Reprinted with permission of the American Entomological Society from:
LaBerge, W. E., Bouseman, J. K. 1970. A revision of the bees of the genus Andrena of the Western Hemisphere. Part III. Tylandrena. Transactions of the American Entomological Society 96: 543-605.
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This species is extremely closely related to A. layiae from which it can be separated in both sexes by the higher vertex, the larger size, and the less punctate metasomal terga. So much like layiae is sublayiae that the description given above for layiae applies to sublayiae except for a few details noted below.
FEMALE. MEASUREMENTS AND RATIOS. — N = 20; length, 12-13 mm; width, 3.5-4.0 mm; wing length, M = 4.64 ± 0.124 mm; FL/FW, M = 0.88 ± 0.004; FOVL/FOVW, M = 4.14 ± 0.063.
INTEGUMENTAL COLOR. — Generally as in layiae but terga slightly less metallic and terga 24 with apical areas usually reddish-brown at least along margins.
STRUCTURE. — Antennal scape about as long as first four flagellar segments or slightly shorter; flagellar segments as in layiae. Eyes as in layiae but inner margins parallel. Malar space, mandibles, and galeae as in layiae. Maxillary palpus as in layiae but segmental ratio about 1.0:0.9:0.8:0.8:0.7:0.8. Labial palpus as in layiae but ratio about 1.0:0.8:0.5:0.5. Clypeus, supraclypeal area, genal area and facial foveae as in layiae. Face above antennal fossae with fine longitudinal rugulae (much less coarse than in layiae) and sparse interrugal punctures. Vertex above lateral ocellus equals one and one-fourth to one and one-half ocellar diameters, sculptured as in layiae.
Pronotum as in layiae. Thoracic sculpturing as in layiae but mesepisternal punctures sparser and smaller. Wing venation as in layiae. Terga 1-5 with apical areas impunctate medially and laterally with punctures obscure. Terga 2-4 with basal area punctures present but shallow and obscure except under high magnification and proper lighting. Tergal surfaces dulled by fine, dense, reticular shagreening. Pygidial plate and sterna as in layiae.
VESTITURE. — Generally as in layiae but usually darker ochraceous especially on thoracic dorsum. MALE. MEASUREMENTS AND RATIOS. — N = 3; length, 11-12 mm; width, 3.0-3.5 mm; wing length, 4.15-4.35 mm; FL/FW, 0.77-0.82; FS1/FS2, 1.50-1.70.
INTEGUMENTAL COLOR. — As in layiae except as follows: tip of mandible rufescent; terga with apical margins reddish-brown.
STRUCTURE. — As in layiae except as follows: vertex above lateral ocellus equals about one and one-half ocellar diameters or very little less; mesoscutum and scutellum slightly shiny medially; terga slightly less punctate, shinier.
Terminalia as figured (Figs. 71-75).
VESTITURE. — As in layiae.
REMARKS. — It should be noted that the ranges, as now known, of A. layiae and A. sublayiae appear to be completely allopatric (Fig. 3). It is quite possible that sublayiae is merely a geographical race of layiae occupying the coastal area of central California. However, no intermediate forms are as yet known and the authors believe it serves to keep this problem in the foreground by naming sublayiae as a species at this time. Three of the sublayiae paratypes (from Salinas, Monterey County, California) are also paratypes of layiae, Timberlake having had a mixed series in the few females he had been able to study before describing layiae.
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