Krombein (1948, Ent. Soc. Amer., Ann. 41: 58-62). Of the liberated species only vemalis Rohwer,

Tiphia popilliavora Rohwer
and asericae Allen and Jaynes established themselves; stemata Parker is
doubtfully estabhshed; biseculata Allen and Jaynes, castaneaevora Parker, chosensis Allen,

Tiphia frater Parker
matura Allen and Jaynes, ywtopolita var. alleyii Roberts, pullivora Allen and
Jaynes, and totopunctata Allen and Jaynes apparently did not become established.

Tiphia femorata Fabricius and T. moyio Fabricius were introduced from Europe and liberated
in New York in 1956 and 1957 for control oi Amphimallon majalis (Raz.), the European chafer;
so far as is known neither species became established.

Only the typical subgenus occurs in the New World.

Revision: Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 1-24, 13 figs, (eastern species).
—Allen, 1971. Amer. Ent. Soc, Trans. 97: 201-359, 147 figs., 13 maps (species of western
North America).

Taxonomy: Allen, 1961. Amer. Ent. Soc, Trans. 87: 1-20 (key to species north of Mexico).
—Evans, 1965. Ent. Soc. Wash., Proc 67: 91 (larva).

Tiphia alamosae Allen
Colo. (Alamosa Co.); Mexico (Chihuahua).

Tiphia alamosae Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 54, figs. 1, 6, 11. ♂, ♀.

Taxonomy: Allen, 1971. Amer. Ent. Soc, Trans. 97: 264, figs. 63-68, map 5. ♂, ♀.

Tiphia andersoni Allen
Ariz, south in Mexico to Michoacan and Morelos.

Tiphia andersoni Allen, 1971. Amer. Ent. Soc, Trans. 97: 288, figs. 100-103, map 8. c?, 9.

Tiphia anguis Allen
Wyo., B. C. to Calif.

Tiphia anguis Allen, 1971. Amer. Ent. Soc, Trans. 97: 278, figs. 94-98, map 7. ♂, ♀.

Tiphia arcuata Allen
Ariz. (Chiricahua Mts.).

Tiphia arcuata Allen, 1971. Amer. Ent. Soc, Trans. 97: 315, figs. 127, 129, 130, map 11. ♀.

Tiphia asensoria Allen
Fla. (Fort Lauderdale).

Tiphia asensoria Allen, 1966. Amer. Ent. Soc, Trans. 92: 317, map 12. ♂.

Tiphia asericae Allen and Jaynes
N. Y., N. J., Pa.; Korea. Introduced from Korea. Host: Autoserica
castanea (Arrow); Serica peregrina Chapin.

Tiphia asericae Allen and Jaynes, 1930. U. S. Natl. Mus., Proc. 76 (17): 74. ♀, ♂.

Biology: Clausen, Jaynes, and Gardner, 1933. U. S. Dept. Agr., Tech. Bui. 366: 45-46.
—Gardner and Parker, 1940. U. S. Dept. Agr., Tech. Bui. 738: 21. —Clausen, 1956. U. S.
Dept. Agr., Tech. Bui. 1139: 131 (life history).

Tiphia atlantis Allen
N. H. to Fla., Mich., 111., Kans., Ariz. ?

Tiphia atlantis Allen, 1934. Amer. Ent. Soc, Trans. 60: 314. ♀.

Taxonomy: Allen, 1961. Amer. Ent. Soc, Trans. 87: 16. 6. —Allen, 1966. Amer. Ent. Soc,
Trans. 92: 336, figs. 23, 32, map 17. ♀, ♂.

Tiphia barberi Allen
Mont., Wyo., Ariz.

Tiphia barberi Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 57, figs. 13, 28, 29. ♂.

Taxonomy: Allen, 1971. Amer. Ent. Soc, Trans. 97: 270, figs. 75-78, map 6. cJ, 9.

Tiphia berbereti Allen
Nebr. (Ainsworth). Host: Phyllophaga anxia (LeC).
Tiphia berbereti Allen, 1970. Ent. Soc. Amer., Ann. 63: 473. ♀, ♂.

Taxonomy: Allen, 1971. Amer. Ent. Soc, Trans. 97: 274, figs. 82-85, map 6. ♀, ♂.

Biology: Berberet and Helms, 1970. Ent. Soc. Amer., Ann. 63: 471-473 (life history).

Tiphia biseculata Allen and Jaynes
Liberated in N. Y. and N. J. but not established. Introduced from
Japan. Host: Anoniala orientalis Waterh.; PopiUia japonica Newm.
Tiphia biseculata Allen and Jaynes, 1930. U. S. Natl. Mus., Proc. 76, Art. 17: 85, figs. 30,
31. ♀, ♂.

Biology: Clausen, Jaynes and Gardner, 1933. U. S. Dept. Agr., Tech. Bui. 366: 38-39 (life
history).

Tiphia boharti Allen
Oreg. (Bend).

Tiphia boharti Allen, 1971. Amer. Ent. Soc, Trans. 97: 287, map 8. ♂.




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Tiphia canamexica Rohwer
Mont., S. Dak., Colo., Utah, Calif., Ariz, south in Mexico to Chiapas and
Yucatan.
Tiphia canamexica Rohwer, 1912. U. S. Natl. Mus., Proc. 41: 452. ♀.
Tiphia dreisbachi AUen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 10. ♂, ♀.
Tiphia canamexicana (!) Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 85. ♀.

Taxonomy: Allen, 1971. Amer. Ent. Soc, Trans. 97: 243, figs. 41-43, map 3.

Tiphia castaneaevora Parker
Liberated in N. J. and Pa. but not established. Introduced from Japan.
Host: Autoserica castanea (Arrow).
Tiphia castaneaevora Parker, 1937. N. Y. Ent. Soc, Jour. 45: 288. ♀, ♂.

Tiphia chosensis Allen
Liberated in Conn, and N. J. but not established. Introduced from Korea.
Host: Anomala orientalis Waterh.
Tiphia {Tiphia) chosensis Allen, 1969. Amer. Ent. Soc, Trans. 95: 359. ♀, ♂.

Taxonomy: Allen and Jaynes, 1930. U. S. Natl. Mus., Proc. 76, Art. 17: 66, figs. 2, 5, 7, 11, 24.
9,6. (Misdet. as bicarinata Cameron).

Biology: Clausen, Jaynes and Gardner, 1933. U. S. Dept. Agr., Tech. Bui. 366: 43-44 (life
history, as bicarinata).

Tiphia cochiseae Allen
N. Mex., Ariz., south in Mexico to Jalisco and Puebla.

Tiphia cochiseae Allen, 1971. Amer. Ent. Soc, Trans. 97: 282, fig. 99, map 7. ♂.

Tiphia coloradensis Allen
Colo.

Tiphia coloradensis Allen, 1971. Amer. Ent. Soc, Trans. 97: 314, fig. 126, map 11. ♀.

Tiphia conformis Malloch
Mass., Mich, and Nebr. south to Fla. and Tex., Ariz.

Tiphia conformis Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 22. ♀, ♂.

Tiphia imitatrix Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 22. ♀.

Taxonomy: Prison, 1927. 111. Nat. Hist. Survey, Bui. 16: 229 (lectotype designation
conformis). —Allen, 1934. Amer. Ent. Soc, Trans. 60: 312. —Allen, 1966. Amer. Ent. Soc,
Trans. 92: 295, fig. 14, map 10. ♂, ♀.

Tiphia convexa Allen
N. H. to N. C, Mich., Kans., Mont.

Tiphia convexa Allen, 1934. Amer. Ent. Soc, Trans. 60: 306. ♂.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 290, map 9. ♂, ♀.

Tiphia crassipunctata Allen
N. C.

Tiphia crassipunctata Allen, 1934. Amer. Ent. Soc, Trans. 60: 307. ♂.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 254, map 3. ♂.

Tiphia dallasae Allen
Tex. (Dallas, Victoria).

Tiphia dallasae Allen, 1966. Amer. Ent. Soc, Trans. 92: 339, fig. 9, map 17. ♀.

Tiphia dawsoni Allen
Nebr., Mont., Idaho, Tex.

Tiphia dawsoni Allen, 1971. Amer. Ent. Soc, Trans. 97: 259, fig. 62, map 5. ♂.

Tiphia dentonae Allen
Tex., Kans.

Tiphia dentonae Allen, 1966. Amer. Ent. Soc, Trans. 92: 300, map 10. ♂.

Tiphia dryophila Krombein
N. J., N. C, Fla., Ohio.

Tiphia dryophila Krombein, 1953. Wasmann Jour. Biol. 10: 301. ♂, ♀.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 299, map 10. ♂, ♀.

Tiphia egregia Viereck
N. H. to S. Dak., south to Fla. and Tex.
Tiphia eyregia (!) Viereck, 1906. Ent. News 17: 303. ♂.
Tiphia egregia Viereck, 1906. Ent. News 17: 350.

Tiphia tuberculata Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 14. ♀, ♂. Preocc.
Tiphia aterrima Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 19. ♀.
Tiphia papillata Roberts, 1930. Canad. Ent. 62: 190. N. name.
Tiphia diminuta Roberts, 1933. Kans. Ent. Soc, Jour. 6: 91. ♀, ♂.

Taxonomy: Prison, 1927. 111. Nat. Hist. Survey, Bui. 16: 229-230 (lectotype designation
aterrima, tuberculata). —Allen, 1934. Amer. Ent. Soc, Trans. 60: 312. —Allen, 1966.
Amer. Ent. Soc, Trans. 92: 325, fig. 34, map 15. ♂, ♀.

Tiphia elaka Allen
Fla.

Tiphia elaka Allen, 1966. Amer. Ent. Soc, Trans. 92: 289, fig. 6, map 9. ♂.




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Tiphia femorata Fabricius
Liberated in N. Y. but not established. Introduced from Europe. Host:
Amphhnallon majalis (Raz.).

Tiphia femoraia Fabricius, 1775. Systema Ent., p. 353. ♀.

Sphex palmipes Schrank, 1781. Enum. Ins. Austriae, p. 384.

Bethylus glabrata Fabricius, 1798. Sup. Ent. System., p. 254.

Betliylus villosa Fabricius, 1804. Systema Piezatorum, p. 236.

Bethylus pilipe7inis Klug, 1808. Mag. Gesell. Naturf. Freunde Berlin 2: 49. ♂.

Bethylus ater Giraud, 1856. Zool.-Bot. Ver., Verhandl. 6: 184.

Tiphia laeviceps Tournier, 1889. Soc. Ent. Belg., Ann. 33: 21. ♀.

Tiphia austriaca Tournier, 1889. Soc. Ent. Belg., Ann. 33: 24. ♀.

Tiphia fenestrata Klug
N. J., N. C, Fla., 111., Miss., La.

Bethylus fenestratus Klug, 1810. In Weber, Beitr. z. Naturk. 2: 193. ♀.

Tiphia confusa Allen, 1934. Amer. Ent. Soc, Trans. 60: 302. ♀.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 253. 9. —Allen, 1972. Ent. Soc. Wash.,
Proc 74: 381-383 (redescription and synonymy).

Tiphia fisheri Allen
Va. (Cape Henry).

Tiphia fisheri Allen, 1966. Amer. Ent. Soc, Trans. 92: 301, map 10. ♂.

