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Andrena heraclei Robertson, 1897
Trachandrena heraclei (Robertson, 1897)

Life   Insecta   Hymenoptera   Apoidea   Andrenidae   Andrena
Subgenus: Trachandrena

Andrena heraclei, Mid-Atlantic Phenology
© Copyright source/photographer · 9
Andrena heraclei, Mid-Atlantic Phenology

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Andrena heraclei, F, Back, WV, Morgan County ---.. ZS PMax
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Andrena heraclei, F, Back, WV, Morgan County ---.. ZS PMax
Andrena heraclei, U, Face, PG county, MD ---.. ZS PMax
© Copyright source/photographer · 5
Andrena heraclei, U, Face, PG county, MD ---.. ZS PMax

Andrena heraclei, f, ---.
© Copyright source/photographer · 5
Andrena heraclei, f, ---.
Andrena heraclei, f, face, ---.
© Copyright source/photographer · 5
Andrena heraclei, f, face, ---.

Andrena heraclei, m, side, ---.
© Copyright source/photographer · 5
Andrena heraclei, m, side, ---.
Andrena heraclei, female, face
© Rebekah Andrus Nelson · 1
Andrena heraclei, female, face

Andrena heraclei F NPS09, ext short malar space
Deana Crumbling · 1
Andrena heraclei F NPS09, ext short malar space
Andrena heraclei F NPS09, facial texture smooth
Deana Crumbling · 1
Andrena heraclei F NPS09, facial texture smooth

Andrena heraclei F NPS09, fovea
Deana Crumbling · 1
Andrena heraclei F NPS09, fovea
Andrena heraclei F NPS09 labral process
Deana Crumbling · 1
Andrena heraclei F NPS09 labral process

Andrena heraclei F NPS09, prop corbicula
Deana Crumbling · 1
Andrena heraclei F NPS09, prop corbicula
Andrena heraclei F NPS09, prop tri areolate outside
Deana Crumbling · 1
Andrena heraclei F NPS09, prop tri areolate outside

Andrena heraclei F NPS09, posterior prop and tranveerse ridge
Deana Crumbling · 1
Andrena heraclei F NPS09, posterior prop and tranveerse ridge
Andrena heraclei F NPS09, scutellum
Deana Crumbling · 1
Andrena heraclei F NPS09, scutellum
Overview
Reprinted with permission of the American Entomological Society from: LaBerge, W. E. 1973. A revision of the bees of the genus Andrena of the Western Hemisphere. Part VI. Subgenus Trachandrena. Transactions of the American Entomological Society 99: 235-371.

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Andrena heraclei is an eastern species closely related to A. spiraeana. The female of heraclei is slightly larger than that of spiraeana, has the facial fovea more widely separated from the inner eye margin (much as in rugosa), has a less punctate, shiny face, has a shiny vertex, and has more punctate terga with longer basal areas. The male of heraclei can be separated from that of spiraeana by the more rugulose lateral genal areas, the usually duller surface of the mesoscutum, and the more densely punctate terga with shorter apical areas. The male of heraclei resembles that of forbesii superficially but can be readily distinguished from the latter by the coarse reticulorugose dorsal and posterior propodeal surfaces (outside of the enclosure) and the slightly longer tergal apical areas.

The males here associated with females of heraclei were not collected with any of the females, hence the association of the sexes is only tentative. However, the structure of these males is such that they are likely to be males of heraclei and nothing about their form or distribution negates the hypothesis. Also, there is no other female with which to associate these males, although they were taken in reasonably well-collected areas.

FEMALE. MEASUREMENTS AND RATIOS. N = 20; length, 9-11 mm; width, 3.0-3.5 mm; wing length, M = 3.96 0.109 mm; FL/FW, M = 1.04 0.004; FOVL/FOVW, M = 3.14 0.039.

