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Heterotheca villosa (Pursh) Shinners
HAIRY FALSE GOLDENASTER
Life   Plantae   Dicotyledoneae   Asteraceae   Heterotheca

Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa

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Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa
Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa

Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa
Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa

Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa
Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa

Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa
Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa

Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa
Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa

Heterotheca villosa
© Copyright Mel Harte 2010 · 3
Heterotheca villosa

Associates · map
FamilyScientific name @ source (records)
Andrenidae  Andrena costillensis @ BBSL (2)

Calliopsis chlorops @ BBSL (1)

Perdita asteris @ BBSL (1)

Perdita fallax @ BBSL (18)

Perdita stottleri @ BBSL (1)
Apidae  Anthophora albata @ BBSL (1)

Bombus bifarius @ RMBL_ENT (11)

Bombus flavifrons @ RMBL_ENT (9)

Bombus frigidus @ RMBL_ENT (1)

Bombus insularis @ RMBL_ENT (17)

Bombus mixtus @ RMBL_ENT (6)

Bombus rufocinctus @ RMBL_ENT (1); BBSL (3)

Bombus sylvicola @ RMBL_ENT (6)

Epeolus mesillae @ BBSL (2)

Epeolus minimus @ BBSL (1)

Holcopasites pulchellus @ BBSL (1)

Melissodes confusa @ RMBL_ENT (1); DART_ENT (1)

Melissodes coreopsis @ BBSL (2)

Melissodes hymenoxidis @ BBSL (3)

Melissodes lutulenta @ BBSL (17)

Melissodes microsticta @ BBSL (3)

Melissodes pallidisignata @ BBSL (1)

Neolarra vigilans @ BBSL (3)

Triepeolus eldredi @ BBSL (1)

Triepeolus helianthi @ DART_ENT (1)
Cantharidae  Chauliognathus basalis @ I_ADG (1)
Colletidae  Colletes lutzi @ BBSL (1)

Colletes phaceliae @ BBSL (5)

Hylaeus wootoni @ BBSL (1)
Halictidae  Agapostemon angelicus @ BBSL (1)

Halictus confusus @ DART_ENT (1); CUIC_ENT (1); RMBL_ENT (2)

Halictus rubicundus @ RMBL_ENT (1)

Halictus virgatellus @ RMBL_ENT (5); DART_ENT (3)

Lasioglossum inconditus @ BBSL (1)

Lasioglossum lazulis @ BBSL (4)

Lasioglossum peraltum @ RMBL_ENT (2)

Lasioglossum pruinosum @ BBSL (1)
Megachilidae  Ashmeadiella californica @ BBSL (2)

Ashmeadiella gillettei @ BBSL (3)

Ashmeadiella pronitens @ DART_ENT (1)

Coelioxys sodalis @ DART_ENT (1)

Heriades cressoni @ BBSL (1)

Hoplitis albifrons @ DART_ENT (1)

Hoplitis fulgida @ RMBL_ENT (4)

Megachile nevadensis @ BBSL (2)

Megachile perihirta @ BBSL (2)

Megachile pugnata @ DART_ENT (1)

Megachile relativa @ RMBL_ENT (1); DART_ENT (2)

Osmia grindeliae @ RMBL_ENT (1); DART_ENT (1)

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Following modified from Plants Database, United States Department of Agriculture
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http://plants.usda.gov/java/profile?symbol=HEVI4 ---> https://plants.usda.gov/java/profile?symbol=HEVI4
&pull 20q v4.662 20091102: Error 501 Protocol scheme 'https' is not supported (LWP::Protocol::https not installed) https://plants.usda.gov/java/profile?symbol=HEVI4

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FNA | Family List | FNA Vol. 20 | Asteraceae | Heterotheca

15. Heterotheca villosa (Pursh) Shinners, Field & Lab. 19: 71. 1951.

Hairy goldenaster

Amellus villosus Pursh, Fl. Amer. Sept. 2: 564. 1813; Chrysopsis villosa (Pursh) Nuttall ex de Candolle; Diplogon villosum (Pursh) Kuntze; Diplopappus villosus (Pursh) Hooker