Tiphia flavipennis Spinola
Calif. (?); Mexico (Guerrero).

Tiphia flavipennis Spinola, 1841. Soc. Ent. France, Ann. 10: 102. ♀.
Tiphia elegans Cameron, 1893. Biol. Cent.-Amer., Hym., v. 2, p. 240. ♀, ♂.
Tiphia ochroptera Dalla Torre, 1897. Cat. Hym., v. 8, p. 139. N. name err. proposed for

Tiphia flavipennis Spinola
not Smith, 1857.

Taxonomy: Turner, 1908. Ann. and Mag. Nat. Hist. (8) 2: 131 (synonymy). —Allen, 1962.

Amer. Ent. Soc, Trans. 88: 38 (redescription elegans female).

Tiphia floridana
***authority mismatch
flavida Allen. Central Fla.

Tiphia floridana flavida Allen, 1966. Amer. Ent. Soc, Trans. 92: 279, map 7. ♀, ♂.

Tiphia floridana
***authority mismatch
floridana Robertson. N. Y. to Fla.

Tiphia floridana Robertson, 1901. Amer. Ent. Soc, Trans. 27: 195. ♀, ♂.

Taxonomy: Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 23. —Allen, 1934. Amer. Ent. Soc,
Trans. 60: 314. — Krombein, 1953. Wasmann Jour. Biol. 10: 305. —Allen, 1966. Amer. Ent.
Soc, Trans. 92: 276, figs. 5, 33, 38, map 1. ♂, ♀.

Tiphia fortistriolata Cameron
B. C. south to Calif, and N. Mex.; Mexico (Michoacan, Morelos,
Veracruz).
Tiphia fortistriolata Cameron, 1907. Invertebrata Pacifica 1: 170. ♂.
Tiphia bakeri Allen, 1961. Amer. Ent. Soc, Trans. 87: 12. ♂.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc 117: 50, 74, figs. 4, 15, 27. 9. —Allen,

1969. Amer. Ent. Soc, Trans. 95: 433. 6. —Allen, 1971. Amer. Ent. Soc, Trans. 97: 229,

figs. 23-27, map 1. ♂, ♀.

Tiphia frater Parker
Liberated in N. J. but not established. Introduced from China.

Tiphia frater Parker, 1935. N. Y. Ent. Soc, Jour. 43: 397. ♀, ♂.

Tiphia fulvicauda Cameron
Nev. (Ormsby Co.).

Tiphia fulvicauda Cameron, 1907. Invertebrata Pacifica 1: 169. ♀.

Taxonomy: Allen, 1962. Amer. Ent. Soc, Trans. 88: 24 (type redescription).

Tiphia gehlsbachi Allen
Tex.

Tiphia gehlsbachi Allen, 1971. Amer. Ent. Soc, Trans. 97: 237, figs 32-34, map 2. ♂.

Tiphia greenei Allen
Md. (Plummers Island).

Tiphia greenei Allen, 1966. Amer. Ent. Soc, Trans. 92: 256, fig. 31, map 3. ♂.

Tiphia greeleyi Allen
Kans. (Greeley Co.).

Tiphia greeleyi Allen, 1971. Amer. Ent. Soc, Trans. 97: 258, map 5. ♂.

Tiphia illinoensis Robertson
N. H., Ont., Mich., 111. and Iowa south to Fla. and Tex.

Tiphia illinoensis Robertson, 1901. Amer. Ent. Soc, Trans. 27: 196. ♀, ♂.

Tiphia waldonii (!) Viereck, 1906. Ent. News 17: 302. ♂.

Tphia brunneicornis Viereck, 1906. Ent. News 17: 303. ♂.

Tiphia waldenii Viereck, 1906. Ent. News 17: 350.




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Tiphia roherfsoni Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 23. ♀.
Tiphia waldeni (!) Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 23.

Taxonomy: Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 24. 6. -Prison, 1927. 111. Nat. Hist.
Survey, Bui. 16: 230 (lectotype designation, robertsoni). —Allen, 1934. Amer. Ent. Soc,
Trans. 60: 313. -Allen, 1961. Amer. Ent. Soc, Trans. 87: 18. -Allen, 1966. Amer. Ent.
Soc, Trans. 92: 280, figs. 4, 27, 35, map 8. ♂, ♀.

Tiphia inaequalis Malloch
Mass. to Nebr., south to Ga., Ala. and Tex.

Tiphia inaequalis Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 22. ♂.

Taxonomy: Prison, 1927. 111. Nat. Hist. Survey, Bui. 16: 229-230 (lectotype designation).
—Allen, 1966. Amer. Ent. Soc, Trans. 92: 293, fig. 29, map 10. ♂.

Tiphia incisurata Malloch
N. C.

Tiphia incisurata Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 17. ♀.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 288, map 9. ♀.

Tiphia indistincta Allen
W. Va. (Kanawha Sta.).

Tiphia indistincta Allen, 1961. Amer. Ent. Soc, Trans. 87: 14. ♀.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 254, map 3. ♀.

Tiphia infossata Allen
Maine to Mich., Wis., Iowa and S. Dak., south to Pla. and Tex., Alta., Mont.,
Colo., Calif.
Tiphia infossata Allen, 1934. Amer. Ent. Soc, Trans. 60: 301. ♀.
Tiphia hollowayi Allen, 1934. Amer. Ent. Soc, Trans. 60: 303. ♂.

Taxonomy: Allen, 1961. Amer. Ent. Soc, Trans. 87: 19 (synonymy). —Allen, 1965. Acad. Nat.
Sci., Phila., Proc 117: 60. -Allen, 1966. Amer. Ent. Soc, Trans. 92: 248, figs. 8, 16, 21, 36,
map 2. 9, 6. -Allen, 1971. Amer. Ent. Soc, Trans. 97: 275, figs. 86-89, map 6. ♀, ♂.

Tiphia inornata Say
Ohio, Pa.

Tiphia inornata Say, 1824. In Keating, Narr. Long's 2nd Exped., v. 2, p. 331 9.

Taxonomy: Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 12. —Allen, 1934. Amer. Ent. Soc,
Trans. 60: 300 (redescription.) —Allen, 1966. Amer. Ent. Soc, Trans. 92: 262. ♀.
(Redescription.)

Tiphia intermedia
***authority mismatch
Malloch. Maine and Que. south to Pla., west to N. Dak. and Calif., south to
Nicaragua. Host: Phyllophaga spp.
Tiphia punctata Robertson, 1901. Amer. Ent. Soc, Trans. 27: 196. ♂. Preocc.
Tiphia clypeolata Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 16. 9, d (in part?).
Tiphia arida Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 20. ♀.
Tiphia punctata var. intermedia Malloch, 1918. 111. Nat. Hist. Survey Bui. 13: 21. ♀.
Tiphia reticulata Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 23. ♀.
Tiphia intermedia var. exitialis Roberts, 1930. Canad. Ent. 62: 189. N. name.

Taxonomy: Prison, 1927. 111. Nat. Hist. Survey, Bui. 16: 230 (lectotype designation, punctata
var intermedia). —Allen, 1934. Amer. Ent. Soc, Trans. 60: 310. — Krombein, 1938. Ent.
News 49: 186. —Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 56. —Allen, 1966. Amer. Ent.
Soc, Trans. 92: 302, figs. 10, 12, 15, 17, 26, 37, 40, map 11. 6, 9. —Allen, 1971. Amer. Ent.
Soc, Trans. 97: 268, figs. 69-72, map 6. ♂, ♀.

Biology: Davis, 1919. 111. Nat. Hist. Survey, Bui. 13: 59-68.

Tiphia irfla Allen
Pla.

Tiphia irfla Allen, 1961. Amer. Ent. Soc, Trans. 87: 10. ♂, ♀.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 314, map 12.

Tiphia jaynesi Allen
Que. and Vt. to Pla., La., Okla., Kans., 111.

Tiphia jaynesi Allen, 1934. Amer. Ent. Soc, Trans. 60: 311. ♀, ♂.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 315, map 13. ♂, ♀.

Tiphia krombeini Allen
N. H. and Ont. to N. C, W. Va., Mich., S. Dak., ? Tex.

Tipkia krombeini Allen, 1966. Amer. Ent. Soc, Trans. 92: 321, map 14. ♀, ♂.

Tiphia letalis Roberts
N. Y. to N. C, west to S. Dak. and Kans., Miss.

Tiphia clypeata Robertson, 1901. Amer. Ent. Soc, Trans. 27: 196. ♂. Preocc.




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Tiphia letalis Roberts, 1930. Canad. Ent. 62: 189. N. name.

Taxonomy: Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 11. 9, 6. —Allen, 1966. Amer. Ent.
Soc, Trans. 92: 257, figs. 1, 3, 11, map i. ♂, ♀.

Tiphia matura Allen and Jaynes
Liberated in N. J. but not established. Introduced from India. Host:
Popillia japonica Newm.
Tiphia matura Allen and Jaynes, 1930. U. S. Natl. Mus., Proc. 76, Art. 17: 81. ♀, ♂.

Tiphia micropunctata Allen
N. S. to Man., Mont, and Calif., south to Fla. and Okla.
Tiphia micropunctata Allen, 1934. Amer. Ent. Soc, Trans. 60: 297. ♀, ♂.
Tiphia dentata Allen, 1934. Amer. Ent. Soc. Trans. 60: 303. ♂.

Taxonomy: Allen, 1961. Amer. Ent. Soc, Trans. 87: 18 (synonymy). —Evans, 1965. Ent. Soc.
Wash., Proc. 67: 90-91, figs. 6-10 (larva). —Allen, 1966. Amer. Ent. Soc, Trans. 92: 243,
figs. 7, 13, 22, 25, map \. 9, S.

Tiphia minor Provancher
B. C. (Vancouver Isl.), ? Wash., ? Oreg.

Tiphia minor Provancher, 1887. Addit. Corr. Faune Ent. Canada Hym., p. 254. ♂.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 65. 3. —Allen, 1971. Amer. Ent.
Soc, Trans. 97: 294, map 8.

Tiphia montana Allen
Sask., Alta., Mont., Utah, Ariz; Mexico (Jalisco).
Tiphia montana Allen, 1961. Amer. Ent. Soc, Trans. 87: 13. ♂.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 53, fig. 22. 3. —Allen, 1971. Amer.
Ent. Soc, Trans. 97: 234, figs. 30, 31, map 2. ♂.

Tiphia morio Fabricius
Liberated in N. Y. but not established. Introduced from Europe. Host:
Amphimallon majalis (Raz.).
Tiphia morio Fabricius, 1787. Mantissa Insectorum, v. 1, p. 280.

Tiphia nebra Allen
Nebr., Ky.

Tiphia nebra Allen, 1966. Amer. Ent. Soc, Trans. 92: 261, map 3. ♀.

Tiphia nevadana Cameron
B. C. south to Calif., Idaho, Wyo., Nev., Colo., Ariz., N. Mex., Tex; Mexico
(Hidalgo).
Tiphia nevadana Cameron, 1906. Invertebrata Pacifica 1: 157. ♂.
Tiphia essigi Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 62, figs. 7, 9, 26. ♀, ♂.