INTEGUMENTAL COLOR. Black except as follows: mandible half to entirely rufescent; flagellar segments 2-10 dark reddish-brown below; pronotum rufescent above; tegula with colorless hyaline margin, translucent summit, otherwise dark red; wing membranes hyaline, yellow, veins reddish-brown; terga 1-4 with rims hyaline, colorless to slightly yellowed, apical areas often rufescent; sterna and tarsi dark rufescent.

STRUCTURE. Antenna, eyes, malar space, mandible and galea as in spiraeana. Maxillary palpus as in sigmundi but segmental ratio about as 0.9:1.0:0.8:0.8:0.7:0.7. Labial palpus as in sigmundi but ratio about as 1.0:0.7:0.6:0.7. Labrum, clypeus and supraclypeal area as in spiraeana but clypeus usually with median impunctate line. Genal area as in spiraeana but posterior half of lateral surface slightly dulled by fine shagreening. Vertex as in spiraeana but punctures smaller, more crowded and surface shiny at least medially. Face above antennal fossae as in shiniest specimen of spiraeana, without rugulae and punctures separated largely by two to three or more puncture widths, shiny. Facial fovea as in spiraeana but greatest space between inner margin eye and fovea usually equals one and one-half to two times width of fovea at that level (much as in rugosa) and upper end of fovea usually separated from lateral ocellus by half an ocellar diameter.

Pronotum as in spiraeana. Mesoscutum as in spiraeana but punctures slightly sparser in anterior half. Scutellum, metanotum, propodeum, mesepisterna, tibial spurs and venation as in spiraeana.

Metasomal tergum 1 medially with apical area indistinct, equal to about half length of tergum, with crowded punctures separated mostly by half a puncture width except in impunctate rim which is usually one-eighth to one-tenth of median tergal length; basal area with crowded punctures in median third, sparser laterally; surface shiny, unshagreened. Tergum 2 with apical area twice as long as basal area medially or more; apical impunctate rim equals about one-fifth median tergal length, otherwise apical area with crowded punctures separated mostly by half a puncture width or slightly more; basal area punctures similar to apical area; surface shiny, unshagreened. Terga 3 and 4 similar to 2 but punctures slightly sparser and less distinct especially on tergum 4. Pygidial plate as in sigmundi but with rounded apex. Sterna as in sigmundi but slightly shinier.

VESTITURE. Generally white but terga 5 and 6 and inner surfaces tarsi yellow. Form and distribution as in sigmundi but generally sparser; terga 2-4 with apical pale fasciae dense and interrupted medially (often worn completely away).

MALE. MEASUREMENTS AND RATIOS. N = 20; length, 8-10 mm; width, 2.0-2.5 mm; wing length, M(11) = 3.59 0.206 mm; FL/FW, M = 1.13 0.012; FS1/FS2, M = 0.66 0.016.

INTEGUMENTAL COLOR. As in female but flagellar segments 2-11 dark brown to black below; tergal apical rims usually more broadly hyaline.

STRUCTURE. Antennae in repose reaching propodeum; scape length equal to first two flagellar segments; flagellar segments as in forbesii; surface dull as in sigmundi. Eyes each about twice as long as broad or slightly longer, inner margins converging slightly toward mandibles. Malar space, mandible and galea as in female. Maxillary palpus as in female but segmental ratio about as 1.0:1.0:0.9:0.9:0.7:0.7. Labial palpus as in female. Labrum as in sigmundi. Clypeus as in female but usually without impunctate median line and punctures slightly smaller and more crowded. Supraclypeal area with crowded punctures, slightly dulled by fine shagreening. Genal area as in spiraeana but punctures more distinct. Vertex as in female but opaque, dulled by dense, fine shagreening. Face above antennal fossae as in sigmundi.

Pronotum as in female. Mesoscutum with deep punctures separated largely by half to one puncture width, sparser posteromedially; surface moderately dulled by coarse reticular shagreening. Scutellum similar but punctures more crowded. Metanotum opaque, densely punctate. Propodeum and mesepisterna sculptured as in spiraeana but propodeum outside of enclosure slightly less coarsely reticulorugose (but more coarse than in sigmundi or forbesii).

