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Superfamily TENTHREDINOIDEA
Family ARGIDAE

This family is cosmopolitan. It is the second largest sawfly family, with over 800 species. The
tropical regions of the world contain the largest number of species, over 100 species of Arge
alone having been described from tropical Africa. Over 300 species of the family have been
described from the Neotropical Region, even though the fauna there is virtually unexplored. In
America north of Mexico, the greater number of species is found in the southwestern United
States. Hosts and habits are various, but most species feed externally on the foliage of the host
plant in the larval stage. Only one North American species is known to be a leafminer,
Schizocerella pilicomis (Holmgren) in purslane.

Taxonomy: Benson, 1938. Roy. Ent. Soc. London, Trans. 87: 371-375 (key to subfamilies,
world). —Malaise, 1941. Ent. Tidskr. 62: 131-140 (key to world genera). —Wong, 1951. Ont.
Ent. Soc, 82nd Ann. Rpt, p. 53 (cocoons). —Benson, 1963. Ann. and Mag. Nat. Hist. (13) 5:
631 (tribes of Arginae and genera of Scobini). —Smith, 1969. Amer. Ent. Soc, Trans. 95:
439-457 (Nearctic genera). —Smith, 1972. Amer. Ent. Soc, Trans. 98: 163-184 (N. Amer.
larvae).

Subfamily ARGINAE

Genus ARGE Schrank

Cryptus Jurine, 1801. Intell. Blatt. Litt.-Ztg. Erlangen, v. 1, p. 163. Suppressed by

Internatl. Comm. Zool. Nomencl., Op. 135, 1936.
Arge Schrank, 1802. Fauna Boica, p. 209.

Type-species: Tenthredo enodis Linnaeus. Desig. by Rohwer, 1911.
Hylotoma Latreille, 1802. Hist. Nat. Crust. Ins., v. 3, p. 302.

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Type-species: Tentkredo rosae Linnaeus. Monotypic.
Corynia Imhof and Labram, 1836. Ins. Schweiz., p. 23.

Type-species: Corynia rosarum Imhof and Labram. Desig. by Rohwer, 1911.
Acanthoptenos Ashmead, 1898. Canad. Ent. 30: 212.

Type-species: Acanthoptenos weithii Ashmead. Orig. desig.
Rhopalospiria Enderlein, 1920. Gesell. f. Naturf. Freunde, Sitzber. 9: 116. N. syn.

Type-species: Rhopalospiria rnbiginosa (Beauvois). Orig. desig.

A large genus not represented only in the Neotropical and Australian regions. This is the most
widespread genus in North America and is found from Canada and Alaska south to northern
Mexico. The larvae feed on the foliage of their hosts. Several of the segregates below are species
complexes, the taxonomy of which is yet to be worked out.

Arge abdominalis (Leach)
Ga., Fla.

Hylotoma abdoyninalis Leach, 1817. Zool. Misc., v. 3, p. 123. ♀.

Arge borealis (Kirby)
N. B., Maine, Mass., Ont. (St. Martin Falls, Hudson's Bay).

Hylotoma borealis Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 66. ♀.

Arge caerulea (Dalla Torre)
Maine to Md.

Hylotoma coenilea Norton, 1864. Ent. Soc. Phila., Proc. 3: 5. ♀. Preocc. by Latreille, 1805.

Hylotoma caerulea Dalla Torre, 1894. Cat. Hym., v. 1, p. 327. N. name.

Arge clavicornis (Fabricius)
Newfoundland to Alaska s. to Ga., N. Mex., Calif.; Holarctic (?).
Ecology: Widespread. Host: Amelanchier, Azalea, Betula, Carpinus, Corylus,
Crataegus, Primus, Salix. A species complex.

Tenthredo clavicoruis Fabricius, 1793. Ent. System., v. 2, p. 108. ♀.

Hylotoma viresceiis Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 6: 296. ♀, ♂.

Hylotoma MacLeayi Leach, 1817. Zool. Misc., v. 3, p. 122. ♀.

Hylotoma analis Leach, 1817. Zool. Misc., v. 3, p. 123. ♂.

Hylotoma mellina Cresson, 1880. Amer. Ent. Soc, Trans. 8: 3. ♀.