Taxonomy: Allen, 1962. Amer. Ent. Soc, Trans. 88: 22 (type redescription). —Allen, 1971.
Amer. Ent. Soc, Trans. 97: 298, figs. 115, 116, map 9. ♂, ♀.

Tiphia nona Allen
Ariz, south in Mexico to San Luis Potosi and Michoacan.
Tiphia nona Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 61. ♂.
Tiphia hurdi Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 75, fig. 24. ♂, ♀.

Taxonomy: Allen, 1971. Amer. Ent. Soc, Trans. 97: 248, figs. 46-48, map 4. ♂, ♀.

notopolita var. alleni Roberts. Liberated in N. Y., but not established. Introduced from Korea
and China. Host: Anomala orientalis Waterh.
Tiphia notopolita var. intermedia Allen and Jaynes, 1930. U. S. Natl. Mus., Proc. 76, Art.

17: 41. 9. Preocc
Tiphia notopolita var. alleni Roberts, 1930. Canad. Ent. 62: 190. N. name.

Biology: Clausen, Jaynes and Gardner, 1933. U. S. Dept. Agr., Tech. Bui. 366: 46 (life
history).

Tiphia occidentata Malloch
Wash., Colo., N. Mex., Ariz, south in Mexico to Oaxaca.
Tiphia occidentata Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 17. ♂.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 52. 6. —Allen, 1971. Amer. Ent.
Soc, Trans. 97: 241, figs. 38-40, map 3. ♂.

Tiphia odontogaster Viereck
N. Mex., Ariz.; Mexico (Durango).

Tiphia odontogaster Viereck, 1903. Amer. Ent. Soc, Trans. 29: 71. ♂.

Taxonomy: Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 13. —Allen, 1965. Acad. Nat. Sci.
Phila., Proc 117: 51, fig. 21. S. —Allen, 1971. Amer. Ent. Soc, Trans. 97: 232, figs. 28, 29,
map 2.




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Tiphia pecosae
***authority mismatch
Alien. Tex. (Pecos River at Sheffield).

Tiphia pecosae Allen, 1971. Amer. Ent. Soc, Trans. 97: 311, fig. 123. ♀.

Tiphia pennsylvanica Allen
N. J. to Fla., La.

Tiphia pennsylvanica Allen, 1961. Amer. Ent. See, Trans. 87: 15. ♀.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 286, map 9. ♀, ♂.

Tiphia popilliavora Rohwer
N. H., Mass., Conn., N. Y., N. J., Pa., Del., Md., Va., Ohio. Introduced
from Japan and Korea. Host: Popillia japonica Newm.
Tiphia popilliavora Rohwer, 1924. Ent. Soc. Wash., Proc. 26: 89. ♀, ♂.

Taxonomy: Allen and Jaynes, 1930. U. S. Natl. Mus., Proc. 76, Art. 17: 51, figs. 3, 21, 22, 27,
28. ♀, ♂.

Biology: Clausen, King, and Teranishi, 1927. U. S. Dept. Agr., Bui. 1429: 33-39. —King and
HoUoway, 1930. Jour. Econ. Ent. 23: 266-274. — Holloway, 1931. N. Y. Ent. Soc, Jour. 39:
555-564. —White, 1943. N. Y. Ent. Soc, Jour. 51: 213-218.

Tiphia portalae Allen
Ariz., Nev.

Tiphia portalae Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 49. ♂.

Taxonomy: Allen, 1971. Amer. Ent. Soc, Trans. 97: 226, figs. 17, 18, map 1. c?, 9.

Tiphia pullivora Allen and Jaynes
Liberated in Pa. but not established. Introduced from India. Host:
Popillia japonica Newm.
Tiphia pullivora Allen and Jaynes, 1930. U. S. Natl. Mus., Proc. 76, Art. 17: 83. ♀, ♂.

Biology: Clausen, Jaynes and Gardner, 1933. U. S. Dept. Agr., Tech. Bui. 366: 39-41 (life
history).

Tiphia puncticeps Cameron
Nev.; ? Mexico (Michoacan).

Tiphia puncticeps Cameron, 1906. Invertebrata Pacifica 1: 158. ♂.

Taxonomy: Allen, 1962. Amer. Ent. Soc, Trans. 88: 23 (type redescription). —Allen, 1971.
Amer. Ent. Soc, Trans. 97: 246, map 4. ♂.

Tiphia pygidialis Allen
Conn., Pa., Md., W. Va., Ky., Iowa, Kans., Okla., Miss.

Tiphia pygidialis Allen, 1966. Amer. Ent. Soc, Trans. 92: 310, map 12. ♀.

Tiphia raui Allen
Mo., Miss.

Tiphia raui Allen, 1934. Amer. Ent. Soc, Trans. 60: 308. ♀.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 269. ♀.

Tiphia relativa Viereck
Maine to Ga., Mich., Iowa, Nebr., Kans. Host: Cyclocephala borealis Arrow;
Lichnathe vulpina (Hentz).
Tiphia relativa Viereck, 1906. Ent. News 17: 304. ♂.
Tiphia winnemanae Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 23. ♂.

Taxonomy: Allen, 1961. Amer. Ent. Soc, Trans. 87: 17 (synonymy). —Allen, 1966. Amer. Ent.
Soc, Trans. 92: 312, map 12. ♂, ♀.

Biology: Adams, 1949. Jour. Econ. Ent. 42: 626. —Franklin, 1950. Mass. Agr. Expt. Sta., Bui.
445, pts. 2-7: 63, 66, 67.

Tiphia roda Allen
N. Mex., Ariz.

Tiphia roda Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 56. ♂.

Taxonomy: Allen, 1971. Amer. Ent. Soc, Trans. 97: 286, map 8. ♂.

Tiphia rodeoensis Allen
N. Mex. south in Mexico to Jalisco and Morelos.

Tiphia rodeoensis Allen, 1971. Amer. Ent. Soc, Trans. 97: 336, fig. 143, map 13. ♀.

rooseveiti Allen. Ariz.

Tiphia rooseveiti Allen, 1971. Amer. Ent. Soc, Trans. 97: 339, figs. 145-147, map 13. ♀.

Tiphia rothi Allen
Ariz. (Cochise Co.).

Tiphia rothi Allen, 1971. Amer. Ent. Soc, Trans. 97: 231, map 1. ♂.
nigulosa Malloch. Maine and Que. to Ga., west to Minn., Iowa and Kans.

Tiphia rugulosa MaUoch, 1918. 111. Nat. Hist. Survey, Bui. 13: 15. ♀, ♂.

Taxonomy: Prison, 1927. 111. Nat. Hist. Survey, Bui. 16: 230 (lectotype designation). —Allen,
1966. Amer. Ent. Soc, Trans. 92: 333, fig. 30, map 16. ♂, ♀.




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Tiphia sayi Allen
N. H., Ont., Wis. and Iowa south to Fla. and Miss.

Tiphia sayi Allen, 1966. Amer. Ent. Soc, Trans. 92: 272, fig. 18, map 6. ♂, ♀.

Tiphia schlingeri Allen
Tex., N. Mex., Ariz., Nev., Calif.

Tiphia schlingeri Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 64, fig. 12. ♂.

Taxonomy: Allen, 1971. Amer. Ent. Soc, Trans. 97: 311, map 10. ♂.

Tiphia sculleni Allen
Ariz. (Cochise and Pima Counties).

Tiphia sculleni Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 60, fig. 25. ♂.

Taxonomy: Allen, 1971. Amer. Ent. Soc, Trans. 97: 291, figs. 104, 105, map 8.

Tiphia similis Malloch
Maine and Que. south to Fla., west to Ont., Mich., 111. and Kans.
Tiphia similis Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 18. ♂.
Tiphia affinis Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 19. ♀, ♂.

Taxonomy: Prison, 1927. 111. Nat. Hist. Survey, Bui. 16: 229-230 (lectotype designations,

similis, affinis). —Allen, 1966. Amer. Ent. Soc, Trans. 92: 318, figs. 20, 28, map 14. ♂, ♀.

Tiphia sinaloae Allen
Okla., Tex., south to Costa Rica.

Tiphia sinaloae Allen, 1971. Amer. Ent. Soc, Trans. 97: 309, map 10. ♂.

Tiphia sternata Parker
Liberated 1933-35 in N. Y., N. J., Pa. but has not been recovered since 1937.
Introduced from Japan. Host: Serica peregrina Chapin.

Tiphia sternata Parker, 1935. N. Y. Ent. Soc, Jour. 43: 399. ♀, ♂.

Tiphia strangulata Allen
Mont., Ariz.; ? Mexico (Puebla).

Tiphia strangulata Allen, 1971. Amer. Ent. Soc, Trans. 97: 261, fig. 73, map 5. ♂.

Tiphia subcarinata Malloch
Mass. to Ga., W. Va., Mich., 111.

Tiphia subcarinata Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 15. ♂.

Tiphia insolita Allen, 1934. Amer. Ent. Soc, Trans. 60: 316. ♂.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 330, map 16. ♂.

Tiphia tarda Say
Ind.

Tiphia tarda Say, 1836. Boston Jour. Nat. Hist. 1: 300. ♂.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 351 (unrecognized).

Tiphia tegulina Malloch
B. C. and Idaho to Calif, and Nev. Host: Phyllophaga errans LeC.
Tiphia tegulina Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 21. ♀.
Tiphia shasteyisis Krombein, 1942. Pan-Pacific Ent. 18: 139. ♀, ♂.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 59. —Allen, 1971. Amer. Ent. Soc,
Trans. 97: 272, figs. 79-81, map 6. ♀, ♂.

Biology: Linsley and Michener, 1942. Pan-Pacific Ent. 18: 154.

Tiphia tempeae Allen
Kans., Colo., Wash., CaHf., Ariz, and N. Mex. south to Costa Rica.
Tiphia tempeae Allen, 1961. Amer. Ent. Soc, Trans. 87: 11. ♂.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 53, 80. 6. -Allen, 1971. Amer. Ent.
Soc, Trans. 97: 263, fig. 74, map 5. ♂, ♀.

Tiphia texensis Malloch
Tex., Kans., Mont.

Tiphia texensis Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 20. ♀.

Taxonomy: Allen, 1961. Amer. Ent. Soc, Trans. 87: 19. —Allen, 1965. Acad. Nat. Sci. Phila.,
Proc 117: 66. ♀.

Tiphia toddi Allen
Ariz.; Mexico (Jalisco, Morelos).

Tiphia toddi Allen, 1971. Amer. Ent. Soc, Trans. 97: 252, fig. 50, map 4. ♂.

Tiphia totopunctata
***authority mismatch
Allen and Jaynes. Liberated in N. Y. but not established. Introduced from
Korea. Host: Anomala orientalis Waterh.
Tiphia totopunctata Allen and Jaynes, 1930. U. S. Natl. Mus., Proc. 76, Art. 17: 35. ♀.

Biology: Clausen, Jaynes and Gardner, 1933. U. S. Dept. Agr., Tech. Bui. 366: 44 (life
history).

Tiphia townesi Allen
N. Y. (Shokan).

Tiphia townesi Allen, 1966. Amer. Ent. Soc, Trans. 92: 338, map 17. ♂.




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Tiphia transversa Say
Vt. to Kans., south to Ga., Ala. and Tex. Host: Phyllophaga spp.
Tiphia transversa Say, 1828. Contrib. Maclur. Lye. Phila. 1: 83 6.

