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Reprinted with permission of the University of Nebraska State Museum from:
LaBerge, W. E. 1967. A revision of the bees of the genus Andrena of the Western Hemisphere. Part I. Callandrena (Hymenoptera: Andrenidae). Bulletin of the University of Nebraska State Museum 7: 1-316.
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This small Mexican species is closely related to A. uyacensis Cockerell from Honduras. The female of sodalis can be told from that of uyacensis by the mesoscutum being entirely opaque with the punctures almost completely obscured by fine dense tessellation. In uyacensis females the mesoscutum has distinct punctures with the surface at least moderately shiny posteromedially. A. sodalis may1 be no more than a race of uyacensis, but this cannot be verified until males of the later have been discovered and compared with the males of sodalis.
FEMALE. MEASUREMENTS AND RATIOS. N = 8; length, 10.5-11.5 mm; width, 3.0-3.5 mm; wing length, M = 3.44 ą 0.147 mm; FL/FW, M = 1.17 ą 0.012; FOVL/FOVW, M = 2.93 ą 0.094.
INTEGUMENTAL COLOR. Black except as follows: mandible with apical third rufescent; flagellar segments 3-10 dark reddish-brown to brownish-black below; tegulae testaceous to translucent brown; wing membranes moderately infumate, brownish, especially apically, veins dark brown; terga 1-4 with apical areas translucent, yellowish at extreme apex, infumate basally; sterna 2-5 hyaline apically, yellow; distitarsi rufescent; hind basitarsi orange at extreme base to orange in basal half; hind tibia usually piceous, reddened in a few specimens.
STRUCTURE. Antennae as in uyacensis. Eye almost four times as long as broad, inner margins parallel. Malar space, mandible and galea as in simulata. Maxillary palpus as in simulata but segmental ratio about 1.2:1.0:0.6:0.7:0.5:0.7. Labial palpus as in simulata but segmental ratio about 3.1:1.0:0.9:1.1. Clypeus as in simulata but distinctly flattened medially, with distinct median impunctate area, coarse lateral punctures slightly elongate, and surface moderately dulled by coarse reticular shagreening. Supraclypeal area and genal yeas as in simulata. Vertex short, above lateral ocellus distinctly but slightly shorter than one ocellar diameter, sculptured as in Uyacensis. Face above antennal fossae and facial fovea as in uyacensis but facial fovea separated from lateral ocellus by one-half to three-fourths of an ocellar diameter.
Thoracic sculpturing as in uyacensis except mesoscutum and scutellum without distinct punctures, surfaces opaque, fine dense tessellation obscuring any punctation and dulling surfaces. Tegulae, parapsidal lines, wing venation, middle basitarsus, claws and spurs in uyacensis.
Metasomal terga sculptured as in uyacensis. Pygidial plate as in uyacensis. Sterna sculptured as in uyacensis.
VESTITURE. Generally as in simulata with the following differences: tergum 2 with apical pale fascia broadly interrupted medially, resulting lateral fasciae each equals one-fourth or less width if tergum; tergum 3 with apical pale fascia narrowly interrupted medially; terga 5 and 6 and sternum 6 occasionally with brown tarsi, including basitarsi, with dark brown hairs; hind tibial hairs occasionally brown in part or entirely. Propodeal corbicula and trochanteral flocculus as in uyacensis. Male. MEASUREMENTS AND RATIOS. N = 5; length, 9-11 mm; width, 2.0-2.5 mm; wing length, M = 3.26 ą 0.473 mm; FL/FW, M = 1.20 ą 0.007; FS1/FS2, M = 1.94 ą 0.086.
INTEGUMENTAL COLOR. Black except as follows: mandible with apical margin and small maculae slightly mesad and below tentorial pits dark brown; flagellar segments 2-11 dark brown below or wholly black; tegulae testaceous to translucent brown; wing membranes hyaline, moderately infumate especially apically, veins dark brown to red; terga 1-5 with apices often broadly hyaline, occasionally infumate except at extreme apices; sterna 2-5 with apices narrowly hyaline, more or less rufescent basally; distitarsi orange to red; hind basitarsi orange; hind tibiae orange with a small oval brown spot just below middle or dark brown with orange near base and at apex, or entirely piceous.
STRUCTURE. Antennae long, surpassing scutellum in repose; scape equal in length to flagellar segments 1 plus 3; segment 1 distinctly shorter than segments 2 plus 3; segment 2 shorter than 3; segments 3-11 longer than broad. Eye slightly less than three times as long as broad, inner margins converging strongly towards mandibles. Malar space linear. Mandible bidentate; outer mandible extends beyond middle of labrum by about one-fourth its length in repose. Galea with lateral portion equal in width to about halt of dorsal portion, turned down evenly at sides, narrow; surface dull, finely tessellate. Maxillary palpus slightly shorter than galea when both extended forward, segmental ratio about 1.0:0.8:0.7:0.7:0.5:0.8. Labial palpus entirely flattened, first segment curved rather abruptly near base, segmental ratio about 2.4:1.0:0.7:0.9. Labial process trapezoidal in outline, about two and one-half times as broad as long, extremely weakly and broadly emarginate, if at all. Clypeus flattened medially, with large median impunctate area, elsewhere with small round punctures irregularly separated by hall to one puncture width, surface shiny and unshagreened except at extreme periphery or with delicate reticular shagreening scarcely dulling surface. Supraclypeal area with minute punctures and fine tessellation dulling surface. Genal area subequal to eye in profile, with small round punctures separated by half to one puncture width or slightly more, dulled by fine reticular shagreening except near eye margin. Vertex above lateral ocellus equals about one ocellar diameter or slightly more, with crowded punctures above ocelli, scattered punctures laterally, surface moderately dulled by tessellation. Face above antennal fossae with longitudinal rugulae, interrugal spaces punctate, surface moderately dulled by fine reticular shagreening.
