Reprinted with permission of the American Entomological Society from:
LaBerge, W. E. 1969. A revision of the bees of the genus Andrena of the Western Hemisphere. Part II. Plastandrena, Aporandrena, Charitandrena. Transactions of the American Entomological Society 95: 1-47.
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This small species from southern California is the same as Plastandrena in having the hind tibial spur strongly curved and with a lamellate expansion in the basal third. It can be distinguished from all the American Plastandrena by having the dorsal propodeal enclosure evenly tessellate and not at all roughened by rugulae. Also, the pterostigma is considerably larger and broader than in Plastandrena and the male terminalia is very distinctive. This species could be confused with two species of Callandrena which have modified posterior hind tibial spurs. However coactipostica females have simple scopal hairs and a carinate ventroposterior border on the hind femur. Both sexes of coactipostica differ from the Callandrena by the broader pterostigma and the long maxillary palpi.
FEMALE. MEASUREMENTS AND RATIOS. — N = 20; length, 9-11 mm; width, 3.0-3.5 mm; wing length, M = 3.52 ± 0.325 mm; FL/FW, M = 0.93 ± 0.006; FOVL/FOVW. M = 2.85 ± 0.046.
INTEGUMENTAL COLOR. — Black except as follows: apical third of mandible dark rufescent; wing membranes hyaline, slightly infumate apically, veins dark reddish-brown; tergal apices slightly reddened or more translucent; sterna 2-5 hyaline apically; distitarsi dark rufescent; tibial spurs pale yellow.
STRUCTURE. — Antennae moderately long; scape length equals first three and three-fourths flagellar segments; flagellar segment 1 about equal to segments 2 plus 3 which are subequal to each other in length and shorter than 4; segments 3-9 quadrate or slightly broader than long. Eye about four times as long as broad, inner margins converging slightly toward mandibles. Malar space linear. Galea moderately dulled by reticular shagreening; lateral surface as broad as dorsal or broader. Maxillary palpus as in crataegi but segmental ratio about 0.8:1.0:0.5:0.5:0.4:0.5. Labial palpus as in crataegi but segmental ratio about 1.0:0.4:0.4:0.5. Labral process as in crataegi; apical labral area concave across labrum, without crista. Clypeus relatively flat, with minute round punctures separated mostly by half to one puncture width, sparser along midline; surface dulled by regular fine tessellation. Supraclypeal area moderately dulled by minute crowded punctures. Genal area distinctly broader than eye in profile, dulled by minute sparse punctures and dense tessellation. Vertex above lateral ocellus equal to half an ocellar diameter or slightly more; with small shallow sparse punctures and dense tessellation, dull. Face above antennal fossae with fine longitudinal rugulae, interrugal space finely punctate, dulled by tessellation. Facial fovea short, just reaching level of posterior margin clypeus, not reaching level of apices of compound eyes, separated from lateral ocellus by slightly more than one ocellar diameter.
Pronotum normal, moderately dulled by fine tessellation and scattered small punctures. Mesoscutum, scutellum and metanotum with extremely small, round punctures separated by half a puncture width or less, fine tessellation dulling surface. Propodeum with dorsal enclosure finely tessellate, margined by slightly raised sutures; dorsolateral and posterior surfaces regularly tessellate with small scattered punctures; corbicular surface moderately shiny, with widely scattered punctures and coarse tessellation. Mesepisternum with small round punctures separated by two to three puncture widths or more, surface dull, tessellate. Metepisternum similar but lower two-thirds impunctate. Pterostigma long, broader than from inner margin prestigma to wing margin; vein 1st m-cu meets second submarginal cell at or beyond middle of cell. Hind femur with posteroventral carinate ridge running from posterior tibiofemoral articulation almost to trochanter.
Metasomal terga 1-5 with small round punctures separated mostly by half a puncture width or slightly more, more crowded in apical areas, surfaces moderately shiny with extremely delicate reticular shagreening. Pygidial plate U-shaped, without distinctly raised internal triangular area. Sterna 2-5 impunctate at extreme base and in narrow apical area, in median area with small round punctures separated mostly by one puncture width, surface moderately shiny, reticularly shagreened.
VESTITURE. — Generally white but vertex of head and thoracic dorsum pale ochraceous, terga 5 and 6 ochraceous or fulvous medially and inner surfaces tarsi dark red. Mesoscutum covered by extremely short, erect, thick hairs with sparse long hairs interspersed, appearing velvety. Terga 2-4 with apical white fasciae but usually narrowly interrupted on tergum 2. Propodeal corbicula incomplete anteriorly, with long simple internal hairs. Trochanteral flocculus complete, dense. Tibial scopal hairs moderately long, simple.
MALE. MEASUREMENTS AND RATIOS: N = 20; length, 7-9 mm; width, 1.5-2.0 mm; wing length. M = 3.20 ± 0.083 mm; FL/FW, M = 0.94 ± 0.008; FS1/FS2, M = 1.26 ± 0.023.
INTEGUMENTAL COLOR. — Generally as in female except as follows: clypeus often with posteromedian third to half or slightly more cream-colored or yellow; terga 1-5 with apical areas hyaline, colorless at least near margins.
STRUCTURE. — Antennae moderately long, reaching metanotum in repose; scape length equals flagellar segments 1 plus 2 or slightly more; flagellar segment 1 equal to one and one-tenth to one and one-third length of second segment, segment 2 shorter than 3, 3 to 10 subequal and each one and a half times as long as broad or slightly less. Eye about three times as long as broad, inner margins converging strongly toward mandibles. Malar space, mandibles and galea as in female. Maxillary palpus as in female but segmental ratio about 1.0:0.8:0.8:0.8:0.5:0.7. Labial palpus as in female but segmental ratio about 1.0:0.4:0.3:0.4. Labrum as in female but process usually bidentate, apical area flat or convex. Clypeus as in female but punctures more crowded and obscured by dense tessellation, Supraclypeal area minutely punctured and densely tessellate. Vertex as in female, opaque, densely tessellate. Genal area broader than eye, obscurely punctate, densely tessellate. Face above antennal fossae with longitudinal rugulae, interrugal spaces punctate and tessellate.
Thorax as in female except as follows: mesoscutum, scutellum and metanotum with punctures obscured by dense regular tessellation, opaque; mesepisternal punctures minute, surface opaque, tessellate; propodeum with lateral surface dull, tessellate. Wing venation and pterostigma as in female. Hind femur with weak posteroventral ridge present at least apically.
Metasomal terga 1-5 with weak sparse punctures obscured by regular tessellation, narrow apical area shiny. Sterna 2-5 sparsely punctate, moderately shiny, reticularly shagreened. Sternum 6 flat, shallowly emarginate apically.
Terminalia as in figures 29-33. Note the following: gonostyli with inner surfaces granular or pebbly; penis valves with well-formed dorsal and ventral lamellae laterally: sternum 7 with long curved basal apodemes, emarginate apically; sternum 8 with apex broadened, truncate to slightly emarginate.
VESTITURE. — Generally white to pale ochraceous, darker on vertex and thoracic dorsum. In general hairs are longer than in female on head, thorax and terga 1-4 basal areas. Terga 2-5 with apical pale fasciae but often narrowly interrupted on 2 and 3. Sterna without distinct subapical fimbriae.
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