Metasomal terga 1-5 with apical rims impunctate (about one-eighth median length of tergum or more). Tergum 1 with apical area indistinct, equal to slightly less than half median length, with small punctures separated mostly by half to one puncture width; basal area punctures separated mostly by one to two puncture widths (more crowded medially); surface shiny, delicately shagreened especially basally. Terga 2-5 similar to 1 but apical areas delicately shagreened. Sterna 2-5 impunctate apically and basally, with small sparse punctures medially; surfaces moderately shiny, with distinct reticular shagreening. Sternum 6 with shallow apical emargination. Terminalia as in sigmundi; see figures 56-57.

VESTITURE. Generally white and distribution and form as in sigmundi; terga 2-5 with apical pale fasciae distinct laterally, interrupted medially by one-fourth to one-third width of tergum.



Reprinted with permission from: Mitchell, T.B. 1960. Bees of the Eastern United States. North Carolina Agricultural Experiment Station Technical Bulletin No. 141.

FEMALE. Length 10 mm.; face very slightly longer than broad, area between antennae and ocelli unusually smooth and shining, very minutely and sparsely punctate; clypeus rather broadly convex, projecting about one-third below suborbital line, smooth and shining between deep, distinct, rather coarse punctures, quite close on each side of a very narrow, slightly elevated, median, impunctate line; facial foveae broad above, occupying most of area between eyes and ocelli, abruptly and very strongly constricted below, the constricted part narrower than the space separating it from eye margin, covered with whitish tomentum; space between margin of vertex and lateral ocelli somewhat greater than their diameter; cheeks subequal to eyes in width, rounded posteriorly, rather smooth, somewhat shining, punctures exceedingly minute and indistinct; malar space very short; basal segment of flagellum considerably shorter than 2nd and 3rd combined; process of labrum quite short, rather broad, slightly narrowed apically to the broadly truncate margin; pubescence of head, of thorax laterally and below, and of basal leg segments whitish, that on dorsum of thorax and on the more apical leg segments pale ochraceous; punctures of scutum deep, distinct, quite coarse, well separated anteriorly becoming very sparse in center posteriorly where surface is somewhat shining, those on scutellum very close, deep and coarse, surface shining, quite flat; pleura coarsely rugose; dorsal area of propodeum oblique, triangle broad, very coarsely and regularly striate; propodeal corbicula short and poorly developed, whitish-ochraceous, with a rather thin, anterior fringe; trochanteral floccus short and poorly developed, whitish; hind tibiae rather broad, apex nearly twice width of basitarsi, scopa dense, hairs rather short, simple, entirely whitish; fore and mid basitarsi nearly equaling their tibiae in width; 2nd submarginal cell considerably shorter than 3rd, receiving 1st recurrent about one-third from apex; abdominal terga somewhat shining between very fine and close but deep and distinct punctures, apical impressed areas broad, occupying medially about two-thirds length of discs, becoming obscurely reddened apically, rims yellowish-hyaline, discal pubescence hardly evident, extremely short, sparse, thin, suberect, apparently entirely pale, apical fasciae poorly developed, evident toward extreme sides, whitish, tergum 5 with a golden-brownish, apical fimbria.

DISTRIBUTION. Kansas to Illinois, Virginia and North Carolina; April.

FLOWER RECORDS. Crataegus. Recorded by Robertson (1929) on Cornus, Heracleum, Ptelea, Salix and Taenidia.

Names
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Hosts · map
FamilyScientific name @ source (records)
Brassicaceae  Lesquerella filiformis @ AMNH_BEE (1)
Fabaceae  Medicago sativa @ BBSL__BBSLID (1)
Rosaceae  Rubus @ UCRC_ENT (6)
Salicaceae  Salix atrocinerea @ AMNH_BEE (1)

Salix @ AMNH_BEE (2)
_  apple @ NLA (1)

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Updated: 2018-09-26 03:31:34 gmt
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