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Andrena reflexa Cresson, 1872
Andrena permitis Cresson, 1872

Life   Insecta   Hymenoptera   Apoidea   Andrenidae   Andrena
Subgenus: Callandrena


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Reprinted with permission of the University of Nebraska State Museum from:
LaBerge, W. E. 1967. A revision of the bees of the genus Andrena of the Western Hemisphere. Part I. Callandrena (Hymenoptera: Andrenidae). Bulletin of the University of Nebraska State Museum 7: 1-316.


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Cresson (1872) named the two sexes from Texas as two species. The male of reflexa is similar to that of A. accepta in the high vertex, broad genal areas, yellow parocular maculae (usually), and sculpturing. The male can be distinguished from that of accepta by the short clypeus, the short labral process, the long mandibles and the reflexed tergal apices. The female of reflexa differs markedly from that of accepta by its shorter vertex, lack of yellow on the clypeus and parocular areas, and short labral process.

FEMALE. MEASUREMENTS AND RATIOS. N = 3; length, about 12 mm; width, about 4 mm; wing length, 4.11-4.44 mm; FL/FW, 0.91-1.04; FOVL/FOVW, 3.00-3.23.

INTEGUMENTAL COLOR. Black except as follows; mandibles largely rufescent; flagellar segments 3-10 red below; tegulae testaceous; wing membranes moderately infumate, darker apically, veins reddish-brown to black; terga with apical thirds hyaline, colorless to slightly rufescent basad; sterna 2-5 more narrowly hyaline apically, clear to yellowish, basally rufescent; tarsi rufescent; tibial spurs yellow.

STRUCTURE. Antennae of moderate length; scape equal to flagellar segments 1-3; flagellar segment 1 equal in length to segments 2-4 plus half of 5, segment 2 equal to 3 and each shorter than segment 4. Eye slightly longer than three and one-half times as long as broad, inner margins parallel or converging slightly towards vertex. Malar space linear. Mandible of moderate length, bidentate, outer mandible in repose extends beyond middle of labrum by one-fifth its length; ventrobasal angle not developed; subgenal coronet well developed. Galea with lateral surface about one-fourth as broad as dorsal; opaque, with scattered obscure punctures and regular coarse tessellation. Maxillary palpus shorter than galea when extended forward, segmental ratio about 1.0:1.0:0.9:0.7: 0.6:0.7. Labial palpus as in accepta but ratio about 1.9:1.0:0.8:0.8. Labral process short, flat, about six times as broad as long not bidentate, median emargination extremely broad and shallow. Clypeus flat, protruding beyond ends of compound eyes by about one-third its own length, with regular round punctures separated mostly by half a puncture width except narrow impunctate midline, surface shiny, shagreening extremely delicate. Supraclypeal area with small punctures and fine shagreening dulling surface. Genal area about one and one-half times as broad as eye in profile, punctures minute, separated by one puncture width except more crowded near eye, surface shiny. Vertex above lateral ocellus equal to almost two ocellar diameters, punctures crowded above ocelli, sparse laterally, surface moderately dulled by reticular shagreening. Face above antennal fossae with well-developed longitudinal rugulae reaching ocelli above but not between lateral ocelli and foveae, interrugal spaces with large punctures and fine shagreening. Facial fovea deep, short, extending little below level of upper margin of antennal fossa, slightly narrowed below, separated from eye margin and converging above, separated from lateral ocellus by slightly less than 1 one ocellar diameter.

Pronotum normal, with small punctures crowded above, scattered at sides and reticular shagreening dulling surface. Mesoscutum, scutellum and metanotum opaque, with crowded round punctures and coarse tessellation dulling surfaces. Parapsidal lines of normal length. Tegulae impunctate. Propodeum with dorsal enclosure tessellate with anastomizing rugulae; dorsolateral and posterior surfaces with round punctures, sparse posteriorly, and fine reticular shagreening, moderately shiny; corbicular area shiny, with scattered punctures and coarse reticular shagreening. Mesepisternum like mesoscutum but punctures separated by one to two puncture widths or more. Metepisternum with crowded punctures above, like corbicular area below. Middle basitarsus about as broad as hind, sides subparallel without apical anterior process. Fore wing with three submarginal cells; vein 1st m-cu meets second sub)marginal cell near middle of cell; pterostigma long, narrower than from inner margin prestigma to wing margin. Claws and tibial spurs normal.

Terga 1-4 with apical areas slightly impressed and apices narrowly reflexed; basal areas with large crowded punctures separated by less than half a puncture width, surfaces shiny; apical area punctures slightly smaller. Pygidial plate V-shaped, usually with blunt apex. Sterna 2-5 impunctate in basal halves, with crowded punctures near apical hyaline margins.

VESTITURE. Generally pale ochraceous, somewhat brighter on vertex and thoracic dorsum. Terga 2-4 with pale apical pubescent fasciae. Tibial plate and tarsi with reddish-brown hairs. Propodeal corbicula incomplete anteriorly, with long, apically plumose, interior hairs in upper half to two-thirds. Trochanteral flocculus complete. Tibial scopal hairs long, plumose throughout.

