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Andrena ceanothi, female, back1 2012-08-07-17.53.44 |
© Copyright Laurence Packer 2014
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Andrena ceanothi FEM CFP comp |
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© Copyright Laurence Packer 2014
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Andrena ceanothi MALE CFP comp |
Mitchell, Bees of the Eastern United States, Vol. I, 1960 · 1
Andrena ceanothi, figure36b |
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© Rebekah Andrus Nelson
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Andrena ceanothi, female, face |
Deana Crumbling · 1
Andrena ceanothi 072375 prop tri |
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Deana Crumbling · 1
Andrena ceanothi female, narrow fovea |
Deana Crumbling · 1
Andrena ceanothi F-a, scutellum pits |
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Overview |
Reprinted with permission of the American Entomological Society from:
LaBerge, W. E. 1973. A revision of the bees of the genus Andrena of the Western Hemisphere. Part VI. Subgenus Trachandrena. Transactions of the American Entomological Society 99: 235-371.
Please report text errors to: leah at discoverlife dot org.
Andrena ceanothi is a relatively common eastern species related to A. heraclei. The female of ceanothi is marked by a sparsely punctate clypeus, the terga having apical margins broadly hyaline and clear, and the facial foveae shaped more or less as in A. forbesii. The female differs from that of heraclei in the longer head and eyes, in the shape of the facial fovea as described below, and in the more crowded and coarser punctures of the face above the antennal fossae. The male of ceanothi is readily confused with that of heraclei but can be distinguished by the more densely punctate terga and sterna and the tergal apical rims usually being more broadly hyaline.
FEMALE. MEASUREMENTS AND RATIOS. — N = 20; length, 8-11 mm; width, 2.5-3.5 mm; wing length, M = 3.84 ± 0.088 mm; FL/FW, M = 1.12 ± 0.004; FOVL/FOVW, M = 4.05 ± 0.060.
INTEGUMENTAL COLOR. — Black except as follows: mandible with apical half rufescent (often basally as well); flagellar segments 2-10 dark reddish-brown below; tegula with colorless margin, translucent to colorless summit, otherwise dark red; wing membranes hyaline, moderately infumate, yellowish-brown, veins dark reddish-brown; terga 1-4 with hyaline colorless rims that on tergum 2 equals in length one-eighth to one-tenth median tergal length; sterna and tarsi dark rufescent.
STRUCTURE. — Antennae, malar space and galea as in spiraeana. Eyes each about four times as long as broad, inner margins parallel. Maxillary palpus as in sigmundi but segmental ratio about as 1.0:1.0:0.7:0.7:0.6:0.7. Labial palpus as in sigmundi but ratio about as 1.0:0.7:0.7:0.7. Labrum as in spiraeana. Clypeus with coarse, round punctures of irregular size and spacing, in median third punctures separated mostly by one-half to one puncture width or more, usually with an irregular median impunctate line or with a subapical impunctate area; surface shiny, unshagreened. Supraclypeal area with smaller round punctures separated largely by one puncture width or less, surface shiny, unshagreened. Genal area in profile only slightly broader than eye; lateral surface with crowded punctures, becoming slightly rugulose near mandibles, surface shiny, unshagreened; ventral surface as in sigmundi. Vertex above lateral ocellus equals about one ocellar diameter, with coarse punctures in one or two rows (rarely three) above lateral ocellus; surface moderately shiny with delicate reticular shagreening. Face above antennal fossae with coarse crowded punctures separated mostly by less than half a puncture width; surface moderately shiny. Facial fovea as in sigmundi but separated from lateral ocellus by less than one ocellar diameter and lower portion exceptionally narrow, distance from inner eye margin to inner foveal margin equals less than from inner foveal margin to margin of antennal fossa at the same level (as in spiraeana); greatest distance from inner eye margin to facial fovea not much greater than width of fovea at that level.
Thoracic form and sculpturing essentially as in spiraeana. Tibial spurs and wing venation as in spiraeana.
Metasomal terga 1-4 as in heraclei except as follows: terga 2-4 with apical areas less than half as long as median tergal length; punctures small and crowded, usually separated by less than half a puncture width except tergum 1 with basal area punctures slightly sparser and apical impunctate rims broader. Pygidial plate and sterna as in heraclei.
VESTITURE. — Generally white but dorsum of thorax often cinereous and terga 5 and 6 and inner surfaces tarsi yellow. Form and distribution much as in sigmundi but usually sparser; terga 2 and usually 3 with apical pale fascia interrupted medially.
MALE. MEASUREMENTS AND RATIOS. — N = 20; length, 8-11 mm; width, 2.0-2.5 mm; wing length, M(19) = 3.35 ± 0.193 mm; FL/FW, M = 1.20 ± 0.005; FS1/FS2, M = 0.73 ± 0.012.