Hylotoma cyra Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 66. ♀.

Hylotoma mentzeliae Cockerell, 1898. Ann. and Mag. Nat. Hist. (7)2: 456. N. syn.

Acanthoptenos weithii Ashmead, 1898. Canad. Ent. 30: 212.

Arge inops Konow, 1906. Ztschr. System. Hym. Dipt. 6: 181. ♂.

Hylotoma nigrescens Rohwer, 1909. Canad. Ent. 41: 13. ♂. N. syn.

Hylotoma onerosa MacGillivray, 1923. Psyche 30: 80. ♀, ♂.

Hylotoma sparta MacGilHvray, 1923. 111. Univ., Bui. 20(50): 18. ♀, ♂.

Arge criddlei Garlick, 1927. Canad. Ent. 59: 182. ♀, ♂.

Arge iynitans Garlick, 1927. Canad. Ent. 59: 182. ♀, ♂.

Biology: Jack, 1891. Psyche 6: 11 (ynacleayi). —Will, 1947. Pa. Acad. Sci., Proc. 16: 47-49

(macleayi). —Will, 1959. Pa. Acad. Sci., Proc. 33: 219 {macleayi).

Arge coccinea (Fabricius)
Conn, to Fla. w. to Tex. Host: Rhiis spp.

Hylotoma coccinea Fabricius, 1804. Systema Piezatorum, p. 24.

Tenthredo nibiginosa Beauvois, 1805. Ins. Afr. Amer., v. 2, p. 98. ♀. N. syn.

Hylotoma miniata Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 6: 298. ♂.

Hylotoma rubra Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 6: 299. ♀.

Hylotoma erythrosoma Leach, 1817. Zool. Misc., v. 3, p. 124. ♀. N. syn.

Hylotoma scutellata Lepeletier, 1823. Monog. Tenthred., p. 47.

Hylotoma sphinx Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 68. ♂. N. syn.

Arge humeralis (Beauvois)
Maine to Fla. w. to Minn., Kans., Tex. Host: Poison-ivy.

Tenthredo humeralis Beauvois, 1805. Ins. Afr. Amer., v. 2, p. 99. ♀.

Hylotoma sanguinea Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 6: 299. ♀.

Hylotoma scutellata Say, 1836. Boston Jour. Nat. Hist. 1: 211. ♀. N. syn.

Hylotoma grandis Rohwer, 1909. Canad. Ent. 41: 14. ♂. N. syn.

Arge ochropa (Gmelin)
Ont.; Europe to Siberia, Middle East. Introduced (?) into N. Amer. Host:
Rosa.

Tenthredo ochropus Gmelin, 1790. In Linnaeus, Syst. Nat., Ed. 13, v. 1, pt. 5, p. 2657.

Taxonomy: Malaise and Benson, 1934. Archiv for Zool. 26A: 7-8, 13 (=rosae of authors, nee
Linnaeus).

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Arge pectoralis (Leach)
Maine to Fla. w. to Ont., Sask., Mich., Ill, Mo., Tex. Host: Alnus, Betula,
Corylus, Crataegus, Salix. Birch sawfly. Probably a species complex.
Hylotoma pectoralis Leach, 1817. Zool. Misc., v. 3, p. 124. ♀.

Hylotoma dulciaria Say, 1824. In Keating, Narr. Long's 2nd Exped., (App.). p. 314. ♀.
Arge salicis Rohwer, 1912. U. S. Natl. Mus., Proc. 43: 206. ♀,♂.

Biology: Schwarz, 1909. Ent. Soc. Wash., Proc. 11: 106-108. -Wong, 1954. Canad. Ent. 86:
154. —Smirnoff, 1971. Ent. Soc. Quebec, Ann. 16: 86-88 (protozoan parasite).

Arge scapularis (Klug)
Maine to Fla. w. to Ont., Minn., Mo., Tex. Host: Alnus, Betula, Quercus,
Ulmus. Probably a species complex.
Hylotoma scapularis Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 6: 299. ♂.
Hylotoma xanthothorax Leach, 1817. Zool. Misc., v. 3, p. 124. ♀.
Hylotoma calcanea Say, 1836. Boston Jour. Nat. Hist. 1: 211.

Biology: Jack, 1891. Psyche 6: 11.

