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© Larry Thompson, 2007-2015
· 10 Colibri thalassinus, Green Violet-ear |
© Larry Thompson, 2007-2015
· 8 Thalurania colombica, Violet-crowned Woodnymph |
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© Larry Thompson, 2007-2015
· 10 Boissonneaua flavescens, Buff-tailed Coronet |
© Larry Thompson, 2007-2015
· 10 Chlorostilbon melanorhynchus, Western Emerald |
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© Larry Thompson, 2007-2015
· 9 Colibri thalassinus, Green Violet-ear |
© Larry Thompson, 2007-2015
· 8 Colibri thalassinus, Green Violet-ear |
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© Larry Thompson, 2007-2015
· 8 Colibri thalassinus, Green Violet-ear |
© Larry Thompson, 2007-2015
· 8 Colibri thalassinus, Green Violet-ear |
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© Larry Thompson, 2007-2015
· 9 Colibri thalassinus, Green Violet-ear |
© Larry Thompson, 2007-2015
· 9 Amazilia luciae, Honduran Emerald |
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© Larry Thompson, 2007-2015
· 10 Amazilia luciae, Honduran Emerald |
© Larry Thompson, 2007-2015
· 8 Coeligena phalerata, White-tailed Starfrontlet |
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© Larry Thompson, 2007-2015
· 6 Coeligena wilsoni, Brown Inca |
© Larry Thompson, 2007-2015
· 10 Coeligena torquata, Collared Inca |
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© Copyright Tom Stephenson, 2006-2011
· 0 Lophornis magnificus |
Following modified from Monterey Bay
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BIRD FAMILIES OF THE WORLD
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a web page by Don Roberson
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HUMMINGBIRDS
Trochilidae
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328 species in the New World
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DR personal total: 196 species (59%), 65 photos
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Colorful, combative, and energetic, hummingbirds are among the world's most popular birds. They are found only in the New World. People across North America attract them to their homes through hummingbird feeders and flowering gardens, and can be rewarded with bright migratory species, such as a male
Rufous Hummingbird
(left). Yet the true center of trochilid diversity is in the Neotropics, home to a dazzling array of species that range from tiny to surprisingly large, and with an assortment of head plumes, tail streamers, and bill shapes. Their ability to hover and fly backwards makes them unique. Further, their interdependence on and co-evolution with flowers provide a fascinating correlation and field for study. Because the sources of nectar are constantly changing as flowers bloom and fade, most hummingbirds live solitary lives, aggressively defending nectar sources from rivals. Despite the strong pressure to gather nectar rapidly, studies have shown that 70% of a hummer's time is spent doing little else than singing, self-preening, and sunbathing (Schuchmann 1999). In the two shots below, a male
White-necked Jacobin
sits near its favored feeder in Trinidad (below left) but immediately reacts when a White-chested Emerald tries to approach. Such interactions can lead to dramatic interspecific aerial chases. |
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A male
Magnificent Hummingbird
sitting in the shade (below left) can look dark and dingy. But when the hummer sits in the sun, and turns its head (below right), the observer is suddenly dazzled by a flash of spectacular glittering color. This species ranges from the southwest United States to Panama, and is a big hit at feeders from Ramsey Canyon, Arizona, to Savegre, Costa Rica. |
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Iridescent color among birds is one of nature's wonders, and hummingbirds display a full spectrum. For many species, iridescence is limited to a throat gorget, or a gorget and crown. North American examples include
Anna's Hummingbird
, the resident species throughout much of California (below, top row left) and
Costa's Hummingbird
of the southwestern deserts (below, top row right). On others the entire body is jeweled with waves of color, like on
Violet Sabrewing
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The Neotropics are inhabited by a host of hummingbirds. Colorful species buzz about rainforest trees and visit flowers at timberline on Andean volcanoes. The Caribbean islands have their own fine assortment, including
Purple-throated Carib
(right), this one on Dominica. In my meanderings around the Neotropics, there have been many wonderful species, such as the Horned Sungem
Heliactin cornuta
recently seen in southern Brazil. Three species have particularly impressed me with their unbelievable beauty, and all have been in the western Andes of Colombia or northwestern Ecuador. I have photos of none of these, but nothing can beat a Velvet-purple Coronet
Boissonneaua jardini
, or a Purple-backed Thornbill
Ramphomicron microrhynchum
, or a Rainbow-bearded Thornbill
Chalcostigma herrani
. Try to see one if you have the chance. The names of hummingbirds alone set one's head spinning: sunangels, comets, pufflegs, fairies, starfrontlets, brilliants, mountain-gems, coquettes .... |
Unusual beaks and strange tails are part of the world of tropical hummingbirds. The bill of
White-tipped Sicklebill
(below left) is adapted best for
Heliconia
flowers. By far the longest bill, proportionate to size, belongs to the ridiculously unbalanced
Sword-billed Hummingbird
(below right) of the high Andes. It is adapted for feeding on exceptionally long tubular flowers. |
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Hummingbirds range in size from the smallest birds on earth to several quite large species in the Andes. Between them, these hummers range in size from 2 to 8.5 inches, and in weight from 1.6 to 19 grams. Among the largest is
Great Sapphirewing
(left), shown here from the Peruvian Andes, and nearby lives the largest of them all, Giant Hummingbird
Patagona gigas
, the size of a swift.
