Reprinted with permission of the University of Nebraska State Museum from:|
LaBerge, W. E. 1967. A revision of the bees of the genus Andrena of the Western Hemisphere. Part I. Callandrena (Hymenoptera: Andrenidae). Bulletin of the University of Nebraska State Museum 7: 1-316.
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This is medium-sized to large bee which is closely related to A. berkeleyi Viereck and Cockerell. The female of utahensis is similar to berkeleyi and differs from gardineri in the high vertex, the lack of the first tergal apical fascia, the broadened middle basitarsus and the sometimes red abdomen. The female of utahensis differs from that of berkeleyi in the shorter and denser mesoscutellar hair (almost as in A. biscutellata Viereck) and the less dense tergal punctures. The male of utahensis differs from that of gardineri in the higher vertex, the larger size and the sixth sternum being reflexed only in the apicolateral angles. The male of utahensis can be distinguished from that of berkeleyi by the less dense tergal punctures and the longer sternal apical fimbriae.
FEMALE. MEASUREMENTS AND RATIOS. — N = 10; length, 11-14 mm; width, 3.0-4.5 mm; wing length, M= 4.62 ±0.194 mm; FL/FW, M = 1.05 ± 0.024; FOVL/FOVW, M = 2.17 ± 0.029.
INTEGUMENTAL COLOR. — Generally as in berkeleyi except as follows: specimens with red abdomens with tergum 1 brown basally and terga 5 and 6 reddish-brown basally; hind basitarsi and tibiae piceous; wing membranes extremely slightly infumate apically.
STRUCTURE. — Antennae as in gardineri. Eye slightly more than four times as long as broad, inner margins converging slightly towards mandibles. Malar space, mandible and galea as in gardineri. Maxillary palpus as in gardineri but segmental ratio about 1.0:1.0:0.7:0.7:.05:0.7. Labial palpus as in gardineri but segmental ratio about 2.1:1.0:0.8:0.9. Labral process as in gardineri. Clypeus slightly flattened medially, protruding beyond lower ends of eves by one-third or less of length of clypeus; with round punctures separated mostly by half a puncture width or slightly more, smaller posteriorly, surface shiny, unshagreened or delicately so near periphery. Supraclypeal area with minute crowded punctures, surface moderately dulled by fine shagreening. Genal area less than one and one, half but more than one times width of eye in profile; sculptured as in gardineri but shiny and unshagreened posteriorly. Vertex above lateral ocellus equals slightly more than one ocellar diameter; with round punctures crowded above ocelli, posteriorly and above facial foveae, surface moderately dulled by reticular shagreening. Face above antennal fossae as in berkeleyi. Facial fovea as in berkeleyi but closer to compound eye in upper half.
Thoracic sculpturing and form as in gardineri except as follows: mesoscutal and scutellar punctures separated mostly by half a puncture width or less except slightly sparser posteromedially on mesoscutum, surfaces shiny to moderately shiny, shagreening delicate or absent; propodeal dorsal enclosure with rugulae fine and obscure. Middle basitarsus distinctly broadened medially, broader than hind basitarsus; without apical spine. Venation as in gardineri but second submarginal cell longer. Claws and tibial spurs normal.
Metasomal terga 1-5 with small round crowded punctures in apical areas except at extreme apices. Terga 1-4 with basal area punctures as in gardineri, surfaces shiny, unshagreened or extremely delicately so. Pygidial plate V-shaped, sides often slightly concave, with truncate apex. Sterna 2-5 as in gardineri but surfaces shiny, shagreening delicate.
VESTITURE. — Generally as in berkeleyi except terga 2-4 with apical pale fasciae uninterrupted medially or at most narrowly interrupted on tergum 2 alone.
MALE. MEASUREMENTS AND RATIOS. — N=9; length, 10-13 mm; Width, 2.5-4.0 mm; wing length, M = 4.28 ± 0.264 mm; FL/FW, M= 1.10 ±0.011; FS1/FS2, M = 2.55 ± 0.087.
INTEGUMENTAL COLOR. — Black except as follows: mandible with apical third rufescent; flagellar segments 3-11 dark reddish-brown below; clypeus yellow except small brown maculae below and mesad of tentorial pits and broad apical margin; parocular area with large yellow macula but not extending up to level of posterior clypeal margin; supraclypeal area usually with small median diffuse yellow spot; tegula dark reddish-brown; wing membranes hyaline, slightly infumate apically, veins dark reddish-brown to black; terga 1-6 broadly hyaline apically, narrowly reddened immediately basad of hyaline apices; sterna 2-5 narrowly hyaline apically, yellow; tarsi piceous; tibial spurs testaceous.
STRUCTURE. — Antennae as in gardineri. Eye about three and one-fourth times as long as broad, inner margins distinctly converging towards mandibles. Malar space, mandible and galea as in female. Maxillary palpus as in female but segmental ratio about 1.9:1.0:0.6:0.7:0.4:0.7. Labial palpus as in female but segmental ratio about 1.5:1.0:0.7:0.6. Labral process bidentate. Clypeus as in female but not flattened medially. Supraclypeal and genal areas as in female. vertex above lateral ocellus equals about one and one-half ocellar diameters, sculptured as in female. Face above antennal fossae as in female but punctures in rows with weak longitudinal rugulae between rows.
Thoracic structure and sculpturing as in female except as follows: propodeal dorsal enclosure with rugulae more distinct at extreme base; propodeal lateral surface dulled by fine tessellation. metasomal terga sculptured as in female but basal area punctures lightly larger and more crowded. Pseudopygidial area not evident. Sterna 2-5 sculptured as in female. Sternum 6 reflexed apicolaterally forming strong lateral teeth, medially almost flat, with broad V-shaped median emargination.
Genitalia and sterna 7 and 8 (Figs. 292-296) similar to those of balsamorhizae and berkeleyi but not sternum 8 with entire apex and longer hairs.
VESTITURE. — Generally white to pale ochraceous, darker on vertex and thoracic dorsum. Mesoscutal hairs moderately long, erect not hiding surface as in female. Terga 1-5 with distinct apical pale fasciae but that of tergum 1 interrupted medially. Inner surfaces hind tarsi yellowish. Sterna 3-5 with moderately long subapical fimbriae much as in gardineri and longer than in berkeleyi.
TYPE MATERIAL. — The holotype (USU) female, allotype (USU) male, and one paratype (USU) female, were collected by G. E. Bohart at Duchesne, Utah, June 11, 1963, on Coreopsis sp. Ten additional female and 8 male paratypes (USU; UCNH; UCB; USNM; INHS; CAS) are as follows: ARIZONA: Flagstaff. 1 male, June 11, 1909, on Iris sp., V. C. Prate. Grand Canyon. 5 females, 1 male, June 5, 1940, G. K. Bohart. Kayenta (28 miles W.). 1 female, June 10, 1933, H. N. Hultgren; (19 miles S.W.) 1 female, June 20, 1933, H. N. Hultgren. Show Low. 2 females, June 2, 1946, R. M. Bohart. COLORADO: Bondad. 1 female, June 27, 1919, F. E, Lutz. UTAH: Duchesne (3 miles W.). 2 males, May 20, 1962, on Geraea sp., R. W. Thorp; 4 males, May 20, 1962, . W. MacSwain. Myton. 1 female, June 11, 1963, Brumly.