Arge spiculata (MacGillivray)
Ariz.; Mexico.

Hylotoma spiculata MacGillivray, 1907. Canad. Ent. 39: 308. ♀.

Subfamily ATOMACERINAE

Genus ATOMACERA Say

Atomacera Say, 1836. Boston Jour. Nat. Hist. 1: 212.

Type-species: Atomacera debilis Say. Desig. by Malaise, 1937.
Micrarge Ashmead, 1898. Canad. Ent. 30: 213.

Type-species: Atoinacera nificollis Norton. Orig. desig.
Braunsiola Konow, 1899. Ent. Nachr. 25: 312.

Type-species: Brauyisiola truculenta Konow. Monotypic.
Spegazziniella Jorgensen, 1913. Buenos Aires Mus. Nac. de Hist. Nat., An. 25: 260.

Type-species: Spegazziniella ovata Jorgensen. Monotypic.
Argina Forsius, 1925. Arkiv for Zool. 17A: 2. Preocc.

Type-species: Argina nubilipennis Forsius. Monotypic.
Arginella Forsius, 1926. Notulae Ent. 5: 207. N. name for Argina.

A genus distributed throughout most of the New World but more highly developed in the
tropics. Larvae feed on the foHage of the host plant.

Revision: Malaise, 1942. Ent. Tidskr. 63: 107-110 (Neotropical species). —Smith, 1969. Amer.
Ent. Soc, Trans. 95: 440-446 (Nearctic species).

Atomacera debilis Say
Ont., Mass. to Fla. w. to Minn., Kans., Ark. Host: Desmodium canadense, D.
gliitinosum, and probably other species; Lespedeza.
Atomacera debilis Say, 1836. Boston Jour. Nat. Hist. 1: 212. ♂.
Atomacera nificollis Norton, 1864. Ent. Soc. Phila., Proc. 3: 6. ♀.
Atoynacera desynodii Dyar, 1900. N. Y. Ent. Soc, Jour. 8: 26. 9, cJ. Larva.

Taxonomy: Smith, 1972. Amer. Ent. Soc, Trans. 98: 167 (larva).

Biology: Dyar, 1900. N. Y. Ent. Soc, Jour. 8: 26. —Weiss and Lott, 1923. Ent. News 34: 167.
— Lindquist and Trinnell, 1965. Canad. Ent. 97: 181-184.

Atomacera decepta Rohwer
Conn, to Fla. w. to Mo., Tex. Host: Hibiscus militaris, H. moscheutos hybrid.
Atomacera decepta Rohwer, 1911. U. S. Natl. Mus., Proc 41: 382. ♀.

Taxonomy: Smith, 1972. Amer. Ent. Soc, Trans. 98: 167 (larva).

Biology: Dyar, 1900. N. Y. Ent. Soc, Jour. 8: 26 {nificollis, nee Norton). — Tippins, 1965.
Jour. Econ. Ent. 58: 161.

Atomacera ebena Smith
S. Ariz.

Atomacera ebena Smith, 1969. Amer. Ent. Soc, Trans. 95: 445. ♀.

Subfamily STERICTIPHORINAE

Genus STERICTIPHORA Billberg

Sterictiphora Billberg, 1820. Enum. Ins. Mus. Billberg, p. 99.

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Type-species: Hylotovia furcata Fabricius. Monotypic.
Schizocerus Berthold, 1827. Nat. Fam. Thierr., p. 441.

Type-species: Hylotoma furcata Villers. Desig. by Westwood, 1840.
Cyphona Dahlbom, 1835. Conspectus Tenthred., Siricid., Oryssin. Scan., p. 8.

Type-species: Hylotoma furcata Villers. Desig. by Rohwer, 1911.

A small Holarctic genus. Larvae feed on the foliage of the host plants most of which appear to
be species of Primus.

Revision: Smith, 1969. Amer. Ent. Soc, Trans. 95: 446-455.

Sterictiphora cruenta Smith
Maine to N. C. w. to Mich. Host: Prunus peimsylvanicus.

Sterictiphora cruenta Smith, 1969. Amer. Ent. Soc, Trans. 95: 450. ♀.