Among the smallest species are the woodstars and the coquettes. The two photos below show two Neotropical coquettes: a female
Tufted Coquette
(below left), taken in Trinidad, and a male
Black-crested Coquette
(below right), from Costa Rica.
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Despite the many species of hummingbirds, they are divided into just two subfamilies (Schuchmann 1999). The Trochilinae include all the colorful species (nearly 300 in all) while the Phaethornithinae is composed of the six genera and 34 species of hermits. Brown, long-tailed hermits forage in the understory; some short-tailed species weigh only 1.6 gram (the same weight as the tiny Bee Hummingbird
Mellisuga helenae
of Cuba, often termed the "world's smallest bird").
All hermits that have been studied display in noisy leks, their clicking dominating a patch of jungle. Hermits often have a species affinity for
Heliconia
plants and it can be rewarding to "stake-out" a patch inside the forest. This
Rufous-breasted
(or Hairy)
Hermit
(right) was sitting right next to 'his'
Heliconia
. Unlike many other hummingbirds, hermits are generally not territorial and few species have iridescence. When present, it is limited to the throat or crown. Nonetheless, it is obvious that hermits are hummingbirds, and even the most jaded "I can't tell one bird from another" non-naturalist can identify a hummingbird as such.
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Of course, not all hummingbirds are tropical, or appear to defy the laws of proportions, nor are all bright and colorful. Females and youngsters are often plainly-colored, particularly among the migratory species of North America This cryptic coloring can be important because only females sit on the nest, and if they were bright then they or the nestlings might more easily become prey. Hummingbird nests are often beautifully spun, like those of my local
Anna's Hummingbird
(above left) or the lichen-covered nest with almost-ready-to-fledge juvenal
Costa's Hummingbirds
(above right; it actual represents the first documented nesting in Monterey County, California).
Schuchmann (1999) summarizes hummingbird breeding biology this way: "Male hummingbirds are polygamous, mating with several females during a reproductive period. In most species studied the male associates with the female only during a short period to fertilize her eggs. All remaining reproductive responsibilities like nest building, incubation, and rearing the young are carried out solely by the female." He goes on to cite a few reports of males participating in incubation or rearing the young, particularly in some tropical species, but generally concludes that there is insufficient evidence to confirm any parental care by males. What then am I to make of this photo I took in Costa Rica? It appears to show a male
Blue-throated Goldentail
incubating a nest. The hummer is certainly this species and seems to be in male plumage with a glittering deep-blue throat. This is a lekking species, so why would a male be incubating? May females occasionally attain a male-like plumage? There are many mysteries here.
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Many North American hummingbirds are migratory, including Rufous Hummingbird shown at the top of this account. Migratory hummers are susceptible to vagrancy, accounting for the variety of species that have reached California, including several
Ruby-throated Hummingbird
, one of which is shown at right (in Brad Schram's yard in San Luis Obispo Co.). Surely the most unexpected was Xantus's Hummingbird
Hylocharis xantusii
from southern Baja California; one female built a nest and laid eggs in Ventura Co. (Hainebach 1992). The only Blue-throated Hummingbird
Lampornis clemenciae
to reach California – in Tulare Co. from Dec 1977-May 1978 — not only built a nest and laid eggs, but fledged three young! These appeared to be Blue-throated X Anna's hybrids (Baldridge et al. 1983).
