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Top-1+1OriginalDatabase of Hymenoptera in America north of MexicoHighertaxa

Superfamily MEGALODONTOIDEA
Family PAMPHILIIDAE

Commonly called webspinning sawflies, the larvae live solitarily or socially in a web or in a
rolled leaf held by silk. The subfamily Cephalciinae is associated with conifers, and the subfamily
Pamphiliinae is associated with deciduous trees or shrubs. Adults of most species are large, dor-
soventrally flattened insects, and are sometimes common around the flowers of their host plants
in the spring.

Taxonomy: Klima, 1937. In Hedicke, Hym. Cat., pt. 3, 84 pp. (catalog). —Benson, 1945. Roy.
Ent. Soc. London, Proc. Ser. B: Taxonomy 14: 25-33 (generic classification, world).




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Subfamily CEPHALCIINAE

Members of this subfamily are associated with conifers. There are two tribes and one, the
Cephalciini, occurs in North America; the other, Caenolydini, is Palaearctic.

Tribe CEPHALCIINI

Genus CEPHALCIA Panzer

Cephalcia Jurine, 1801. Intell. Blatt Litt.-Ztg. Erlangen 1: 163. Suppressed, Internatl.

Comm. Zool. Nomencl. Op. 135, 1939.
Cephalcia Panzer, 1805. Faunae Ins. German, heft 86, pis. 8, 9.

Type-species: Cephalcia arvensis Panzer. Desig. by Rohwer, 1911.
Cephalcia Jurine, 1807. Nouv. Meth. Class. Hym. Dipt. 67, t. 2, f. 7.
Liolyda Ashmead, 1898. Canad. Ent. 30: 209.

Type-species: Lyda frontalis Westwood. Orig. desig.

Revision: Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 132-149. -Eidt, 1969. Canad. Ent.

Soc, Mem. No. 59, 56 pp. (taxonomy, morphology, larval descriptions, life histories for

most of North American species).

Cephalcia californica Middlekauff
Alta., Mont., Wyo., B. C, Calif. Host: Pinus contorta, P. ponderosa.

Cephalcia californica Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 135. ♀.
distincta (MacGillivray). Newfoundland to N. Y., w. to Ont. and Mich. Host: Abies balsamea,
Tsuga canadensis.

Cephalcia distincta MacGillivray, 1912. Canad. Ent. 44: 296. ♂.

Cephalcia rossi Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 147. ♀.

Cephalcia fascipennis (Cresson)
Que., N. S., Maine, N. H. w. to B. C. Host: Picea engelmaymi, P. glauca,
P. mariana, P. pungens, P. rubens.

Lyda fascipennis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 31. ♀.

Pamphilius fasciipenyiis Dalla Torre, 1894. Cat. Hym., v. 1, p. 427. Emend.

Cephalcia criddlei MacGillivray, 1912. Canad. Ent. 44: 296. ♀.

Cephalcia dissipator MacGillivray, 1923. 111. Univ., Bui. 20(50): 8. ♂.

Biology: Eidt, 1965. Canad. Ent. 97: 148-153.

Cephalcia frontalis (Westwood)
Ont. and Mich. s. to Ga. Host: Pinus resinosa, probably other pines.

Lyda frontalis Westwood, 1874. Thesaurus Ent. Oxon., p. 110, pi. 20, fig. 7. ♂.

Caenolyda onekama MacGillivray, 1923. 111. Univ., Bui. 20(50): 8. ♀.

Cephalcia fulviceps (Rohwer)
Que., N. B., N. H., Conn., N. J., Ont., Ark. Host: Pi^ius resi^wsa, P.
banksiana.

Cephalcia fulviceps Rohwer, 1910. Canad. Ent. 42: 90. ♀.

Cephalcia hopkinsi (Rohwer)
Ariz. Host: Pinus ponderosa(f). An adult collection record.

Cephalcia hopkinsi Rohwer, 1910. Canad. Ent. 42: 217. ♂.