Sterictiphora maura (Cresson)
Utah, Nev., Oreg., Cahf. Host: Prunus

Schizocera maura Cresson, 1880. Amer. Ent. Soc, Trans. 8: 3. ♂.
pninivora (Dyar). Maine s. to N. Y., Pa. Host: Prunus, Amelanchier.

Schizocerus pninivonis Dyar, 1897. N. Y. Ent. Soc, Jour. 5: 23. larva.

Sterictiphora sericea (Norton)
Maine to N. J., Pa. w. to 111.

Schizocerus sericeus Norton, 1867. Amer. Ent. Soc, Trans. 1: 55. ♀.

Sterictiphora serotina Smith
Maine to Va. Host: Prunus serotina.

Sterictiphora serotina Smith, 1969. Amer. Ent. Soc, Trans. 95: 452. ♀, ♂.

Taxonomy: Smith, 1972. Amer. Ent. Soc, Trans. 98: 169 (larva).

Sterictiphora transversa Smith
Md., Alta., Alaska.

Sterictiphora transversa Smith, 1969. Amer. Ent. Soc, Trans. 95: 454. ♀.

Genus APROSTHEMA Konow

Aprosthema Konow, 1899. Ent. Nachr. 25: 149.

Type-species: Hylotoma brevicomis Fallen. Desig. by Rohwer, 1911.
Lyrola Ross, 1937. 111. Biol. Monog. 15(2): 55.

Type-species: Schizocera brunniventris Cresson. Orig. desig.

A small Holarctic genus with two Nearctic representatives, both of which are found in
western North America.

Revision: Smith, 1971. Amer. Ent. Soc, Trans. 97: 546-549.

Aprosthema breviantennatum Smith
Oreg., Calif.

Aprosthema breviantennatum Smith, 1971. Amer. Ent. Soc, Trans. 97: 546. ♀, ♂.

Aprosthema brunniventre (Cresson)
Wash., Oreg., Nev., Calif. Host: Hosackia.

Schizocera brunniventris Cresson, 1880. Amer. Ent. Soc, Trans. 8: 2. ♀, ♂.

Schizocera tristis Cresson, 1880. Amer. Ent. Soc, Trans. 8: 52. ♀, ♂.

Schizocera tristis \ar.fumipen7iis Dyar, 1893. Canad. Ent. 25: 195. 9, 6, larva.

Taxonomy: Smith, 1972. Amer. Ent. Soc, Trans. 98: 170 (larva).
Biology: Dyar, 1893. Canad. Ent. 25: 195.

Genus NEOPTILIA Ashmead

Neoptilia Ashmead, 1898. Canad. Ent. 30: 213.

Type-species: Neoptilia mexicana Ashmead. Orig. desig.
Pseudocyphona Ashmead, 1898. Canad. Ent. 30: 211.

Type-species: Pseudocyphona mexicana Ashmead. Orig. desig.
Rhagonyx Konow, 1903. Ztschr. System. Hym. Dipt. 3: 108.

Type-species: Rhagonyx lituratus Konow. Monotypic.

A small genus known only from southwestern United States south to Ecuador.

Revision: Smith, 1971. Amer. Ent. Soc, Trans. 97: 537-543.

Neoptilia malvacearum (Cockerell)
Tex., N. Mex., Ariz.; Mexico. Host: Althaea rosea, Malvastrum,
Malva, Sphaeralcea.
Nematoneura malvacearum Cockerell, 1894. U. S. Dept. Agr., Insect Life 7: 253. ♀, ♂.

Taxonomy: Smith, 1972. Amer. Ent. Soc, Trans. 98: 179 (larva).

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Biology: Cockerell, 1894. U. S. Dept. Agr., Insect Life 7: 251-253.

Neoptilia tora Smith
S. Tex.

Neoptilia tora Smith, 1971. Amer. Ent. Soc, Trans. 97: 540. ♀, ♂.

Genus PTENUS Kirby

Pteyios Norton, 1872. Amer. Ent. Soc, Trans. 4: 77. Preocc. by Gray, 1843.

Type-species: Ptenos niger Norton. Desig. by Rohwer, 1911.
Ptenus Kirby, 1882. List Hym. Brit. Mus., v. 1, p. 51. N. name.
Ptenellus Malaise, 1937. Ent. Tidskr. 58: 56.

