9.
Pediocactus despainii
S. L. Welsh & Goodrich, Great Basin Naturalist. 40: 83, fig. 5. 1980.
San Rafael cactus, Despain footcactus, Despain's pincushion cactus
Pediocactella bradyi
(L. D. Benson) Doweld var.
despainii
(S. L. Welsh & Goodrich) Doweld;
Pediocactus bradyi
L. D. Benson subsp.
despainii
(S. L. Welsh & Goodrich) Hochstätter;
P. bradyi
var.
despainii
(S. L. Welsh & Goodrich) Hochstätter
Plants
typically unbranched.
Stems
subglobose to obovoid, 3.8-6 × 3-9.5 cm; areoles circular to oval, villous to glabrate.
Spines
smooth, relatively hard, all radial, 9-15 per areole, spreading, white, 2-6 mm.
Flowers
1.5-2.5 × 1.8-2.5 cm; scales and outer tepals minutely toothed to entire and undulate; outer tepals yellow-bronze to peach-bronze or pink with purple midstripes, 4-10 × 3-6 mm; inner tepals yellow-bronze to peach-bronze (rarely pink), 6-12 × 4-6 mm.
Fruits
green, drying reddish brown, turbinate, 9-11 × 10-12 mm.
Seeds
black, 3.5 × 2.5 mm, shiny, papillate and rugose.
Flowering spring. Desert pavements of cobble or pebble in pinyon-juniper woodlands; of conservation concern; 1500-1800 m; Utah.
Population-level studies of chloroplast DNA (J. M. Porter et al. unpubl.) confirm a close relationship between
Pediocactus despainii
and
P. winkleri
. Genetic relationships among populations correspond to geographic features of the region: populations with the San Rafael Swell represent one group of related populations, and populations south of the San Rafael Swell represent another group of related populations. This pattern conflicts in some ways with morphology. While all of the populations south of the San Rafael Swell are classified as
P. winkleri
, within the Swell some populations are
P. despainii
, several are
P. winkleri
, and a few are morphologically intermediate.
