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Andrena clarkella (Kirby, 1802)
Melitta clarkella Kirby, 1802; Andrena dispar Zetterstedt, 1838

Life   Insecta   Hymenoptera   Apoidea   Andrenidae   Andrena
Subgenus: Andrena



Andrena clarkella, female
Jelle Devalez · 9
Andrena clarkella, female

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Andrena clarkella, female
Jelle Devalez · 9
Andrena clarkella, female
Andrena clarkella, male
Jelle Devalez · 9
Andrena clarkella, male

Andrena clarkella MALE CFP comp
© Copyright Laurence Packer 2014 · 7
Andrena clarkella MALE CFP comp
Andrena clarkella, F, face, Hancock co
© Copyright source/photographer · 5
Andrena clarkella, F, face, Hancock co

Andrena clarkella, F, back, Hancock co
© Copyright source/photographer · 5
Andrena clarkella, F, back, Hancock co
Andrena clarkella, F, side, Hancock co-Recovered
© Copyright source/photographer · 5
Andrena clarkella, F, side, Hancock co-Recovered

Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 5
Andrena clarkella
Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella

Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella
Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella

Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella
Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella

Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella
Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella

Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella
Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella

Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella
Andrena clarkella
© Copyright Malcolm Storey 2011-2118 · 3
Andrena clarkella

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Andrena clarkella – A northern mining bee that specializes on willows (Salix sp.) noted for bright yellow to orange hind tibia and tarsi. Females have a pale scopae but tergal hairs are dark.

Overview
Reprinted with permission of the American Entomological Society from: LaBerge, W. E. 1980. A revision of the bees of the genus Andrena of the Western Hemisphere. Part X. Subgenus Andrena. Transactions of the American Entomological Society 106: 395-526.

Please report text errors to: leah at discoverlife dot org.

Andrena clarkella is a colorful, holarctic species ranging across northern North America and Eurasia; At one time the author was of the opinion that clarkella was, perhaps, a bee imported into North America from Europe; However, its North American range seems to deny this hypothesis; This bee is closely related to A; frigida but can be distinguished readily in both sexes by the orange to red hind tarsi and tibiae; Basal mandibular teeth are lacking in both sexes and the vestitural color of the female is very much like the darkest specimens of frigida.

FEMALE. MEASUREMENTS AND RATIOS. N = 20; length, 11-13 mm; width, 3-4 mm; wing length, M = 4.69*0.172 mm; FL/FW, M = 0.95 0.004; FOVL/FOVW, M = 2.85 0.050.

INTEGUMENTAL COLOR. Black except as follows: mandible with apical third to half rufescent; flagellum below dark brown; tegula translucent, reddish-brown; wing membranes hyaline, slightly infumate, veins reddish-brown to dark brown; tergal apical areas and sternal apical areas translucent, dark reddish-brown; hind basitarsi and tibiae bright red to orange; hind distitarsi dark rufescent; tibial spurs testaceous.

STRUCTURE. Antennal scape length equals first two and one-half flagellar segments or slightly more; flagellar segments as in frigida; Eyes each almost four times as long as broad, inner margins parallel; Mandibles, malar space and galea as in frigida; Maxillary palpus as in frigida but segmental ratio about as 1.0:1.0:0.9:0.9:0.8:0.6; Labial palpus as in frigida but ratio about as 1.0:0.7:0.5:0.5; Labral process and labrum, clypeus, supraclypeal area, face above antennal fossae, vertex, facial fovea and genal areas as in frigida, Thoracic form and sculpturing as in frigida but propodeum with dorsal enclosure finely tessellate, not roughened at base.

Metasomal terga 1-4 as in frigida but basal area punctures extremely minute, usually not visible except on terga 3 and 4; Pygidial plate as in frigida but apex broadly rounded to truncate; Sterna as in frigida.

VESTITURE. � Head hairs dark brown to black except vertex often with hairs brown or pale ochraceous mixed with brown and lower genal area may have some paler hairs; Thoracic hairs dark brown to black except dorsum of thorax pale ochraceous or pale and dark mixed; Leg hairs brown to dark brown except hind tibiae and basitarsi with outer surface hairs (scopal) white to pale ochraceous and inner surfaces with hairs pale yellow; Metasomal vestiture dark brown to black except tergum 1 and often base of tergum 2 with pale ochraceous hairs; Pollen-collecting hairs, form and distribution of vestiture as in frigida.

MALE. MEASUREMENTS AND RATIOS. N = 6; length, 9-10 mm; width, 2.0-2.5 mm; wing length, 4.07 0.406 mm; FL/FW, M = 0.92 0.012; FS1/FS2 (4), M = 1.89 0.157.

INTEGUMENTAL COLOR. Black except as follows: mandible with apical third or less rufescent; flagellum below dark brown to reddish-brown; tegula translucent, reddish-brown; wing membranes hyaline, veins reddish-brown to dark brown; hind basitarsi and tibiae red to bright orange, tergal and sternal apical areas translucent, reddish-brown (sterna 2-5 often narrowly colorless apically); tibial spurs testaceous.

