basse Terre.

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Madoniella antennatra Opitz, 2010. Type locality: Honduras: Yoro, Parque Nacional
Pico Pijol. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Honduras: Yoro.

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Madoniella apotoma Opitz, 2010. XX. Type locality: Honduras: Comayagua: 7 km E
Siguatepeque. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Honduras: Comayagua.

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Madoniella apsis Opitz, 2010. XX. Type locality: M‚xico: Chiapas: El Aguacero. Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: M‚xico: Chiapas.

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Madoniella avina Opitz, 2010. XX. Type locality: Argentina: Salta: El Rey National
Park, Rio La Selva. Primary type: holotype ? (CMNC).
Figure (holotype).
Distribution: Argentina: Salta.

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Madoniella basilaris Opitz, 2010. XX. Type locality: Dominican Republic: La Vega, 1
km NW Manaboa. Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: Dominican Republic: La Vega: Independencia: Pedernales.

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Madoniella basilia Opitz, 2010. XX. Type locality: Cuba: Granma: Loma del Gato,
Sierra Maestra. Primary type: holotype ? (INHS).
Figure (holotype).
Distribution: Cuba: Granma.

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Madoniella bilineata (Chevrolat). Type locality: Cuba. Primary type: holotype ?
(GCHC).
Figure (nontype specimen).
Distribution: Cuba: Granma.

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Madoniella bullalis Opitz, 2010. XX. Type locality: Dominican Republic: Monte Cristi,
8.6 km N Villa Elisa. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Dominican Republic: Monte Cristi, Pedernales.

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Madoniella cardinalis Opitz, 2010. Type locality: Costa Rica: Heredia: Estaci¢n
Biologica La Selva. Primary type: holotype ? (INBC).
Figure (holotype).
Distribution: Costa Rica: Heredia: Cartago, Guanacaste, Puntarenas. Panam :
Panam .

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Madoniella careorita Opitz, 2010. Type locality: Costa Rica: Heredia: Estaci¢n
Biologica La Selva. Primary type: holotype ? (INBC).
Figure (holotype).
Distribution: Costa Rica: Alajuela, Heredia: Cartago, Guanacaste.

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Madoniella cavina Opitz, 2010. Type locality: Dominican Republic: Pedernales: 23.5 N
Cabo Rojo, 18ø06'N 71ø30'W. Primary type: holotype ? (CMNH).
Figure (holotype).
Distribution: Dominican Republic: Pedernales.

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Madoniella cerviculina Opitz, 2010. Type locality: Dominican Republic: Pedernales: 3.3
km NE Los Arroyos, 18ø15'N 71ø45'W. Primary type: holotype ? (CMNH).
Figure (holotype).
Distribution: Dominican Republic: Pedernales.

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Madoniella chiricahua Opitz, 2010. Type locality: Arizona: Cochise County, Chiricahua Mountains. Primary type: holotype ? (FMNH).
Figure (paratype).
Distribution: United States of America: Arizona. M‚xico: Chiricahua, Durango,
Sinaloa.

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Madoniella collata Opitz, 2010. Type locality: Brazil: Tocatins: Chapada. Primary type:
holotype ? (CMNH).
Figure (holotype).
Distribution: Brazil: Tocatins: Chapada.


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Madoniella corporaali Pic, 1935: 11. Type locality: Guadeloupe: Guadeloupe. Trois-
Rivers. Primary type: lectoype ? (MNHN).
Figure (lectotype).
Distribution: Guadeloupe: Basse Terre, Guadeloupe. Dominica: St. Peter, St.
Paul.

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Madoniella cracentis Opitz, 2010. Type locality: Brazil: Amazonas: 1 km W Taruma
Falls. Primary type: holotype ? (MCNZ).
Figure (holype).
Distribution: Colombia: Putumayo. Venezuela: Bolivar. Per£: Hu nuco. Brazil:
Amazonas

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Madoniella crinis Opitz, 2010. Type locality: M‚xico: Tlaxcala: Temetzontla. Primary
type: holotype ? (FSCA).
Figure (holotype).
Distribution: M‚xico: Michoac n, Morelos, Tlaxcala.

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Madoniella cymatilis Opitz, 2010. Type locality: Haiti: Nord: Cap du Nord. Primary
type: holotype ? (INHS).
Figure (holype).
Distribution: Haiti: Nord.

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Madoniella dariensis Opitz, 2010. Type locality: Panam : Darien: Pirre, Estaci¢n Rancho
Frio Primary type: holotype ? (MIUP).
Figure (paratype).
Distribution: Panam : Darien.

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Madoniella disjuga Opitz, 2010. XX. Type locality: M‚xico: San Luis Potosi: 40 km W
Xilitla. Primary type: holotype ? (CMNC).
Figure (holotype).
Distribution: M‚xico: San Luis Potosi.

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Madoniella dislocata (Say). Type locality: United States of America:
Primary type: neotype ? (MCZC).
Figure (nontype specimen).
Distribution: Canada: New Brunswick, Nova Scotia, Quebec, Ontario. United
States of America: Alabama, Conecticut, Florida, Georgia, Illinois,
Indiana, Iowa, Kansas, Kentucky, Lousiana, Maine, Maryland,
Massachusetts, Michigan, Mississippi, Missouri, New Hampshire,
New Jersey, New York, North Carolina, Ohio, Oklahoma,
Pennsylvania, Rhode Island, Tennessee, Texas, Virginia, West
Virginia.

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Madoniella displicata Opitz, 2010: XX. Type locality: Venezuela: Aragua: Ocumare:
Primary type: lectotype ? (FSCA).
Figure (holotype).
Distribution: Venezuela: Aragua.

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Madoniella ebena Opitz, 2010: XX. Type locality: Dominican Republic: Hato Major:
Parque Loa Haitises, 3 km W Cueva de arena, 19ø04'N 69ø29'W. Primary type:
lectotype ? (CMNH).
Figure (holotype).
Distribution: Venezuela: Aragua.

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Madoniella emblema Opitz, 2010: XX. Type locality: Costa Rica: Heredia: 11 km SE La
Virgen, 10ø20'N 84ø04'W. Primary type: lectotype ? (INBC).
Figure (holotype).
Distribution: Costa Rica: Heredia.

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Madoniella erythrocephala (Gorham), 1882: 167. Type locality: Panam : Chiriqu¡:
Volc n de Chiriqu¡. Primary type: lectotype ? (INBC).
Figure (homotype).
Distribution: Panam : Chiriqu¡.

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Madoniella extensiva Opitz, 2010. Type locality: Puerto Rico: Maricao: Maricao Forest.
Primary type: holotype ? (MCZC).
Figure (holotype).
Distribution: Dominican Republic: Pedernalis. Puerto Rico: Maricao.

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Madoniella facis Opitz, 2010. Type locality: Guyana: Rupununi: Iwokrama Research
Forest, 6.4 km N of Kurupukari.
Primary type: holotype ? (CASC).
Figure (holotype).
Distribution: Guyana: Rupunui.

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Madoniella fonteboa Opitz, 2010. Type locality: Brazil: Amazonas: Fonteboa.
Primary type: holotype ? (ZMHB).
Figure (holotype).
Distribution: Brazil: Amazonas.