Cephalcia marginata Middlekauff
N. S. and Que. s. to Va., W. Va. Host: Pinus resi^wsa, P. strobus.

Cephalcia marginata Middlekauff, 1953. Pan-Pacific Ent. 29: 133. ♀, ♂.

Biology: Daviault, 1956. Canad. Ent. 88: 488-492.
Morphology: Rivard, 1955. Canad. Ent. 87: 382-399.

Cephalcia nigra Middlekauff
N. S., N. B., Maine, Mich. Host: Picea glauca.

Cephalcia nigra Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 143. ♀.

Cephalcia provancheri (Huard)
Newfoundland s. to N. C, w. to Yukon, Alaska and s. in w. mountains to
Colo., Utah, Nev., and Calif. Ecology: Widespread. Host: Picea engelmanni, P. glauca,
P. mariana, P. mbens, P. sitchensis.
Lyda provancheri Huard, 1879. Nat. Canad. 11: 148. ♂.
Lyda nigropectus Cresson, 1880. Amer. Ent. Soc, Trans. 8: 32. ♀.
Pamphilius mathematicus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 348. ♂.
Cephaleia punctata Rohwer, 1908. Canad. Ent. 40: 177. ♀.

Cephalcia semidea (Cresson)
Que., N. B., Maine, N. H., Conn., N. Y., Ont, Mich. Host: Picea glauca, P.
ynariana.
Lyda semidea Cresson, 1880. Amer. Ent. Soc, Trans. 8: 31. ♀.




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Caenolyda nortoni Rohwer, 1910. Canad. Ent. 42: 91. ♀.
Cephaleia jenseni MacGillivray, 1912. Canad. Ent. 44: 297. ♀.

Genus ACANTHOLYDA Costa

Revision: Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 1-132.

Genus ACANTHOLYDA Subgenus ACANTHOLYDA Costa

Acantkocnema Costa, 1859. In Costa, Fauna Regno. Napoli, Imen. v. 3, p. 2. No included
species. Suppressed, Internatl. Comm. Zool. Nomencl. Op. 290, 1954.

Type-species: Tenthredo erythrocephala Linnaeus. Desig. by Benson, 1945.
Acantholyda Costa, 1894. Prosp. Imen. Ital., v. 3, p. 232.

Type-species: Tenthredo erythrocephala Linnaeus. Desig. by Rohwer, 1910.

Acantholyda apicalis (Westwood)
N. C, Ga., Fla., Miss. Host: Pinus taeda.

Lyda apicalis Westwood, 1874. Thesaurus Ent. Oxon. p. 11, pi. 20, fig. 8. ♂.

Acantholyda atripes (Cresson)
N. J., Pa., Md., N. C, Ga., Ark.

Lyda atripes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 27. ♀.

Acantholyda erythrocephala (Linnaeus)
Conn., N. Y., N. J., Pa., Ont; Europe to Japan and Korea. Introd.,
first found in Pa. in 1925. Host: Pinus austriaca, P. densiflora, P. montana, P. mughus,
P. nigra, P. pungens, P. resinosa, P. strobus, P. sylvestris. Pine false webworm.
Tenthredo erythrocephala Linnaeus, 1758. Syst. Nat. Ed. 10, v. 1, p. 558.

Biology: Griswold, 1939. Jour. Econ. Ent. 32: 467-468. — Eidt and McPhee, 1963. Canad. Dept.
For., For. Ent. Path. Branch, Bimonthly Prog. Rpt. 19: 2.

Acantholyda marginiventris (Cresson)
Mass., N. Y., N. J., Va., N. C.

Lyda marginiventris Cresson, 1880. Amer. Ent. Soc, Trans. 8: 29. ♀.

Acantholyda pini Rohwer
Que., Ont., and Mich. s. to Ga. Host: Pinus resinosai?). An adult collection
record.
Acantholyda pini Rohwer, 1911. U. S. Natl. Mus., Proc. 41: 377. ♀.
Acayitholyda modesta MacGilHvray, 1923. Brooklyn Ent. Soc, Bui. 18: 53. ♀.