Type-species: Ptenos nigripectusil) Norton. Orig. desig.

This genus is found only in the New World and is distributed from southwestern United
States to northern South America.
Revision: Smith, 1970. Amer. Ent. Soc, Trans. 96: 79-101.

Ptenus apulatus Greenbaum
Ariz.

Ptenus apulatus Greenbaum, 1975 (1974). Pan-Pacific Ent. 50: 423-425. ♀.

Ptenus bicolor Smith
Tex.; Mexico. Host: Acacia faniesiana.

Ptenus bicolor Smith, 1970. Amer. Ent. Soc, Trans. 96: 83. ♀, ♂.

Taxonomy: Smith, 1972. Amer. Ent. Soc, Trans. 98: 176 (larva).

imus Smith. Tex.; Mexico.

Ptejuis ivius Smith, 1970. Amer. Ent. Soc, Trans. 96: 84. ♀.

Ptenus jocus Smith
Ariz. (Patagonia Mts.).

Ptenus jocus Smith, 1970. Amer. Ent. Soc, Trans. 96: 86. ♀.

Ptenus magnus Smith
W. Tex., N. Mex., Ariz.

Ptenus magnus Smith, 1970. Amer. Ent. Soc, Trans. 96: 86. ♀, ♂.

Ptenus modestius Smith
Tex., Ariz., s. Calif. Host: Adults collected from mesquite.

Ptenus modestius Smith, 1970. Amer. Ent. Soc, Trans. 96: 88. ♀, ♂.

Ptenus nigropectus (Norton)
Miss., La., Tex. Host: Acacia angustissima, Desmayithusi?)-

Ptenos nigropectus Norton, 1872. Amer. Ent. Soc, Trans. 4: 77. ♂.

Ptenus parvus Smith
W. Tex., Ariz. Host: Adults collected from Acacia.

Ptenus parvus Smith, 1970. Amer. Ent. Soc, Trans. 96: 92. ♀, ♂.

Ptenus patulus Smith
Ariz.

Ptenus patulus Smith, 1970. Amer. Ent. Soc, Trans. 96: 94. ♀, ♂.

Ptenus texanus (Norton)
Tex.; Mexico.

Ptilia texana Norton, 1869. Amer. Ent. Soc, Trans. 2: 367. ♂.
Ptenos niger Norton, 1872. Amer. Ent. Soc, Trans. 4: 77. ♂.

Ptenus torridus Smith
Ariz.

Ptenus torridus Smith, 1970. Amer. Ent. Soc, Trans. 96: 96. ♀.

Ptenus vanus Smith
N. Mex., Ariz.; Mexico.

Ptenus vanus Smith, 1970. Amer. Ent. Soc, Trans. 96: 96. ♀, ♂.

Ptenus vargus Smith
Tex.; Mexico.

Pte7ius vargiis Smith, 1970. Amer. Ent. Soc, Trans. 96: 98. ♀.

Genus SCHIZOCERELLA Forsius

Schizocerella Forsius, 1927. Notulae Ent. 7: 19.

Type-species: Schizocerella axillaris Forsius. Orig. desig.
Sofus Malaise, 1937. Ent. Tidskr. 58: 137.

Type-species: Scbizocera pilicomis Holmgren. Orig. desig.
Sterictiphora subg. Leston Ross, 1937. 111. Biol. Monog. 15(2): 54.

Type-species: Schizocerus zabriskei Webster and Malley. Orig. desig.

Taxonomy: Benson, 1963. Ann. and Mag. Nat. Hist. (13) 5: 631. —Smith, 1971. Ar.
Soc, Trans. 97: 543-546.

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Schizocerella pilicornis (Holmgren)
N. Y. and Ont. to Fla. w. to Wash., Oreg., Calif, and to Argentina.
Ecology: Leafminer. A larva of this species may mine a Poiinlaca leaf, leave it, and
then mine a succession of leaves before becoming full grown. The mature larva pupates
in the soil. Host: Portulaca. Introduced into Australia.

Schizocera piliconiis Holmgren, 1868. Eug. Resa I, Zool. I, Ins. p. 391. ♀, ♂.

Schizoceros lateralis Konow, 1899. Buenos Aires Mus. Nac. de Hist. Nat., An. 6: 404. ♀.

