Diploplectron beccum Parker
Calif., Nev., Utah, Ariz., southwest. Tex.

Diploplectron beccum Parker, 1972. Ent. Soc. Amer., Ann. 65: 1195, figs. 5, 19, 32. ♂, ♀.

Diploplectron brunneipes (Cresson)
Idaho, Wyo., Colo., Calif. Ecology: Makes a nest of 2-3 cells in

hard-packed sandy clay, stores 6 prey per cell. Prey: Uhleriola floralis (Uhler), nymphs.
Predator: Philanthus pacificus Cr.
Liris i brunneipes Cresson, 1881. Amer. Ent. Soc, Trans. 9: Proc p. iii. 9 (misdet. in

part), 6.
Diploplectron bidentatns Ashmead, 1899. Ent. News 10: 56. "♀"=♂.
Diploplectron foxii Ashmead, 1899. Ent. News 10: 56. ♀.
Diploplectron bidentatiformis Rohwer, 1909. Amer. Ent. Soc, Trans. 35: 121. ♂.




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Taxonomy: Krombein, 1939. Brooklyn Ent. Sec, Bui. 34: 138 (synonymy). —Evans, 1975.
Great Basin Nat. 35: 123-125, 5 figs, (larva).

Biology: Evans, 1975. Great Basin Nat. 35: 123 (nest, prey).

Diploplectron californicum Parker
Calif., Oreg., Idaho. Prey: Rhyparochromus califomicus Van D. adults,
Emblethis vicarius Horv. ? nymphs, Megalonotus chiragnis (F.) adult.
Diploplectron californicum Parker, 1972. Ent. Soc. Amer., Ann. 65: 1199, figs. 1, 13, 25. ♂.

9.

Biology: Williams, 1946. Hawaii. Ent. Soc, Proc. 12: 648 (prey).

Diploplectron diablense Williams
Coast Range and Sierra Nevada Mts. of Calif.

Diploplectron diablensis Williams, 1951 (1950). Wasmann Jour. Biol. 8: 363, fig. 1. ♂.

Taxonomy: Williams, 1959. Wasmann Jour. Biol. 17: 303, fig. 5. ♀.

Diploplectron ferrugineum Ashmead
Idaho, Colo., N. Mex., Ariz., Calif.

Diploplectron ferrugineus Ashmead, 1899. Ent. News 10: 56. "♀"=♂.

Diploplectron ashmeadi Rohwer, 1909. Amer. Ent. Soc. Trans. 35: 122. ♂.

Diploplectron cressoni Rohwer, 1909. Amer. Ent. Soc, Trans. 35: 123. ♀.

Diploplectron relativus Rohwer, 1909. Amer. Ent. Soc, Trans. 35: 123. ♀.

Diploplectron fossor Rohwer
B. C, Wyo., Colo., Utah, Nev., Calif. Prey: Labops sp. nymphs. Predator:
Philanthus pacificus Cr.

Diploplectron fossor Rohwer, 1909. Amer. Ent. Soc, Trans. 35: 120. ♀.

Diploplectron rufoantennatus Rohwer, 1909. Amer. Ent. Soc, Trans. 35: 120. ♂.

Biology: Parker, 1972. Ent. Soc. Amer., Ann. 65: 1193 (prey).

Diploplectron irwini Parker
South Ariz.

Diploplectron irwini Parker, 1972. Ent. Soc. Amer., Ann. 65: 1200, figs. 7, 12. ♂.

Diploplectron kantsi Pate
Southwest. Tex., south. N. Mex.; Mexico (Tamaulipas).

Diploplectron kantsi Pate, 1941. Ent. News 52: 6. ♂.

Diploplectron peglowi Krombein
N. Y., N. W. T., Wyo., Colo., Utah, Idaho, Nev., Calif. Ecology: Nests in

sand, making either a unicellular nest or 3-4 cells per nest and storing 2-6 prey per cell.
Parasite: Hedychridium sp. ? Prey: Sphaerobius iyisigyiis (Uhl), Lygaeus sp., Emblethis
vicarius Horv., Sphragisticus nebulosus (Fall); Aufeius impressicollis Stal. All prey
specimens were nymphs.
Diploplectron peglowi Krombein, 1939. Brooklyn Ent. Soc, Bui. 34: 136. ♀, ♂.