Taxonomy. Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 11. —Allen, 1934. Amer. Ent. Soc,
Trans. 60: 297. -Allen, 1966. Amer. Ent. Soc, Trans. 92: 270, fig. 19, map 6. ♂, ♀.

Biology: Davis, 1919 111. Nat. Hist. Survey, Bui. 13: 68.

Tiphia treherni Allen
B. C. to Calif.

Tiphia treherni Allen, 1971. Amer. Ent. Soc, Trans. 97: 329, figs. 35, 36, map 12. ♀.

Tiphia truncata Cameron
Nev. (Ormsby Co.).

Tiphia truncata Cameron, 1906. Invertebrata Pacifica 1: 156. ♂.

Taxonomy: Allen, 1962. Amer. Ent. Soc, Trans. 88: 22 (type unlocated).

Tiphia unica Allen
N. C. to Fla., Ala., Tenn., Kans.

Tiphia unica Allen, 1934. Amer. Ent. Soc, Trans. 60: 309. ♀.

Taxonomy: Allen, 1966. Amer. Ent. Soc, Trans. 92: 267, fig. 2, map 5. ♀, ♂.

Tiphia uvaldae Allen
Tex. (Uvalde Co.).

Tiphia uvaldae Allen, 1971. Amer. Ent. Soc, Trans. 97: 313, fig. 124, map 11. ♀.

Tiphia vallis Allen
Calif. (Kern and El Dorado Counties).

Tiphia vallis Allen, 1971. Amer. Ent. Soc, Trans. 97: 334, map 12. ♀.

Tiphia veracruzae Allen
Southern Ariz, to Panama.

Tiphia veracruzae Allen, 1965. Acad. Nat. Sci. Phila., Proc 117: 82, fig. 5. ♂, ♀.
Tiphia litoris Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 84. cJ, 9.

Taxonomy: Allen, 1971. Amer. Ent. Soc, Trans. 97: 295, figs. 108-114, map 9. ♂, ♀.

Tiphia vernalis Rohwer
Mass., N. H., R. I., Conn., N. Y., N. J., Pa., Del., Md., D. C, Va., N. C, W. Va.,
Ohio; Orient. Introduced from Japan, Korea, and China. Host: Popillia japonica Newm.
Tiphia vernalis Rohwer, 1924. Ent. Soc Wash., Proc 26: 91. ♀, ♂.

Taxonomy: Allen and Jaynes, 1930. U. S. Natl. Mus., Proc 76, Art. 17: 78, figs. 6, 16, 19, 25,
29.

Biology: Clausen, King, and Teranishi, 1927. U. S. Dept. Agr., Bui. 1429: 40-41. — Balock,
1934. Jour. Econ. Ent. 27: 491-496. —White, 1943. N. Y. Ent. Soc, Jour. 51: 213-218.

Tiphia virotha Allen
Ariz. (Cochise Co.); ? Mexico (Hidalgo).

Tiphia virotha Allen, 1971. Amer. Ent. Soc, Trans. 97: 338, map 13. ♀.

Tiphia vulgaris Robertson
111., Wis., Ind., Iowa, S. Dak., Kans., Mo., Tex. Host: Phyllophaga spp.
Tiphia vulgaris Robertson, 1901. Amer. Ent. Soc, Trans. 27: 195. ♀, ♂.

Taxonomy: Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 12. —Allen, 1934. Amer. Ent. Soc,
Trans. 60: 305. —Allen, 1961. Amer. Ent. Soc, Trans. 87: 19. —Allen, 1966. Amer. Ent.
Soc, Trans. 92: 263, fig. 39, map 5. ♂, ♀.

Biology: Davis, 1919. 111. Nat. Hist. Survey, Bui. 13: 69.

Tiphia yosemitensis Allen
Nebr., Mont., Calif.

Tiphia yosemitensis Allen, 1971. Amer. Ent. Soc, Trans. 97: 330, figs. 137-139, map 12. ♀.

Krombeinia chiricahua (Pate)
Ariz. (Chiricahua).

Neotiphia chiricahua Pate, 1939. Ent. News 50: 221. ♂, ♀.

Taxonomy: Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 233. ♂, ♀.

Krombeinia cristata (Allen)
Ariz., N. Mex.

Neotiphia cristata Allen, 1935. Amer. Ent. Soc, Trans. 61: 59. ♀.

Taxonomy: Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 238, figs. 3, 29. ♀.




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Krombeinia jonesi Allen and Krombein
Ariz. (Douglas, Chiricahua, Huachuca Mts., Oracle).

Krombeinia jonesi Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 239, figs. 6, 65.
6, 9.

Krombeinia robusta (Allen)
Tex.

Neotiphia ocellata Allen, 1935. Amer. Ent. Soc, Trans. 61: 56. ♀.

Neotiphia robusta Allen, 1935. Amer. Ent. Soc, Trans. 61: 64. ♂.

Taxonomy: Krombein, 1951. U. S. Dept. Agr., Monog. 2: 739 (synonymy). —Allen and
Krombein, 1964. Amer. Ent. Soc, Trans. 89: 229, figs. 2, 30. cJ, 9.

Neotiphia acuta Malloch
Tex.

Neotiphia acuta Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 9. ♀, ♂.

Taxonomy: Prison, 1927. 111. Nat. Hist. Survey, Bui. 16: 229 (lectotype designation). —Allen

and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 259, figs. 5, 46, 67. ♂.

Neotiphia anima Allen and Krombein
N. Mex., Ariz.

Neotiphia anima Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 262. ♀.

Neotiphia barbata Allen and Krombein
Ariz. (Continental, Tucson, Douglas).

Neotiphia barbata Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 264. ♂.

Neotiphia cockerelli Allen
Tex., N. Mex.

Neotiphia cockerelli Allen, 1935. Amer. Ent. Soc, Trans. 61: 62. ♂.

Taxonomy: Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 255, fig. 50. ♂, ♀.

Neotiphia crawfordi Allen
Tex.

Neotiphia crawfordi Allen, 1935. Amer. Ent. Soc, Trans. 61: 60. ♀.
Neotiphia conspicua Allen, 1935. Amer. Ent. Soc, Trans. 61: 61. ♂.

Taxonomy: Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 257, figs. 28, 41, 42. ♀, ♂.

Neotiphia flavipennis Allen and Krombein
Ariz., N. Mex. south in Mexico to Mexico and Hidalgo.
Neotiphia flavipennis Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 263. ♀.
Neotiphia pamia Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 267, figs. 7, 54. ♂.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc 117: 87 (synonymy).

Neotiphia luteipennis (Cresson)
Colo.

Tiphia luteipennis Cresson, 1865. Ent. Soc. Phila., Proc 4: 445. ♀.

Taxonomy: Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 10. — AUen and Krombein, 1964.
Amer. Ent. Soc, Trans. 89: 260. ♀.

Neotiphia novomexicana Allen
N. Mex., Ariz.

Neotiphia novomexicana Allen, 1935. Amer. Ent. Soc, Trans. 61: 63. ♀.
Neotiphia carinata Krombein, 1938. Ent. Soc. Amer., Ann. 31: 59. ♂.

Taxonomy: Allen and Krombein, 1964. Amer: Ent. Soc, Trans. 89: 251, figs. 4, 9, 11, 18, 21,
25,38,49,55,60,68. ♀, ♂.

Neotiphia rioverdei Allen and Krombein
Ariz. (Del Rio Verde).

Neotiphia rioverdei Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 263. ♀.

Neotiphia rostrata Allen
Ariz.

Neotiphia rostrata Allen 1935. Amer. Ent. Soc, Trans. 61: 64. ♂.

Taxonomy: Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 265, figs. 14, 53. cj.

Neotiphia sulcata (Roberts)
Mich., Iowa and S. Dak. south to Tex. and Ariz.; Mexico (Jalisco).
Tiphia canaliculata Malloch, 1918. 111. Nat. Hist. Survey, Bui. 13: 10. ♂. Preocc.
Tiphia sulcata Roberts, 1930. Canad. Ent. 62: 190. N. name.
Tiphia mexicana Allen, 1935. Amer. Ent. Soc, Trans. 61: 57. ♀.




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Taxonomy: Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 245, figs. 1, 8, 12, 17, 20,
23, 24, 27, 34, 44, 48, 56, 58, 59, 66. ♂, ♀.

Neotiphia waltoni Allen
Kans. and Colo, to Tex., N. Mex. and Ariz.; Mexico (Chihuahua, Durango).
Neotiphia waltoni Allen 1935. Amer. Ent. Soc, Trans. 61: 62. ♀.
Neotipkia puna Pate, 1939. Ent. News 50: 245. cJ, (9 misdet).

Taxonomy: Allen and Krombein, 1964. Amer. Ent. Soc, Trans. 89: 248, 254, fig. 15, 36, 37, 47,
62. 9,6 (treat waltoni and piyna as discrete species). —Allen, 1965. Acad. Nat. Sci. Phila.,
Free. 117: 87 (synonymy).

Paratiphia albilabris (Spinola)
Calif.

Tiphia albilabris Spinola, 1841. Soc. Ent. France, Ann. 10: 102. ♂.

Taxonomy: Allen, 1968. Amer. Ent. Soc, Trans. 94: 107 (type not located).

Paratiphia asotinae Allen
Wash, and Nebr. south to Calif., Kans. and N. Mex.
Paratiphia asotinae Allen, 1963. Ent. News 74: 222. ♂, ♀.

Taxonomy: Allen, 1968. Amer. Ent. Soc, Trans. 94: 57, map 4. cJ, 9.
beifragei Allen. Nebr. and Idaho to Tex. and Calif., south in Mexico to Hidalgo and Jalisco.
Paratiphia beifragei Allen, 1963. Ent. Soc. Amer., Ann. 56: 575, figs. 1, 2 c, d. d, 9.
Paratiphia insueta Allen, 1963. Ent. Soc. Amer., Ann. 56: 576, figs. 1, 2 e. 6.

Taxonomy: Allen, 1968. Amer. Ent. Soc, Trans. 94: 66, figs. 21, 23, 24, map 6.

Paratiphia bridwelli Allen
Kans., Tex., Ariz.; Mexico (Tamaulipas).

Paratiphia bridwelli Allen, 1963. Amer. Ent. Soc, Trans. 88: 231, figs. 1, 6, 11, 19. ♂.

Taxonomy: Allen, 1968. Amer. Ent. Soc, Trans. 94: 44, map 2. ♂.

Paratiphia cincta Allen
Calif., Nev., Ariz.; Mexico (Baja California).
Paratiphia cincta Allen, 1962. Ent. News 73: 262. ♂.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 98, 106, fig. 2. 6. -Allen, 1968.
Amer. Ent. Soc, Trans. 94: 78, figs. 13, 34, map 5. ♂, ♀.

Paratiphia bradleyi Allen
Tex. to Ariz., south in Mexico to Baja California and Veracruz.

Paratiphia bradleyi Allen, 1963. Amer. Ent. Soc, Trans. 88: 231, figs. 2, 7, 12. ♂.
Paratiphia pachucae Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 101, fig. 17. ♂.