Acantholyda poeppigii (Brischke and Zaddach)
Pa., N. C, Ga.

Lyda Poppigii Brischke and Zaddach, 1865. Phys. Okonom. Gesell. Konigsberg., Schr. 6:
123. ♀.

Acantholyda runcinata Middlekauff
Tex., Colo., N. Mex., Ariz.

Acantholyda (Acantholyda) runcinata Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 89. ♀, ♂.

Acantholyda tesselata (Klug)
Que. and Mass. s. to Ga. and w. to Minn. Host: Pinus banksiana, P. resinosa.
Lyda tesselata Klug, 1808. Mag. Gesell. Naturf. Freunde Berlin 2: 276. ♀.
Lyda abdominalis Norton, 1862. Ent. Soc. Phila., Proc. 1: 199. ♀.

Genus ACANTHOLYDA Subgenus ITYCORSIA Konow

Itycorsia Konow, 1897. K. K. Naturhist. Hofmus., Ann. 12: 13.

Type-species: Tenthredo hieroglyphica Christ. Desig. by Rohwer, 1910.

Acantholyda aequorea Middlekauff
Calif.

Acantholyda (Itycorsia) aequorea Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 93. ♀, ♂.

Acantholyda albomarginata (Cresson)
Mich. w. to Colo., Alaska and CaHf. Host: Pinus monophylla, P.
muricata, P. ponderosa.
Lyda albomarginata Cresson, 1880. Amer. Ent. Soc, Trans. 8: 30. ♀.
Itycorsia Kincaidi Rohwer, 1910. Canad. Ent. 42: 91. ♀.

Acantholyda angulata (MacGillivray)
N. B. and Que. s. to N. C. and w. to Minn. Host: Pinus austriaca, P.
banksiana, P. densiflora, P. rigida, P. strobus.
Itycorsia angulata MacGillivray, 1912. Canad. Ent. 44: 295. ♀.

Acantholyda atrata (Cresson)
Idaho, Nev., B. C, Wash., Oreg., Calif. Host: Tsuga heterophylla, Abies
concolori^).
Lyda atrata Cresson, 1880. Amer. Ent. Soc, Trans. 8: 30. ♂.
Itycorsia balata MacGillivray, 1923. 111. Univ., Bui. 20(50): 18. ♀.




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Acantholyda aurigera Middlekauff
Calif.

Acantholyda (Itycorsia) aurigera Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 100. ♀.

Acantholyda balanata (MacGillivray)
Alaska, B.C., Oreg., Calif. Host: Pseudotsiiga menziesii, Picea
sitche7isis, Picea sp., Abies sp., Tsiiga sp.

Itycorsia balanata MacGillivray, 1923. 111. Univ., Bui. 20(50): 18. ♀.

Acantholyda bicolorata (Norton)
Maine, N. Y., Md., Ark.

Lyda bicolorata Norton, 1869. Amer. Ent. Soc, Trans. 2: 332. ♀.

Acantholyda brunnicans (Norton)
Mont., Wyo., and Colo. w. to B. C. and Calif. Host: Pijius contorta, P.
ponderosa, P. radiata, P. sabiniana.

Lyda brunnicans Norton, 1864. Ent. Soc. Phila., Proc. 3: 15. ♀.

Lyda discolor Cresson, 1880. Amer. Ent. Soc, Trans. 8: 26. ♀.

Lyda morrisoni Cresson, 1880. Amer. Ent. Soc, Trans. 8: 27. ♀.

Lyda nevadensis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 28. ♂.

Pamphilius hnmyieicans Dalla Torre, 1894. Cat. Hym., v. 1, p. 423. Emend.

Acantholyda brunniceps (Cresson)
N. B., Que., Mass., N. Y., Ont., Minn.

Lyda brunniceps Cresson, 1880. Amer. Ent. Soc, Trans. 8: 29. ♀.