Perennials, (5—)16—40(—70) cm; taprooted. Stems 1—50+, decumbent to erect (sometimes brown or reddish brown, sometimes whitish distally, sometimes ± brittle), sparsely to densely hispido-strigose, sparsely to abundantly long-hispid, eglandular or sparsely to densely stipitate-glandular. Leaves generally not crowded; proximal cauline petiolate, blades oblanceolate, (90—)220—400(—600) × (2—)4—8(—13) mm, bases cuneate, margins flat, usually entire, rarely with 1—2 apical teeth, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; distal sessile, blades usually lanceolate or oblanceolate to oblong, rarely ovate or lanceolate-triangular, (4—)15—28(—42) × (1.5—)3.5—7(—12.5) mm, bases attenuate to convex-cuneate to rounded, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular. Heads 1—16(—42), in usually open, corymbiform, rarely paniculiform arrays. Peduncles (4—)15—41(—98) mm, sparsely to densely hispido-canescent, eglandular or sparsely to densely stipitate-glandular; bracts 1—7+, usually linear-oblanceolate, rarely leaflike and linear-oblanceolate, usually greatly reduced, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; rarely 1—2 leaflike, oblanceolate bracts subtending heads. Involucres narrowly cylindric to campanulate, (5—)6—9.5(—13) mm. Phyllaries in 4—5 series, lanceolate to linear-lanceolate or triangular-lancelate, unequal (outer 1 / 5 — 1 / 3 length of inner), margins scarious, sometimes reddish purple distally, ciliate distally or apically, faces sparsely to densely strigose, eglandular or sparsely to moderately stipitate-glandular. Ray florets (5—)10—20(—38); laminae (3.5—)6.5—11(—20) × 1—2(—3) mm. Disc florets (10—)20—50(—85); corollas barely ampliate, (4—)5—6(—8) mm, glabrous or glabrate (few, minute hairs), lobes 0.4—0.75(—1) mm, glabrous or glabrate (hairs 0.1—0.35 mm). Cypselae monomorphic, obconic, compressed, 1.7—2.7(—3.4) mm, ribs 4—8(—10) (rarely brownish), faces sparsely to moderately strigose; pappi off-white, outer of linear scales 0.25—1 mm, inner of 30—45 bristle (4—)5—6.5(—8.5) mm, longest weakly clavate (usually equaling or longer than corollas). 2 n = 18, 36.

Varieties 9 (9 in the flora): w North America.

Heterotheca villosa is variable in stem height, leaf base shape, stem and leaf indument, number of heads, and number of florets. Thus, the species is difficult to circumscribe because each variety has a suite of diagnostic traits and a distribution that is restricted to a portion of the range of the species. Generally, var. minor (glandular), and to a lesser extent, var. foliosa (eglandular) are the glue holding the other varieties together in a widely distributed polymorphic species; this is comparable to the situation in H. sessiliflora , in which var. echioides is the glue. Variety minor hybridizes with all other varieties. Numerous local races occur that are sometimes quite distinct when extreme, but they intergrade with one or more other races, especially in var. minor (J. C. Semple 1996). The species is divided here on the basis of indument features, leaf shape, and stem height, paralleling the infraspecific treatments of H. fulcrata and H. sessiliflora . H. A. Gleason and A. Cronquist (1991) and Cronquist (1994) acknowledged the variability of the species and the existence of many local races, but lumped most of these into var. villosa and var. hispida of Chrysopsis villosa , in which they also included H. camporum, H. canescens, H. fulcrata, H. pumila, H. stenophylla var. angustifolia, H. viscida , and H. zionensis . Diploid races are usually distinct from each other, but each has given rise to one or more tetraploid lines that are less distinct. The treatment here is based on the detailed presentation in Semple.