The field identification of female North American hummingbirds is one of our most difficult problems. Stiles (1971, 1972), Baltosser (1987), Jones (1992), Heidcamp (1997), and, particularly Howell (2002) are useful sources.
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While some hummingbirds are reasonably widespread (e.g.,
Scaly-breasted Hummingbird
(left, in display), many hummingbirds are endemics, limited to single Caribbean islands, or isolated mountain chains..
Some of these are threatened with extinction: 29 species are listed as endangered or vulnerable by Birdlife International.
Surely the most sought-after of them all – rare, endangered, spectacular, and limited to a very small range in northern Peru – is
Marvelous Spatuletail
, shown below in a collage. I've taken two of my photos from Huembo Reserve and combined them so that both side of this wonderful species can be shown in one photo.
A separate page
focused on this prized species, created long before my visit, has photos by James Hecht, taken at a time before the Reserve was created.
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From
Green Violetear
(left), widespread from Mexico to Bolivia, to
Bearded Mountaineer
(right), endemic to the high Andes of central Peru, hummingbirds are fascinating. I could go on and on about them. Elsewhere on this website, check out:
Hummingbirds are reason enough to be a birdwatcher. Here's another favorite:
Rufous-crested Coquette
from Peru (below).
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Photos
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The male
Rufous Hummingbird
Selasphorus rufous
was in my Pacific Grove front yard in California in March 2006. The
White-naped Jacobin
Florisuga mellivora
was at Asa Wright Nature Reserve on Trinidad in Dec 2006. Both shots of
Magnificent Hummingbird
Eugenes fulgens
were at Savegre, Costa Rica, on 24 Dec 2007. The male
Anna's Hummingbird
Calypte anna
was on her nest at Carmel R. mouth, Monterey Co., Calif., 22 Mar 2007, and the male
Costa's Hummingbird
Calypte costae
was in Anza-Borrego SP, Calif., in April 2006. The
Violet Sabrewing
Campylopterus hemileucurus
was at Mirador Cinchona, Costa Rica, on 19 Dec 2007. The
Purple-throated Carib
Eulampis jugularis
was on Dominica in March 2000. The
White-tipped Sicklebill
Eutoxeres aquila
was roosting over the Rio Pizore, Osa Peninsula, Costa Rica, on 26 Dec 2007. The
Sword-billed Hummingbird
Ensifera ensifera
was at Owlet Lodge, Peru, 22 Nov 2014. The female
Great Sapphirewing
Pterophanes cyanopterus
was below Abre Malaga, Dept. Cusco, Peru, on 15 June 1987. The female
Tufted Coquette
Lophornis ornatus
was at Asa Wright Nature Reserve, Trinidad, in Dec 2006. The
Black-crested Coquette
Lophornis helenae
was near Braulio Carrillo NP, Costa Rica, on 18 Dec 2007. The
Rufous-breasted Hermit
Glaucis hirsuta
was at Asa Wright Nature Reserve, Trinidad, in Dec 2006. The female
Anna's Hummingbird
Calypte anna
was on her nest at Soberanes Pt., Monterey Co., Calif., in May 1991. The nest of
Costa's Hummingbird
Calypte costae
was in Bixby Canyon, Monterey Co., on 30 June 1981. The incubating
Blue-throated Goldentail
Hylocharis eliciae
was near the Rio Rincon, Osa Peninsula, Costa Rica, on 27 Dec 2007. The male
Ruby-throated Hummingbird
Archilochus colubris
was at Arroyo Grande, Calif., on 4 Sep 2014. The displaying male
Scaly-breasted Hummingbird
Amazilia boucardi
was at Rio Rincon, Osa Peninsula, Costa Rica, on 27 Dec 2007. The
Marvelous Spatuletail
Loddigesia mirabilis
collage is from photos at Huemba Reserve, Amazonas, Peru, in Nov 2014. The
Green Violetear
Colibri thalassinus
was at Savegre, Costa Rica, on 24 Dec 2007. The
Bearded Mountaineer
Oreonympha nobilis
was at Uripichancha, Peru, on 13 June 1987.