Biology: Williams, 1946. Hawaii. Ent. Soc, Proc. 12: 648 (prey, nest). —Parker, 1972. Ent.
Soc. Amer., Ann. 65: 1193 (nest, prey). — Kurczewski, 1972. Ent. Soc. Wash., Proc. 74:
385-397, 4 figs, (nest, prey hunting and transport). —Kurczewski, 1975. Ent. Soc. Wash.,
Proc. 77: 97-99 (nest, prey, egg).

Diploplectron reticulatum Williams
South. Calif, and Ariz.

Diploplectron reticulatus WilUams, 1946. Hawaii. Ent. Soc, Proc. 12: 648. ♂.

Taxonomy: Williams, 1951 (1950). Wasmann Jour. Biol. 8: 365, fig. 2. ♀.

Diploplectron secoense Parker
Calif, in Coast Range and Sierra Nevada Mts.

Diploplectron secoense Parker, 1972. Ent. Soc. Amer., Ann. 65: 1201, figs. 7, 20, 29. ♂, ♀.

Diploplectron sierrense Parker
Sierra Nevada Mts. of Calif, and Nev.

Diploplectron sierrense Parker, 1972. Ent. Soc. Amer., Ann. 65: 1199, figs. 5, 15, 30, 36, 37.
6, 9.

Diploplectron vierecki Pate
South. Ariz, and N. Mex., southwest. Tex. Prey: Microporus obliquus Uhl.
nymph.
Diploplectron vierecki Pate, 1941. Ent. News 52: 4. ♂.

Dimorpha Panzer, 1806. Krit. Rev. Insektenf. Deutschlands, p. 126.
Type-species: Tiphia abdominalis Panzer. Monotypic.

The species of Astata build multicellular nests in soU of varying types ranging from sandy to
hard-packed with included gravel. The cells may be in short series in some species separated by
earthen partitions. They prey upon Hemiptera, principally Pentatomidae and Lygaeidae, and
usually the nymphal stages.

Revision: Fox, 1892. Canad. Ent. 24: 232-235 (N. Amer. spp.). —Fox, 1894 (1893). Acad. Nat.
Sci. Phila., Proc. 45: 539-548 (N. Amer. spp.). —Parker, 1962. Ent. Soc. Amer., Ann. 55:
643-659, 42 figs. (spp. north of Mexico). —Parker, 1964. Ent. Soc. Amer., Ann. 57: 552-559,
21 figs. (Mexican and Cent. Amer. spp.).

Astata bakeri Parker
Ont., Sask. to B. C, south to Tex. and Calif.; Mexico (Chihuahua, Jalisco,

Guanajuato, Morelos, Puebla, Chiapas). Prey: Nysius raphanus How. adult and nymph,
Lygaeus bicrucis Say ? nymph. Predator: Philanthus crabroniformis Sm.
Astata bakeri Parker, 1962. Ent. Soc. Amer., Ann. 55: 647, text fig. 1, figs. 11, 21. ♂, ♀.

Biology: Williams, 1946. Hawaiian Ent. Soc, Proc. 12: 647 (prey; rec. as Astata sp.).

Astata bechteli Parker
Desert areas. Wash, and Colo., south to CaUf. and Tex.; Mexico (Sonora).

Astata nubecula bechteli Parker, 1962. Ent. Soc. Amer., Ann. 55: 649. ♂, ♀.

Astata bicolor Say
Southern Canada and U. S. east of Rocky Mts.; Mexico (Chihuahua, Sonora,

Durango, San Luis Potosi, Jalisco, Guanajuato, Mexico). Ecology: Nests in hard-packed
sand, sometimes beneath overhanging vegetation, provides up to 5 cells per nest with
6-8 nymphs per cell. Prey: Pentatomidae spp.
Astata bicolor Say, 1823. West. Quart. Rptr. 2: 78. (J, 9.
Astata terminata Cresson, 1872. Amer. Ent. Soc, Trans. 4: 218. ♂.
Astatus pygidialis Fox, 1892. Canad. Ent. 24: 234. ♀.

Taxonomy: Evans, 1959. Amer. Ent. Soc, Trans. 85: 165-166 (larva).

