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Andrena milwaukeensis Graenicher, 1903
Andrena hitei Cockerell, 1907; Andrena jacobaea Cockerell, 1915

Life   Insecta   Hymenoptera   Apoidea   Andrenidae   Andrena
Subgenus: Andrena

Andrena milwaukeensis FEM comp
© Copyright Laurence Packer 2014 · 7
Andrena milwaukeensis FEM comp

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Andrena milwaukeensis MALE mm .x f
© Copyright Laurence Packer 2014 · 7
Andrena milwaukeensis MALE mm .x f
Andrena milwaukeensis
© Copyright Micheal Veit 2010 · 6
Andrena milwaukeensis

Andrena milwaukeensis
© Copyright Micheal Veit 2010 · 6
Andrena milwaukeensis
Andrena milwaukeensis, F, back, Hancock co., Brooklin ---.. ZS PMax
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Andrena milwaukeensis, F, back, Hancock co., Brooklin ---.. ZS PMax

Andrena milwaukeensis, F, Face, Hancock co., Brooklin ---.. ZS PMax
© Copyright source/photographer · 5
Andrena milwaukeensis, F, Face, Hancock co., Brooklin ---.. ZS PMax
Andrena milwaukeensis, F, Side, Hancock co., Brooklin ---.. ZS PMax
© Copyright source/photographer · 5
Andrena milwaukeensis, F, Side, Hancock co., Brooklin ---.. ZS PMax

Andrena milwaukeensis, Female, Side, WV, Randolph Co ---.. ZS PMax
© Copyright source/photographer · 5
Andrena milwaukeensis, Female, Side, WV, Randolph Co ---.. ZS PMax
Andrena milwaukeensis, Milwaukee Andrena
© Copyright John Ascher, 2006-2014 · 4
Andrena milwaukeensis, Milwaukee Andrena

Andrena milwaukeensis, Milwaukee Andrena
© Copyright John Ascher, 2006-2014 · 4
Andrena milwaukeensis, Milwaukee Andrena
Andrena milwaukeensis, Milwaukee Andrena
© Copyright John Ascher, 2006-2014 · 4
Andrena milwaukeensis, Milwaukee Andrena

Andrena milwaukeensis, Milwaukee Andrena
© Copyright John Ascher, 2006-2014 · 4
Andrena milwaukeensis, Milwaukee Andrena
Andrena milwaukeensis
Karl Hillig · 1
Andrena milwaukeensis

Andrena milwaukeensis, back
Karl Hillig · 1
Andrena milwaukeensis, back
Andrena milwaukeensis, Barcode of Life Data Systems
Barcode of Life Data Systems · 1
Andrena milwaukeensis, Barcode of Life Data Systems
Overview
Reprinted with permission of the American Entomological Society from: LaBerge, W. E. 1980. A revision of the bees of the genus Andrena of the Western Hemisphere. Part X. Subgenus Andrena. Transactions of the American Entomological Society 106: 395-526.

Please report text errors to: leah at discoverlife dot org.

Andrena milwaukeensis is a widespread, vernal, polylectic bee related to A. vicinoides, though not closely; There are two types of females of milwaukeensis, a western form in which metasomal terga 1-4 or 5 have abundant, long, fox-red hair and an eastern form in which only terga 1 and 2 have long ochraceous to red hair; A broad zone of intergradation occurs across northern United States and Canada and subspecies are not here recognized; The female of milwaukeensis in typical (eastern) form is similar to that of vicinoides in having basal mandibular teeth, dark tergal hairs, dark scopal hairs and pale thoracic dorsum; However, the milwaukeensis females have a longer clypeus with sparser punctures, an apically rounded pygidial plate, the pale vestiture usually fox-red, and the labral process less sharply bidentate; The male of milwaukeensis differs from that of vicinoides by having the sixth sternum flat (not reflexed along apical margin) and is marked by a relatively long first flagellar segment and short rounded basal mandibular teeth.

FEMALE. MEASUREMENTS AND RATIOS. — N = 20; length, 9-13 mm; width, 2.5-3.5 mm; wing length, M = 4.70 ± 0.226 mm; FL/FW, M = 1.03 ± 0.039; FOVL/FOVW, M = 3.24 ± 0.035.

INTEGUMENTAL COLOR. — As in vicinoides but tibial spurs usually testaceous.

