16.
Thelypodium wrightii
A. Gray, Smithsonian Contr. Knowl. 3(5): 7. 1852.
Stanleyella wrightii
(A. Gray) Rydberg;
Thelypodium wrightii
subsp.
oklahomensis
Al-Shehbaz
Biennials;
slightly glaucous, glabrous.
Stems
branched distally, (6.5-)9.5-22.5(-28) dm.
Basal leaves
(and proximal cauline): petiole (2-)4.5-8.5 (-13.7) cm; blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5-)9.5-22.5(-28) cm × 28-55(-75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate).
Cauline leaves
petiolate; blade lanceolate to linear-lanceolate, 3.5-8.5(-13.5) cm × 7-14(-23) mm, margins usually entire or dentate, rarely lobed.
Racemes
somewhat lax, considerably elongated in fruit.
Fruiting pedicels
horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5-)7-13(-17) mm, slightly flattened at base.
Flowers:
sepals spreading or ascending oblong to linear, (3-)4-6(-7) × (0.8-)1-1.3(-1.5) mm; petals usually white, rarely lavender, linear or oblong, 4-7.5(-9) × 1-1.8(-2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base); nectar glands confluent; filaments equal, (3-)3.5-6.5(-8.5) mm; anthers (1-)1.5-2.5(-3) mm, circinately coiled; gynophore 0.2-2(-5) mm.
Fruits
horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5-)3.8-7.4(-9) cm × 1-1.2(-1.5) mm, (replum not constricted between seeds); ovules 76-128 per ovary; style usually clavate to subclavate, rarely cylindrical, (0.5-)0.8-2(-3) mm.
Seeds
0.7-1.3 (-1.5) × 0.5-0.8 mm.
Flowering (Mar-)Jun-Oct. Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands; 1200-2300 m; Ariz., Colo., N.Mex., Tex., Utah; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo).
Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp.
oklahomensis
and subsp.
wrightii
, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993).