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Madoniella gonia Opitz, 2010. XX. Type locality: M‚xico: Drango: 4.8 km E El Salto.
Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: M‚xico: Durango.

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Madoniella guana Opitz, 2010. XX. Type locality: Guana Island. Primary type:
holotype ? (MCZC).
Figure (holotype).
Distribution: British Virgin Island: Guana Island. Puerto Rico: Gu nica, San Juan.

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Madoniella howdenorum Opitz, 2010. XX. Type locality: M‚xico: Chiapas: 2.6-6 km S
La Trinitaria. Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: M‚xico: Chiapas. Oaxaca.

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Madoniella ignis Opitz, 2010. XX. Type locality: Guatemala: Zacapa: San Lorenzo.
Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Guatemala: Zacapa.

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Madoniella infula Opitz, 2010. XX. Type locality: Dominican Republic: La Vega:
Cordillera Central, 4.1 km SW El Convento. Primary type: holotype, gender not
known (CMNH).
Figure (holotype). Dominican Republic: La Vega.
Distribution: Dominican Republic: La Vega.

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Madoniella insignis Opitz, 2010. XX. Type locality: Brazil: Bahia: Santo Antonio de
Barra. Primary type: holotype ? (ZMHB).
Figure (holotype).
Distribution: Brazil: Bahia.

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Madoniella knullorum Opitz, 2010. XX. Type locality: United States of America: Texas:
Jeff Davis Co., Davis Mountains. Primary type: holotype ? (FMNH).
Figure (paratype).
Distribution: United States of America: Texas.

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Madoniella khlorum Opitz, 2010. Type locality: Nicaragua: Matagalpa. Primary type:
holotype ? (FSCA).
Figure (paratype).
Distribution: M‚xico: Chiapas. Belize: Cayo. Honduras: Francisco Moraz n,
Olancho, Comayagua. Nicaragua: Granada, Matagalpa. Costa Rica:
San Jos‚.

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Madoniella latinopsis Opitz, 2010. XX. Type locality: Venezuela: Aragua: El Lim¢n.
Primary type: holotype ? (IZAV).
Figure (holotype).
Distribution: Venezuela: Aragua.

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Madoniella leona Opitz, 2010. XX. Type locality: M‚xico: Nuevo Leon: 14.4 km west
Iturbide. Primary type: holotype ? (TAMU).
Figure (holotype).
Distribution: M‚xico: Nuevo Leon.

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Madoniella linea Opitz, 2010. XX. Type locality: Jamaica: Portland.: Hardwar Gap.
Primary type:
holotype ? (CNCI).
Figure (paratype).
Distribution: Jamaica: Saint Andrew: Saint Thomas

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Madoniella lineola Opitz, 2010. XX. Type locality: Guatemala: Zacapa: 2 Km S San
Lorenzo. Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: M‚xico: Chiapas: Guatemala: Baja Verapaz El Progresso,
Sacatep‚quez. Zacapa: Honduras: Comayagua.

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Madoniella lurida Opitz, 2010. XX. Type locality: Dominican Republic: Pedernales, La
Abeja, 38 km NNW Cabo Rojo, 18ø09' 71ø38'. Primary type: holotype ?
(CMNH).
Figure (holotype).
Distribution: Dominican Republic: Pedernales.

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Madoniella magdalena Opitz, 2010. Type locality: Colombia: Magdalena: PNN Tayrona
Pueblito, 11ø20'N 74ø02': Primary type: holotype ? (IAVH).
Figure (holotype).
Distribution: Colombia: Magdalena.

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Madoniella maxicornis Opitz, 2010. XX. Type locality: M‚xico: Vera Cruz: Primary
type: holotype ? (USNM).
Figure (holotype).
Distribution: M‚xico: San Luis Potosi, Vera Cruz. Guatemala: Alta Vera Paz.

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Madoniella merga Opitz, 2010. Type locality: M‚xico: Durango: 32 km W El Salto. Primary type: holotype ? (CNCI).
Figure (holotype).
Distribution: M‚xico, Durango, Sinaloa

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Madoniella melina Opitz, 2010. Type locality: Honduras: Yoro: 8 km N La Habana. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Honduras: Yoro.

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Madoniella minor Pic, 1935: 10. Type locality: Guadeloupe. Primary type: lectotype ?
(MNHN). Lepesme, 1947: 169. Corporaal, 1950: 306. Opitz, 1997: 62; 2002: 278.
Figure (nonype specimens).
Distribution: Guadeloupe: Basse-Terre. Montserrat. Dominica: St. Peter.

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Madoniella nana Opitz, 2010. XX. Type locality: United States of America: Texas:
Cameron Co., Sabal Palm Grove.
Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: United States of America: Texas.

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Madoniella nebulosa (Chevrolat) Type locality: Cuba. Primary type: lectotype, gender
unknown (MNHN).
Figure (homotype).
Distribution: Cuba.

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Madoniella orientalis, Zayas, 1988: 60. Type locality: Cuba
Primary type: syntype series not available (FDZC).
Figure (nontype specimen).
Distribution: Cuba: Cienfuego, Granma, Guant namo, Pinar del Rio.

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Madoniella orosiensis Opitz, 2010. XX. Type locality: Costa Rica: Guanacaste: Estaci¢n Maritza, eastern slope of Volc n Orosi. Primary type: holotype ? (INBC).
Figure (homotype).
Distribution: M‚xico: Oaxaca, Chiapas, Comayagua, Quintana Roo, Yucat n.
Honduras: Atlantida, El Para¡so, Francisco Moraz n, Lampira,
Olancho, Santa Barbara. Nicaragua: Granada, Matagalpa. Costa
Rica: Guanacaste, Heredia, Puntarenas, San Jos‚. Panam : Chiriqu¡,
Panam .

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Madoniella patula Opitz, 2010. XX. Type locality: M‚xico: Chiapas: 54 km S Ocosingo.
Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: M‚xico: Chiapas, Oaxaca, Queretaro, Veracruz, Yacat n.
Guatemala: Suchitip‚quez. Belize: Belize, Cayo. Honduras:
Comayagua. Cop n, Cortez, Ocotepeque.

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Madoniella pedalis Opitz, 2010. XX. Type locality: Cuba: La Habana, Cayamas. Primary
type: holotype ? (USNM).
Figure (holotype).
Distribution: Cuba: La Habana.

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Madoniella pellis Opitz, 2010. XX. Type locality: Cuba: Holguin: Sierra de Nipe, 23 km
S Mayari, Pinares de Mayari. Primary type: holotype ? (USNM).
Figure (paratype).
Distribution: Cuba: Holguin, La Habana.

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Madoniella penninsularis Opitz, 2010. XX. Type locality: M‚xico: Oaxaca: Baja
California, Cabo San Lucas. Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: M‚xico: Oaxaca.

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Madoniella pici Lepesme. Type locality: Guadeloupe: Guadeloupe: Trois Rivieres
Primary type: lectotype ? (MNHN).
Figure (lectotype).
Distribution: Guadeloupe: Guadeloupe: Trois Rivieres

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Madoniella pinicola Opitz, 2010. Type locality: United States of America: Arizona:
Cochise Co., Chiricahua
Mountains. Primary type: holotype ? (FMNH).
Figure (paratype).
Distribution: United States of America: Arizona. M‚xico: M‚xico.