STRUCTURE. — Antennal scape length equals first two and one-half flagellar segments or slightly more; flagellar segment 1 as long as following two and one-half segments, segment 2 about as long as broad and slightly shorter than 3; Eyes each about three and three-fourths times as long as broad, inner margins parallel; Mandible as in vicinoides; Malar space moderately long, minimum length one-third as long as width of mandibular base or slightly less; Galea as in frigida; Maxillary palpus as in frigida but segmental ratio about as 0.9:1.0:0.8:0.8:0.7:0.7; Labial palpus as in frigida but ratio about as 1.0:0.7:0.6:0.6; Labral process trapezoidal to squarish, slightly reflexed apically, emarginate with blunt teeth apically, basally with transverse depression; labrum apical to process with weak median crista, flat, shiny; Clypeus long, rounded from side to side, punctures of irregular size and spacing, separated by half to one or two puncture widths, smaller near base; surface shiny except basal fourth or more shagreened; Supraclypeal area as in vicinoides; Face above antennal fossae as in vicinoides but rugulae often coarser especially near facial foveae; Facial fovea as in vicinoides; Vertex above lateral ocellus usually equal to slightly less than one ocellar diameter, sculptured as in vicinoides but terga 2 and 3 with basal area punctures denser, separated mostly by half to one puncture width; Pygidial plate V-shaped with rounded apex, broad with internal raised triangular area; Sterna as in vicinoides.

VESTITURE. — Head entirely black except face and posteriorly on vertex often with red or ochraceous hair; Thorax black except dorsum with fox-red to ochraceous hair (usually red).

Metasomal terga black except terga 1 and 2 or 1 to 4 with long fox-red to ochraceous hair except at extreme lateral surfaces of terga 2 to 4; sternal hairs black; Leg hairs entirely black; Form and distribution of pollen-collecting hairs as in vicinoides.

MALE. MEASUREMENTS AND RATIOS. — N = 20; length, 8-12 mm; width, 1.5-2.5 mm; wing length, M = 3.63 ± 0.243 mm; FL/FW, M = 1.00 ± 0.006; FS1/FS2, M = 1.53 ± 0.031; INTEGUMENTAL COLOR. — As in vicinoides but tibial spurs testaceous and tergal apical areas usually dark.

STRUCTURE. — Scape length usually less than first two flagellar segments; flagellar segment 1 terete, one and one-third to two times as long as second segment and usually longer than segment 3, segment 2 usually longer than broad and shorter than segment 3, median segments one and one-half times as long as broad; Eyes about three times as long as broad, inner margins parallel; Mandibles long, decussate, as in vicinoides but basal inferior tooth small, rounded; Malar space, galea and maxillary palpus as in female; Labial palpus as in female but segmental ratio about as 1.0:0.6:0.6:0.5; Labral process as in female but usually smaller; labrum apical to process usually without crista, shiny; Clypeus moderately flattened, punctures separated mostly by half a puncture width, slightly sparser apicomedially, impunctate midline usually narrow, weak or absent; surface shiny; Supraclypeal area and face above antennal fossae as in female; Vertex as in female but above lateral ocellus usually as tall as one ocellar diameter and in large specimens taller; Genal area broad, as in vicinoides; Thoracic form and sculpturing as in vicinoides.

Metasomal terga 1-5 sculptured as in vicinoides; Pseudopygidial area absent or extremely narrow; Sterna 2-5 sculptured as in vicinoides; Sternum flat apically, not reflexed.

Terminalia as in figures 102-106.

VESTITURE. — Generally ochraceous to fox-red except as follows: face between upper ends of compound eyes and along inner margins eyes black; genal area entirely black to black in upper fourth or less; tergum 2 apically or tergum 3 apically and terga 4 and 5 entirely dark brown to black; sternal hairs brown or largely so; tarsi with inner surfaces pale yellow.




Reprinted with permission from: Mitchell, T.B. 1960. Bees of the Eastern United States. North Carolina Agricultural Experiment Station Technical Bulletin No. 141.

FEMALE — Length 11 mm.; face very slightly longer than broad; clypeus moderately convex, slightly broader than long, smooth, with widely spaced, shallow and rather fine punctures; facial foveae broad above, covered with brownish tomentum; space between margin of vertex and lateral ocelli subequal to their diameter; cheeks considerably broader than eyes, with minute, well separated punctures; malar space equal to about one-third the basal width of mandible; basal segment of flagellum longer than 2nd and 3rd combined; process of labrum extensive, much broader than long, the tip truncate with a very small median emargination; mandibles rather short, with a small subapical inner tooth; pubescence of head rather copious, fuscous in general but with some ochraceous pubescence on vertex and around antennae; thoracic integument dull and tessellate, very obscurely punctate; enclosure of propodeum smooth; thoracic pubescence long, dense and erect, pale ochraceous above, becoming slightly more fuscous beneath; propodeal corbicula rather well developed, pale ochraceous; trochanteral floccus well developed, fuscous; tibial scopa short, dense, of simple hairs, fuscous; fore and hind basitarsi slightly narrower than their tibiae, the mid basitarsi subequal to their tibiae; pubescence of legs blackish; 2nd submarginal cell much shorter than 3rd, receiving 1st recurrent toward tip; abdominal terga smooth but dull, punctures exceedingly minute and obscure, but rather close, apical margins hardly at all depressed, pubescence rather short but dense and erect, ochraceous on terga 1 and 2, fuscous on the remainder.

