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Andrena scutellinitens Viereck, 1917
Andrena (Andrena) scutellinitens Viereck, 1917

Life   Insecta   Hymenoptera   Apoidea   Andrenidae   Andrena
Subgenus: Cnemidandrena

Andrena scutellinitens FEM CFP
© Copyright Laurence Packer 2014 · 7
Andrena scutellinitens FEM CFP

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Andrena scutellinitens MALE CFP
© Copyright Laurence Packer 2014 · 7
Andrena scutellinitens MALE CFP
Andrena scutellinitens, face
Smithsonian Institution, Entomology Department · 1
Andrena scutellinitens, face

Andrena scutellinitens, side
Smithsonian Institution, Entomology Department · 1
Andrena scutellinitens, side
Andrena scutellinitens, top
Smithsonian Institution, Entomology Department · 1
Andrena scutellinitens, top

Andrena scutellinitens, face
Smithsonian Institution, Entomology Department · 1
Andrena scutellinitens, face
Andrena scutellinitens, side
Smithsonian Institution, Entomology Department · 1
Andrena scutellinitens, side

Andrena scutellinitens, top
Smithsonian Institution, Entomology Department · 1
Andrena scutellinitens, top
Overview
Reprinted by permission of the Regents of the University of California from: Donovan, B. J. 1977. A revision of North American bees of the subgenus Cnemidandrena (Hymenoptera: Andrenidae). University of California Publications in Entomology: 1-107.

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Andrena scutellinitens can be distinguished from other species in this group by the much more protuberant clypeus, and by the metasomal apical fasciae which are often light yellow, rather than white as in A. robervalensis and A. specularia.

FEMALE. MEASUREMENTS AND RATIOS. N = 20; length, 7.8-8.9 mm; width, 2.2-2.8 mm; wing length, M = 2.54 0.104 mm; FL/FW, M = 1.09 0.104; FOVL/FOVW, M = 3.10 0.211.

INTEGUMENTAL COLOR. Black except as follows: antenna medium brown ventrally beyond second flagellar segment; mandible with apical half bright red; tegula, wing veins and pterostigma light yellow-brown; wing membrane hyaline except for medium darkening apically, darkening most pronounced just beyond marginal cell; tibial spurs nearly white; tarsal claws with basal half yellow, apical half red; metasomal terga and sterna with median width of apical margin hyaline; pygidial plate red.

STRUCTURE. Scape equal in length to flagellar segments 1 to 3; flagellar segment 1 equal to flagellar segments 2 plus 3. Eye less than 4 times longer than wide with inner margin diverging above except for upper end incurved. Malar space 7 times wider than long. Mandible in repose extends for one-fourth its length beyond middle of labrum; notch of mandible tooth slightly less than half way between middle of labrum and mandible apex. Galea moderately tessellate, lateral surface equal in width to dorsal surface. Maxillary palpus reaching just to apex of galea, segmental ratio 0.8:0.9:0.9:0.8:0.7:0.7. Labial palpus with basal segment moderately flattened laterally, curved, apical segment attached subapically, segmental ratio 1.3:0.8:0.7:0.7. Labral process flat, 3 times wider than long, median emargination broad, depth one-third length of labral process. Clypeus strongly protuberant, extending below eyes for less than one-half its length, dorsal and lateral periphery shagreened, remaining area with large punctures separated by less than 1 diameter except for large, ventral median area with punctures widely separated, leaving an irregular impunctate area; non-shagreened area shiny. Supraclypeal area densely shagreened with crowded, medium-sized punctures. Genal area 1.5 times wider than eye, with small punctures separated by 3 diameters, one-fourth of area near eye smooth, shiny, remainder shagreened, dull. Vertex above lateral ocellus equals more than 1 ocellar diameter, with moderate, irregular tessellation obscuring medium-sized, deep punctures separated by 1 diameter or less. Supra-antennal area with longitudinal rugulae and irregular punctures between the rugulae. Facial fovea rounded above and below, broad above, narrowing below, reaching above to a line above median ocellus, below to a line equidistant between upper limit of clypeus and lower edge of antennal fossa.