Taxonomy: Allen, 1968. Amer. Ent. Soc, Trans. 94: 49, map 3. 6, 9. — Betrem and Allen,
1970. Ent. Soc Wash., Proc. 72: 71.

Paratiphia claripennis Cameron
B. C, Idaho, Nev., Calif, to Tex., south in Mexico to Baja California,
Sinaloa and Puebla.
Paratiphia claripennis Cameron, 1905. Invertebrata Pacifica 1: 108. ♂.
Paratiphia varipunctata Cameron, 1905. Invertebrata Pacifica 1: 106. ♀.
Paratiphia fuscinerva Cameron, 1905. Invertebrata Pacifica 1: 107. ♀.
Paratiphia bakeri Cameron, 1905. Invertebrata Pacifica 1: 109. ♂.
Paratiphia ormsbyensis Cameron, 1907. Invertebrata Pacifica 1: 175. ♂.




Page 1264 • Top • -1 • +1 • Original • Database of Hymenoptera in America north of Mexico • Highertaxa

Taxonomy: Cameron, 1907. Invertebrata Pacifica 1: 172. 9. —Allen, 1962. Amer. Ent. Soc,
Trans. 88: 29-30. -Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 96. 6. —Allen, 1968. Amer.
Ent. Soc, Trans. 94: 75, fig. 31, map 7. ♂.

Paratiphia dreisbachi Allen
Colo., N. Mex.

Paratiphia dreisbachi Allen, 1968. Amer. Ent. Soc, Trans. 94: 83, fig. 32, map 6. ♀.

Paratiphia ephippiata Allen
Kans. and Wyo. to Tex. and Ariz, south in Mexico to Zacatecas and San Luis
Potosi.
Paratiphia ephippiata Allen, 1963. Ent. Soc. Amer., Ann. 56: 574, figs. 1, 2 a, b, f. S, 9.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc 117: 93, fig. 19. Amer. Ent. Soc, Trans. 94: 50, figs. 6-10, map S. 6 , 9 .

Paratiphia fossata Allen
N. Mex. and Ariz, south in Mexico to Veracruz and Oaxaca.
Paratiphia fossata Allen, 1962. Ent. News 73: 259. ♂.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 98, 103. 6. —Allen, 1968. Amer. Ent.

Soc, Trans. 94: 64, figs. 18-20, map 4. ♂, ♀.

Paratiphia halli Allen
Calif. (Riverside Co.).

Paratiphia halli Allen, 1968. Amer. Ent. Soc, Trans. 94: 68, map 6. ♂.

Paratiphia huachucae Allen
Ariz. (Huachuca Mts.).

Paratiphia huachucae Allen, 1968. Amer. Ent. Soc, Trans. 94: 47, map 2. ♂.

Paratiphia irwini Allen
Ariz. (Chiricahua Mts. ).

Paratiphia irwini Allen, 1968. Amer. Ent. Soc, Trans. 94: 74, map 6. ♂.

Paratiphia knowltoni Allen
Utah., Nev.

Paratiphia knowltoni Allen, 1968. Amer. Ent. Soc, Trans. 94: 82, fig. 35, map 6. ♀.

Paratiphia knulli Allen
Tex.

Paratiphia knulli Allen, 1968. Amer. Ent. Soc, Trans. 94: 60, fig. 16, map 5. ♂.

Paratiphia lagosae Allen
Tex. and Calif, south in Mexico to Michoacan and Hidalgo.
Paratiphia lagosae Allen, 1965. Acad. Nat. Sci. Phila., Proc 117: 101. ♂.

Taxonomy: Allen, 1968. Amer. Ent. Soc, Trans. 94: 37, map 1. ♂, ♀.

Paratiphia magna Roberts
Idaho (Pocatello).

Paratiphia ynagna Roberts, 1929. Psyche 36: 358. ♀.

Taxonomy: Allen, 1968. Amer. Ent. Soc, Trans. 94: 84, map 6. ♀.

Paratiphia mitchelli Allen
Tex. to Ariz, south in Mexico to Puebla.

Paratiphia mitchelli Allen, 1963. Amer. Ent. Soc, Trans. 88: 232, figs. 3, 8, 13, 18. ♂.

Taxonomy: Allen, 1968. Amer. Ent. Soc, Trans. 94: 46, map 2. ♂, ♀.

Paratiphia neomexicana Cameron
Oreg. to Nebr. south to Calif, and Tex.; Mexico (Sonora, Chihuahua).
Paratiphia neomexicana Cameron, 1907. Invertebrata Pacifica 1: 175. ♂.

Taxonomy: Allen, 1962. Amer. Ent. Soc, Trans. 88: 31 (type redescription). —Allen, 1965.
Acad. Nat. Sci. Phila., Proc. 117: 94. cj, 9. —Allen, 1968. Amer. Ent. Soc, Trans. 94: 54,
figs. 11, 12, 14, map 4. cJ, 9.

Paratiphia nevadensis Cameron
B. C. to Calif, and Tex., south in Mexico to Baja California and Puebla.
Paratiphia nevadensis Cameron, 1905. Invertebrata Pacifica 1: 108. ♂.
Paratiphia parvula Cameron, 1907. Invertebrata Pacifica 1: 176. ♂.
Paratiphia intermedia Cameron, 1907. Invertebrata Pacifica 1: 176. ♂.

Taxonomy: Allen, 1962. Amer. Ent. Soc, Trans. 88: 27, 32 (synonymy, type redescription).
—Allen, 1963. Amer. Ent. Soc, Trans. 88: 229, figs. 4, 9, 14, 17 (redescription). —Allen,
1965. Acad. Nat. Sci. Phila., Proc. 117: 91, 95. 6, 9. —Allen, 1968. Amer. Ent. Soc, Trans.
94: 39, fig. 5, map 2. ♂, ♀.

Paratiphia occidentalis Cameron
Nev. (Ormsby Co.).

Paratiphia occidentalis Cameron, 1907. Invertebrata Pacifica 1: 173. ♂.

Taxonomy: Allen, 1968. Amer. Ent. Soc, Trans. 94: 108 (type not located).

Paratiphia robusta Cameron
B. C, Idaho and Nebr. to Calif, and Tex.; Mexico (Baja California, Nayarit).
Paratiphia robusta Cameron, 1905. Invertebrata Pacifica 1: 106. ♀.
Paratiphia fortistriolata Cameron, 1907. Invertebrata Pacifica 1: 172. ♂.




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Taxonomy: Allen, 1962. Amer. Ent. Soc, Trans. 88: 33 (type redescribed, synonymy). —Allen,

1965. Acad. Nat. Sci. Phila., Proe. 117: 92. 9, 6. -Allen, 1968. Amer. Ent. Soc, Trans. 94:

70, figs. 22, 25-30, 41, map 6. ♀, ♂.

Paratiphia sculleni Allen
Oreg., Utah, Ariz., N. Mex.

Paratiphia sculleni Allen, 1968. Amer. Ent. Soc, Trans. 94: 52, map 3. ♂, ♀.

Paratiphia texana Cameron
Ont. and Mass. to Mont, south to Fla. and Tex.

Paratiphia texana Cameron, 1907. Invertebrata Pacifica 1: 174. ♀, ♂.

Paratiphia algonguina Viereck, 1909. Ent. Soc. Wash., Proc 11: 45. ♀, ♂.

Paratiphia algonquina (!) Smith, 1910. N. J. State Mus., Ann. Rpt. for 1909, p. 666.

Taxonomy: Allen, 1962. Amer. Ent. Soc, Trans. 88: 25 (redescription texana). —Allen, 1965.
Acad. Nat. Sci. Phila., Proc 117: 97. 9, d. -Allen, 1968. Amer. Ent. Soc, Trans. 94: 62,
figs. 15, 17, map 5. ♂, ♀.

Paratiphia verna Allen
Calif., Nev., Ariz.; Mexico (Baja California).
Paratiphia verna Allen, 1963. Ent. News 74: 219. ♂, ♀.

Taxonomy: Allen, 1965. Acad. Nat. Sci. Phila., Proc. 117: 99. 6. — AUen, 1968. Amer. Ent.
Soc, Trans. 94: 80, fig. 36, map 7. ♂, ♀.

Unplaced Taxon of Paratiphia Sichel

Paratiphia fuscipennis Cameron
No locality given (title says n. spp. from "Nevada and Southern
California").
Paratiphia fuscipennis Cameron. 1905. Invertebrata Pacifica 1: 106. 9 (in key).

Epomidiopteron julii Romand
Southern Ariz, to Peru and Argentina.

Epoynidiopteron Julii Romand, 1836. Soc Ent. France, Ann. 4: 653. ♀.
Paratiphia 12-maculata Cameron, 1904. Amer. Ent. Soc, Trans. 30: 94. ♂.
Scoliphia spilota Banks, 1912. Canad. Ent. 44: 201. ♀, ♂.
Epimodiopteron (!) spilota Bridwell, 1919. Hawaii. Ent. Soc, Proc. 4: 119.

Taxonomy: Allen, 1966. Ent. News 77: 206, 2 figs. 9, 6 (synonymy and redescription).
—Allen, 1972. Smithsn. Contrib. Zool. 113: 3.

Myzinum berlyi
***authority mismatch
berlyi (Brimley). S. C, 111., Ky, Wis, Kans, Miss, Tex.
Elis berlyi Brimley, 1927. Ent. News 38: 238. ♀.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 249-251. ♀, ♂.

Myzinum berlyi
***authority mismatch
parks! Krombein. La., Tex.

Myzine (!) berlyi jmrksi Krombein, 1938. Amer. Ent. Soc, Trans. 64: 251. ♀, ♂.

Myzinum berlyi
***authority mismatch
patei Krombein. N. J., N. C, Ga, Fla.

Myzine (!) berlyi patei Krombein, 1938. Amer. Ent. Soc, Trans. 64: 254. ♀, ♂.

Myzinum carolinianum
***authority mismatch
carolinianuin (Panzer). Va. to Fla., west to Tex.

Tiphia caroliniana Panzer, 1806. Krit. Revis. Insektenf. Deutschlands, v. 2, p. i, pi. 1, figs.

a-c. 9.
Myzine {\) flavipes Olivier, 1811. Encycl. Meth., v. 8, p. 136. ♀.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 260-262, fig. 7. ♀, ♂.
carolinianuin collare (Say). Ohio west to S. Dak., south to La. and N. Mex.
Meria collaris Say, 1837. Boston Jour. Nat. Hist. 1: 362. ♀.
Myzine (!) thoracica Fox, 1893. Canad. Ent. 35: 113. 9. Preocc
Myzine (!) illinoisensis Dalla Torre, 1897. Cat. Hym., v. 8, p. 124. N. name.
Elis atriventris Gahan, 1913. U. S. Natl. Mus, Proc 40: 431. ♀, ♂.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 262-265. ♀, ♂.

Myzinum confluens Cresson
Utah, Ariz., south in western Mexico to Oaxaca.

Myzine (!) confluens Cresson, 1865. Ent. Soc. Phila., Proc. 4: 443. ♀.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 287-288. ♀.