Pamphilius brunneiceps Dalla Torre, 1894. Cat. Hym., v. 1, p. 423. Emend.

Acantholyda bucephala (Cresson)
Mont., B. C, Wash., Oreg., Calif. Host: Pseudotsuga menziesii, Abies
concolori?).

Lyda bucephala Cresson, 1880. Amer. Ent. Soc, Trans. 8: 29. ♂.

Lyda olympia MacGillivray, 1893. Canad. Ent. 25: 243. ♂.

Acantholyda burkei Middlekauff
Calif. Host: Pinus radiata.

Acantholyda (Itycorsia) burkei Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 108. ♀, ♂.

Acantholyda chicoutimiensis (Huard)
Ont., Que. Host: Picea mariana; Pinus banksiana{?).

Lyda chicoutimiensis Huard, 1879. Nat. Canad. 11: 149. ♀.

Taxonomy: Comeau, 1957. Nat. Canad. 84: 29 (nomenclature).

Acantholyda circumcincta (Klug)
N. B., Que., Pa., Ga., Fla. Host: Piyius baiiksiana, P. clausa.

Lyda circumcijicta Klug, 1808. Mag. Gesell. Naturf. Freunde Berlin 2: 279. ♀.

Acantholyda crocina Middlekauff
Idaho, Calif. Host: Pinus muricata, P. sabiniana.

Acantholyda (Itycorsia) crocina Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 113. ♀, ♂.

Acantholyda depressa Middlekauff
Ariz.

Acantholyda (Itycorsia) depressa Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 115. ♂.

Acantholyda floridana Greenbaum
North. Fla. Host: Pinus elliottii var. elliottii, P. clausa(f).

Acantholyda (Itycorsia) floridana Greenbaum, 1975. Fla. Ent. 58: 47-51. 9, larva.

Acantholyda luteomaculata (Cresson)
N. B. to Conn. w. to Ohio and Minn. Host: Pinus banksiana, P.
strobus, P. sylvestris.

Lyda luteomaculata Cresson, 1880. Amer. Ent. Soc, Trans. 8: 28. ♀.

Acantholyda maculiventris (Norton)
Labrador s. to N. C, w. to Wyo., B. C. and Calif. Ecology:

Widespread. Host: Abies balsamea, Picea glauca, P. mariana(^), Pinus strobus(?).

Lyda maculiventris Norton, 1869. Amer. Ent. Soc, Trans. 2: 333. ♂.

Itycorsia nivea Rohwer, 1910. Canad. Ent. 42: 92. ♀.

Acantholyda ferruginea Felt, 1917. Canad. Ent. 49: 192. ♂.

Acantholyda nigripes
***authority mismatch
(Cresson). Nev., Oreg., Calif.

Lyda nigripes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 28. ♂.

Acantholyda nigrita (Cresson)
Nev., Calif.

Lyda nigrita Cresson, 1880. Amer. Ent. Soc, Trans. 8: 30. ♂.

Acantholyda ochrocera (Norton)
Que. to Pa. w. to Wis.

Lyda ochrocera Norton, 1869. Amer. Ent. Soc, Trans. 2: 332. ♀, ♂.

Acantholyda ruficeps (Harrington)
Que. s. to N. C. and w. to Minn.; B. C. Host: Pinus banksiana, P.
contorta, P. resiyiosa.

Pamphilius ruficeps Harrington, 1893. Canad. Ent. 25: 62. ♀.

Acantholyda terminalis (Cresson)
Colo., Nev., Wash., Oreg., Calif.

Lyda tenninalis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 29. ♀.

Itycorsia ballista MacGillivray, 1923. 111. Univ., Bui. 20(50): 19. ♀.




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Acantholyda thalictra Middlekauff
Tex., N. Mex. Host: Pinus ediilisin.

Acantholyda (Itycorsia) thalictra Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 126. ♀.

Acantholyda verticalis (Cresson)
Minn., Nebr., and Colo. w. to B. C. and Calif. Host: Piniis contorta, P.
monticola, P. ponderosa, P. radiata.