Biology: Peckham and Peckham, 1898. Wis. Geol. Nat. Hist. Survey, Bui. 2: 92-97, 217, pis. 1,
4, 9 (nest, prey). —Peckham and Peckham, 1905. Wasps Social and Solitary, pp. 287-289
(orientation flight). — Mickel, 1918 (1917). Nebr. Univ., Studies 17: 108 (prey). —Evans,
1962. N. Y. Ent. Soc, Jour. 70: 30-31 (nest, prey).

Astata bigeloviae Cockerell
Western Tex. to Calif.; Mexico (Jalisco).

Astatus bigeloviae Cockerell, 1897. In Cockerell and Fox, Acad. Nat. Sci. Phila., Proc. 49:
138. ♀.

Astata boharti Parker
N. Mex., Ariz.

Astata boharti Parker, 1962. Ent. Soc. Amer., Ann. 55: 650, fig. 25.

Astata clypeata Parker
East slope of Rocky Mts. in U. S.; Mexico (western mts.).

Astata clypeata Parker, 1962. Ent. Soc. Amer., Ann. 55: 651, figs. 15, 26. cJ, 9.

Astata femorata Parker
Ariz. (Pena Blanca in Santa Cruz Co.).

Astata femorata Parker, 1963. Pan-Pacific Ent. 39: 185, figs. 1-3, 6.

Astata leuthstromi Ashmead
Alaska, Canada, northern U. S., south in mts. to Ariz. Ecology: Nests in
soil, sometimes beneath overhanging vegetation, provides 2 cells per nest with up to 5
nymphal prey per cell. Prey: Acrostemum hilare Say, Cosmopepla bimaculata Thom.
Astata leuthstromi Ashmead, 1897. Psyche 8: 129. ♀.

Biology: Peckham and Peckham, 1898. Wis. Geol. Nat. Hist. Survey, Bui. 2: 98 (nest, prey).
—Evans, 1958 (1957). N. Y. Ent. Soc, Jour. 65: 176-177 (nest, prey transport). —Evans,
1962. N. Y. Ent. Soc, Jour. 70: 31-32 (nest, prey).

Astata mexicana Cresson
Canada and U. S. west of Rocky Mts.; Mexico south to El Salvador.

Astata mexicana Cresson, 1881. Amer. Ent. Soc, Trans. 9: Proc p. v. S ■

Astata nevadica Cresson
Western Canada and U. S.; Mexico (Chihuahua, Sonora, Durango, Zacatecas,
Guanajuato, Mexico).
Astata nevadica Cresson, 1881. Amer. Ent. Soc, Trans. 9: Proc. p. v. 9.

Astata nubecula Cresson
N. W. T., northern U. S. south to Va. in east and along mts. in Great Basin
and in Calif, to San Diego. Ecology: Nests in hard, bare stony soil or in sand, provides
up to 8 cells per nest, each cell provisioned with 2-4 nymphs. Parasite: Senotainia




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trilineata Wulp, Hilarella hilarella Zett. ? Prey: Chlorochroa uhleri Stal ?, Thyanta sp.,
prob. casta Stal or pallidovirens Stal, Pentatomidae sp. Predator: Philanthus zebratus
nitens (Bks.), P- crabroniformis Sm.

Astata nubecula Cresson, 1865. Ent. Soc. Phila., Proc. 4: 466. ♂.

Astata nigropilosa Cresson, 1881. Amer. Ent. Soc, Trans. 9: Proc. p. iv. ♂, ♀.

Biology: Parker, 1962. Ent. Soc. Amer., Ann. 55: 649 (nest, prey). —Evans, 1970. Mus.

Compar. Zool., Bui. 140: 487-488 (male territoriality, nest, prey, parasites).

Astata occidentalis Cresson
South. Canada, U. S.; Mexico (Chihuahua, Baja California, Guanajuato,
Morelos, Michoacan, Puebla). Ecology: Nests in hard-packed clay devoid of vegetation,
makes up to 14 cells per nest, and stores 3-9 usually adult prey per cell. Parasite:
Senotainia trilmeata Wulp; Diptera sp. Prey: Hymenarcys nervosa Say, Thyanta
calceata (Say), T. p. pallidovirens Stal, T. p. accerra McAtee, T. brevis Van D., T.
custator P., T. punctiventris Van D., Euschistus variolarius (Beauv.), Holcostethus
limbolaritis (Stal), Banasa calva (Say), Perillus bioculatus (P.), Trichopepla sp.

