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© Copyright source/photographer
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Andrena wheeleri, f, face, Crayson Co., VA |
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Andrena wheeleri, f, left, Crayson Co., VA |
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© Copyright Laurence Packer 2014
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Andrena wheeleri FEM CFP comp |
© Copyright Laurence Packer 2014
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Andrena wheeleri MALE CFP comp |
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© Copyright source/photographer
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Andrena wheeleri, M, back, North Carolina, Buncombe County |
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Andrena wheeleri, M, back, North Carolina, Buncombe County |
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© Copyright source/photographer
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Andrena wheeleri, M, face, North Carolina, Buncombe County |
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Andrena wheeleri, M, side, North Carolina, Buncombe County |
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Andrena wheeleri, M, back, North Carolina, Buncombe County |
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Andrena wheeleri, F, back, North Carolina, Buncombe County |
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© Copyright source/photographer
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Andrena wheeleri, F, face, North Carolina, Buncombe County |
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Andrena wheeleri, F, side, North Carolina, Buncombe County |
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© Copyright source/photographer
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Andrena wheeleri, M, side, North Carolina, Buncombe County |
© David Biddinger, 2024
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Andrena wheeleri |
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Overview |
Reprinted with permission of the American Entomological Society from:
LaBerge, W. E. 1989. A revision of the bees of the genus Andrena of the Western Hemisphere. Part XIII. Subgenera Simandrena and Taeniandrena. Transactions of the American Entomological Society 115: 1-56.
Please report text errors to: leah at discoverlife dot org.
Andrena wheeleri Graenicher is a small species readily confused with A. nasonii Robertson. The female of wheeleri can be distinguished from that of nasonii by the shiny, punctate clypeus, the longer scopal hairs, and the lack of tergal fasciae. The male of wheeleri can be separated from that of nasonii by the slightly longer first flagellar segment and the lack of tergal fasciae. Both sexes of wheeleri have a flat mesopleural signum, unlike the tuberculate signum of nasonii.
FEMALE. MEASUREMENTS AND RATIOS. — N = 20; length, 7-9 mm; width, 2.5-3.0 mm; wing length, M = 3.04 ± 0.125 mm; FL/FW, M = 1.04 ± 0.008; FOVL/FOVW, M = 2.88 ± 0.032.
INTEGUMENTAL COLOR. — As in nasonii except as follows: wing veins often reddish brown; metasomal terga with apical areas slightly translucent, never hyaline apically.
STRUCTURE. — Antennal scape as long as first four and one-fourth flagellar segments; flagellar segment 1 as long as segments 2 and 3 or slightly shorter, segment 2 slightly shorter than 3, segment 3 about as long as broad. Eyes, malar space, mandibles and galeae as in nasonii. Maxillary palpus with segmental ratio about as 1.0:1.0:0.7:0.7:0.6:0.6. Labial palpus with segmental ratio about as 1.0:0.6:0.4:0.4. Labral process usually rounded, occasionally feebly emarginate apically, shiny; labrum apical to process with several cristae. Clypeus with round punctures separated by one to two or three puncture widths, midline impunctate, shiny except along posterior border. Supraclypeal area dull, with minute longitudinal rugulae. Face above antennal fossae, vertex and genal area as in nasonii.
Pronotum as in nasonii. Mesoscutum with punctures more sparse than in nasonii, separated mostly by three or more puncture widths, surface dulled by fine dense shagreening. Scutellum and metanotum similar. Propodeum sculptured as in nasonii but rugulae tend to be linear, parallel and more coarse, without transverse rugulae at apex. Wing venation as in nasonii.
Metasomal tergum 1 basal area punctures small, separated by about one puncture width, obscured by dense shagreening; apical area impunctate, shagreened. Terga 2-4 with basal area punctures indistinct, obscured by shagreening, sparse; apical areas impunctate, shagreened. Pygidial plate V-shaped, apex not rounded unless worn, internal raised triangular area distinct. Sterna as in nasonii.
VESTITURE. — Generally white to pale ochraceous, often darker on thoracic dorsum. Thoracic dorsum with hairs sparse; metasomal tergal fasciae absent; scopal hairs moderately long, only in apicomedian fourth of scopa or less are hairs extremely short; scopal hairs without brown or only in small area just below pygidial plate.
MALE. MEASUREMENTS AND RATIOS. — N = 20; length, 6.5-8.0 mm; width, 1.5-2.5 mm; wing length, M = 2.76 ± 0.130 mm; PL/ FW, M = 1.08 ± 0.005; FS1/FS2, M = 1.26 ± 0.026.
INTEGUMENTAL COLOR. — As in female but flagellar segments often reddish brown below.
STRUCTURE. — Antennal scape as long as first two and two-thirds flagellar segments; flagellar segment 1 slightly longer than segment 2 which is about as long as broad and shorter than segment 3. Mandibles, malar space, and galeae as in female. Maxillary palpus as in female but segmental ratio about as 1.0:1.0:0.7:0.7:0.7:0.6. Labial palpus as in female but ratio about as 1.0:0.6:0.5:0.6. Labral process as in female but usually emarginate slightly. Clypeus as in female but occasionally entirely shagreened but not completely obscuring punctures, usually moderately shiny at least in apical half. Supraclypeal area, face above antennal fossae, vertex and genal area as in female.