Lyda verticalis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 26. ♀.

Lyda situilaris Cresson, 1880. Amer. Ent. Soc, Trans. 8: 27. ♀.

Lyda montivaga Cresson, 1880. Amer. Ent. Soc, Trans. 8: 28. ♂.

Lyda rufiventris Cresson, 1880. Amer. Ent. Soc, Trans. 8: 29. ♂.

Itycorsia luteopicta Rohwer, 1910. Canad. Ent. 42: 92. ♀.

Acantholyda zappei (Rohwer)
Que. to N. J. w. to Mich. Host: Pinus aiistriaca, P. rigida, P. strobus.

Itycorsia zappei Rohwer, 1920. U. S. Natl. Mus., Proc 57: 209. ♀, ♂.

Subfamily PAMPHILIINAE

Members of this subfamily are associated with deciduous trees and shrubs. The Nearctic spe-
cies of Neurotomini are confined to plants of the family Rosaceae. There are two tribes, and
both tribes are Holarctic.

Tribe NEUROTOMINI

Genus NEUROTOMA Konow

Neurotoma Konow, 1897. K. K. Naturhist. Hofmus., Ann. 12: 18.

Type-species: Tenthredo flaviventris Retzius. Desig. by Rohwer, 1910.
Neurotoma subg. Gongylocorsia Konow, 1897. K. K. Naturhist. Hofmus., Ann. 12: 19.

Type-species: Lyda mandibularis Zaddach. Monotypic
Gongylocorisa{\) Ashmead, 1898. Canad. Ent. 30: 208.

Revision: Middlekauff, 1940. Canad. Ent. 72: 201-206 (adults and larvae). —Middlekauff,
1958. Calif. Univ., Pubs., Ent. 14: 149-158.

Neurotoma crataegi Middlekauff
Mass., N. Y., Mich., 111., Wis., Man. Host: Crataegus succulenta, C.
brainerdi.
Neurotoma crataegi Middlekauff, 1940. Canad. Ent. 72: 202. ♀.

Neurotoma fasciata (Norton)
Que. to Fla. w. to Wis. and Mo. Host: Pnimis serotina, P. pennsylvanica.
Lydafasciata Norton, 1862. Ent. Soc Phila., Proc. 1: 200. ♀.

Biology: Pearsall, 1902. Canad. Ent. 34: 214-216.

Neurotoma inconspicua (Norton)
Que. and Mass. to Pa. w. to S. Dak., Mont, and B. C, Ark. and Kans.

Host: Primus americana, P. cerasus, P. besseyi, P. davidiayia, P. domestica, P. na7ia, P.
nigra, P. pennsylvanica, P. virginiana, and probably other species of Prunus. Plum
webspinning sawfly.
Lyda inconspicua Norton, 1869. Amer. Ent. Soc, Trans. 2: 350. ♀.
Lyda spoliana Williams, 1896. S. Dak. Agr. Expt. Sta., Bui. 48: 14-17. ♀, ♂.
Lyda nifipes Marlatt, 1896. In Williams, S. Dak. Agr. Expt. Sta., Bui. 48: 17. ♀, ♂.

Biology: Severin, 1920. S. Dak. State Ent. Tech. Bui. 1, 53 pp. — Severin, 1920. S. Dak. Agr.
Expt. Sta., Bui. 190: 224-251.
Willi Middlekauff. N. Y., Pa. Host: Crataegus sp.(?). An adult collection record.
Neurotoma willi Middlekauff, 1958. Calif. Univ., Pubs., Ent. 14: 158. ♂.

Taxonomy: Middlekauff, 1965. N. Y. Ent. Soc, Jour. 73: 238 (female, distribution, host).

Tribe PAMPHILIINI
Genus PAMPHILIUS Latreille

Pamphilius Latreille, 1802. Hist. Nat. Crust. Ins., v. 8, p. 303.

Type-species: Tenthredo sylvatica Linnaeus. Monotypic.
Lyda Fabricius, 1804. Systema Piezatorum, p. 43.