Shizocenis zabriskei Webster and Malley, 1900. Canad. Ent. 32: 51.

Schizocenis sericeifonnis Rohwer, 1908. N. Y. Ent. Soc, Jour. 16: 112. ♂.

Schizocenis lineaius Rohwer, 1909. Canad. Ent. 41: 12. ♀.

Schizocenis collaris Rohwer, 1909. Canad. Ent. 41: 13. ♀.

Schizocenis johnsoiii MacGillivray, 1909. Canad. Ent. 41: 403. ♀.

Schizocerella axillaris Forsius, 1927. Notulae Ent. 7: 19.

Schizocerella scapularis Forsius, 1927. Notulae Ent. 7: 19. ♀.

Taxonomy: Smith, 1972. Amer. Ent. Soc, Trans. 98: 171 (larva).

Biology: Webster and Malley, 1900. Canad. Ent. 32: 51-54. — Garlick, 1922. Canad. Ent. 54:
240. -Force, 1965. Calif. Dept. Agr. Bui, v. 54, pp. 157-160.

Genus SPHACOPHILUS Provancher

This genus is found only in the New World, where most of its representatives are found from
southwestern United States to South America. A few species extend north through the central
plains region to central Canada.

Revision: Smith, 1971. Amer. Ent. Soc, Trans. 97: 549-580.

Genus SPHACOPHILUS Subgenus SPHACOPHILUS Provancher

Sphacophiiiis Provancher, 1889. Addit. Corr. Faune Ent. Canada Hym., Sup., p. 427.
Type-species: Sphacophiiiis craivii Provancher. Monotypic

Sphacophilus crawii Provancher
Idaho, Oreg. s. to N. Mex., Ariz., Calif.

Sphacophiiiis crawii Provancher, 1889. Addit. Corr. Faune Ent. Canada Hym., Sup., p. 427. ♀.

Sphacophilus invitus (Cresson)
Wash. s. to Colo., Ariz., Cahf.

Schizocera iiivita Cresson, 1880. Amer. Ent. Soc, Trans. 8: 3. ♀.

Genus SPHACOPHILUS Subgenus LITOCOLUS Smith

Sphacophiiiis subg. Litocolus Smith, 1971. Amer. Ent. Soc, Trans. 97: 556.
Type-species: Atomacera celiularis Say. Grig, desig.

Known hosts for this subgenus are members of the plant family Convolvulaceae.

Sphacophilus apiculus Smith
S. Ariz.

Sphacophiiiis {Litocolus) apiculus Smith, 1971. Amer. Ent. Soc, Trans. 97: 556. ♀.
celiularis (Say). Conn, to Fla. w. to Ont., Minn., Nebr., Ark. Host: Ipomoea, Convolvulus.

Atoviacera celiularis Say, 1836. Boston Jour. Nat. Hist. 1: 213. ♀.

Schizocenis ebenus Norton, 1867. Amer. Ent. Soc, Trans. 1: 55. ♂.

Schizocenis privatus Norton, 1867. Amer. Ent. Soc, Trans. 1: 56. ♀.

Taxonomy: Smith, 1972. Amer. Ent. Soc, Trans. 98: 174 (larva).

Biology: Riley and Howard, 1888. U. S. Dept. Agr., Insect Life 1: 43-45. — Marlatt, 1892. U.
S. Dept. Agr., Insect Life 5: 24-27.

Sphacophilus orthius Smith
Ariz.

Sphacophiiiis (Litocolus) orthius Smith, 1971. Amer. Ent. Soc, Trans. 97: 558. ♀.

Sphacophilus orus Smith
Ariz.

Sphacophiiiis (Litocolus) onis Smith, 1971. Amer. Ent. Soc, Trans. 97: 559. ♀, ♂.

Sphacophilus partitus Smith
Ariz. (Chiricahua Mts.).

Sphacophiiiis (Litocolus) partitus Smith, 1971. Amer. Ent. Soc, Trans. 97: 560. ♀.

Sphacophilus quixus Smith
Tex., Ariz. Host: IpomoeaU).

Sphacophiiiis (Litocolus) quixus Smith, 1971. Amer. Ent. Soc, Trans. 97: 561. ♀, ♂.