Pronotum as in female. Mesoscutum with punctures separated mostly by about three puncture widths, somewhat more distinct than in female, surface shagreened. Scutellum and metanotum similar. Propodeum sculptured as in female but rugulae usually less distinct. Metasomal terga virtually impunctate, dull, shagreened, basal area punctures absent or extremely shallow and sparse. Sternal basal areas with no punctures except in line near apical areas, shagreened. Sternum 6 shallowly emarginate apically. Terminalia as in Figures 11-15.
VESTITURE. — Pale to bright ochraceous, usually somewhat darker than female. Clypeal hairs dense but not entirely hiding surface; dorsal thoracic hairs more dense and longer than female; tergal fasciae absent; sternal fimbriae absent.
Reprinted with permission from: Mitchell, T.B. 1960. Bees of the Eastern United States. North Carolina Agricultural Experiment Station Technical Bulletin No. 141.
FEMALE. — Length 9 mm.; length and breadth of face subequal; clypeus broadly convex, projecting about one-third below suborbital line, somewhat shining medially but becoming dull and tessellate laterally, punctures rather fine and shallow, scattered and rather sparse medially, becoming quite close laterally; facial foveae quite broad, occupying above most of area between eyes and ocelli (as in nasonii, fig. 54), covered with silvery tomentum; space between margin of vertex and lateral ocelli nearly equal to their diameter; cheeks subequal to eyes in width, rounded posteriorly, rather dull, punctures obscure; malar space lacking; basal segment of flagellum subequal to 2nd and 3rd combined; process of labrum rather large, semicircular, median length about half the basal width; pubescence of head, thorax and legs ochraceous; thoracic integument dull, rather densely tessellate, but scutum somewhat shining toward posterior middle, punctures very fine, shallow and rather sparse, those on scutellum much closer, but very fine and obscure; pleura subrugose anteriorly, but otherwise impunctate; dorsal area of propodeum oblique, triangle very finely granular; propodeal corbicula rather short but well separated, with a distinct anterior fringe, pale ochraceous, trochanteral floccus well developed, of moderate length, more whitish-ochraceous; hind tibiae broadened apically, apex fully twice width of basitarsi, scopa quite dense, hairs short and simple, pale ochraceous; front and middle basitarsi very slightly narrower than their respective tibiae; 2nd submarginal cell considerably shorter than 3rd, receiving 1st recurrent about at middle; abdominal terga smooth, but rather dull, microscopically tessellate, very minutely and obscurely punctate, punctures rather close on discs, but lacking on the shallow, apical, impressed areas which occupy medially about one-third length of discs, becoming brownish-hyaline along rims, discal pubescence very sparse, obscure and entirely pale, fasciae not developed, tergum 5 with a brownish, apical fimbria.
MALE. — Length 7.5 mm.: clypeus broadly convex, about one-third below suborbital line, surface medially somewhat shining, becoming dull laterally, punctures rather close, deep and distinct on each side of an obscure, median, impunctate area, pubescence quite dense, obscuring surface details; space between margin of vertex and lateral ocelli slightly less than their diameter; cheeks subequal to eyes in width, rounded posteriorly, rather dull, punctures very obscure beneath rather dense pubescence; malar space lacking; basal segment of flagellum subequal to 2nd segment; process of labrum rather large, length about half basal width. apex rather broadly truncate; mandibles short, with a distinct, subapical, inner tooth, overlapping but very slightly; pubescence of head, thorax and legs entirely pale ochraceous; thoracic integument dull, quite densely tessellate, punctures hardly visible even on dorsum of thorax; dorsal area of propodeum more oblique, short, triangle very finely and obscurely granular; all basitarsi slender and elongate, considerably narrower than their respective tibiae; 2nd submarginal cell slightly shorter than 3rd, receiving 1st recurrent somewhat basad of middle; abdominal terga smooth but rather dull, microscopically tessellate, punctures exceedingly minute and obscure, hardly visible, apical depressed areas rather narrow and shallow, becoming brownish-hyaline along rims, discal pubescence thin and obscure, erect, entirely pale, fasciae not developed; apical portion of sternum 8 gradually narrowed from base, slightly broadened at the subtruncate apex, pubescence rather long and dense; penis valves very slender, very slightly broadened toward base, gonocoxites rather short, slightly dilated apically, gonocoxal lobes only very slightly produced, rather narrowly rounded
DISTRIBUTION. — Minnesota and Colorado to Maine, south to North Carolina; April, May and June.
FLOWER RECORDS. — Chrysanthemum leucanthemum and Rubus. Collected by Lovell on Sedum acre. Recorded also by Brittain & Newton on Centaurea, Cornus and Rhododendron.
This was thought by Graenicher to be oligotropic on the Umbelliferae. Cockerell described the subspecies pallidior as a race distinct from typical wheeleri and extending from Maine to Alberta. Specimens from Wisconsin and Colorado he assigned to the typical form. The chief differences he cited were the more converging facial foveae in pallidior and the much less pubescent (plumose?) hair. A considerable series of females from the North Carolina highlands seem to agree with the description of pallidior, as does also a specimen from New Hampshire. One of the North Carolina specimens was compared with the type of wheeleri and was thought to be identical. The desirability of further study of the material is indicated.
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Updated: 2024-04-24 04:17:05 gmt
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