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Overview |
Main identification features
- all blackish
- ll: 2 overlapping parts
- c: pointed, confluent with dorsal fin and a
- pelvic - 1 filament under operculum
Dorsal rays 95-115; anal rays 76-91; pectoral rays 24-29; pelvic fins below rear part of head, each consisting of a single filamentous ray; dorsal and anal fins confluent with pointed tail; palate teeth absent; operculum with 1 short spine; preoperculum with 2 teeth; body covered with small scales, about 100-115 in midlateral series, head partly scaleless; scattered papillae on head; origin of dorsal fin above middle part of pectoral fins; lateral line in 2 parts overlapping.
Dark purplish to nearly black, including fins.
Size: grows to about 20 cm.
Inhabits rocky crevices.
Depth: 5-20 m.
Southern California to the central Gulf of California to Ecuador, the Revillagigedos and Galapagos.
Attributes
Abundance: Common.
Cites: Not listed.
Climate Zone: North Temperate (Californian Province &/or Northern Gulf of California); Northern Subtropical (Cortez Province + Sinaloan Gap); Northern Tropical (Mexican Province to Nicaragua + Revillagigedos); Equatorial (Costa Rica to Ecuador + Galapagos, Clipperton, Cocos, Malpelo).
Depth Range Max: 20 m.
Depth Range Min: 5 m.
Diet: mobile benthic crustacea (shrimps/crabs); bony fishes.
Eastern Pacific Range: Northern limit=33; Southern limit=-2; Western limit=-118; Eastern limit=-78; Latitudinal range=35; Longitudinal range=40.
Egg Type: Live birth; Pelagic larva.
Feeding Group: Carnivore.
FishBase Habitat: Reef Associated.
Global Endemism: Tropical Eastern Pacific (TEP) endemic; East Pacific endemic; All species.
Habitat: Reef associated (reef + edges-water column & soft bottom); Rocks; Reef (rock &/or coral); Reef only.
Inshore Offshore: Inshore; Inshore Only.
IUCN Red List: Not evaluated / Listed.
Length Max: 20 cm.
Regional Endemism: Island (s); Continent; Continent + Island (s); TEP endemic; All species.
Residency: Resident.
Salinity: Marine; Marine Only.
Water Column Position: Bottom; Bottom only;
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Names |
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Links to other sites |
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References |
- Béarez, P., 1996., Lista de los Peces Marinos del Ecuador Continental., Revista de Biologia Tropical, 44:731-741.
- Cohen, D.M., 1964., A review of the ophidioid fish genus Oligopus with the description of a new species from the West Africa., Proc. U.S. Nat. Mus., 116:1-22.
- Eschmeyer , W. N. , Herald , E. S. and Hamman, H., 1983., A field guide to Pacific coast fishes of North America from the Gulf of Alaska to Baja California. Peterson Field Guide Ser. 28., Houghton Mifflin:336pp.
- Findley, L.T., Hendrickx, M.E., Brusca, R.C., van der Heiden, A.M., Hastings, P.A., Torre, J., 2003., Diversidad de la Macrofauna Marina del Golfo de California, Mexico., CD-ROM versión 1.0. Projecto de la Macrofauna del Golfo . Derechos reservados de los autores y Conservación Internacional.
- Galván-Magaña, F., Gutiérrez-Sánchez, F., Abitia-Cárdenas, L.A., Rodríguez-Romero, J., 2000., The distribution and affinities of the shore fishes of the Baja California Sur lagoons. In Aquatic Ecosystems of Mexico: Status and Scope. Eds. M. Manuwar, S.G. Lawrence, I.F. Manuwar & D.F. Malley. Ecovision World Monograph Series., Backhuys Publishers:383-398.
- Heller , E. and Snodgrass, R.E., 1903., Papers from the Hopkins Stanford Galapagos expedition, 1898-1899. XV. New fishes., Proc. Wash. Acad. Sci., 5:189-229.
- Lopez , M. I. and Bussing, W. A., 1982., Lista provisional de los peces marinos de la Costa Rica., Revista de Biologia Tropical, 30(1):5-26.
- Love, M.S., Mecklenburg, C.W., Mecklenburg, T.A., Thorsteinson, L.K., 2005., es of the West Coast and Alaska: a checklist of North Pacific and Artic Ocena species from Baja California to the Alaska-Yukon border., U.S. Department of the Interior, U.S. Geological Survey, Biological Resources Division, 288pp.
- Snodgrass , R. E. and Heller, E., 1905., Papers from the Hopkins Stanford Galapagos expedition, 1898-1899. XVII. Shorefishes of the Revillagigedo, Clipperton, Cocos and Galapagos Island., Proc. Wash. Acad. Sci., 6:333-427.