Type-species: Tenthredo sylvatica Linnaeus. Desig. by Curtis, 1831.
Anoplolyda Costa, 1894. Prosp. Imen. Ital., v. 3, p. 233.

Type-species: Lyda alternans Costa. Desig. by Rohwer, 1910.


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Kelidoptera Konow, 1897. K. K. Naturhist. Hofmus., Ann. 12: 20.

Type-species: Lyda ynaculipejinis Stein. Monotypic.
Bactroceros Konow, 1897. K. K. Naturhist. Hofmus., Ann. 12: 21.

Type-species: Tenthredo vafer Linnaeus. Desig. by Rohwer, 1910.
Pseudocephaleia Zimgiebl, 1937. Embrik Strand 60. Geburtst. Festschr. (Riga) 3: 339.

Type-species: Pseudocephaleia brachycenis Zirngiebl. Monotypic.
Pamphilius subg. Oiiycholyda Takeuchi, 1938. Tenthredo 2: 218.

Type-species: Pamphilius sulphureipes Kirby. Orig. desig.

Since this section was completed, Benes (1972) defined the genera of the Pamphiliini. He
recognized four genera: Pseudocephaleia Zirngiebl, Kelidoptera Konow, Pamphilius Latreille,
and Onycholyda Takeuchi. The latter two genera are found in North America, and, of the spe-
cies listed below in Pamphilius, the following would belong in Onycholyda: amplechis, ex-
cavatus, luteiconiis, ynultisignatus, 7iigriiibialis, quebecensis, rufofasciatus, and sitkensis.
These species that belong in Onycholyda are the ones that are found ir. couplets 2 to 9 and 24 to
31 in Middlekauffs (1964) key.

Revision: Middlekauff, 1964. Calif. Univ., Pubs., Ent. 38: 1-80.

Taxonomy: Benes, 1972. Acta Ent. Bohem. 69: 378-395 (generic classification of the

Pamphiliini). —Benes, et a/., 1973. Bui. Zool. Nomencl. 30: 95-96 (request for designation

of Pamphilius viriditibialis Takeuchi as type-species of Onycholyda Takeuchi).

Pamphilius amplectus (Fabricius)
Maine to Fla. w. to 111. and Tex. Host: Rubus.

Lyda aniplecta Fabricius, 1804. Systema Piezatorum, p. 46.

Lyda plagiata Klug, 1808. Mag. Gesell. Naturf. Freunde Berlin, 2: 278. ♀.

Lyda insigjiis Brischke and Zaddach, 1865. Phys. Okonom. Gesell. Konigsberg., Schr. 6:
179. ♂.

Pamphilius burquei (Provancher)
Que., Ont., Va.

Lyda burquei Provancher, 1878. Nat. Canad. 10: 204. ♀.

Pamphilius cinctus Harrington
Que., N. Y., Ont., Mich., Wis.

Pamphilius cinctus Harrington, 1893. Canad. Ent. 25: 62. ♀.

Pamphilius excavatus (Norton)
Que. to Va. w. to Minn.; B. C. Host: Conius stolonifera.

Lyda excavata Norton, 1869. Amer. Ent. Soc, Trans. 2: 337. ♂.

Pamphilius greenei Rohwer
Conn., Pa., Mich.

Pamphilius greenei Rohwer, 1913. U. S. Natl. Mus., Proc. 45: 273. ♀.

Pamphilius infuscatus Middlekauff
Maine, Que., Ont., Alta., B. C. Host: Populns tremuloides.

Pamphilius infuscatus Middlekauff, 1964. Calif. Univ., Pubs., Ent. 38: 24. ♀, ♂.

Pamphilius luteicornis (Norton)
N. S. and N. B. s. to Fla. w. to 111., Minn, and Alta. Host: Rubiis.

Lyda luteiconiis Norton, 1869. Amer. Ent. Soc, Trans. 2: 339. ♀.