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Madoniella plenita Opitz, 2010. Type locality: Costa Rica: Cartago: Turrialba. Primary type: holotype ? (INBC).
Figure (holotype).
Distribution: Costa Rica: Cartago.

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Madoniella pumilis Opitz, 2010. XX. Type locality: Colombia: Magdalena: Pueblito
Primary type: holotype ? (IAVH).
Figure (holotype).
Distribution: Colombia: Magdalena..

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Madoniella punctata Opitz, 2010. XX. Type locality: Guatemala: El Quich‚: Quich‚
Mountains. Primary type: holotype ? (BMNH).
Figure (nontype specimen).
Distribution: M‚xico: Chiapas. Guatemala: El Quich‚.

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Madoniella quintana Opitz, 2010. XX. Type locality: M‚xico: Quintana Roo, 66 km E
Xpujil
Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: M‚xico: Quintana Roo.

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Madoniella redacta Opitz, 2010. Type locality: Costa Rica: Heredia: Estaci¢n
Biologica La Selva, 3 km S Pto. Viejo,10ø26' N 84ø01'W. Primary type: holotype
? (INBC).
Figure (holotype).
Distribution: Costa Rica: Cartago.

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Madoniella rectangularis Opitz, 2010. Type locality: United States of America: Texas:
Hidalgo Co.Primary type: holotype ? (FMNH).
Figure (paratype).
Distribution: United States of America: Texas. M‚xico: Coahila, Nuevo Leon,
Tamaulipas.

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Madoniella rubidia Opitz, 2010. Type locality: Brazil: Bahia. Primary type: holotype
? (MNHN).
Figure (holotype).
Distribution: Brazil: Bahia.

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Madoniella storea Opitz, 2010. XX. Type locality: Jamaica: Portland: Hardwar Gap. Primary type: holotype ? (CNCI).
Figure (paratype).
Distribution: Jamaica: Portland, Saint Andrew, Saint Thomas; Trelawney, Saint
Catherine.

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Madoniella thomasi Opitz, 2010. Type locality: Bahamas: Andros Island, Maidenhair
Coppica. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Bahamas: Andros Island.

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Madoniella tegetis Opitz, 2010. Type locality: Honduras: Cop n: 6 km NW San Agustin.
Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Honduras: Cop n.

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Madoniella texis Opitz, 2010. Type locality: Costa Rica: Cartago: Turrialba. Primary type: holotype ? (INBC).
Figure (paraype).
Distribution: Costa Rica: Alajuela, Cartago, Cocl‚, Guanacaste, Heredia, Puntarenas. Panam : Chiriqu¡, Panam .

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Madoniella vogti Opitz, 2010. Type locality: United States of America: Texas: South-east
Hidalgo Co.Primary type: holotype ? (FMNH).
Figure (holotype).
Distribution: United States of America: Texas. M‚xico: San Luis Potosi,
Tamaulipas.

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Madoniella welderi Opitz, 2010. Type locality: United States of America: Texas: San
Patricio Co., Welder Wildlife Refuge.
Primary type: holotype ? (FMNH).
Figure (holotype).
Distribution: United States of America: Texas.

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Madoniella zonula Opitz, 2010. Type locality: Panam : Col¢n. Primary type: holotype ?
(MIUP).
Figure (holype).
Distribution: Panam : Col¢n.


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MEGAPHLOEUS OPITZ, 2009: XX

Type species: Megaphloeus setulosus Thomson, 1860: 60.
Most recent generic revision: Opitz, W. 2009. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VIII. The genus Megaphloeus, new genus. Bulletin of the American Museum of Natural History XXX
Post Corporaal (1950) citations: Mawdsley, 1992: 129 (Epiphloeus). Opitz, 1997: 53 (Epiphloeus). Ekis, 1975: 47. Opitz, 2008b: 2.
Distribution: M‚xico to Argentina.
Number of species: 27.

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Megaphloeus absentis, Opitz, 2009: XX. Type locality: BelizeM‚xico: Puebla: 24 km E Teziutlan. Primary type: holotype ? (CMNC).
Figure (paratype).
Distribution: M‚xico: Puebla: Veracruz.

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Megaphloeus animosus (Wolcott), 1927: 92. Type locality: Brazil: Costa Rica: Cartago Primary type: holotype (FMNH).
Figure (nontype specimen).
Distribution: Ecuador: M‚xico: Veracruz. Honduras: Atl ntida. Nicaragua: Zelaya. Costa Rica: Cartago, Guanacaste, Heredia, Lim¢n, Puntarenas. Panam : Bocas del Toro, Chiriqu¡, Col¢n, Darien, Veraguas. Colombia: Amazonas, Choc¢, Caquet . Venezuela: Tachira. Ecuador: Napo. Per£: Jun¡n..Brazil: Rond“nia.

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Megaphloeus bulatus Opitz, 2009: XX. Type locality: Costa Rica: Puntarenas, Estaci¢n
Agujas. Primary type: holotype ? (INBC).
Figure (paratype).
Distribution: Nicaragua: Chontales. Costa Rica: Alajuela. Heredia, Lim¢n, Puntarenas. Panam : Col¢n, Panam .

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Megaphloeus cartus, Opitz, 2009: XX. Type locality: Belize: Orange Walk, Rio Bravo,
La Milpa. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Belize: Orange Walk.

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Megaphloeus circinus, Opitz, 2009: XX. Type locality: Colombia: Vaup‚s, R. N. Mosiro- Itajura (capar£), Centro Abiental, 01ø04'S 69ø31'W. Primary type: holotype ? (IAVH).
Figure (paratype).
Distribution: Colombia: Amazonas, Vaup‚s.

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Megaphloeus fucoaquilus Opitz, 2009: XX. Type locality: Brazil: Rond“nia: 62 km SE
Ariquemes. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Brazil: Rond“nia.

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Megaphloeus lividipes (Chevrolat), 1874: 320. Type locality: Venezuela. Primary type:
lectotype ? (CMNC).
Figure (homotype).
Distribution: Venezuela: Aragua, Carabobo..

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Megaphloeus longius, Opitz, 2009: XX. Type locality: Honduras: Olancho: Parque Nacional La Muralla. Primary type: holotype ? (CMNC).
Figure (paratype).
Distribution: Honduras: Cort‚s, Olancho.

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Megaphloeus marginipes (Chevrolat), 1874: 320. Type locality: M‚xico: Veracruz. Primary type: lectotype ? (MNHN).
Figure (nontype specimen).
Distribution: M‚xico: Veracruz. Belize: Cayo, Orange Walk, Toledo. Guatemala:
Izabal. Honduras: Atl ntida. Olancho. Costa Rica: Alajuela,
Guanacaste, Heredia, Puntarena. Pan ma: Chiriqu¡, Col¢n, Panam .

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Megaphloeus megasensibilis Opitz, 2009: XX. Type locality: Brazil: Rond“nia: 62 km
SE Ariquemes. Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: Brazil: Mato Grosso, Rond“nia.