MALE. MEASUREMENTS AND RATIOS. N = 4; length, 11-13 mm; width, 3.5-4.0 mm; wing length, M = 4.16 0.663 mm; FL/FW, M = 0.77 0.023; FS1/FS2, M = 2.97 0.145.

INTEGUMENTAL COLOR. Black except as follows: clypeus yellow except apical margin, lateral areas to and including tentorial pits (about mediobasal one-half or slightly more is yellow); parocular area with small yellow macula at apex (absent in one specimen, very large in another); flagellar segments 3-11 slightly reddened below; tegulae brownish-red; wing membranes moderately infumate especially apically, veins dark brown; terga and sterna broadly hyaline apically, colorless to slightly yellowed; tarsi, tibiae and usually femora in part somewhat rufescent.

STRUCTURE. Antennae short, barely reaching tegulae; scape subequal to or slightly shorter than flagellar segments 1-3; flagellar segment 1 subequal in length to 2 through 4, segments 2 and 3 subequal to one another, each shorter than 4. Eye more than three times as long as broad, subparallel or converging slightly towards vertex. Malar space short, about five times as broad as long, slightly broader below. Mandible short, extends beyond midline of labrum in repose by about one-fourth its length, broad near apex, obscurely tridentate, with large cutting edge along inner margin in apical third. Galea as in accepta. Maxillary palpus as in accepta bin ratio about 1.2:1.0:0.9:0.8:0.8:1.2. Labial palpus as in accepta, but ratio about 1.8:1.0:0.6:0.9. Labrum small, no wider than one-third distance between mandibular bases, process not well developed, obscurely bidentate. Hat, short. Clypeus short, not protruding beyond lower level of eyes, rounded medially, with round punctures separated mostly by half a puncture width except sparser medially, surface shiny. Supraclypeal area with small distinct punctures separated by one-half to one puncture width, surface somewhat dulled by reticular shagreening. Genal areas broad, almost two times width of eye in profile, with small round punctures separated by one-half to one puncture width (sparser posteriorly), surface shiny. Vertex tall, above lateral ocellus equals about 4 ocellar diameters, with small round punctures separated by one puncture width at apex and sparser laterally, surface dulled above by reticular shagreening. Face punctate up to level of ocelli, without longitudinal rugae below ocelli.

Pronotum as in accepta. Mesoscutum as in accepta but surface dulled by fine, dense, reticular shagreening. Scutellum as in accepta. Propodeum as in accepta but enclosure without transverse rugulae, either granular or with irregular, extremely fine, largely longitudinal rugulae. Episterna as in accepta. Wings as in accepta but vein 1st m-cu meets second submarginal cell at or before middle of cell. Legs and spurs normal.

Metasomal terga with apical areas moderately strongly impressed with apices reflexed; terga with punctures separated by one-half to one puncture width, smaller but no more crowded in apical areas except narrow reflexed border impunctate; surface shiny. Sterna 2-5 with scattered punctures except apical areas impunctate and slightly reflexed. Sternum 6 flat with apical margin straight or gently concave.

Genital capsule and sterna 7 and 8 (Figs. 55-59) similar to those of accepta but note: emarginate apex of sternum 7, broad and only slightly emarginate apex of sternum 8, hairs of gonoforceps.

VESTITURE. Hair generally ochraceous (one specimen rather reddish on thoracic dorsum). Distribution and type of hairs as in accepta.

TYPE MATERIAL. The lectotype (PANS) male (No. 2159) of reflexa and the lectotype (PANS) female (No. 2158) of permitis were collected in Texas.

DISTRIBUTION. A. reflexa has been taken only a few times in Texas and Mexico. The collection data are given below in lull.

TEXAS: Fort Davis. 1 male, October 7, 1930, G. F. Englehardt. Lampasas (8 miles N. of highway U.S. 190 on the Lampasas River). 1 female, October 8, 1955, Mathews. Six females and four males (paratypes) labeled "Tex" and "Belfrage collection." Mexico. CHIHUAHUA: Minaca (22 miles S.). Males, August 23, 1950, Ray F. Smith. JALISCO: Guadalajara (15 miles N.W.). 1 male, September 13, 1938, L. J. Lipovsky. MICHOACAN: Carapan. 1 male September 1, 19622, on Crotolaria longirostrata, D. H. Janzen.

Variation. The male from Fort Davis differs from the Mexican specimens by having large yellow parocular areas (extending up to the antennal fossae), the clypeus almost entirely yellow, and the mandible yellow in median half. The Guadalajara males and the Carapan males differ from die other three in lacking parocular yellow areas. All of the males studied are very similar in structure and sculpturing.

Names
Scientific source:

References
Andrena reflexa Cresson, 1872, Transactions of the American Entomological Society, vol. 4, p. 256; Howard, 1905, The Insect Book, pi. Ill, fig. 2.

Andrena permitis Cresson, 1872, Transactions of the American Entomological Society, vol. 4, p.

257 (new synonymy).

Andrena (Pterandrena) reflexa: Lanham, 1949, University of California Publications in Entomology, vol. 8, p. 200.

Andrena (Pterandrena) permitis: Lanham, 1949, University of California Publications in Entomology, vol. 8, p. 200.

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