INTEGUMENTAL COLOR. — As in female except as follows: tegulae often entirely translucent, reddened only near base; tergal hyaline apical rims longer, in length equal to one-sixth to one-eighth median tergal length.
STRUCTURE. — Antennae in repose not quite reaching propodeum, usually not reaching metanotum; scape equal in length to first two flagellar segments or slightly shorter; flagellar segments as in forbesii; surface dull as in sigmundi. Eyes each about three and one-half times as long as broad or slightly shorter, converging slightly toward mandibles. Malar space, mandible and galea as in female. Maxillary palpus as in female but segmental ratio about as 1.0:0.9:0.7:0.7:0.7:0.8. Labial palpus as in female but ratio about as 1.0:0.5:0.5:0.6. Labrum as in sigmundi. Clypeus as in female but punctures slightly more crowded and usually without median impunctate line. Supraclypeal area punctures small, crowded, surface moderately dull. Genal area as in spiraeana but punctures more discrete. Vertex as in female but opaque to moderately dull. Face above antennal fossae with coarse contiguous punctures, rugae absent or weak, surface dulled by fine shagreening.
Thoracic sculpturing as in spiraeana. Tibial spurs and wing venation as in forbesii.
Metasomal terga generally as in heraclei with following exceptions: tergum 1 basal area with punctures separated mostly by less than one puncture width, apical area punctures more crowded, separated by less than half a puncture width especially laterally; terga 2-4 similar but apical areas progressively less punctate from tergum 3 posteriorly. Sterna as in heraclei but punctures larger and denser in median area of each sternum. Sternum 6 with dish-shaped apicomedian emargination. Terminalia as in sigmundi; see figures 58-59.
VESTITURE. — White to ochraceous, generally darker on vertex and thoracic dorsum; distribution and form as in sigmundi but tergum 2 with pale apical fascia dense and broadly interrupted medially, tergum 3 with dense apical fascia usually narrowly interrupted medially, and terga 4 and 5 with apical pale fasciae usually complete.
Reprinted with permission from: Mitchell, T.B. 1960. Bees of the Eastern United States. North Carolina Agricultural Experiment Station Technical Bulletin No. 141.
FEMALE. — Length 10 mm.; face very slightly longer than broad; clypeus broadly convex, projecting about one-third below suborbital line, smooth and shining, with a rather indefinite median impunctate line, irregularly but rather deeply punctate on each side, punctures well separated toward center, becoming close at extreme sides; facial foveae rather broad above, occupying about two-thirds of space between eyes and ocelli, lower half abruptly and strongly constricted to a width about one-fourth that of the upper part, covered with whitish tomentum; space between margin of vertex and lateral ocelli somewhat wider than their diameter; cheeks slightly broader than eyes, rounded posteriorly, somewhat shining, deeply, distinctly, closely and rather finely punctate, punctures becoming sparse on vertex laterally just above eyes and foveae; malar space very short; basal segment of flagellum slightly shorter than 2nd and 3rd combined; process of labrum quite broad, rather short, slightly narrowed to the broadly truncate apical margin; pubescence of head very short and sparse, entirely whitish, that on thorax whitish laterally and beneath, more greyish above, thin and sparse, that on legs whitish on the more basal segments, becoming somewhat brownish on tarsi; scutum and scutellum somewhat shining between coarse and deep punctures, which are quite close on scutum anteriorly, becoming rather sparse in center posteriorly, those on scutellum very close and coarse; pleura coarsely rugose, becoming more finely so posteriorly; dorsal area of propodeum quite narrow, oblique, triangle broad, coarsely and evenly striate; propodeal corbicula quite short, poorly developed, whitish, without an anterior fringe; trochanteral floccus short and poorly developed, whitish; hind tibiae somewhat broadened apically, apex not quite twice width of basitarsi, scopa rather dense, hairs rather short, fine, entirely whitish; front and mid basitarsi slightly narrower than their respective tibiae; 2nd submarginal cell considerably shorter than 3rd, receiving 1st recurrent just beyond middle; abdominal terga somewhat shining between close, deep and rather coarse punctures, apical impressed areas broad and shallow, occupying medially fully half length of discs, anterior margin indicated by a slightly bowed, transverse, nearly impunctate line, becoming narrowly yellowish-hyaline along rims, discal pubescence extremely short, thin and suberect, apparently entirely pale, apical, white fasciae quite dense laterally on terga 2-4, more or less widely interrupted medially, tergum 5 with a pale brownish, apical fimbria.