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Megaphloeus mucoreus (Klug), 1842: 371. Type locality: Brazil. Primary type: holotype ? (FSCA). Opitz, 1997: 68.
Figure (homotype).
Distribution: Colombia: Amazonas, Magdalena. Venezuela: Bol¡var. Guyana:
Grosso do Sul, Potaro. Ecuador: Napo, Santiago Morona. Per£: Huanuco, Madre de Dios, San Martin.Brazil: Acre, Alag“as, Amazonas, Bahia, Sear , Esp¡rito Santo, Goi , Guanabara, Sao Paulo, Par , Mato Grosso, Mato Grosso, Paran , Rio Grande do Sul, Rond“nia, Santa Catarina. Paraguay: Itapua, San Pedro. Argentina: Entre R¡os. Misiones.

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Megaphloeus nubilus (Klug). Type locality: Brazil.
Primary type: holotype ? (FSCA).
Figure (homotype).
Distribution: French Guiana: Cayenne. Brazil: Paran , Esp¡tu Santo.

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Megaphloeus parvulus (Schenkling). Type locality: Brazil: Goi s
Primary type: lectotype ? (DEIG).
Figure (holotype).
Distribution: Brazil: Bahia, Goi s, Mato Grosso, Mato Grosso do Sul, Paran , Rio
de Janeiro, Rond“nia, Santa Catarina, Sao Paulo.

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Megaphloeus pectilus Opitz, 2009: XX. Type locality: Guyana: Demerara: 1 km W
Kurupukari, Iwokrama Field Station. Primary type: holotype ? (SEMC).
Figure (paratype).
Distribution: Guyana: Demerara. Brazil: Par .

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Megaphloeus platyglenus Opitz, 2009: XX. Type locality: Ecuador Orellana: Ethnica,
Waorani, I km S Onko, Gare camp. Primary type: holotype ? (USNM).
Figure (holotype).
Distribution: Ecuador: Orellana.

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Megaphloeus rictocaliginus Opitz, 2009: XX. Type locality: Brazil: Rond“nia, 62 km SE
Ariquemes. Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: Costa Rica: Cartago, Guanacaste. Panam : Col¢n. Venezuela:
Bolivar, Barinas. Guyana: Essequibo. French Guiana. Bolivia:
Chapar‚, Brazil: Par , Rond“nia..

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Megaphloeus setulosus (Thomson), 1860: 60. Type locality: M‚xico. Primary type:
lectotype ? (MNHN).
Figure (lectotype).
Distribution: M‚xico: Chiapas, Guerrero, Jalisco, Nayarit, Oaxaca, Quintana Roo,
Sinaloa, Tabasco, Veracruz, Yucat n. Guatemala: Alta Verapaz,
Baja Verapaz, Quetzaltenango. Belize: Belize. Honduras: Atlantida,
El Paraiso, Olancho. El Salvador: Moraz n. Nicaragua: Chontales. Costa Rica: Alajuela, Cartago, Guanacaste, Heredia, Lim¢n, San Jos‚, Puntarenas. Panam : Chiriqu¡, Col¢n, Dari‚n, Panam , Colombia: Cundinamarca, Magdalena,, Putumayo. Ecuador: Napo, Tachira. Bolivia: Cavinas, Santa Cruz. Per£: Cuzco. Brazil: Mato
Grosso, Rond“nia.

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Megaphloeus sexplagiatus (Kuwert), 1893: 494. Type locality: Per£: Amazones. Primary type: holotype, gender not known (MNHN).
Figure (homotype).
Distribution: Belize: Stann Creek. Panam : Colon, Darien. Colombia: Magdalena.
Venezuela: Bolivar, Tachira. Guyana: Demerara Ecuador: Napo. Per£: Hu nuco, San Mart¡n, Madre de Dios. Brazil: Amazonas, Par , Rond“nia, Tocatins.

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Megaphloeus terzonatus (Gorham), 1877: 242. Type locality: Brazil: Amazon: Tef‚. Primary type: lectotype ? (BMNH).
Figure (homotype).
Distribution: Ecuador: Esmeraldas, Pichincha. Per£: Jun¡n, Madre de Dios. Bolivia: Cochabamba. Brazil: Amazonas, Chapare, Mato Grosso, Par , Rond“nia.

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Megaphloeus tigrinus Opitz, 2009: XX. Type locality: Colombia: Putumayo, PNN La
Paya, Finca Charapa, 0ø8''S 74ø57øW. Primary type: holtotype ? (IAVH).
Figure (paratype).
Distribution: Colombia: Putumayo, Caquet , Magdalena.

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Megaphloeus tricolor (Kuwert), 1893: 493. Type locality: Brazil: Amazonas. Primary
type: lectotype, gender not known (MNHN).
Figure (lectotype).
Distribution: Costa Rica: Cartago, Heredia, Puntarenas. Panam : Col¢n, Darien,
Panam , Colombia: Amazonas, Caquet , Putumayo, Vaup‚s.
Venezuela: Aragua. Ecuador: Zemora-Chinchipe, Napo. Bolivia:
Beni, Cochabamba, La Paz, Santa Cruz. Brazil: Amazonas, Mato
Grosso, Rond“nia.

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Megaphloeus ustafinis Opitz, 2009: XX. Type locality: Brazil: Mato Grosso. Primary type: holootype ? (FSCA).
Figure (paratype).
Distribution: Brazil: Amazonas, Mato Grosso, Rond“nia.

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Megaphloeus ustus Opitz, 2009: XX. Type locality: Brazil: Santa Catarina: Nota Teutonia, 27ø11'S52ø23'W. Primary type: holotype ? (FSCA).
Figure (paratype).
Distribution: Brazil: Santa Catarina.

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Megaphloeus variegatus (Klug), 1842: 371. Type locality: Brazil: Par . Primary type:
lectotype ? (ZMHB).
Figure (homotype).
Distribution: Colombia: Amazonas, Magdalena, Meta, Nari¤o, Putumayo.
Venezuela: Delta Amacura. Guyana: Demerara. Ecuador: Napo.
Per£: Huanuco, Lareto, San Mart¡n, Santipo. Brazil: Amazonas, Mato Grosso, Par , Paranhao.

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Megaphloeus velutinus (Kuwert), 1893: 493. Type locality: Brazil: Amazonas. Primary
type: lectotype, ? (BMNH).
Figure (homotype).
Distribution: Panam : Col¢n, Darien, Panam . Colombia: Amazonas, Caquet , Putumayo. Suriname: Saramacca. Per£: Lareto. Bolivia: Chapar‚.
Brazil: Amazonas, Mato Grosso, Par , Rond“nia.

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Megaphloeus vitellinus Opitz, 2009: XX. Type locality: Brazil: Bahia: Encruzilhada.
Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Brazil: Bahia.

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MEGATRACHYS OPITZ, 1997: 61.

Type species: Megatrachys paniculus Opitz, 1997: 61.
Most recent generic revision: Opitz, W. 2008. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nem‚sio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus, new genus. Zootaxa 1754: 1-40.
Post Corporaal (1950) citations: Opitz, 1997: 61. Opitz, 2008: 21.
Distribution: M‚xico to Guatemala.
Number of species: 3.

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Megatrachys bibara Opitz, 2008c: 21. Type locality: Guatemala: Zacapa: 2 km San
Lorenzo. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Guatemala: Zacapa.

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Megatrachys paniculus Opitz, 1997: 61. Type locality: M‚xico: Chiapas: 8 km W San Crist¢bal (= San Cristobal de Las Casas). Primary type: holotype ?.
(CNCI). Opitz, 2008: 23.
Figure (holotype).
Distribution: M‚xico: Chiapas.