Following modified from CalPhotos
   Top | See original


CalPhotos     Photo Database

 

Number of matches : 4
Query: SELECT * FROM img WHERE ready=1 and taxon like "Heterotheca villosa var. villosa%" and (lifeform != "specimen_tag" OR lifeform != "Plant") ORDER BY taxon

Click on the thumbnail to see an enlargement

Heterotheca villosa var. villosa
Heterotheca villosa var. villosa
Golden Aster
ID: 9189 3301 3549 0063 [detail]
Robert Potts
© 2001 California Academy of Sciences

Heterotheca villosa var. villosa
Heterotheca villosa var. villosa
ID: 0000 0000 0506 2119 [detail]
© 2006 Louis-M. Landry

Heterotheca villosa var. villosa
Heterotheca villosa var. villosa
ID: 0000 0000 0506 2120 [detail]
© 2006 Louis-M. Landry

Heterotheca villosa var. villosa
Heterotheca villosa var. villosa
ID: 0000 0000 0506 2121 [detail]
© 2006 Louis-M. Landry

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2016 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT      University of California, Berkeley

Following modified from CalPhotos
   Top | See original


CalPhotos     Photo Database

 

Number of matches : 9
Query: SELECT * FROM img WHERE ready=1 and taxon like "Heterotheca villosa var. pedunculata%" and (lifeform != "specimen_tag" OR lifeform != "Plant") ORDER BY taxon

Click on the thumbnail to see an enlargement

Heterotheca villosa var. pedunculata
Heterotheca villosa var. pedunculata
Hairy Golden Aster
ID: 0000 0000 1112 0665 [detail]
© 2012 Richard Spellenberg

Heterotheca villosa var. pedunculata
Heterotheca villosa var. pedunculata
Hairy Golden Aster
ID: 0000 0000 1112 0666 [detail]
© 2012 Richard Spellenberg

Heterotheca villosa var. pedunculata
Heterotheca villosa var. pedunculata
Hairy Golden Aster
ID: 0000 0000 1112 0667 [detail]
© 2012 Richard Spellenberg

Heterotheca villosa var. pedunculata
Heterotheca villosa var. pedunculata
Hairy False Goldenaster
ID: 0000 0000 1214 0640 [detail]
© 2014 Richard Spellenberg

Heterotheca villosa var. pedunculata
Heterotheca villosa var. pedunculata
Hairy False Goldenaster
ID: 0000 0000 1214 0641 [detail]
© 2014 Richard Spellenberg

Heterotheca villosa var. pedunculata
Heterotheca villosa var. pedunculata
Hairy False Goldenaster
ID: 0000 0000 1214 0642 [detail]
© 2014 Richard Spellenberg

Heterotheca villosa var. pedunculata
Heterotheca villosa var. pedunculata
Hairy False Goldenaster
ID: 0000 0000 1115 1336 [detail]
© 2015 Richard Spellenberg

Heterotheca villosa var. pedunculata
Heterotheca villosa var. pedunculata
Hairy False Goldenaster
ID: 0000 0000 1115 1337 [detail]
© 2015 Richard Spellenberg

Heterotheca villosa var. pedunculata
Heterotheca villosa var. pedunculata
Hairy False Goldenaster
ID: 0000 0000 1115 1547 [detail]
© 2015 Richard Spellenberg

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2016 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT      University of California, Berkeley

Following modified from CalPhotos
   Top | See original


CalPhotos     Photo Database

 

Number of matches : 1
Query: SELECT * FROM img WHERE ready=1 and taxon like "Heterotheca villosa var. nana%" and (lifeform != "specimen_tag" OR lifeform != "Plant") ORDER BY taxon

Click on the thumbnail to see an enlargement

Heterotheca villosa var. nana
Heterotheca villosa var. nana
Hairy False Goldenaster
ID: 0000 0000 1115 1335 [detail]
© 2015 Barry Breckling

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2016 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT      University of California, Berkeley

Following modified from CalPhotos
   Top | See original


CalPhotos     Photo Database

 

Number of matches : 91
Query: SELECT * FROM img WHERE ready=1 and taxon like "Heterotheca villosa%" and (lifeform != "specimen_tag" OR lifeform != "Plant") ORDER BY taxon

Click on the thumbnail to see an enlargement

Heterotheca villosa
Heterotheca villosa
Hairy Goldenaster
ID: 0000 0000 1101 0296 [detail]
© 1978 Lee Dittmann

Heterotheca villosa
Heterotheca villosa
Hairy False Goldaster
ID: 0024 3291 1998 0020 [detail]
Charles Webber
© 2000 California Academy of Sciences