Myzinum dubiosum Cresson
D. C. south to Fla., Ala., 111. south to La., Colo., Ariz., south in Mexico to
Veracruz, Sonora and Baja California.
Myzine (!) dubiosa Cresson, 1872. Amer. Ent. Soc, Trans. 4: 200. ♀.
Myziiie (!) texana Cresson, 1872. Amer. Ent. Soc, Trans. 4: 200. ♀.
Plesia fulvinervis Cameron, 1908. Amer. Ent. Soc, Trans. 34: 239. ♀.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 281-284, figs. 1,10. ♀, ♂.

Myzinum frontale Cresson
Kans., Tex., N. Mex., Colo., Ariz., Nev, Calif., south in Mexico to Baja
California, Sonora and Chihuahua.
Plesia marginata Say, 1823. West. Quart. Rptr. 2: 75. ♀. Preocc.
Myziyie {\) frontalis Cresson, 1875. Rpt. Geog. Geol. Expl. and Survey west of 100th
Meridian, p. 711. ♀.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 285-287, fig. 9. ♀, ♂.

Myzinum maculatum (Fabricius)
Mass. to Fla., west to Nev. and Ariz., Calif, south to Guatemala and El

Salvador.
Tiphia maculata Fabricius. 1793. Ent. System., p. 224. ♀.
Sapyga majorta Panzer, 1806. Krit. Rev. Insektenf. Deutschlands, v. 2, p. iv, pi. 2, figs, d,

e. 6.
Tiphia interrupta Say, 1824. In Keating, Narr. Long's 2nd Exped., v. 2, p. 322. ♀.
Myzine (!) hamatus Say, 1836. Boston Jour. Nat. Hist. 1: 300. ♂.
Meria costata Say, 1837. Boston Jour. Nat. Hist. 1: 360. ♀.
Myzine (!) hyalina Cresson, 1865. Ent. Soc. Phila., Proc. 4: 442. ♂.
Myzine (!) maiorta (!) Cresson, 1887. Amer. Ent. Soc, Trans., Sup. Vol., p. 268.
Elis floridanus Rohwer, 1920. Ent. Soc. Wash., Proc. 22: 54. ♀, ♂.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 274-281, fig. 11. 9, cJ. —Krombein,
1949. U. S. Natl. Mus., Proc. 100: 55-56, figs. 1-3 (gynandromorph).

Myzinum marginatum (Klug)
Ga.

Tiphia marginata Klug, 1810. In Weber, Beitr. z. Naturk. 2: 184. ♂.

Myzinum namea
***authority mismatch
fulviceps Cameron. South. Ariz, to Guatemala.

Myzine (I) fulviceps Cameron, 1893. Biol. Cent.-Amer., Hym., v. 2, p. 248, pi. 12, fig. 20. ♀.

Myzinum namea
***authority mismatch
namea (Fabricius). Va, N. C, S. C, Fla., Ala., Tex.

Tiphia namea Fabricius, 1805. Systema Piezatorum, p. 233. ♀.
Elis propodealis Rohwer, 1920. Ent. Soc. Wash., Proc. 22: 56. ♀.




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Myzine (!) magna Krombein, 1938. Amer. Ent. Soc, Trans. 64: 256. ♂.
Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 256-259, fig. 5. ♂, ♀.

Myzinum navajo Krombein
Ariz, south to El Salvador.

Myzine (!) navajo Krombein, 1938. Amer. Ent. Soc, Trans. 64: 272. ♀, ♂.

Myzinum obscurum (Fabricius)
N. Y. to Fla., west to 111. and Tex., ? Ariz. Host: Dyscinetus
trachypygus (Burm.). Parasite: Dasymutilla mittata (BL).
Tiphia obscura Fabricius, 1805. Systema Piezatorum, p. 233. ♀.
Plesia nigripes Guerin, 1838. Rev. Zool. 1: 57. ♀.
Plesia fuliginosa Lepeletier, 1845. Hist. Nat. Ins. Hym., v. 3, p. 574. ♂.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 246-249, fig. 3. ♀, ♂.

Myzinum quinquecinctum (Fabricius)
Que. to Fla., west to Wyo. and N. Mex., Calif.; Mexico (Coahuila).
Host: Phyllophaga spp. Parasite: Dasymutilla quadriguttata (Say).
Tiphia 5-cincta Fabricius, 1775. Systema Ent., p. 353. ♀.
Tiphia cingulata Klug, 1810. In Weber, Beitr. z. Naturk. 2: 185. ♂.
Sapyga subulata Say, 1823. West. Quart. Rptr. 2: 75. ♂.
Myzine (!) proxima Guerin, 1837. Diet. Pitt. Hist Nat., p. 581. ♂.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 265-272, fig. 6. 9, d. —Evans, 1965.
Ent. Soc Wash., Proc 67: 92, figs. 11-14 (larva).

Biology: Flint and Sanders, 1912. Jour. Econ. Ent. 3: 490. —Davis, 1919. 111. Nat. Hist.
Survey, Bui. 13: 73.

Morphology: Snodgrass, 1941. Smithsn. Inst., Misc. Collect. 99, no. 14, p. 44, pi. 14, figs. M-S
(male genitalia).

Myzinum serenum (Fabricius)
S. C.

Tiphia serena Fabricius, 1805. Systema Piezatorum, p. 234. ♀.

Myzinum spilonotum (Cameron)
Tex., N. Mex., Colo., Utah, Ariz., south in western Mexico to Oaxaca.

Plesia spilonota Cameron, 1908. Amer. Ent. Soc, Trans. 34: 240. ♀.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 288-290, fig. 12. ♀, ♂.

Pterombrus rufiventris
***authority mismatch
hyalinatus Krombein. West. Tex., Ariz., Calif.

Pterombrus rufiventris hyalinatus Krombein, 1949. Pan-Pacific Ent. 25: 88. ♀, ♂.

Pterombrus rufiventris
***authority mismatch
rufiventris (Cresson). Va. south to Ga. to east. Tex.

Myzine rufiventris Cresson, 1872. Amer. Ent. Soc, Trans. 4: 201. ♂.

Myzine cressonii Dalla Torre, 1897. Cat. Hym., v. 8, p. 122. N. name for Myzine rufiventris
Cresson, not Meria rufiventris Klug.

Taxonomy: Krombein, 1938. Amer. Ent. Soc, Trans. 64: 236-238, fig. 2. 9 , 6 . —Krombein,
1949. Pan-Pacific Ent. 25: 88.

lusa Pate. Idaho, Wash, to Calif.

Lalapa lusa Pate, 1947. N. Y. Ent. Soc, Jour. 55: 128. ♀, ♂.

Brachycistis arenivaga Bradley
Sonoran desert of Calif, and Ariz.; Mexico (Baja California, Sonora).
Brachycistis (Brachycistis) arenivaga Bradley, 1917. Amer. Ent. Soc, Trans. 43: 269. ♂.
Brachycistis (Brachycistis) eriopis Bradley, 1917. Amer. Ent. Soc, Trans. 43: 266. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 20-21, figs. 23, 40, 95, 96, map 1. ♂.
iinsleyi Wasbauer. Mojave, Sonoran and Chihuahuan deserts, Calif., Nev., Ariz., N. Mex. and
Tex.; Mexico (Baja California, Chihuahua).
Brachycistis Iinsleyi Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 21, figs. 16, 17, 41, 97, 98,
map 2. ♂.

Brachycistis nitida (Cresson)
Great Basin and Rocky Mt. grasslands, Nebr., Kans., Tex., Mont., Utah, N.
Mex., Ariz.
Agama nitida Cresson, 1875. Rpt. Geog. Geol. Expl. and Survey west of 100th Meridian, p.

710. ♂.
Photopsis sobrinns Blake, 1886. Amer. Ent. Soc, Trans. 13: 268. ♂.
Photopsis lepidus Blake, 1886. Amer. Ent. Soc, Trans. 13: 269. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 23-25, figs. 43, 44, 99, 100, map 3. ♂.

Brachycistis texana Malloch
Chihuahuan desert of Tex.; Mexico (Coahuila, Nuevo Leon).

Brachycistis (Brachycistis) texana Malloch, 1926. U. S. Natl. Mus., Proc 68 (3): 12. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 25-27, figs. 18, 19, 45, 46, 91, 101, 102,
map 5 (not 4). ♂.




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Brachycistis verticalis Malloch
Sonoran desert of Ariz.; Mexico (Sonora).

Brachycistis (Brachycistis) verticalis Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 19. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 27-30, map 4 (not 5). ♂.

Species Group Petiolata
Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 30-31.

Brachycistis elegantula Cockerell and Casad
Southern Mojave, Sonoran and Chihuahuan deserts of Calif.,
Ariz., N. Mex. and Tex.; Mexico (Baja California, Sonora).
Brachycistus (!) elegantulus Cockerell and Casad, 1894. Ent. News 5: 295. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 31-33, figs. 35, 47, 48, 103, 104, map 6.
6.

Brachycistis imitans Malloch
Central Valley, Mojave and Sonoran deserts of Calif., Ariz.; Mexico (Baja
California).
Brachycistis (Brachycistis) imitans Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 23. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 33-35, figs. 39, 49, 50, 92, 105, 106,
map 7. 6. -Wasbauer, 1969 (1968). Pan-Pacific Ent. 44: 298-299. ♀.

Brachycistis juncea Wasbauer
Calif. (San Bernardino Co.).

Brachycistis juncea Wasbauer, 1966. Univ. Calif. Pubs., Ent. 43: 35, figs. 36, 51, 52, 107,
108. ♂.

Brachycistis lacustris
***authority mismatch
hurdi Wasbauer. Sonoran desert of Calif., Ariz., ? Tex.

Brachycistis lacustris hurdi Wasbauer, 1966. Univ. Calif. Pubs., Ent. 43: 38, map 8. ♂.

Brachycistis lacustris
***authority mismatch
lacustris Malloch. Mojave desert of Calif., Nev.

Brachycistis (Brachycistis) lacustris Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 24. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 37, map 8. ♂.

Brachycistis longula Wasbauer
Calif. (Riverside Co.).

Brachycistis longula Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 39, figs. 37, 55, 56, 111,
112. ♂.

Brachycistis petiolata Fox
Mojave and Sonoran deserts of Calif., Nev., Ariz., also Utah, Idaho; Mexico
(Baja California, Sonora).
Brachycistis petiolata Fox, 1893. Calif. Acad. Sci., Proc. (2) 4: 8. ♂.
Brachycistis gaudii Cockerell, 1901. Canad. Ent. 33: 340. ♂.
Brachycistis (Brachycistis) parva Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 14. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 40-42, figs. 8, 27, 57, 58, 113, 114, map

9. ♂.

Species Group Atrata

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 43.

Brachycistis alcanor (Blake)
Alta., Mont, and S. Dak. south to Ariz, and Tex.; Mexico (Chihuahua, Coahuila,
Durango, Jalisco, Mexico, Puebla, Zacatecas).
Agama ^/ca?io?- Blake, 1871. Amer. Ent. Soc, Trans. 3: 264. ♂.
Brachycistis cremastogaster Melander, 1903. Amer. Ent. Soc, Trans. 29: 329. ♂.
Brachycistis crematogaster (!) Krombein, 1951. U. S. Dept. Agr., Monog. 2: 743.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 46-48, 75, figs. 30, 61, 62, 117, 118,
map 11. ♂.

Brachycistis ampla (Blake)
Colo., Ariz., Nev., Calif., Wash.