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Megatrachys truncatia Opitz, 2008c: 23. Type locality: M‚xico: Chiapas: 47.5 km NW Comit n. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: M‚xico: Chiapas.


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OPITZIA NEMSIO, 2006: 77.


Type species: Arenaria chiapas Opitz, 1997: 57.

Generic synonyms: None.

Most recent generic revision: Opitz, W. 2008. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nem‚sio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus, new genus. Zootaxa 1754: 1-40.
Post Corporaal (1950) citations: Opitz, 1997: 61(Arenaria). Nem‚sio, 2006: 77 (Opitzia). Opitz, 2008: 24 (Opitzia).
Distribution: M‚xico and Bolivia.
Number of Species: 2.

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Opitzia apicula Opitz, 2008c: 25. Type locality: Bolivia: Santa Cruz: Amboro road,

above Achira Campo. Primary type: holotype ? (UASC).

Figure: (holotype).

Distribution: Bolivia: Santa Cruz.

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Opitzia chiapas (Opitz), 1997: 57. Type locality: M‚xico: Chiapas: 39 km NW Comit n.
Primary type: holotype ? (MCMC). Nem‚sio, 2006: 77 (Opitzia). Opitz, 2008: 25 (Opitzia).
Figure: (holotype).

Distribution: M‚xico: Chiapas. Guatemala: El Progresso: Zacapa.



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OPITZIUS BARR, 2006: 215.


Type species: Opitzius thoracicus Barr, 2006: 215.

Generic synonyms: None.
Most recent generic revision: Opitz, W. 2008. Classification, natural history, and
evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VI. The genera Epiphlaeus
Spinola and Opitzius Barr. Annales Zoologoci (Warszawa) 58(1): 1-34.

Post Corporaal (1950) citations: Opitz, 2008: 29.
Distribution: Brazil.
Number of Species: 1

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Opitzius thoracicus Barr, 2006: 215. Type locality: Brazil: Bahia: Encruzilhada. Primary type: holotype ? (MZSP).
Figure (holotype)

Distribution. Brazil: Bahia: Esp¡rito Santo: Guanabara: Paran : Rio de Janeiro.



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PARVOCHAETUS OPITZ, 2006: 109.

Type species: Parvochaetus fucolatus Opitz, 2006: 113.
Generic synonyms: None.

Most recent generic revision: Opitz, W. 2006. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part III. The genera Parvochaetus, n. gen., Amboakis, n. gen., and Ellipotoma Spinola. Insecta Mundi 20(3-4): 97-164.
Post Corporaal (1950) citations: None.
Distribution: Panam  to Brazil.
Number of Species: 5.

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Parvochaetus albicornis Opitz, 2006: 112. Type locality: Brazil: Mato Grosso: Sinop,

Coordenadas. Primary type: holotype ? (MCNZ).
Figure (holotype).

Distribution: Brazil: Mato Grosso.

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Parvochaetus froeschneri Opitz, 2006: 112. Type locality: Brazil: Rio de Janeiro:

Rio Grande (= Riograndina). Primary type: holotype ? (DZUP).
Figure (holotype).

Distribution: Brazil: Mato Grosso: Rio de Janeiro..

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Parvochaetus fucolatus Opitz, 2006: 113. Type locality: Brazil: Rio de Janeiro:

Rio Grande (= Riograndina). Primary type: holotype ? (IZAV).

Figure (holotype).

Distribution: Brazil: Rio de Janeiro

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Parvochaetus linearis (Gorham), 2006: 116. Type locality: Panam : Bugaba. Primary

type: holotype ? (BMNH).

Figure

Distribution: Panam : Bugaba.

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Parvochaetus sandaracus Opitz, 2006: 117. Type locality: Brazil: Nova Teutonia: Santa

Catarina, 27ø11'S 52ø23'W. Primary type: holotype ? (AMNH).

Figure (paratype).

Distribution: Brazil: Nova Teutonia: Paran .



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PENNASOLIS OPITZ, 2008: 27.


Type species: Phyllobaenus merkeli Horn, 1896: 374.

Most recent generic revision: Opitz, W. 2008. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nem‚sio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus, new genus. Zootaxa 1754: 1-40.
Post Corporaal (1950) citations: Opitz, 2008: 27.
Distribution: United States of America.
Number of species: 2.

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Pennasolis californica (Van Dyke), 1923: 47 (Phyllobaenus). Type locality: Arizona. Primary type: holotype ? (CASC). Opitz, 2008c: 29 (Pennasolis).
Figure (nontype specimen).
Distribution: United States of America: Arizona: California.

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Pennasolis merkeli (Horn), 1896: 374 (Phyllobaenus). Type locality: California: Yosemite Valley. Primary type: lectotype ? (CASC).Opitz, 2008c: 29 (Pennasolis).
Figure (nontype specimen).
Distribution: United States of America: Arizona: California: Colorado: New Mexico: Texas: Utah.

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PERICALES OPITZ, 2008: 32.

Type species: Pericales albogilvus Opitz, 2008: 32.
Most recent generic revision: Opitz, W. 2008. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nem‚sio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus, new genus. Zootaxa 1754: 1-40.
Post Corporaal (1950) citations: Opitz, 2008: 32.
Distribution: Haiti and the Dominican Republic.
Number of species: 1.

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Pericales albogilvus Opitz, 2008: 32. Type locality: Haiti: Sud-Oueste (=Sud-Est), Massif de La Selle, Morne d? Enfer. Primary type: holotype ? (FSCA).
Figure
Distribution: Haiti: Sud-Est. Dominican Republic: Pedernales

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PLOCAMOCERA SPINOLA, 1844a: 17.


Type species: Plocamocera sericella Spinola, 1844a: 19.

Generic synonyms: None.

Most recent generic revision: Opitz, 2004. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part II. The genera Chaetophloeus Opitz and Plocamocera Spinola. Bulletin of the American Museum of Natural History 280: 1-82.
Post Corporaal (1950) citations: Opitz, 2004: 24. Mecke, et al, 2001: 119
[Plocamocera sp. should read Silverasia hispida (Opitz)].

Distribution: M‚xico to Brazil.

Number of species: 35.

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Plocamocera aliguantula Opitz, 2004: 38 locality: Costa Rica: Lim¢n: Revantaz¢n, Hamburg

Farm. Primary type: holotype ? (USNM).

Figure (paratype)

Distribution: Costa Rica: Lim¢n.

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Plocamocera ambra Opitz, 2004: 60 locality: Costa Rica: Puntarenas: Estaci¢n Agujas. Primary type: holotype ? (INBC).
Figure (paratype)

Distribution: Costa Rica: Alajuela; Puntarenas.

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Plocamocera argentea Opitz, 2004: 55 locality: Bolivia: Santa Cruz: Provincia del Sara.
Primary type: holotype ? (CMNH).
Figure (paratype).

Distribution: Bolivia: Santa Cruz.

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Plocamocera aspera Opitz, 2004: 52 locality: Brazil: Amazonas: Paraque, 30 km E of

Manaus. Primary type: holotype ? (MZSP).

Figure. (holotype)

Distribution: Brazil: Amazonas.