- Thomson , D.A. , Findley , L.T. and Kerstitch, A.N., 2000., Reef fishes of the Sea of Cortez., University of Texas Press(Revised Ed.):353.
- Van der Heiden , A. M. and Findley, L. T., 1988., Lista de los peces marinos del sur de Sinaloa, México., Anales del Centro de Ciencias del Mar y Limnologia de la Universidad Autonoma Nacional de Mexico, 15:209-224.
- Villareal-Cavazos, A., Reyes-Bonilla, H., Bermúdez-Almada, B. and Arizpe-Covarrubias, O., 2000., Los peces del arrecife de Cabo Pulmo, Golfo de California, México: Lista sistemática y aspectos de abundancia y biogeografía., Rev. Biol. Trop., 48:413-424.
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Acknowledgements |
I thank Ashley MacDonald and John Pickering, University of Georgia, for technical support in building this page.
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http://www.fishbase.org/Summary/speciesSummary.php?genusname=Grammonus&speciesname=diagrammus ---> http://192.134.151.83/Summary/speciesSummary.php?genusname=Grammonus&speciesname=diagrammus http://192.134.151.83/Summary/speciesSummary.php?genusname=Grammonus&speciesname=diagrammus ---> https://fishbase.mnhn.fr/Summary/speciesSummary.php?genusname=Grammonus&speciesname=diagrammus https://fishbase.mnhn.fr/Summary/speciesSummary.php?genusname=Grammonus&speciesname=diagrammus ---> https://fishbase.mnhn.fr/summary/Grammonus-diagrammus.html
Grammonus diagrammus, Purple brotula
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Grammonus diagrammus
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Teleostei (teleosts) >
Ophidiiformes
(Cusk eels) >
Bythitidae
(Livebearing brotulas)
Etymology:
Grammonus:
Greek, gramma = signal, mark + Greek, onos = hake
.
Environment: milieu / climate zone / depth range / distribution range
Ecology
Marine; reef-associated; depth range 6 - 18 m (Ref.
36488
). Subtropical
Eastern Pacific: San Clemente Island in south California, USA to Panama and the Galapagos Islands.
Size / Weight / Age
Maturity: L
m
 
?
  range ? - ? cm
Max length : 20.3 cm TL male/unsexed; (Ref.
4930
)
Dorsal
spines
(total): 0;
Dorsal
soft rays
(total): 95-115;
Anal
spines
: 0;
Anal
soft rays
: 75 - 91;
Vertebrae
: 60 - 63. Branchiostegal rays: 8 (Ref.
36488
).
Uncommon species (Ref.
34024
). Inhabits rocky areas of shallow waters (Ref.
2850
). Hides in dark caverns and crevices during the day, active at night (Ref.
2850
). Gives birth to live young (Ref.
28023
).
Life cycle and mating behavior
Maturity
|
Reproduction
|
Spawning
|
Eggs
|
Fecundity
|
Larvae
Ovoviviparous (Ref.
36488
).
Nielsen, J.G., D.M. Cohen, D.F. Markle and C.R. Robins
, 1999. Ophidiiform fishes of the world (Order Ophidiiformes). An annotated and illustrated catalogue of pearlfishes, cusk-eels, brotulas and other ophidiiform fishes known to date. FAO Fish. Synop. 125(18):178p. Rome: FAO. (Ref.
34024
)
IUCN Red List Status (Ref.
130435
)
Least Concern (LC)
; Date assessed:
29 April 2008
CITES
Not Evaluated
Not Evaluated
Threat to humans
Harmless
Human uses
Fisheries: of no interest
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Zoological Record
Estimates based on models
Preferred temperature (Ref.
123201
): 21.3 - 29.1, mean 26.7 °C (based on 204 cells).
Phylogenetic diversity index (Ref.
82804
): PD
50
= 0.5005 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00457 (0.00179 - 0.01169), b=3.10 (2.87 - 3.33), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref.
93245
).
Trophic level (Ref.
69278
): 3.5 ±0.50 se; based on food items.
Resilience (Ref.
120179
): High, minimum population doubling time less than 15 months (Preliminary K or Fecundity.).
Fishing Vulnerability (Ref.
59153
): Low vulnerability (10 of 100).
Nutrients (Ref.
124155
): Calcium = 52.9 [24.3, 133.5] mg/100g; Iron = 0.662 [0.352, 1.248] mg/100g; Protein = 18 [15, 21] %; Omega3 = 0.163 [0.086, 0.306] g/100g; Selenium = 12 [5, 30] μg/100g; VitaminA = 67.1 [20.4, 228.7] μg/100g; Zinc = 0.994 [0.611, 1.562] mg/100g (wet weight);
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Updated: 2024-06-15 21:42:43 gmt
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