Agama ampla Blake, 1879. Amer. Ent. Soc, Trans. 7: 252. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 48, figs. 63, 64, 119, 120, map 12. ♂.

Brachycistis atrata (Blake)
B. C, Wash, and Idaho south to Calif, and Ariz.; Mexico (Baja California,
Sonora).
Agama atrata Blake, 1879. Amer. Ent. Soc, Trans. 7: 253. ♂.
Mutilla agama Dalla Torre, 1897. Cat. Hym., v. 8, p. 7. N. name for Agama atrata Blake,

not Mutilla atrata Linnaeus.
Brachycistis nudus Fox, 1899. Amer. Ent. Soc, Trans. 25: 281. ♂.
Brachycistis 7iigritus Fox, 1899. Amer. Ent. Soc, Trans. 25: 282. ♂.




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Brachycistis contigmis Fox, 1899. Amer. Ent. Soc, Trans. 25: 282. ♂.
Glyptometopa francisca Mickel and Krombein, 1942. Amer. Midland Nat. 28: 656. ♀.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 43-46, figs. 6, 21, 34, 59, 60, 89, 115,
116, map 10. 6. —Wasbauer, 1969 (1968). Pan-Pacific Ent. 44: 297-298 (female synonymy).

Brachycistis carinata Fox
South, Calif, coastal mts.; Mexico (Baja California).

Brachycistis carinatus Fox, 1899. Amer. Ent. Soc, Trans. 25: 283. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 49-51, figs. 20, 65, 121, 122, map 13.
6. —Wasbauer, 1971. Pan-Pacific Ent. 47: 212. ♀.

Brachycistis convexa Wasbauer
Calif. (Imperial Co.).

Brachycistis convexa Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 51, figs. 31, 66, 67, 94,
123, 124. ♂.

Brachycistis cruenta Wasbauer
Tex. (Nueces Co.); Mexico (Tamaulipas).

Brachycistis cruenta Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 53, figs. 68, 69, 125, 126.
6.

Brachycistis curvata Malloch
Sonoran desert of Calif., Ariz.; Mexico (Baja California).

Brachycistis (Brachycistis) curvata Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 16. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 54-56, figs. 70, 71, 127, 128, map 14.
6.

Brachycistis davidi Wasbauer
Tex.; Mexico (Coahuila).

Brachycistis davidi Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 56, figs. 22, 32, 72, 73, 129,
130. ♂.

Brachycistis glabrella (Cresson)
Alta., Oreg., Idaho, Wyo. and Nebr. to Calif, and Tex., south in Mexico to
Oaxaca.
Mutilla glabrella Cresson, 1865. Ent. Soc. Phila., Proc. 4: 441. 6-
Brachycistis (Brachycistis) dentata Bradley, 1917. Amer. Ent. Soc, Trans. 43: 263. ♂.
Brachycistis (Brachycistis) micrommata Bradley, 1917. Amer. Ent. Soc, Trans. 43: 265. ♂.
Brachycistis (Brachycistis) utahensis Malloch, 1926. U. S. Natl. Mus., Proc 68 (3): 17. ♂.
Brachycistis (Brachycistis) melanocephala Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 19.

S.
Brachycistis (Brachycistis) dakotensis Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 20. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 57-59, figs. 25, 33, 74, 75, 131, 132,
map 15. ♂.

Brachycistis inaequalis Fox
South. Calif., Nev., Ariz, and Tex.; Mexico (Baja California).
Brachycistis inaeqrialis Fox, 1899. Amer. Ent. Soc, Trans. 25: 282. ♂.
Brachycistis nevadensis Fox, 1899. Amer. Ent. Soc, Trans. 25: 283. ♂.
Brachycistis nocticola Bradley, 1917. Amer. Ent. Soc, Trans. 43: 289, figs. 19, 39, 40. ♂.

Taxonomy: Krombein, 1954. Ent. Soc. Wash., Proc. 56: 85-86 (synonymy of nocticola).
—Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 59-61, figs. 29, 76, 77, 133, 134, map 16. ♂.

Brachycistis indiscreta Fox
Nebr. to Tex., N. Mex., Ariz.; Mexico (Chihuahua).

Brachycistis indiscreta Fox, 1899. Amer. Ent. Soc, Trans. 25: 284. ♂.
Brachycystis (!) stictinotus Viereck, 1906. Amer. Ent. Soc, Trans. 32: 190. 3-

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 61-63, figs. 26, 38, 78, 135, 136, map
17. ♂.

Brachycistis ioachinensis Bradley
Wash, and Mont, to Calif, and Tex. south in Mexico to Baja California

and Sonora.
Brachycistis (Brachycistis) ioachinensis Bradley, 1917. Amer. Ent. Soc, Trans. 43: 267. ♂.
Brachycistis (Brachycistis) rugosa Malloch, 1926. U. S. Natl. Mus., Proc 68 (3): 11. ♂.
Brachycistis (Brachycistis) siibcarijiata Malloch, 1926. U. S. Natl. Mus., Proc 68(3): 11. ♂.
Brachycistis (Brachycistis) nigrifrons Malloch, 1926. U. S. Natl. Mus., Proc 68 (3): 16. ♂.
Brachycistis (Brachycistis) washingtona Malloch, 1926. U. S. Natl. Mus., Proc 68 (3): 21.

6.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 65-68, figs. 1, 12, 13, 79, 80, 137, 138,
map 18. ♂.




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Brachycistis semiatra Malloch
Wash, to northeast. Calif., to Idaho and Utah.

Brachycistis (Brachycistis) semiatra Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 24. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 68-70, figs. 81, 82, 139, 140, map 19.
6.

Brachycistis subquadrata Fox
Calif. (San Diego Co.).

Brachycistis subquadratus Fox, 1899. Amer. Ent. Soc, Trans. 25: 282. ♂.

Brachycistis (Brachycistis) intermedia Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 18. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 64-65. ♂.

Brachycistis timberlakei Wasbauer
Calif., Nev., Utah, Ariz., N. Mex., Tex.

Brachycistis timberlakei Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 70, figs. 90, 141, 142,
map 20. ♂.

Brachycistis triangularis Fox
South. Calif., Nev., Ariz., N. Mex., Tex.; Mexico (Baja California, Sonora,
Chihuahua, Durango).
Brachycistis triangularis Fox, 1899. Amer. Ent. Soc, Trans. 25: 284. ♂.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 72-73, figs. 24, 83, 84, 93, 143, 144,
map 21. ♂.

Unplaced Taxa of Brachycistis Fox

Brachycistis alutacea (Mickel and Krombein)
Tex., N. Mex.

Stilbopogon alutacea Mickel and Krombein, 1942. Amer. Midland Nat. 28: 659. ♀.

Brachycistis americana (Ashmead)
Calif. (Alameda Co.).

Giyptometopa ainericayia Ashmead, 1898. Psyche 8: 251. ♀.

Brachycistis convergens (Mickel and Krombein)
Calif. (Santa Monica).

Giyptometopa convergens Mickel and Krombein, 1942. Amer. Midland Nat. 28: 657. ♀.

Brachycistis emarginata (Mickel and Krombein)
Tex. (Valentine).

Astigmometopa emarginata Mickel and Krombein, 1942. Amer. Midland Nat. 28: 668. ♀.

Taxonomy: Wasbauer, 1971. Pan-Pacific Ent. 47: 212.

Brachycistis protracta Bradley
Ariz. (Phoenix).

Brachycistis (Brachycistis) protracta Bradley, 1917. Amer. Ent. Soc, Trans. 43: 270. ♂.
Brachycistis (Brachycistis) protractor (!) Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 7.

Taxonomy: Wasbauer, 1966. Univ. Calif. Pubs. Ent. 43: 74. ♂.

Colocistis brevis (Fox)
Ariz., Nev., Calif.

Brachycistis brevis Fox, 1899. Amer. Ent. Soc, Trans. 25: 285. ♂.

Colocistis castanea (Cresson)
Tex., N. Mex., Colo., Ariz., Nev., Calif.

Mutilla castanea Cresson, 1865. Ent. Soc. Phila., Proc. 4: 388. ♂.

Colocistis crassa (Bradley)
Tex., Colo., Ariz., Nev., Calif.

Brachycistis (Brachycistis) crassa Bradley, 1917. Amer. Ent. Soc, Trans. 43: 277. ♂.

Colocistis eremi (Bradley)
Ariz., Nev., Calif.

Brachycistis (Brachycistis) eremi Bradley, 1917. Amer. Ent. Soc, Trans. 43: 279. ♂.

Colocistis pilosa Krombein
Calif. (San Diego).

Colocistis pilosa Krombein, 1942. Ent. Soc. Wash., Proc. 44: 66. ♂.

Colocistis stygia (Bradley)
Ariz. (Nogales).

Brachycistis (Brachycistis) stygia Bradley, 1917. Amer. Ent. Soc, Trans. 43: 276. ♂.

Colocistis thermarum (Bradley)
Ariz.

Brachycistis (Brachycistis) thermarum Bradley, 1917. Amer. Ent. Soc, Trans. 43: 274. ♂.

Type-species: Acanthetropis lamellatus Wasbauer. Orig. desig.

Taxonomy: Wasbauer, 1968. Pan-Pacific Ent. 44: 197, figs. 20, 24.

Acanthetropis aequalis (Fox)
Nebr., Kans., Colo., Idaho, Nev., Ariz.

Brachycistis aequalis Fox, 1899. Amer. Ent. Soc, Trans. 25: 284. ♂.

Acanthetropis idiotes
***authority mismatch
(Coclierell). N. Mex., CaUf.

Brachycistis idiotes Cockerell, 1895. Ent. News. 6: 63. ♂.

Acanthetropis noctivaga (Bradley)
Utah, Nev., Ariz., Calif.; Mexico (Sonora, Baja California).

Brachycistis {Brachycistis) noctivaga Bradley, 1917. Amer. Ent. Soc, Trans. 43: 272. ♂.

Acanthetropis normalis (Malloch)
S. Dak., Colo., Ariz., Calif.

Brachycistis normalis Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 8. ♂.

Brachycistina acuta (Malloch)
Ariz., Nev., Calif.

Brachycistis {Brachycistina) acuta Malloch, 1926. U. S. Natl. Mus., Proc. 68 (3): 25. ♂.

Hadrocistis bicolor Wasbauer
Calif. (Imperial Co.).

Hadrocistis bicolor Wasbauer, 1968. Pan-Pacific Ent. 44: 187, figs. 4-6, 14, 18. ♂.

Hadrocistis slanskyae Wasbauer
Calif. (Imperial Co.).

Hadrocistis slanskyae Wasbauer, 1968. Pan-Pacific Ent. 44: 188, figs. 1-3, 13, 17. ♂.

Brachycistellus figitiformis Baker
Calif. (Colusa Co., Claremont).

Brachycistellus figitiformis Baker, 1907. Invertebrata Pacifica 1: 178. ♂.

Taxonomy: Wasbauer, 1968. Pan-Pacific Ent. 44: 195-196, figs. 10-12, 16, 19. ♂.

Quemaya arenicola Wasbauer
Calif. (Imperial Co.).

Quemaya arenicola Wasbauer, 1967. Biol. Soc. Wash., Proc. 80: 169, figs. 2-6. ♂.