STRUCTURE. Antennae as in frigida but scape length equals first two flagellar segments or slightly less; Eyes narrow, each about three and one-fourth times as long as broad, inner margins parallel; Mandible, malar space and galea as in frigida; Maxillary palpus as in frigida but segmental ratio about as 1.0:1.0:0.9:0.9:0.5:0.5; Labial palpus as in frigida but radio about as 1.0:0.5:0.5:0.5; Labral process as in frigida, labrum without crista; Clypeus, supraclypeal area, face above antennal fossae and genal area as in frigida; Vertex above lateral ocellus equals about one ocellar diameter, dulled by dense fine tessellation and punctures.

Pronotum as in frigida; Thoracic sculpture as in frigida but propodeum with dorsal enclosure without basal roughening or restricted to extreme base and weak.

Metasomal terga sculptured as in female; Tergum 7 with weak, extremely narrow, pseudopygidial area; Sterna 2-5 sculptured as in frigida; Sternum 6 flat, apical margin broadly and extremely shallowly emarginate.

Terminalia much as in frigida but sternum 8 not emarginate apically (see Figs; 22 to 26).

VESTITURE. Generally white to cinereous except as follows: face alongside compound eyes, vertex and often genal areas near compound eyes with abundant long dark brown to black hairs; posterior mesepisternum, metepisternum, lateral surface of propodeum often with dark brown hairs mixed with pale; inner surfaces hind basitarsi and tibiae pale yellow; Form and distribution of hairs as in frigida.




Reprinted with permission from: Mitchell, T.B. 1960. Bees of the Eastern United States. North Carolina Agricultural Experiment Station Technical Bulletin No. 141.



FEMALE Length 12 mm.; clypeus slightly convex, broader than long, surface rather dull, punctures rather fine and close; facial foveae broad, fuscous tomentose; space between margin of vertex and lateral ocelli slightly exceeding their diameter; cheeks slightly broader than eyes, minutely punctate; malar space short but distinct; basal segment of flagellum equal in length to 2nd and 3rd combined; process of labrum subtriangular, the tip truncate and slightly emarginate; mandibles of ordinary length, with a small inner subapical tooth; pubescence of head long, dense and erect, fuscous or blackish except for some pale hairs on vertex; thoracic integument dull and densely tessellate, punctures inevident except along lower portions of pleura; enclosure of propodeum smooth, pubescence long, dense, and erect, that on scutum, scutellum and upper portion of propodeum pale ochraceous or whitish, otherwise fuscous to black; propodeal corbicula and trochanteral floccus well developed, black; tibial scopa rather long and dense, orange; fore and hind basitarsi slightly narrower than their respective tibiae, the mid basitarsi subequal to their tibiae; hind tibiae and tarsi bright ferruginous, legs otherwise black; 2nd submarginal cell much shorter than 3rd, receiving 1st recurrent slightly beyond middle; abdominal terga rather dull but smooth, apical third slightly depressed and slightly paler in color than the remainder of the disc, pubescence long, dense and erect, entirely fuscous except for some pale pubescence on basal segment.



MALE Length 10 mm.; clypeus slightly convex, much broader than long, closely and finely punctate; space between margin of vertex and lateral ocelli slightly exceeding their diameter; cheeks broad and flat, subangulate, angle opposite middle of eye; malar space well developed but short; process of labrum only slightly broader than long, truncate; mandibles rather slender and slightly flexed, overlapping about one-third; pubescence of head long, dense and erect, whitish on cheeks below and on clypeus and vertex, otherwise fuscous; thoracic integument dull, densely tessellate and impunctate; propodeal enclosure smooth; thoracic pubescence long, dense and erect, whitish in general, but with some fuscous hairs on pleura above; basitarsi narrow and elongate, much narrower than their respective tibiae; hind tibiae and tarsi ferruginous, the others dark; pubescence of legs largely pale; 2nd submarginal cell much shorter than 3rd, receiving 1st recurrent beyond middle; abdominal terga smooth but dull, the punctures exceedingly minute and obscure, apical third of discs slightly depressed, lighter in color than remainder of discs, pubescence rather long, thin and erect, entirely pale, with no trace of fasciae; apical portion of sternum 8 gradually narrowed toward the tip, with a very low, subapical, dorsal ridge, tip beyond this ridge slightly rounded, slightly longer than broad; penis valves broadly expanded basally, excavated beneath this expansion, gonocoxites moderately expanded apically, the gonocoxal lobes produced and narrowly rounded.



DISTRIBUTION Holarctic, being recorded from Europe, Alaska, eastern Canada, and in the United States from Minnesota to New York, Massachusetts and Maine; April to July.

Identification
Extracted from Joel Gardner. "The mining bees of Minnesota (unpublished manuscript)"

Andrena clarkella has orange hind tibiae with light scopal hairs, is also only found in northern Minnesota

Names
Scientific source:
References
Melitta Clarkella Kirby, 1802. Monog. Apum Angliae, 2, p. 130. F.

Andrena clarkella Sladen, 1919. Canad. Ent. 51, p. 124. F M.

Andrena (Andrena) clarkella Lanham, 1949. Univ. Calif. Pubs. Ent. 8, p. 205. (classification)

Hosts · map
FamilyScientific name @ source (records)
Salicaceae  Salix sp @ CUIC_ENT (3)

Salix @ AMNH_BEE (10)

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Updated: 2024-10-04 23:55:24 gmt
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