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Genus SPHACOPHILUS Subgenus CEOCOLUS Smith

Sphacophilus subg. Ceocoius Smith, 1971. Amer. Ent. See, Trans. 97: 562.
Type-species: Schizocenis abdoniinalis Cresson. Orig. desig.

Known hosts for species of this subgenus are members of the plant family Leguminosae.
These sawflies appear to be most common in the prairie regions of central North America.

Sphacophilus abdominalis (Cresson)
N. C; S. Dak., 111. w. to Colo., N. Mex. Host: P.wralea.

Schizocenis abdoininalis Cresson, 1865. Ent. Soc. Phila., Proc. 4: 243. ♂.

Sphacophilus albicosta Smith
Idaho, Wash., Oreg. Host: Psoralea.

Sphacophilus (Ceocoius) albicosta Smith, 1971. Amer. Ent. Soc, Trans. 97: 563. ♀, ♂.

Sphacophilus apios (Ross)
N. C, 111., Kans. Host: Desniodiurn, Psoralea.

Sterictiphora apios Ross, 1933. Ent. Soc. Wash., Proc. 35: 13. 9, d, larva.

Sterictiphora apios form atresce^is Ross, 1933. Ent. Soc. Wash., Proc. 35: 14. ♀, ♂.

Biology: Ross, 1933. Ent. Soc. Wash., Proc. 35: 13-19.

Sphacophilus argutus Smith
Ariz. (Tucson Mts.).

Sphacophilus {Ceocoius) argidus Smith, 1971. Amer. Ent. Soc, Trans. 97: 566. ♀.

Sphacophilus crenus Smith
W. Tex.

Sphacophilus (Ceocoius) crenus Smith, 1971. Amer. Ent. Soc, Trans. 97: 567. ♀, ♂.

Sphacophilus dissensus Smith
Ariz. (Indian Pine).

Sphacophilus (Ceocoius) dissensus Smith, 1971. Amer. Ent. Soc, Trans. 97: 568. ♀.

Sphacophilus hamus Smith
S. Ariz.

Sphacophilus (Ceocoius) hannis Smith, 1971. Amer. Ent. Soc, Trans. 97: 568. ♀, ♂.

Sphacophilus memmonius Smith
S. Calif. (Algadones Sand Dune, Imperial Co.).

Sphacophilus (Ceocoius) n)enimonius Smith, 1971. Amer. Ent. Soc, Trans. 97: 570. ♀.

Sphacophilus nigriceps (Konow)
Man., Minn., 111. w. to Alta., Wyo., Colo., N. Mex., Tex. Host: Petalostemum.

Brachyphafnus nigriceps Konow, 1906. Ztschr. System. Hym. Dipt. 6: 251. ♀.

Schizocenis nortoni Rohwer, 1909. Canad. Ent. 41: 11. ♂.

Taxonomy: Smith, 1972. Amer. Ent. Soc, Trans. 98: 175 (larva).

Sphacophilus oblatus Smith
N. Mex., Ariz.

Sphacophilus (Ceocoius) oblatus Smith, 1971. Amer. Ent. Soc, Trans. 97: 576. ♀.

Sphacophilus plumiger (Klug)
N. C, Ga., Fla. Host: Petalostemum.

Hylotoma plumiger Klug, 1814. Mag. Gesell. Naturf. Freunde Berlin 6: 306. 9. "New

York" (!).
Cry plus Klugii Leach, 1817. Zool. Misc., v. 3, p. 125. ♀, ♂.

Taxonomy: Smith, 1973. Ent. Soc. Wash., Proc. 75: 28-29.

Sphacophilus precarius Smith
Ga., Fla.

Sphacophilus (Ceocoius) precarius Smith, 1971. Amer. Ent. Soc, Trans. 97: 578. ♀.

Sphacophilus rallus Smith
Ariz. (Santa Catalina Mts.).

Sphacophilus (Ceocoius) rallus Smith, 1971. Amer. Ent. Soc, Trans. 97: 578. ♀.

Sphacophilus triangularis Smith
Ariz. (Douglas).

Sphacophilus (Ceocoius) triangularis Smith, 1971. Amer. Ent. Soc, Trans. 97: 580. ♀.

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