Astata occidentalis Cresson, 1881. Amer. Ent. Soc, Trans. 9: Proc. p. iii. ♂.

Astata apicipennis Cameron, 1890. Biol. Cent.- Amer., Hym., v. 2, p. 66., pi. 5, fig. 1. ♀.

Astata tinctipennis Cameron, 1890. Biol. Cent.- Amer., Hym. v. 2, p. 67. ♂.

Astatus sayi Fox, 1894 (1893). Acad. Nat. Sci. Phila., Proc. 45: 542. ♀.

Taxonomy: Evans, 1958. Amer. Ent. Soc, Trans. 84: 110-111, figs. 1-8 (larva). —Parker, 1964.
Ent. Soc. Amer., Ann. 57: 559 (synonymy).

Biology: Evans, 1958 (1957). N. Y. Ent. Soc, Jour. 65: 168-176, figs. 3-8 (nest, prey transport,
parasite). —Powell and Burdick, 1960. Pan-Pacific Ent. 36: 25-30 (nest, prey transport,
parasite).

Astata unicolor Say
Southern Canada, U. S.; Mexico to El Salvador, Cuba, Jamaica. Ecology: Nests

usually beneath overhanging vegetation in hard-packed sand or heavier soil containing

gravel, makes 12-14 cells in a complete nest, and stores 2-4 nymphal prey per cell.

Parasite: Chrysis sp. Prey: Podisus modestus F., P. maculiventris (Say), Etischistus

tristigmus Say, E. euschistoides (VoU.) ?
Astata unicolor Say, 1824. In Keating, Narr. Long's 2nd Exped. v. 2, p. 337. ♀.
Astata insularis Cresson, 1865. Ent. Soc. Phila., Proc. 4: 140. ♀.
Astata rufiventris Cresson, 1872. Amer. Ent. Soc, Trans. 4: 218. ♀.

Taxonomy: Evans, 1958. Amer. Ent. Soc, Trans. 84: 111-112, figs. 46-48 (larva).

Biology: Peckham and Peckham, 1898. Wis. Geol. Nat. Hist. Survey, Bui. 2: 88-92, pis. 9, 11,
12 (nest, prey, parasite). —Peckham and Peckham, 1905. Wasps Social and Solitary, pp.
289-290 (orientation flight). —Evans, 1958 (1957). N. Y. Ent. Soc, Jour. 65: 160-168, figs. 1,
2 (nest, prey hunting and transport, male behavior, life cycle).

Morphology: Snodgrass, 1941. Smithsn. Inst., Misc. Collect. 99 (14): pi. 20, figs. L, M (male
genitalia).

Astata williamsi Parker
Oreg., Idaho, Utah, Nev., Calif.

Astata williamsi Parker, 1962. Ent. Soc. Amer., Ann. 55: 653, figs. 14, 29, 35, 36. cJ, 9.

Dryudella bella (Cresson)
Idaho, Nev., Calif., Ariz., N. Mex.; Mexico.

Astata bella Cresson, 1881. Amer. Ent. Soc, Trans. 9: Proc. p. vi. ♂.
caerulea (Cresson). Wash., Idaho, Colo., Nev., Calif., Ariz., N. Mex.; Mexico south to Puebla.
Ecology: Nests in sand under vegetation, 3 cells per nest, with 4-6 prey per cell. Prey:
Hyalmenus tarsatus (F.) nymphs; Euschistus conspersus Uhl. nymphs.
Astata caerulea Cresson, 1881. Amer. Ent. Soc, Trans. 9: Proc. p. iv. ♂.




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Biology: Alcock, 1973. Wasmann Jour. Biol. 31: 328-329, fig. 4 (nest, prey).
clegans (Cresson). Idaho, Wyo., Utah, Nev., Calif., Ariz.

Astata elegans Cresson, 1881. Amer. Ent. Soc, Trans. 9: Proc. p. vi. ♂, ♀.