Lyda harringtoni Provancher, 1886. Addit. Corr. Faune Ent. Canada Hym. p. 17. ♀.

Pamphilius ocellatus Rohwer, 1910. Canad. Ent. 42: 218. ♀.

Painphilins rubi Rohwer, 1910. Canad. Ent. 42: 219. ♀.

Pamphilius dentatus MacGillivray, 1912. Canad. Ent. 44: 297. ♀, ♂.

Pamphilius fletcheri MacGillivray, 1912. Canad. Ent. 44: 298. ♀, ♂.

Biology: Fletcher, 1900. Rpt. Ent. Bot. Dom. Expt. Farm Ottawa, Sess. Paper 8a: 180-181

(multisignata, nee Norton). — Walden, 1913. Conn. (State) Agr. Expt. Sta., Ann. Rpt, 36:

236-240 (dentatus). — Britton, 1913. Jour.Econ. Ent. 6: 197-199 (dentatus).

Pamphilius multisignatus (Norton)
Ont., Mich., 111. w. to Colo., N. Mex., and B. C.

Lyda 7nnltisignata Norton, 1864. Ent. Soc. Phila., Proc. 3: 15. ♀.

Lyda cavifrons Cresson, 1865. Ent. Soc. Phila., Proc. 4: 246. ♂.

Lyda ynulticinctaV.) Norton, 1869. Amer. Ent. Soc, Trans. 2: 332.

Bactroceros pugnax Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 103. 9. Preocc

Anoplolyda saeva Rohwer, 1910. Canad. Ent. 42: 218. N. name.

Pamphilius nigritibialis Rohwer
Que. to N. C. w. to S. D., Colo., and Alta.

Pamphilius iiigritibialis Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 205. ♂.

Paynphilius transversus MacGillivray, 1912. Canad. Ent. 44: 297. "♀"=♂.

Pamphilius ochreipes (Cresson)
Que. to N. C. and Ala., w. to Alaska and B. C. Host: Viburnum opulus.

Lyda ochreipes Cresson, 1880. Amer. Ent. Soc, Trans. 8: 30. ♀.


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Lyda perplexa Cresson, 1880. Amer. Ent. Soc, Trans. 8: 31. ♀.

Biology: Lindquist and Harnden, 1970. Canad. Ent. 102: 95-97.

Pamphilius ocreatus (Say)
Que. to N. C. w. to 111., Man., Minn. Host: Coryhis, Comus{i).
Lyda ocreata Say, 1836. Boston Jour. Nat. Hist. 1: 222. ♀.

Biology: Dyar, 1895. Ent. News 6: 199-200 (with larva description).
paciflcus (Norton). Colo., Idaho, w. to B. C. and Calif. Host: Rosa califomica.
Lyda pacifica Norton, 1869. Amer. Ent. Soc, Trans. 2: 338. ♀.
Lyda melliventris Cresson, 1880. Amer. Ent. Soc, Trans. 8: 32. ♂.

Biology: Middlekauff, 1964. Calif. Univ. Pubs., Ent. 38: 41-42.

Pamphilius palachei (Ashmead)
Alaska (Kodiak).

Bactroceros palachei Ashmead, 1902. Wash. Acad. Sci., Proc 4: 254. ♀.

Pamphilius pallimaculus (Norton)
Que. to Ga., w. to Colo, and B. C. Host: Rosa{?). An adult collection
record.
Lyda pallimacula Norton, 1869. Amer. Ent. Soc, Trans. 2: 337. ♀.
Lyda pallidirnaaila Dalla Torre, 1894. Cat. Hym., v. 1, p. 435. Emend.

Pamphilius persicum MacGillivray
Que. to Fla. w. to Minn., Nebr., Tex. Ecology: A pest of peach. Host:
PruJius persica, Primus serotina.
Pamphilius persicum MacGillivray, 1907. Canad. Ent. 39: 308. ♀.
Pamphilius perseae Rohwer, 1926. In Leonard, Cornell Univ. Agr. Expt. Sta., Mem. 101:

872. Emend.
Pamphilius persicus Klima, 1937. Hym. Cat., pt. 3, p. 45. Emend.
Pamphilius unalatus MacGillivray, 1920. Brooklyn Ent. Soc, Bui. 15: 112. ♀.