Quemaya inermis (Malloch)
Ariz. (Higley).

Brachycistus (!) (Brachycistellus) inermis Malloch, 1924. Brooklyn Ent. Soc, Bui. 19: 23.
d.

Quemaya marcida (Bradley)
Calif. (Calexico).

Brachycistis {Brachycistellus) marcida Bradley, 1917. Amer. Ent. Soc, Trans. 43: 283. ♂.

Quemaya paupercula (Bradley)
Calif., Nev.

Brachycistis {Brachycistellus) paupercula Bradley, 1917. Amer. Ent. Soc, Trans. 43: 282.

Quemaya perpunctata (Cockerell)
N. Mex., Calif.

Brachycistis perpunctatus Cockerell, 1896. Amer. Ent. Soc, Trans. 22: 291. ♂.

Taxonomy: Wasbauer, 1968. Biol. Soc. Wash., Proc. 80: 170, fig. 1.




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Glyptacros angustior Mickel and Krombein
Ariz.

Glyptacros angustior Mickel and Krombein, 1942. Amer. Midland Nat. 28: 660. ♀.

Glyptacros ashmeadi Mickel and Krombein
No locality given.

Glyptacros ashmeadi Mickel and Krombein, 1942. Amer. Midland Nat. 28: 661. ♀.

Xeroglypta egregia
***authority mismatch
Mickel and Krombein. Calif. (Palm Springs).

Xeroglypta egregia Mickel and Krombein, 1942. Amer. Midland Nat. 28: 663. ♀.

Aulacros latior Mickel and Krombein
Calif. (Palm Springs).

Aulacros latior Mickel and Krombein, 1942. Amer. Midland Nat. 28: 665. ♀.

Bruesiella furtivus (Mickel and Krombein)
Ariz. (Tucson, Avondale Ranch).

Eurycros furtivus Mickel and Krombein, 1942. Amer. Midland Nat. 28: 667. ♀.

Aglyptacros eureka (Banks)
Ariz. (Palmerlee).

Glyptometopa eureka Banks, 1912. Canad. Ent. 44: 202. ♀.

Aglyptacros paxillatus Mickel and Krombein
Colo. (La Junta).

Aglyptacros paxillatus Mickel and Krombein, 1942. Amer. Midland Nat. 28: 673. ♀.

Aglyptacros segredentatus Mickel and Krombein
Ariz. (Benson).

Aglyptacros segredentatus Mickel and Krombein, 1942. Amer. Midland Nat. 28: 671. ♀.

Aglyptacros sulcatus Mickel and Krombein
Tex. (Valentine).

Aglyptacros sulcatus Mickel and Krombein, 1942. Amer. Midland Nat. 28: 674. ♀.

Two of our native species have been reared from larvae of tiger beetles, the known host of
several extrahmital species.
Only the typical subgenus occurs in North America.

Morphology: Reid, 1941. Roy. Ent. Soc. London, Trans. 91: 389-390, figs. 26-28 (female, male
thorax).

Methocha californica Westwood
Calif., Nev., Wash. Host: Cicindela senilis Horn ?; cicindelid sp.; reared
in laboratory on Omus califomicus Esch.
Mefhoca (!) californica Westwood, 1881. Ent. Soc. London, Trans., p. 133. ♀.
Methoca (!) nigrior Fox, 1899. Amer. Ent. Soc, Trans. 25: 288. ♂.

Biology: Bridwell, 1912. Pacific Coast Ent. Soc, Proc. 1: 46-48 (host). — Burdick and

Wasbauer, 1959. Wasmann Jour. Biol. 17: 75-88, 6 figs, (prey capture, oviposition, life

history, egg, larva).

Methocha formosa Krombein
Fla. (Arcadia).

Methocha (Met hoc ha) formosa Krombein, 1954. Amer. Ent. Soc, Trans. 80: 2, figs. 1, 3. ♀.

Methocha impolita Krombein
W. Va., D. C, N. Y.

Methocha (Methocha) impolita Krombein, 1958. Ent. Soc. Wash., Proc. 60: 58. ♀.

Methocha stygia (Say)
General in south. Canada and U. S. Host: Cicindelid larvae.

Mutilla (Methoca (!)) pacalis Harris, 1835. In Hitchcock, Rpt. Geol. Mineral. Bot. Zool.
Mass., p. 587. Nom. nud.

Tengyra stygia Say, 1836. Boston Jour. Nat. Hist. 1: 299. ♂.

Methoca (!) bicolor Say, 1836. Boston Jour. Nat. Hist. 1: 299. ♀.

Methoca (!) cayiadensis Smith, 1855. Cat. Hym. Brit. Mus., v. 3, p. 67. ♂.

Taxonomy: Evans, 1965. Ent. Soc Wash., Proc 67: 88-90, figs. 1-5 (larva).
Biology: Williams, 1916. Psyche 23: 121-125, pi. 12.

Myrmosa blakei Bradley
N. Y., Va.

Myrmosa (Myrmosa) blakei Bradley, 1917. Amer. Ent. Soc, Trans. 43: 251. ♀.

Taxonomy: Krombein, 1940. Amer. Ent. Soc, Trans. 65: 438. ♀.

Myrmosa bradleyi Roberts
Wash., Calif.

Myrmosa bradleyi Roberts, 1929. Psyche 36: 362. ♂.

Taxonomy: Krombein, 1940. Amer. Ent. Soc, Trans. 65: 436-438, fig. 2. ♂, ♀.

Biology: Linsley, 1960. Pan-Pacific Ent. 36: 36 (mating behavior).

Myrmosa peculiaris Krombein
Kans. (Onaga).

Myrmosa (Myrrnosa) peculiaris Krombein, 1940. Amer. Ent. Soc, Trans. 65: 438. ♀.

Myrmosa unicolor Say
N. B. south to N. C, west to B. C. and Ariz. Host: Tiphia sp.; Lindenius
columbianus errans (Fox) ?; Dialictus pruinosus (Robt.); D. inconspicuus (Sm.).
Myrmosa unicolor Say, 1824. In Keating, Narr. Long's 2nd. Exped., v. 2, p. 331. ♂.




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Ischioceras mgosa Provancher, 1882. Nat. Canad. 13: 8. 9, ( Mutilla thoracica Blake, 1886. Amer. Ent. Soc, Trans. 13: 204. Preocc.
Mutilla erythronota Dalla Torre, 1897. Cat. Hym., v. 8, p. 36. N. name.
Myrmosa {Mynnosa) banksi Bradley, 1917. Amer. Ent. Soc, Trans. 43: 249. ♀, ♂.
Myrmosa dakotensis Weber, 1934. Psyche 41: 57. ♀.

Taxonomy: Krombein, 1940. Amer. Ent. Soc, Trans. 65: 430-436, fig. 6.

Biology: Melander and Brues, 1903. Biol. Bui. 5: 7. —Krombein, 1940. Amer. Ent. Soc, Trans.
65: 435-436. —Krombein, 1956. Ent. Soc. Wash., Proc 58: 154 (mating behavior).

Myrmosa nocturna
***authority mismatch
nocturna Krombein. Md., La., Tex., Kans.

Myrmosa (Myrmosina) nocturna nocturna Krombein, 1940. Amer. Ent. Soc, Trans. 65:

454. ♂.

Myrmosa nocturna
***authority mismatch
rufigastra Krombein. Tex., Calif.

Myrmosa {Myrmosina) nocturna ncfigastra Krombein, 1940. Amer. Ent. Soc, Trans. 65:

455. ♂.

Myrmosa texana Krombein
Dak., Tex.

Myrmosa (Myrmosina) texana Krombein, 1940. Amer. Ent. Soc, Trans. 65: 453. ♂.

Leiomyrmosa spilota Wasbauer
Calif. (Blythe in Riverside Co.).

Leiomyrmosa spilota Wasbauer, 1974 (1973). Pan-Pacific Ent. 49: 326, figs. 1, 2, 19, 23. ♀.

Myrmosula boharti Wasbauer
Calif. (Thousand Palms in Riverside Co.).

Myrmosula boharti Wasbauer, 1974 (1973). Pan-Pacific Ent. 49: 331, figs. 13, 14. ♀.
exaggerata (Krombein). Calif.

Myrmosa (Myrmosula) exaggerata Krombein, 1940. Amer. Ent. Soc, Trans. 65: 460. ♀.

Taxonomy: Mickel, 1940. Pan-Pacific Ent. 16: 133.

Myrmosula nasuta Wasbauer
Southern Calif, and Ariz. Ecology: Visits mat Euphorbia and has been
collected in a kangaroo rat burrow.
Myrmosula nasuta Wasbauer, 1974 (1973). Pan-Pacific Ent. 49: 329, figs. 7, 8, 20. ♀.

Myrmosula pacifica (Mickel)
Calif. (Antioch). Possibly a synonym of exaggerata Krom.

Myrmosa (Myrmosula) pacifica Mickel, 1940. Pan-Pacific Ent. 16: 134. ♀.

Myrmosula parvula (Fox)
D. C. south to Ala., west to Mont, and Tex. Host: Dialictus imitatus (Sm.), D.
zephynis (Sm.); Augochlorella striata (Prov.)(?), A. persimilis (Vier.)(?).
Myrmosa parvula Fox, 1893. N. Y. Ent. Soc, Jour. 1: 53. ♂.
Mutilla antisemitica Dalla Torre, 1897. Cat. Hym., v. 8, p. 10. N. name for Myrmosa

Myrmosula parvula
***authority mismatch
fox, not Mutilla parvula Fabricius.
Brachycistis bimaculatus Fox, 1899. Amer. Ent. Soc, Trans. 25: 285. ♀.

Taxonomy: Krombein, 1940. Amer. Ent. Soc, Trans. 65: 457-459, figs. 5, 10. ♂, ♀.

Biology: Michener and Wille, 1961. Univ. Kans. Sci. Bui. 42: 1130 (host record). — Ordway,
1964. Kans. Ent. Soc, Jour. 37: 149 (host records). — Batra, 1965. Kans. Ent. Soc, Jour. 38:
386-387 (behavior in host nest). —Brothers, 1972. Univ. Kans. Sci. Bui. 50: 23 (host record).




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Myrmosula peregrinatrix (Krombein)
Alta., Ariz., Tex.

Mijnnosa (Myrmosula) peregrinatrix Krombein, 1946. Ent. Soc. Wash., Proc. 48: 247. ♀.

Myrmosula rufiventris Blake
Nev., Oreg.

Myrmosa rufiventris Blake, 1879. Amer. Ent. Soc, Trans. 7: 254. ♂.
Mutilla erythrogastra Dalla Torre, 1897. Cat. Hym., v. 8, p. 36. N. name for Mymiosa

Myrmosula rufiventris Blake
not Mutilla rufiventris Klug nor M. rufiventris Smith.

Taxonomy: Krombein, 1940. Amer. Ent. Soc, Trans. 65: 459-460. ♂.

Myrmosula rutilans (Blake)
Calif. Host: No7nadopsis scutellaris (Fwlr.)

Mutilla rutilans Blake, 1879. Amer. Ent. Soc, Trans. 7: 248. ♀.

Taxonomy: Krombein, 1940. Amer. Ent. Soc, Trans. 65: 461-462, fig. 12. ♀.


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