MALE — Length 9 mm.; face somewhat longer than broad; clypeus rather flat, only slightly broader than long, surface smooth and shining, with rather close, fine punctures; space between margin of vertex and lateral ocelli slightly exceeding their diameter; cheeks much broader than eyes, subangulate, angle opposite middle of eye (as in tridens, fig. 21) quite sparsely and minutely punctate; malar space short but distinct; process of labrum short and broad, the apical margin broadly and slightly emarginate; mandibles exceedingly long and slender, tip of one about reaching base of the other, without a subapical inner tooth, with a ventral, basal, triangular projection; pubescence of head rather long and dense, ochraceous except for some fuscous hairs around bases of antennae and on vertex laterally; thoracic integument dull, densely tessellate, punctures inevident; propodeal enclosure smooth; thoracic pubescence long, dense, and erect, reddish-ochraceous; basitarsi narrow and elongate, much narrower than their respective tibiae; 2nd submarginal cell much shorter than 3rd, receiving 1st recurrent near middle; abdominal terga shining, nearly impunctate, punctures being exceedingly minute and obscure, apical margins only very slightly depressed, slightly reddened, pubescence rather long and dense on terga 1 and 2 where it is entirely pale, much shorter and darker on the following terga; apical portion of sternum 8 narrow, sides nearly parallel, tip beyond the subapical ridge very short, very slightly incised medially; penis valves broadly expanded basally, excavated beneath this expansion, gonocoxites rather broadly expanded apically, gonocoxal lobes produced and broadly rounded.

DISTRIBUTION — Minnesota to the New England states, Quebec and Nova Scotia, south to North Carolina and Georgia; April to July.

FLOWER RECORDS — Aruncus, Ilex and Viburnum. Brittain and Newton (1934) record it also on Cornus, Pyrus malus and Rubus.


Identification
Extracted from Joel Gardner. "The mining bees of Minnesota (unpublished manuscript)"

A. milwaukeensis have extremely long, dense, orange and black hair, and resemble small bumble bees. The tibiae and scopa are all dark. Is found statewide in Minnesota.


Names
Scientific source:

Supported by

Hosts · map
FamilyScientific name @ source (records)
Adoxaceae  Viburnum lentago @ AMNH_BEE (1)

Viburnum sp @ CUIC_ENT (3)

Viburnum @ AMNH_BEE (1)
Anacardiaceae  Rhus aromatica @ BBSL (1)
Asteraceae  Taraxacum campylodes @ CUIC_ENT (2)

Taraxacum officinale @ BBSL (1)

Tussilago sp @ CUIC_ENT (1)
Berberidaceae  Berberis thunbergii @ AMNH_BEE (2)

Berberis @ AMNH_BEE (2)
Boraginaceae  Hydrophyllum virginianum @ AMNH_BEE (1)
Brassicaceae  Barbarea vulgaris @ CUIC_ENT (16)

Dentaria sp @ CUIC_ENT (1)
Cornaceae  Cornus racemosa @ CUIC_ENT (1)

Cornus stolonifera @ CUIC_ENT (1)
Ericaceae  Pieris japonica @ UCMS_ENT (2)

Vaccinium sp @ CUIC_ENT (1)

Vaccinium @ AMNH_BEE (2); CUIC_ENT (1)
Euphorbiaceae  Euphorbia esula @ UCMS_ENT (2)
J. rykken  1028 @ JRYB__SHEN (1)
Rosaceae  Amelanchier arborea @ CUIC_ENT (6)

Malus pumila @ BBSL (10); CUIC_ENT (1)

Prunus armeniaca @ BBSL (1)

Prunus avium @ BBSL (5)

Prunus maritima @ AMNH_BEE (2)

Prunus virginiana @ BBSL (2)

Prunus @ AMNH_BEE (2); UCRC_ENT (1)

Pyrus communis @ BBSL (26)

Rosa sp @ CUIC_ENT (7)

Rubus sp @ CUIC_ENT (7)

Rubus Ă—paracaulis @ BBSL (1)

Rubus @ I_JSA (4)
Salicaceae  Salix sp @ CUIC_ENT (1)

Salix @ AMNH_BEE (2)
Sapindaceae  Acer rubrum @ CUIC_ENT (1)
Scrophulariaceae  Penstemon cyaneus @ BBSL (1)
_  Withheld @ BBSL (340); BBSL__ZION (6)

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Updated: 2018-09-20 01:39:30 gmt
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