Biology: Walden, 1907. U. S. Dept. Agr., Bur. Ent. Bui. 67: 85-87. — Walden, 1908. Conn.
(State) Agr. Expt. Sta., Ann. Rpt., pp. 285-300.

Morphology: Crampton, 1919. Ent. Soc. Wash., Proc. 21: 219 (genitalia).

Pamphilius phyllisae Middlekauff
N. B. to N. C. w. to 111. and Minn. Host: Quercus rubra.
Pamphilius phyllisae Middlekauff, 1964. Calif. Univ., Pubs., Ent. 38: 49. ♀, ♂.

Biology: Eidt and Nichols, 1970. Canad. Ent. 102: 53-63.

Pamphilius pullatus (Cresson)
Newfoundland to Va., w. to 111., Mo. Host: Viburnum dentatum.

Lyda pidlata Cresson, 1880. Amer. Ent. Soc, Trans. 8: 31. ♀.

Pamphilius quebecensis (Provancher)
Que. to N. C, w. to Mich.

Lyda quebecensis Provancher, 1878. Nat. Canad. 10: 205. ♀.

Pamphilius rileyi (Cresson)
Conn, to W. Va., w. to Mich., Iowa, Mo. Host: Amelanchier.

Lyda rileyi Cresson, 1880. Amer. Ent. Soc, Trans. 8: 32. ♀.

Pamphilius subcavifrons Rohwer, 1910. Canad. Ent. 42: 219. ♂.

Pamphilius rufofasciatus (Norton)
Newfoundland to Ga. w. to Minn., Man., Colo., Alta.; Siberia.

Lyda rufo-fasciata Norton, 1869. Amer. Ent. Soc, Trans. 2: 340. ♀.

Lyda rxifocincta Cresson, 1880. Amer. Ent. Soc, Trans. 8: 32. ♂.

Taxonomy: Benes, 1972. Acta Ent. Bohem. 69: 388 (in Onychohjda; a holarctic sp.).

Pamphilius semicinctus (Norton)
Que. to N. C. w. to Tenn. and Ind. Host: Amelanchier canadensis,

Amelanchier sp., Pyrus malus, Prunus virginiana. Some may not represent true hosts.
Lyda semicincta Norton, 1862. Ent. Soc Phila., Proc. 1: 144. ♀.

Pamphilius sitkensis (Kincaid)
Idaho, Nev., Alaska, B. C, Wash., Oreg., Calif. Host: Rubus parviflorus, R.
vitifolius, and probably other species of Rubus.
Bactroceros sitkensis Kincaid, 1900. Wash. Acad. Sci., Proc 2: 344. ♀.
Pamphilius ftilvifrons Rohwer, 1910. Canad. Ent. 42: 218. ♀.
Pamphilius fortuitus MacGillivray, 1923. 111. Univ., Bui. 20(50): 27. ♀.

Biology: Middlekauff, 1964. Calif. Univ., Pubs., Ent. 38: 62-63.

Pamphilius uniunguis Middlekauff
N. C, Man., Minn.

Pamphilius uniunguis Middlekauff, 1964. Calif. Univ., Pubs., Ent. 38: 65. ♀.

Pamphilius vernalis Middlekauff
Man., Nebr.

Pamphilius vernalis Middlekauff, 1964. Calif. Univ., Pubs., Ent. 38: 66. ♀.




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Unplaced Taxa of Pamphiliidae

Lyda cayiadensis Norton, 1869. Amer. Ent. Soc, Trans. 2: 336. 6. No locality given.
Tarpa scripta Say, 1824. In Keating, Narr. Long's 2nd Exped., v. 2, (App.), p. 312. N. W. T. and
Ark.


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