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Plocamocera aura Opitz, 2004: 46 locality: Bolivia: La Paz: Tumupasa. Primary type:
holotype ? (USNM).
Figure (holotype).

Distribution: Bolivia: La Paz.

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Plocamocera auratilis Opitz, 2004: 55 locality: Panam : Panam : Pan American Highway,

30 km E of Canita. Primary type: holotype ? (AMNH).

Figure (paratype).

Distribution: Costa Rica: Alajuela. Panam .

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Plocamocera baria Opitz, 2004: 46 locality: Venezuela: Amazonas: Rio Negro, Rio Baria,

0ø55'N 66ø10'W. Primary type: holotype ? (IZAV).

Figure (holotype).

Distribution: Venezuela: Amazonas.

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Plocamocera bispina Opitz, 2004: 53 locality: Brazil: Mato Grosso: Rio Caraguata, 21ø48'S

52ø27'W. Primary type: holotype ? (FMNH).

Figure (holotype).

Distribution: Brazil: Mato Grosso.

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Plocamocera bolivari Opitz, 2004: 59 locality: Venezuela: Trujillo: Betijoque, 09ø19'N

70ø24'W. Primary type: holotype ? (IZAV).

Figure (holotype).

Distribution: Venezuela: Trujillo.

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Plocamocera buenavista Opitz, 2004: 46 locality: Bolivia: Santa Cruz: 3.7 km SSE

Buenavista, Hotel Flora & Fauna. Primary type: holotype ? (FSCA).

Figure (holotype).

Distribution: Bolivia: Santa Cruz.

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Plocamocera carnegei Opitz, 2004: 54 locality: Brazil: Guanabara: Rio de Janeiro. Primary
type: holotype ? (CMNH).
Figure (holotype).
Distribution: Brazil: Guanabara.

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Plocamocera castanea Opitz, 2004: 42 locality: Brazil: Go¡as: Jatai Go¡as. Primary type: holotype ? (MZSP).
Figure (paratype).

Distribution: French Guiana: Cayenne. Ecuador: Napo. Bolivia:

Cochabamba. Brazil: Go¡as.

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Plocamocera coactilis Opitz, 2004: 51 locality: Brazil: Mato Grosso: Mato Grosso. Primary
type: holotype ? (MZSP).
Figure (paratype).

Distribution: Brazil: Amazonas; Goias; Mato Grosso; Par .

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Plocamocera confrater Kuwert, 1893: 496 locality: Per£: Amazon. Primary type: lectotype ? (MNHN).
P. impressicollis Pic, 1942: 3 (synonymized by Opitz, 2004: 48).

Figure (nontype specimen)

Distribution: Guyana: Bartica. French Guiana: Cayenne. Ecuador: Napo. Per£:

Madre de Dios. Bolivia: Beni. Brazil: Amazonas; Goi s, Mato

Grosso; Par ; Rond“nia.

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Plocamocera insula Opitz, 2004: 60 locality: Trinidad: Saint George Parsh: Arima. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Trinidad.

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Plocamocera iota Opitz, 2004: 38 locality: Brazil: Mato Grosso: Rio Caraguata, 21ø48'S

52ø27'W. Primary type: holotype ? (FMNH).

Figure (holotype).

Distribution: Brazil: Mato Grosso.

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Plocamocera jayhawkalis Opitz, 2004: 45 locality: Suriname: Saramacca: 175 km WSW

Zanderij Airport, 04ø59'06?N 56ø18'48?W. Primary type: holotype, gender not known (SEMC).
Figure (holotype)

Distribution: Suriname: Saramaca.

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Plocamocera lucis Opitz, 2004: 40 locality: Brazil: Coordenadas: Mato Grosso: Sinop. Primary type: holotype ? (MZSP).
Figure (paratype).

Distribution: Brazil: Mato Grosso.

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Plocamocera manausensis Opitz, 2004: 33 locality: Brazil; Amazonas: Manaus: 1 km W

Taruma Falls. Primary type: holotype ? (MZSP).

Figure (paratype).

Distribution: Brazil: Amazonas.

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Plocamocera minima Opitz, 2004: 38 locality: Panam : Panam : Barro Colorado Island,

08ø10'N 79ø50'W. Primary type: holotype ? (AMNH).

Figure (paratype).

Distribution: Panam : Barro Colorado Island; Canal Zone.

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Plocamocera onorei Opitz, 2004: 53 locality: Ecuador: Napo: Yasuni Research Station. Primary type: holotype ? (QCAZ).
Figure (holotype).

Distribution: Ecuador: Napo.

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Plocamocera paris Opitz, 2004: 52 locality: Brazil: Distrito Federal: Parque Nacional. Primary type: holotype ? (CNCI).
Figure (holotype).

Distribution: Brazil: Distrito Federal.

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Plocamocera procera Opitz, 2004: 51 locality: Colombia: Amazonas: P.N.N. Amacayacu,

03'23'S 70'06'W. Primary type: holotype ? (LACM).

Figure (holotype).

Distribution: Colombia: Amazonas.

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Plocamocera prolixa Opitz, 2004: 39 locality: Alajuela: 20 km S Upsula. Primary type: holotype ? (INBC).
Figure (holotype).

Distribution: Costa Rica: Alajuela.

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Plocamocera pupula Opitz, 2004: 42 locality: Brazil: Mato Grosso: Sinop. Primary type:
holotype ? (MZSP).
Figure (paratype).

Distribution: Brazil: Mato Grosso.

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Plocamocera quadrula Opitz, 2004: 44 locality: Brazil: Mato Grosso: Sinop. Primary type:
holotype ? (AMNH).
Figure (holotype).

Distribution: Brazil: Mato Grosso.

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Plocamocera salasis Opitz, 2004: 33 locality: Brazil; Amazonas: Manaus: 1 km W

Taruma Falls. Primary type: holotype ? (MZSP).

Figure (holotype)

Distribution: Brazil: Amazonas.

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Plocamocera santa Opitz, 2004: 54 locality: Brazil: Santa Catarina: Nova Teutonia. Primary type: holotype ? (IZAN).
Figure (holotype)

Distribution: Brazil: Santa Catharina.

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Plocamocera selva Opitz, 2004: 61 locality: Bolivia: Santa Cruz: 5 km ESE Warnes, Hotel

Rio Selva, Primary type: holotype ? (FSCA).

Figure (holotype)

Distribution: Bolivia: Santa Cruz.

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Plocamocera sericella Spinola, 1844: 19 locality: Panam : Panam : Madden Forest. Primary
type: neotype ? (AMNH).
Epiphloeus byssinus Erichson, 1847: 86.

Plocamocera latefasciata Pic, 1942: 3.
Figure (nontype specimen).
Distribution: M‚xico: Campeche; Chiapas; Nayarit; Oaxaca; Quintana Roo;
Tamaulipas; Veracruz. Belize: Belize. Honduras: Comayagua. Costa Rica: Alajuela; Guanacaste; Lim¢n. Panam : Barro Colorado Island; Canal Zone; C¢lon. Colombia: Cundilamarca; Magdalena. Venezuela: Aragua; Bolivar. French Guiana: Nouvea Chantier. Ecuador: Napo; Pichincha. Per£: Loretto; Sam Martin. Bolivia:
Cochabamba; Santa Cruz. Brazil: Amazonas; Goi s; Mato Grosso; Rond“nia.