Pronotum with humeral angle well developed, humeral ridge moderately developed below, but broadly depressed medially where crossed by pronotal suture; pronotum with moderate, reticular shagreening throughout. Scutum with light tessellation in anterior third, remainder nearly smooth with large, shallow punctures separated by 1 to 2 diameters, punctures tessellate internally. Scutellum with anterior two-thirds polished, shiny, remainder lightly tessellate, dull. Metanotum, dorsoposterior face of propodeum with moderate tessellation obscuring medium-sized punctures, punctures crowded on posterior margin of metanotum, 3 diameters apart on dorsoposterior face of propodeum. Propodeal enclosure with irregular rugulae except for a small, narrowed posterior area with moderate tessellation. Mesepisternum densely shagreened with medium-sized punctures separated by 2 to 3 diameters. Metepisternum with very light, reticular shagreening, shiny. Corbicular area of propodeum with reticular shagreening and widely spaced, medium-sized punctures in anterodorsal third. Metabasitarsus widest in proximal half, 3.5 times longer than wide, outer edge nearly straight, inner edge convex. Mesobasitarsus 2.3 times longer than wide, somewhat less than one-third wider than metabasitarsus. Metatibia nearly 4 times longer than wide, more than two-thirds wider than metabasitarsus. Wing with pterostigma 4 times longer than wide, vein 1st m-cu meets second submarginal cell one-third distance from apex.

Metasomal terga 1 to 4 with discs finely tessellate anteriorly, becoming shagreened posteriorly; hyaline area nearly shagreened basally, smooth apically. Tergum 1 with disc with small punctures separated by 3 diameters except closer posteriorly; terga 2 to 4 with very small, indistinct punctures on disc separated by 3 to 4 diameters, becoming longer and separated by 1 diameter posteriorly. Tergum 5 with weak, median longitudinal ridge; with reticular shagreening and large punctures separated by 1 diameter or less. Pygidial plate recurved, rounded apically, whole surface shagreened, median longitudinal third and sides raised. Sternum 1 with apical margin with a very shallow, median emargination; sterna 1 to 6 shagreened except for large, nearly smooth, central basal areas on sterna 3 to 6. Sternum 1 with medium-sized punctures separated by 2 diameters, sterna 2 to 6 with small punctures on shagreened areas separated by 2 to 3 diameters.

VESTITURE. Facial hair except for facial fovea light yellow; clypeus broadly naked medioventrally, hairs becoming more numerous toward dorsolateral periphery; facial fovea with white-brown hairs. Mesosoma with yellow hairs dorsally, hairs more white ventrally; scutum with short hairs medially, scutellum with anterior half naked; propodeal corbicula with long, widely spaced hairs in anterodorsal third. Trochanteral floccus imperfect. Legs with hairs to femur white except metafemur with apical fimbria dark brown; tibial and tarsal hairs golden, except outer basal hairs of tibial scopa light brown to black.

Metasomal terga 1 to 4 with apical fasciae of dense, nearly appressed white-yellow hairs, obscuring less than half terga; fascia of tergum 1 narrower with hairs less appressed than fasciae of terga 2 to 4. Tergum 1 with long, erect, yellow hairs laterally and anteriorly on disc; median area nearly naked; tergum 2 with apical fascia wider medially with anterior hairs erect but much shorter than those on tergum 1. Terga 3 to 4 with very fine, short, sparse, pale hairs in about median thirds. Tergum 2 with extremely short, erect, brown-black hairs laterally, tergum 3 similar but brown-black hairs longer but still short and present on about lateral thirds. Tergum 5 obscured by long, reclining, brown to black hairs. Sternum 1 with erect, yellow hairs throughout; sterna 2 to 5 with subapical fimbriae of long, nearly erect, white-yellow hairs in about lateral thirds, and shorter brown hairs laterally on discs. Sterna 2 to 6 with apical fringes of short, nearly appressed, white hairs.

MALE. MEASUREMENTS AND RATIOS. N = 20; length, 5.9-7.8 mm; width, 1.7-2.2 mm; wing length, M = 2.14 0.134 mm; FL/FW, M = 1.09 0.110;FSl/FS2, M = 1.43 0.060.

INTEGUMENTAL COLOR. Black except as follows: antenna slightly lighter ventrally beyond first flagellar segment; mandible with apical third deep translucent red; tibial spurs nearly white; tarsi brown-yellow, basal half of tarsal claws yellow, apical half red; metasomal terga broadly hyaline apically; sterna more narrowly hyaline.

