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Taken from: A Revision of the North American Ammophila (hymenoptera, Sphecidae). Arnold S. E. Menke, 1965.
Ammophila (Ammophila) pictipennis Walsh
(Fig. 21)
Ammophila pictipennis Walsh, 1869. Amer. Natur. 1:164. female, “southern Illinois” (types destroyed).
Ammophila anomala Taschenberg, 1869. Zeitscbr. Ges. Naturwiss. Halle 34:434. Syntypes male, female, Illinois (Zoologisches Institut, Halle). New synonymy.
Sphex nigropilosus Rohwer, 1912, Proc. U. S. Nat. Mus. 41:465. Holotype female, Distrito Federal, Mexico (United States National Museum, Washington). Tentative new synonymy.
Ammphila placida of authors, not Smith.
Ammophila xanthoptera of Fernald 1934 (in part), not Cameron.
The penis valve (fig. 101) of pictipennis will separate males of this species from its sibling, placida. Females cannot readily be separated except by association males.
Ammophila pictipennis has generally been considered a synonym of placida. Taschenberg’s description of anomala was published in late 1869 (November-December) while Walsh’s name apparently appeared in the Spring of the same year, hence pictipennis has priority over anomala. The types of pictipennis have been destroyed. Fernald (1934) studied Taschenberg’s syntypes and in one male the third submarginal cell was lacking. The type of nigropilosus appears to be synonymous with pictipennis but it could be placida.
Ammophila pictipennis occurs over most of the United States east of the Rocky Mountains. It ranges south along the eastern half of Mexico as far as Mexico City.
Ammophila nigricans Group
Diagnosis. Primary characters: Free margin of male clypeus essentially truncate apically, truncation sometimes broadly, shallowly emarginate; hypostoma in male without a process; collar and scutum without ridges; propodeal enclosure usually with broadly spaced ridges, which are uniform across width of enclosure, interspaces granulate, surface dull (figs. 20-21); preëpisternal sulcus usually long, ending in ventral region, (sometimes Indistinct below pronotal lobe or lacking in apicalis, pictipennis, extremitata, placida and nigricans males and both sexes of con); penis valve head usually with a basal spine-like process (except in apicalis, extremitata and pictipennis) base of gonoforceps sot dorsoventrally elongate. Secondary characters metapleural flange not lamellate; forewing with three submarginal cells.
Included North American species. Ammophila extremitata Cresson, fernaldi (Murray), femurrubra Fox, mcclayi Menke, nigricans Dahlbom, pictipennis Walsh; placida Smith, and xanthoptera Cameron. Other New World species in the group are: apicalis Guérin-Mêneville (West Indies), centralis Cameron, cora Cameron and gaumeri Cameron (Mexico and Central America).
Discussion. The granulate appearance of the propodeal enclosure is diagnostic for the group. In addition, the mesosoma in most North American species is largely devoid of appressed silver hair and the integument is mostly dull black or opaque. Most species in the nigricans group have bluish micropubescence on the head and mesosomal dorsum, and in a few species it coven the entire body. Females of six species (apicalis, cora, extremitata. femurrubra, pictipennis and usually have a minute tooth on the inside edge of the claw near the base (fig. 30). The other species sometimes have a poorly defined tooth on the claw.
The male genitalia, especially the aedeagus and gonoforceps, display excellent specific differences in most species of this group. The shape of the subgenital plate is also of importance. The presence or absence of appressed silver hair on the face, pronotal lobe and other mesosomal areas is of considerable value in species discrimination. Wing color is often diagnostic when used with other characters.
The nigricans group is restricted to North and Central America but appears closely allied with the fallax group of the Old World.
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Updated: 2024-05-04 14:40:22 gmt
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