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Plocamocera sericellopsis Opitz, 2004: 59 locality: Brazil: Rond“nia: 62 km SE Ariquenes.
Primary type: holotype ? (MZSP).
Figure (holotype).

Distribution: Brazil: Rond“nia.

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Plocamocera sesquipedalis Opitz, 2004: 40 locality: Guyana: Demerara: Ikuribisi. Primary type: holotype ? (BMNH).
Figure (paratype).

Distribution: Guyana: Bartica; Demerara. Per£: Loreto. Brazil

Rond“nia.

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Plocamocera similis Opitz. 2004: 41 locality: Ecuador: Pichincha: 18 km S Tinalandia. Primary type: holotype ? (QCAZ).
Figure (holotype).

Distribution: Ecuador: Pichincha.

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Plocamocera specula Klug, 1842: 372 locality: Brazil. Primary type: holotype ? (ZMHB).

Figure

Distribution: Brazil.

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Plocamocera taruma Opitz, 2004: 47 locality: Brazil: Amazonas: Manaus: 1 km W

Taruma Falls. Primary type: holotype ? (MZSP).

Figure (holotype).

Distribution: Brazil: Amazonas.


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PTEROFERUS OPITZ, 2008c: 33.

Type species: Pteroferus zolnerowichi Opitz, 2008c: 33.
Most recent generic revision: Opitz, W. 2008. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nem‚sio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus, new genus. Zootaxa 1754: 1-40.
Post Corporaal (1950) citations: Opitz, 2008: 33.
Distribution: Brazil.
Number of species: 1.

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Pteroferus zolnerowichi Opitz, 2008c: 33. Type locality: Brazil: Santa Catarina: Nova Teutonia. Primary type: holotype ? (FSCA).
Figure (holotype).
Distribution: Brazil: Santa Catarina.


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PYTICEROIDES KUWERT, 1894: 7, 9.


Type species: Pyticeroides arrogans Kuwert, 1894: 7.

Generic synonyms: Neichnea Wolcott & Chapin, 1918: 103.

Most recent generic revision: Opitz, 2007. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part IV. The genera Pyticeroides Kuwert and Diapromeces Opitz. Entomologica Basiliensia et Collectionis Frey 29: 77-166.
Post Corporaal (1950) citations: Opitz, 1997: 59.
Distribution: Canada to Argentina.
Number of species: 32

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Pyticeroides acaris Opitz, 2007: 113 locality: Colombia: Magdalena: PNN Tyrona Palangana, 11ø20'N 74ø02'W. Primary type: holotype ? (IAVH).
Figure: (holotype).
Distribution: Colombia: Magdalena.

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Pyticeroides arrogans Kuwert. 1894: 7 locality: Per£: Amozonas. Primary type: lectotype ? (MNHN).
Figure (holotype).
Distribution: Per£: Amazonas.

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Pyticeroides caecoatus Opitz. 2007: 135 locality: M‚xico: Chiapas: 2.6-6 km S La Trinitaria.
Primary type: holotype ? (AMNH).
Figure (holotype).
Distribution: M‚xico: Chiapas.

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Pyticeroides caligneus Opitz, 2007: 115 locality: Brazil: Paran : Rondon. Primary type:
holotype ? (FMNH).
Figure (paratype).
Distribution: Brazil: Paran .

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Pyticeroides chiriquianum (Gorham), 1883: 182 locality: Panam : Chiriqu¡: Volcan de Chiriqu¡. Primary type: lectotype ? (BMNH).
Figure (homotype).
Distribution: Costa Rica: Guanacate: Puntarenas. Panam : Chiriqu¡: Col¢n: Panam . Colombia: Bolivar: Magdalena. Ecuador: Guayas.

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Pyticeroides cinericius Opitz, 2007: 119 locality: Brazil: Mato Grosso: Rio Caraguata, 21ø48'S 52ø27'W. Primary type: holotype ? (FMNH).
Figure (paratype).
Distribution: Brazil: Mato Grosso.

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Pyticeroides decurialis Opitz, 2007: 111 locality: Brazil: Esp¡rito Santo, Linhares. Primary
type: holotype ? (MCNZ).
Figure (holotype).
Distribution: Brazil: Esp¡rito Santo.

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Pyticeroides enormis Opitz, 2007: 105 locality: Bolivia: Cochabamba. Primary type:
holotype ? (MNHN).
Figure (holotype).
Distribution: Bolivia: Cochabamba.

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Pyticeroides eurides Opitz, 2007: 116 locality: Brazil: Mato Grosso: Diamantino, Fazenda
Rio Arinos. Primary type: holotype ? (MCNZ).
Figure (holotype).
Distribution: Brazil: Mato Grosso.

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Pyticeroides fustis Opitz, 2007: 125 locality: Brazil: Mato Grosso: Sinop, Coordenadas.
Primary type: holotype ? (MCNZ).
Figure (holotype).
Distribution: Brazil: Mato Grosso: Esp¡ritu Santo.

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Pyticeroides hansoni Opitz, 2007:120 locality: Ecuador: Napo: Yasuni Research Station.
Primary type: holotype ? (QCAZ).
Figure (holotype).
Distribution: Ecuador: Napo.

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Pyticeroides ichnopsis Opitz, 2007: 129 locality: Brazil: Bahia: Salobro. Primary type:
holotype ? (MNHN).
Figure (holotype).
Distribution: Brazil: Bahia.

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Pyticeroides inconscriptus Opitz, 2007: 134 locality: M‚xico: Chiapas: Parque Nacional Montebello. Primary type: holotype ? (AMNH).
Figure (holotype).
Distribution: M‚xico: Chiapas.

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Pyticeroides inexilis Opitz, 2007: 107 locality: Brazil: Sao Paulo: Fazenda de Alho Itu.
Primary type: holotype ? (MZSP).
Figure (holotype).
Distribution: Brazil: Sao Paulo: Paran .

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Pyticeroides iscus Opitz, 2007: 108 locality: Brazil: Par : Tucurui. Primary type: holotype ?
(MCNZ).
Figure (holotype).
Distribution: Brazil: Par .

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Pyticeroides jubaris Opitz, 2007: 109 locality: Brazil: Sao Paulo: Rio Caraguata, 21ø48'S 52ø27'W. Primary type: holotype ? (FMNH).
Figure (holotype).
Distribution: Brazil: Sao Paulo.

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Pyticeroides laticornis (Say), 1835: 164 locality: United States of America: North Carolina.
Primary type: lectotype, gender not known (MCZC).
Figure (holotype).
Distribution: Canada: Ontario. United States of America: Alabama, Arkansas, Colorado, Connecticut, District of Columbia, Florida, Georgia, Illinois, Indiana, Kansas, Kentucky, Maryland, Massachusetts, Michigan, Missouri, Mississippi, New Jersey, New York, North Carolina, Oklahoma, Ohio, Pennsylvania, Tennessee, Texas, Virginia, West Virginia.

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Pyticeroides luteolus Opitz, 2007: 121 locality: Brazil: Santa Catarina: Nova Teutonia,
27ø11' S 52ø23'W. Primary type: holotype ? (FMNH).
Figure (paratype).
Distribution: Brazil: Santa Catarina.