Pronotum normal, with sparse minute punctures and coarse reticular shagreening dulling surface. Mesoscutum opaque, with extremely shallow punctures obscured by coarse regular tessellation; parapsidal line shorter than from its posterior end to margin oĢ scutum. Tegula normal, impunctate. Scutellum sculptured like mesoscutum. Metanotum opaque, dulled by minute contiguous punctures and fine tessellation. Propodeum with dorsal enclosure dulled by fine tessellation and slightly roughened basally and medially by irregular, indistinct rugulae; dorsolateral, posterior and upper part of lateral surfaces with large, extremely shallow, obscure punctures and regular coarse tessellation; lower lateral surfaces moderately shiny with coarse reticular shagreening. Mesepisternum with large, round, shallow, obscure punctures and coarse tessellation dulling surface. Metepisternum with lower portion like lower lateral propodeal surface, upper third minutely punctate and tessellate. Fore wing venation as in simulata but in one paratype second submarginal cell receives vein 1st m-cu near middle. Claws and tibial spurs normal.
Metasomal terga 1-5 with extremely narrow apices impunctate, basally with small round punctures which on tergum 1 separated by one to two puncture widths, on succeeding terga by hall to one puncture width; surfaces dulled by moderately coarse, reticulotransverse shagreening except extreme apices. Pseudopygidial area bare, extremely narrow, V-shaped. Sterna 2-5 with hyaline margins impunctate, impunctate basally and with scattered punctures in area near hyaline margins. Sternum 6 flat, shallowly emarginate medially. Genitalia and sterna 7 and 8 (Figs. 214-218) similar to those of simulata except as follows: gonoforceps longer, blunter, less hairy, inner-apical margins truncate; penis valves with basad lamellae larger, raised above general level of valves; dorsal lobe of gonocoxite much as in tegularis; sternum 7 with more distinct apical lobes; sternum 8 with apex enlarged, suboval, shallowly emarginate.
VESTITURE. Generally ochraceous, brighter on thoracic dorsum and vertex (orange in one male). Terga 2-5 with weak apical fasciae of pale decumbent hairs; tergum 2 with fascia interrupted medially. Sterna 3-5 with well-developed subapical fimbriae of long suberect pale hairs. Inner surfaces basitarsi with hairs pale golden.
TYPE MATERIAL. The holotype female (KMNH No. 17-a-2663) of sodalis was collected at Oaxaca, Mexico.
DISTRIBUTION. A. sodalis is known only from central Mexico. Nine female and thirteen male specimens were examined in addition to the holotype. The collection data is given in full for these specimens: HIDALGO: El Chico. 1 male, September 23, 1938, L. J. Lipovsky. Ozumbilla. 1 male, October 30, 1957, R. and K. Dreisbach. Pachuca. 1 male, October 30, 1957, R. and K. Dreisbach. MEXICO: Chapingo. 1 female. Mexico City. 1 female, 1 male, September 25, 1957, R. and K. Dreisbach; 1 male, September 7, 1957 (40 miles E.), H. A. Scullen; 1 male, September 10, 1957 (18 miles S.), H. A. Scullen. Atlacomulco. 1 male, August 15, 1964 (22 miles N.), C. D. Michener and party. MICHOACAN; Morelia. 1 male, September 22, 1957, R. and K. Dreisbach; 1 female, February 12, 1952. PUEBLA: Amecameca. 1 male, September 25, 1957, R. and K. Dreisbach. TLAXCALA: Apizaco. 4 males, on Argemone sp., August 19, 1962 (10 miles E.), R. B. Roberts. VERA CRUZ: Rio Blanco. 6 females, November 13, 1957, R. and K. Dreisbach.
REMARKS. The female from Morelia, Michoacan, is darker than the other females. It has entirely brown scopal hairs and brown hairs on terga 5 and 6 and sternum 6. It is possible that this dark female represents another species but it is included here as a color variant of A. sodalis as no structural differences have been found to differentiate it. This dark specimen is in the collection of Mr. P. H. Timberlake.
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Andrena sodalis Smith, 1879, Descriptions of New Species of Hymenoptera in the British Museum, p. 52; Morice and Cockerell, 1901, Canadian Entomologist, vol. 33, p. 152; Cockerell, 1906, Psyche, vol. 13, p. 8.
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Updated: 2024-04-29 18:32:14 gmt
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