MALE. — Length 8 mm.; face somewhat longer than broad; clypeus somewhat protuberant, strongly convex, projecting about one-third below suborbital line; smooth and shining between deep and rather close and coarse punctures; space between margin of vertex and lateral ocelli subequal to their diameter; cheeks subequal to eyes in width, rounded posteriorly, rather dull, obscurely and very finely striate, with some barely evident, fine punctures along margins of eyes; malar space short; basal segment of flagellum short, only slightly longer than apical width, considerably shorter than 2nd and following segments; process of labrum quite broad, rather short, slightly narrowed to the broadly truncate apical margin; mandibles of moderate length, with a distinct, inner, subapical tooth, overlapping about one-third; pubescence of head, thorax and legs entirely whitish-ochraceous; scutum and scutellum somewhat shining between deep, distinct and rather coarse punctures, close over anterior half of scutum, becoming slightly more sparse in center posteriorly, those on scutellum slightly more coarse and very close; pleura rather shallowly rugose; dorsal area of propodeum oblique, triangle rather coarsely striate; all basitarsi slender and elongate, considerably narrower than their respective tibiae: 2nd submarginal cell much shorter than 3rd, receiving 1s; recurrent slightly beyond middle; abdominal terga somewhat shining between deep, distinct and rather coarse and close punctures, apical impressed areas rather shallow, occupying medially about one-third length of discs, becoming narrowly yellowish-hyaline along rims, discal pubescence short, thin and erect, entirely pale, terga 2-4 with white apical fasciae evident laterally, widely interrupted medially; apical portion of sternum 8 very narrow and slender, very slightly broader toward apex, which is truncate, clothed beneath with rather dense, long pubescence; penis valves rather slender apically, somewhat dilated basally, excavated laterally, gonocoxites abruptly and very broadly dilated apically, upper surface being rather strongly concave, gonocoxal lobes strongly produced, narrow, tips just reaching apical lobes (similar to hippotes, fig. 37).
DISTRIBUTION. — Minnesota to the New England states and Nova Scotia, south to Georgia; April to July.
FLOWER RECORDS. — Amelanchier, Ilex, Pyrus, Rhus, Salix, Sassafras, Symplocos, Vaccinium and Viburnum. Recorded by Brittain and Newton (1934) on Diervilla, Prunus and Pyrus malus.
Described using the synonymous name: Andrena compacta Mitchell
MALE. — Length 9 mm.; face somewhat longer than distance between eyes above; eyes slightly convergent below; vertex quite broad, lateral ocelli separated from its hind margin by considerably more than their diameter; cheeks subequal to eyes in width; clypeus convex, projecting somewhat less than one-half below suborbital line, smooth and shining between rather close, deep, distinct, and uniformly distributed punctures; punctures over remainder of head shallow and obscure but rather coarse and close, surface quite densely tessellate; malar space very short; process of labrum broad and short. broadly truncate; basal segment of flagellum much shorter than segment 2, very slightly longer than broad; mandibles rather short, with a distinct, inner, subapical tooth only slightly removed from tip; pubescence of entire head, thorax and legs quite long and erect, but rather thin, entirely creamy; dorsum of thorax somewhat shining between quite coarse, deep and distinct punctures, these close on anterior half of scutum, becoming slightly separated in center posteriorly, those on scutellum more irregular, slightly separated along anterior margin; pleura coarsely rugoso-punctate; triangle of propodeum coarsely striate, lateral and posterior surfaces of propodeum coarsely rugose or reticulate, the propodeal corbicula rather well developed; legs blackish, becoming somewhat reddened on apical tarsal segments; spurs pale yellow; wings subhyaline, veins and stigma brownish-ferruginous, 2nd submarginal cell quite short, receiving 1st recurrent toward apex; tegulae yellowish-hyaline posteriorly, becoming piceous anteriorly; abdominal terga closely, deeply, distinctly and rather coarsely punctate, apical impressed areas occupying slightly less than half the median length of each plate, becoming narrowly yellowish-hyaline along rims, the punctures gradually becoming slightly finer toward these rims, discal pubescence very short, erect, thin, entirely whitish, apical fasciae evident toward sides of segments 2-4; sternum 7 only slightly emarginate medially (as in pseudobscura, figure 36); sternum 8 also quite similar to pseudobscura, but apical process more slender and elongate; genital armature as in hippotes (Figure 37).
TYPES. — Holotype: Male, Roscommon Co., Mich., May 26, 1957. Paratype: 1 M, topotypical, May 31, 1957 (both R. & K. Dreisbach) [Dreisbach].
Compacta evidently is closely related to pseudobscura, and the 7th and 8th sterna and the genital armature are nearly identical. The chief differences are the somewhat shorter apical impressed area of the abdominal terga, with the punctures considerably coarser in compacta, in comparison with pseudobscura. In the latter, the impressed area occupies fully half the median length of each tergal plate. There is also a difference in color of pubescence, that in pseudobscura varying from markedly fulvous to ochraceous on the dorsum of the thorax, while in compacta it is pale ochraceous to whitish or creamy.
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Updated: 2024-04-26 14:34:23 gmt
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