Heterotheca villosa
Heterotheca villosa
Hairy Goldenaster (chrysopsis)
ID: 8253 3202 4147 0032 [detail]
Beatrice F. Howitt
© 1999 California Academy of Sciences

Heterotheca villosa
Heterotheca villosa
Hairy False Goldaster
ID: 0000 0000 0905 0007 [detail]
© 2005 Robert Sivinski

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 1009 0173 [detail]
© 2009 Barry Breckling

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 1010 1188 [detail]
© 2010 Barry Breckling

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 1010 1189 [detail]
© 2010 Barry Breckling

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 1010 1190 [detail]
© 2010 Barry Breckling

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 0912 0742 [detail]
© 2012 Zoya Akulova

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 0912 0743 [detail]
© 2012 Zoya Akulova

Heterotheca villosa
Heterotheca villosa
Golden Aster
ID: 0000 0000 1012 0225 [detail]
© 2012 Jean Pawek

Heterotheca villosa
Heterotheca villosa
Golden Aster
ID: 0000 0000 1012 0226 [detail]
© 2012 Jean Pawek

Heterotheca villosa
Heterotheca villosa
Golden Aster
ID: 0000 0000 1012 1003 [detail]
© 2012 Jean Pawek

Heterotheca villosa
Heterotheca villosa
Golden Aster
ID: 0000 0000 1012 1004 [detail]
© 2012 Jean Pawek

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 1213 1518 [detail]
© 2013 John Doyen

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 1213 1519 [detail]
© 2013 John Doyen

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 1213 1520 [detail]
© 2013 John Doyen

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 0114 2319 [detail]
© 2014 John Doyen

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 0114 2320 [detail]
© 2014 John Doyen

Heterotheca villosa
Heterotheca villosa
Hairy False Goldenaster
ID: 0000 0000 0715 2465 [detail]
© 2015 Barry Breckling

Heterotheca villosa
Heterotheca villosa
Hairy False Goldenaster
ID: 0000 0000 0715 2466 [detail]
© 2015 Barry Breckling

Heterotheca villosa
Heterotheca villosa
Hairy False Goldenaster
ID: 0000 0000 0715 2467 [detail]
© 2015 Barry Breckling

Heterotheca villosa
Heterotheca villosa
Hairy False Goldenaster
ID: 0000 0000 0815 0512 [detail]
© 2015 Barry Breckling

Heterotheca villosa
Heterotheca villosa
ID: 0000 0000 0815 0678 [detail]
© 2015 Zoya Akulova

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2016 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT      University of California, Berkeley

Following modified from CalPhotos
   Top | See original


CalPhotos     Photo Database

 

Number of matches : 20
Query: SELECT * FROM img WHERE ready=1 and taxon like "Heterotheca villosa var. minor%" and (lifeform != "specimen_tag" OR lifeform != "Plant") ORDER BY taxon

Click on the thumbnail to see an enlargement

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Bristly Hairy Goldenaster
ID: 0000 0000 0114 0613 [detail]
© 2014 Dana York

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Bristly Hairy Goldenaster
ID: 0000 0000 0114 0614 [detail]
© 2014 Dana York

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Bristly Hairy Goldenaster
ID: 0000 0000 0114 0616 [detail]
© 2014 Dana York

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Bristly Hairy Goldenaster
ID: 0000 0000 0114 0617 [detail]
© 2014 Dana York

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Bristly Hairy Goldenaster
ID: 0000 0000 0114 0618 [detail]
© 2014 Dana York

Heterotheca villosa var. minor
Heterotheca villosa var. minor
ID: 0000 0000 0615 2972 [detail]
© 2015 Steve Matson

Heterotheca villosa var. minor
Heterotheca villosa var. minor
ID: 0000 0000 0615 2973 [detail]
© 2015 Steve Matson

Heterotheca villosa var. minor
Heterotheca villosa var. minor
ID: 0000 0000 0615 2974 [detail]
© 2015 Steve Matson

Heterotheca villosa var. minor
Heterotheca villosa var. minor
ID: 0000 0000 0615 2975 [detail]
© 2015 Steve Matson