The
Rufous-crested Coquette
Lophornis delattrei
was at Waqanki, Peru, in Nov 2014.
All photos © Don Roberson; all rights reserved.
Bibliographic note
: Although hummingbirds have inspired hundreds of books, there is no single family book. Such a work would be a difficult undertaking, since hummingbirds are the second largest family in the New World (after flycatchers) with over 330 species. Between 1849 and 1861, John Gould published a five volume
Hummingbirds
series totaling 418 lithographed plates capturing the beauty and aerial artistry of many South American hummers (now reprinted as Gould 1990). For many years, this was the major work on the family until Crawford Greenewalt (1960) published a spectacular set of color "stop-action" photographs of South American hummingbirds (Greenewalt's pioneering volume is now rare and expensive). Numerous other such books followed using strobe-flash technology to capture hummers in mid-flight, including Scheithauer (1966) and Dunning (1970) who included incredible Andean species, or Keppelman (1988) on southeast Arizona. I have found that by far the most useful for detailed identification purposes for North American species is Tyrell & Tyrell (1985), not because of the text (which is very general) but because the excellent photo quality is often sufficient to determine points like wing formula and remige or rectrix shape.
While not considered a "family book" per se, the Schuchmann (1999) chapter on the Trochilidae in the
Handbook of the Birds of the World
series now serves as the next best thing. At 213 pages, which include a field guide style painting of every species in the world plus perhaps the most spectacular set of photos of wild birds ever brought together, it is a major achievement. The text includes not only a solid introduction to the natural history of the family but 328 species accounts featuring 18 authors.
Literature cited
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Baldridge, F. A., Kiff, L. F., Baldridge, S. K., and Hansen, R. B. 1983. Hybridization of a Blue-throated Hummingbird in California. West. Birds 14: 17-30.
Baltosser, W. H. 1987. Age, species, and sex determination of four North American hummingbirds. North Am. Bird Bander 12: 151-166.
Dunning, J. S. 1970. Portraits of Tropical Birds. Livingston Publ., Wynnewood, PA.
Gould, J. 1990. John Gould's Hummingbirds. Wellfleet Press, Secaucus, N.J.
Greenewalt, C. 1960. Hummingbirds. Dover Publ., New York.
Hainebach, K. 1992. First records of Xantus' Hummingbird in California. West. Birds 23: 133-136.
Heidcamp, A. 1997.
Selasphorus
hummingbirds. Birding 29: 19-29.
Howell, S.N.G. 2002. Hummingbirds of North America: the Photographic Guide. Academic Press, San Diego, CA.
Jones, E. G. 1992. Color variation in maturing male Rufous Hummingbirds. North Amer. Bird Bander 17: 119-120.
Keppelman, T. 1988. Hummingbirds. Little, Brown & Co., New York.
Scheithauer, W. 1966. Hummingbirds. Thomas Y. Crowell, New York.
Schuchmann, K. L. 1999. Family Trochilidae (Hummingbirds)
in
del Hoyo, J., Elliott, A., & Sargatal, J., eds. Handbook of the Birds of the World. Vol. 5. Lynx Edicions, Barcelona.
Stiles, F. G. 1971. On the field identification of California hummingbirds. Calif. Birds 2: 41-54.
Stiles, F. G. 1972. Age and sex determination in Rufous and Allen Hummingbirds. Condor 74: 24-32.
Tyrell, E. Q., and R. A. Tyrell. 1985. Hummingbirds: Their Life and Behavior, a Photographic Study of North American Species. Crown Publ., New York.
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page created 11-13 Dec 1999, extensively revised 2–3 Feb 2008, revised slightly 16 Jun 2016
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all text & photos © Don Roberson, except as otherwise indicated; all rights reserved
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Following served from Phaethornis syrmatophorus, H. C. Kyllingstad, Animal Diversity Web |
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Following modified from Animal Diversity Web
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&pull 20q v5.145 20180528: Error 301 Moved Permanently http://animaldiversity.ummz.umich.edu/site/accounts/information/Trochilidae.html |
Updated: 2024-04-28 21:31:39 gmt
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