Dryudella immigrans (Williams)
Wash., Idaho and Wyo. south to Calif, and southwest. Tex.; Hawaii;
Mexico south to Chiapas including Baja California. Prey: Nysius caenosulus Stal
nymphs. Adventive in Hawaii.
Astata immigrans Williams, 1946. Hawaii. Ent. Soc, Proc. 12: 641. ♂, ♀.

Biology: Williams, 1946. Hawaii. Ent. Soc, Proc. 12: 642-648 (prey, nesting under lab.
conditions).
millsi Cockerell. Colo. (Canad. Zone of Long's Peak).

Dryudella millsi Cockerell, 1914. Ent. News 25: 32. ♀.

Dryudella montana (Cresson)
Wash, to Wyo. south to Calif, and Colo. Ecology: Nests in flat, bare sand,
makes 2-3 cells per nest. Parasite: Senotainia trilineata (Wulp). Prey: Zelus or
Pselliopus sp.; Eurygaster altematus Say.; Corim.elaena montana Van D.; Alydus sp.;
Aufeius impressicollis Stal; Euschistus conspersus Uhl. All observed prey were
nymphs.
Astata montana Cresson, 1881. Amer. Ent. Soc, Trans. 9: Proc p. v. 9.
Diploplectron florissantensis Rohwer, 1909. Amer. Ent. Soc, Trans. 35: 122. ♀.

Biology: Evans, 1963. Ent. News 74: 236 (nest, prey). —Parker, 1969. Ent. Soc. Amer., Ann.
62: 965 (prey). —Alcock, 1973. Wasmann Jour. Biol. 31: 324-326, figs. 2, 3, 7, 8 (nest, prey
transport, parasite).

Dryudella pernix Parker
South. Calif, to southwest. Tex.; Mexico (Hidalgo).

Dryudella pernix Parker, 1969. Ent. Soc. Amer., Ann. 62: 972, figs. 13, 22, 25, 40, 54, 66. ♂.
9.

Dryudella picta (Kohl)
N. W. T., Oreg., Calif., Idaho, Nev., Utah, Colo., Ariz.; Mexico south to Morelos.
Prey: Liorhyssus hyalinus (F.) adult; Lygus sp. nymphs in laboratory.
Astatus pictus Kohl, 1888. Zool.-Bot. Gesell. Wien, Verhandl. 38: 146. ♂.
Astata Kohli Cameron, 1890. Biol. Cent.- Amer., Hym., v. 2, p. 68, pi. 5, figs. 2, 2a. 9 .
Astatus asper Fox, 1894 (1893). Acad. Nat. Sci. Phila., Proc. 45: 546. ♂, ♀.
Astata aspera Dalla Torre, 1897. Cat. Hym., v. 8, p. 651. Emend.
Astata asperiformis Rohwer, 1909. Ent. News 20: 371. ♂.

Biology: Parker, 1969. Ent. Soc. Amer., Ann. 62: 965 (prey). — Steiner, 1973. Quaestiones Ent.
9: 23 (mating flight).

Dryudella pinguis (Dahlbom)
Alaska, N. W. T., Colo.; northern Europe and Asia. Prey: Trapezonotus
arenarius (L.), Drymus sylvaticus (F.), Nysius thymi (Wolff) ? in Europe.
Larra pinguis Dahlbom, 1832. Exercit. Hym., fasc. 4, p. 50. ♀.
Astata jaculator Smith, 1845. Zoologist 3: 1157. ♀.

Biology: Verhoeff, 1951. Zool. Meded. 31: 162 (prey in Europe).

Dryudella rhimpa Parker
B. C. to Calif., and Idaho and Wyo. to N. Mex.; Mexico south to Queretaro.
Ecology: Nests in compact soil containing gravel. Prey: Thyanta punctiventris Van D.
nymphs and adult; Lygaeus sp. nymph; Leptocoris trivittatus (Say) nymph.
Dryudella rhimpa Parker, 1969. Ent. Soc. Amer., Ann. 62: 970, figs. 5, 8, 17, 30, 44, 55, 62.
6, 9.

Biology: Parker, 1969. Ent. Soc. Amer., Ann. 62: 965 (nest, prey).




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