STRUCTURE. Scape twice as long as flagellar segment 1; flagellar segment 2 two-thirds length of flagellar segment 1. Eye less than 3.5 times longer than wide, with inner margins diverging above except for upper ends subparallel. Malar space, mandible and galea as in female except mandible with out ventrobasal lamella. Maxillary palpus with almost whole of apical segment projecting beyond apex of galea, segmental ratio 0.6:0.9:0.7:0.5:0.7:0.7. Labial palpus as in female except segmental ratio 1.3:0.6:0.7:0.9. Labral process moderately recurved, twice as wide as long, median emargination deep, depth half length of labral process. Clypeus moderately protuberant, extending below eyes for half its length, with large punctures, crowded on upper periphery but separated by 2 or more diameters medioventrally, surface shiny. Supraclypeal area as in female except shagreening weak. Gena 1.3 times wider than eye, rounded posteriorly to weakly angulate opposite mid-point of eye, with median punctures separated by 1 diameter; posterior half shagreened; anterior half shiny. Vertex as in female. Supra-antennal and parocular areas with longitudinal rugulae and large, irregular interrugal punctures.

Pronotum with humeral angle and ridge moderately developed, humeral ridge not depressed where reached by weak pronotal suture; posterior margin above pronotal suture expanded laterally; whole area strongly shagreened except shagreening weak and surface nearly shiny below pronotal suture. Remainder of thorax as in female except as follows: unshagreened areas of scutum, scutellum, shinier; propodeum except propodeal enclosure densely and roughly shagreened, with medium-sized punctures separated by 1 diameter. Mesepisternum with large, deep punctures separated by 1 diameter, whole area with dense shagreening, dull. Metepisternum impunctate, polished.

Metasomal terga 1 and 2 moderately shagreened; terga 3 to 5 smooth, terga 1 to 5 with small punctures separated by 2 diameters, punctures slightly larger on tergum 5. Sternum 1 with deep, V-shaped, median apical emargination; sterna 1 to 6 shagreened throughout, sternum 1 with medium-sized punctures separated by 1 diameter, sterna 2 to 6 with smaller punctures separated by 1 to 2 diameters. Sternum 7 (fig. 48) with basal apodemes narrowing very slightly and rounded basally, apex with small, shallow, median emargination and areas lateral to emargination with long hairs. Sternum 8 (fig. 49) with apex and neck region much longer than basal region; apex not markedly expanded; neck with long hairs. Gonoforceps (figs. 45 to 47) in dorsal view ridged longitudinally. Penis valves in dorsal and ventral views with a very weak, lateral expansion; apex rounded, not lobed, in lateral view strongly bent downwards.

VESTITURE. Yellow-white throughout except as follows: clypeus with abundant pubescence but pubescence less abundant medially. Scutum with moderately long, erect, sparse hairs, scutellum naked except long, erect hairs on posterior margin. Apical fascia of tergum 1 of long, yellowish hairs appressed posteriorly, nearly erect anteriorly; fascia less compact than remaining fasciae; disc of tergum 1 with long, erect hairs. Terga 2 to 5 with fasciae of short, nearly appressed yellowish hairs, obscuring about one-half to one-third of terga. Disc of tergum 2 with short, erect, white hairs, discs of terga 3 to 5 with short, erect, brown hairs, tergum 6 with yellow-brown, posteriorly inclined pubescence obscuring whole surface. Sternum 1 with long, erect hairs, sterna 1 to 5 with prominent, dense, subapical fimbriae of long hairs, remainder of sterna with shorter, erect hairs.

VARIATION. In females the scutum varies from moderately tessellate throughout to the posterior half smooth, shiny. Females from the southern half of the range of this species have the facial foveal hairs white, while females from the northern half have the facial foveal hairs brown-white or brown. In both sexes terga 2 to 4 may have numerous brown, erect hairs, or brown hairs may be almost absent. In females the vestiture of tergum 5 varies from black to light brown-yellow, but most specimens are brown.

Names
Scientific source:

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FamilyScientific name @ source (records)
Asteraceae  Achillea millefolium @ BBSL (1)

Anaphalis margaritacea @ BBSL (1)

Aster foliaceus @ BBSL (1)

Aster integrifolius @ BBSL (2)

Erigeron speciosus @ BBSL (14)

Senecio serra @ BBSL (1)

Solidago sp @ BBSL (1)

Solidago @ AMNH_BEE (4)
_  Withheld @ BBSL__YOSE (8); BBSL (5)

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Updated: 2021-05-13 13:33:44 gmt
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