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Pyticeroides maesi Opitz, 2007: 124 locality: Nicaragua: Granada: Volc n Mombacho, Finca
San Joaquin. Primary type: holotype ? (MCNZ).
Figure (paratype).
Distribution: Nicaragua: Granada.

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Pyticeroides manni Chapin, 1927:8 locality: Bolivia: Beni: Cavinas, Rio Beni. Primary type:
Holotype, gender not known (USNM).
Figure (nontype specimen).
Distribution: Bolivia: Beni: Cochabamba. Brazil: Rond“nia.

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Pyticeroides notialis Opitz, 2007: 126 locality: Argentina: Tucum n: Las Tipas. Primary
type: holotype ? (MLPA).
Figure (paratype).
Distribution: Argentina: Salta: Tucum n. Santa del Estero: Missiones.

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Pyticeroides petilus Opitz, 2007: 108 locality: Bolivia: Cochabamba. Primary type: holotype
? (MNHN).
Figure (holotype).
Distribution: Bolivia: Cochabamba.

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Pyticeroides plautus Opitz, 2007: 135 locality: M‚xico: Oaxaca: highway 131, 75 km S
Oaxaca. Primary type: holotype ? (CNCI).
Figure (paratype).
Distribution: M‚xico: Oaxaca.

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Pyticeroides quadratus Opitz, 2007: 132 locality: Brazil: Goias: Jatai. Primary type: holotype
? (DEIG).
Figure
Distribution: Brazil: Goias.

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Pyticeroides rifkindi Opitz, 2007: 127 locality: Costa Rica: San Jos‚: Cerro de Escazu,
Mtinilla. Primary type: holotype ? (LACM).
Figure (holotype).
Distribution: Costa Rica: San Jos‚.

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Pyticeroides roseicollis Opitz, 2007: 123 locality: Costa Rica: Guanacaste: 3 km SE Rio
Naranjo. Primary type: holotype ? (INBC).
Figure (holotype).
Distribution: Costa Rica: Guanacaste. Panam : Panam .

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Pyticeroides similis Opitz, 2007: 113 locality: Brazil: Rondonia: 62 km SE Ariquemes.
Primary type: holotype ? (MCNZ).
Figure (holotype).
Distribution: Brazil: Rond“nia.

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Pyticeroides stenotes Opitz, 2007: 128 locality: Venezueal: Aragua: El Lim¢n. Primary type:
holotype ? (IZAV).
Figure (holotype).
Distribution: Venezuela: Aragua.

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Pyticeroides strictus Opitz, 2007: 117 locality: Bolivia: Cochabamba.
Figure (holotype).
Distribution: Bolivia: Cochabamba.

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Pyticeroides tessares Opitz, 2007: 126 locality: Honduras: Olancho: La Muralla. Primary
type: holotype ? (FSCA).
Figure (holotype).
Distribution: Honduras: Olancho. Guatemala: Izabal.

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Pyticeroides trilineatum (Chevrolat) 1874: 324 locality: M‚xico: Jalisco. Primary type:
lectotype ? (MNHN).
Figure (lectotype).
Distribution: M‚xico: Jalisco: Veracruz.

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Pyticeroides vichadium Opitz, 2007: 117 locality: Colombia: Vichada: PNN Tuparro, Cerro
Tomas, 5ø21'N67ø51'W. Primary type: holotype ? (IAVH).
Figure (holotype).
Distribution: Colombia: Vichada.

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SILVEIRASIA NEMSIO

Type species: Chaetophloeus hispidus (Opitz), 2004: 21
Generic synonyms: None.
Most recent generic revision: Nem‚sio, A. 2006. Two homonomies in Epiphloeinae (Coleoptera: Cleridae): new generic names for Arenaria Opitz and Chaetophloeus Opitz. Lundiana 7(1): 77.
Distribution: Brazil: Rio Grande do Sul.
Number of species: 1.

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Silveirasia hispidus (Opitz), 2004: 23. Type locality: Brazil: Rio Grande do Sul: Sao
Francisco de Paula. Primary type: holotype ? (MCNZ). Mecke, et al, 2001: 119 (
misidentified as Plocamocera sp.)
Figure (paratype).
Distribution: Brazil: Rio Grande do Sul: Sao Francisco de Paula, forest reserve
Pr¢-Mata.

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STEGNOCLAVA OPITZ, 2009: .

Type species: Ichnea fumigata Gorham, 1877: 414.

Generic synonyms: None.

Most recent generic revision: Opitz, W. 2009. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VIII. The genera Acanthocollis, new genus, Stegnoclava, new genus, and Ichnea Laporte. The Coleopterist Bulletin XXXX
Post Corporaal (1950) citations: Opitz, 2009: XX
Distribution: Argentina, Costa Rica, Per£, Brazil.
Number of Species: 3.

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Stegnoclava fumigate (Gorham), 1877: 414. Type locality: Brazil: Amazonia.
type: lectotype ? (BMNH).
Figure (homotype).
Distribution: Per£: Loreto, Madre de Dios. Brazil: Amazonia, Par 

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Stegnoclava velatis Opitz, 2009. XX. Type locality: Brazil: Mato Grosso: Sinop.
Primary type: lholotype ? (FSCA).
Figure (holotype).
Distribution: Brazil: Mato Grosso.

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Stegnoclava zorropsis Opitz, 2009. Type locality: Costa Rica: Heredia
type: holotype ? (INBC).
Figure (holotype).
Distribution: Costa Rica: Heredia

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TURBOPHLOEUS

Type species: Epiphlaeus simplex, 1900: 397.
Generic synonyms: None.
Most recent generic revision: Opitz, W. 2008. Classification, natural history, and evolution of the Epiphloeinae (Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nem‚sio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus, new genus. Zootaxa 1754: 1-40.
Distribution: Colombia, Bolivia.
Number of species: 1.

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Turbophloeus simplex (Schenkling), 1900: 397. Type locality: Bolivia. Primary type: lectotype ? (DEIG).
Figure (homotype).
Distribution: Colombia: Boyac , Caquet , Huila, Norte de Santander, Risaralda.
Bolivia: Cochabamba, Santa Cruz.



REFERENCES




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Hespenheide, H.A. 1973. A novel mimicry complex: Beetles and flies. Journal of
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Opitz, W. 2008a. Classification, natural history, and evolution of the Epiphloeinae
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from Hispaniola with description of its four new species. American Museum
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Opitz, W. 2008b. Classification, natural history, and evolution of the Epiphloeinae
(Coleoptera: Cleridae). Part VI. The genera Epiphlaeus Spinola and Opitzius Barr.
Annales Zoologoci (Warszawa) 58: 1-34.
.
Opitz, 2008c. Classification, natural history, and evolution of the Epiphloeinae
(Coleoptera: Cleridae). Part VII. The genera Hapsidopteris Opitz, Iontoclerus Opitz, Katamyurus Opitz, Megatrachys Opitz, Opitzia Nem‚sio, Pennasolis Opitz, new genus, Pericales Opitz, new genus, Pteroferus Opitz, new genus, and Turbophloeus, new genus. Zootaxa 1754: 1-40.

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Zayas, F. 1988. Entomofauna Cubana, orden Coleoptera, seperata descripcion de nuevas
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