Heterotheca villosa var. minor
Heterotheca villosa var. minor
ID: 0000 0000 0615 2977 [detail]
© 2015 Steve Matson

Heterotheca villosa var. minor
Heterotheca villosa var. minor
ID: 0000 0000 0615 2978 [detail]
© 2015 Steve Matson

Heterotheca villosa var. minor
Heterotheca villosa var. minor
ID: 0000 0000 0615 2979 [detail]
© 2015 Steve Matson

Heterotheca villosa var. minor
Heterotheca villosa var. minor
ID: 0000 0000 0615 2980 [detail]
© 2015 Steve Matson

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Hairy False Goldenaster
ID: 0000 0000 0717 1735 [detail]
© 2017 Steven Perry

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Hairy False Goldenaster
ID: 0000 0000 0717 1736 [detail]
© 2017 Steven Perry

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Hairy False Goldenaster
ID: 0000 0000 0717 1737 [detail]
© 2017 Steven Perry

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Hairy False Goldenaster
ID: 0000 0000 0717 1738 [detail]
© 2017 Steven Perry

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Hairy False Goldenaster
ID: 0000 0000 0717 1739 [detail]
© 2017 Steven Perry

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Hairy False Goldenaster
ID: 0000 0000 0717 1740 [detail]
© 2017 Steven Perry

Heterotheca villosa var. minor
Heterotheca villosa var. minor
Hairy False Goldenaster
ID: 0000 0000 0717 1741 [detail]
© 2017 Steven Perry

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2016 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT      University of California, Berkeley

Following modified from CalPhotos
   Top | See original


CalPhotos     Photo Database

 

Number of matches : 9
Query: SELECT * FROM img WHERE ready=1 and taxon like "Heterotheca villosa var. ballardii%" and (lifeform != "specimen_tag" OR lifeform != "Plant") ORDER BY taxon

Click on the thumbnail to see an enlargement

Heterotheca villosa var. ballardii
Heterotheca villosa var. ballardii
Hairy False-golden-aster
ID: 0000 1111 1234 0742 [detail]
© 2016 Keir Morse

Heterotheca villosa var. ballardii
Heterotheca villosa var. ballardii
Hairy False-golden-aster
ID: 0000 1111 1234 0743 [detail]
© 2016 Keir Morse

Heterotheca villosa var. ballardii
Heterotheca villosa var. ballardii
Hairy False-golden-aster
ID: 0000 1111 1234 0744 [detail]
© 2016 Keir Morse

Heterotheca villosa var. ballardii
Heterotheca villosa var. ballardii
Hairy False-golden-aster
ID: 0000 1111 1234 0745 [detail]
© 2016 Keir Morse

Heterotheca villosa var. ballardii
Heterotheca villosa var. ballardii
Hairy False-golden-aster
ID: 0000 1111 1234 0746 [detail]
© 2016 Keir Morse

Heterotheca villosa var. ballardii
Heterotheca villosa var. ballardii
Hairy False-golden-aster
ID: 0000 1111 1234 0747 [detail]
© 2016 Keir Morse

Heterotheca villosa var. ballardii
Heterotheca villosa var. ballardii
Hairy False-golden-aster
ID: 0000 1111 1234 0748 [detail]
© 2016 Keir Morse

Heterotheca villosa var. ballardii
Heterotheca villosa var. ballardii
Hairy False-golden-aster
ID: 0000 1111 1234 0749 [detail]
© 2016 Keir Morse

Heterotheca villosa var. ballardii
Heterotheca villosa var. ballardii
Hairy False-golden-aster
ID: 0000 1111 1234 0750 [detail]
© 2016 Keir Morse

Using these photos: A variety of organizations and individuals have contributed photographs to CalPhotos. Please follow the usage guidelines provided with each image. Use and copyright information, as well as other details about the photo such as the date and the location, are available by clicking on the [detail] link under the thumbnail. See also: Using the Photos in CalPhotos .   


Copyright © 1995-2016 UC Regents. All rights reserved.

CalPhotos is a project of BSCIT      University of California, Berkeley

Updated: 2017-07-23 21:00:56 gmt
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