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Coelioxys rufitarsis Smith, 1854
Coelioxys rufitarsus Smith, 1854, species description; Coelioxys rufitarsis Smith, 1854, index; Coelioxys dubitata Smith, 1854; Coelioxys comstockii Cresson, 1878; Coelioxys coloradensis Cresson, 1878; Coelioxys rufitarsis rhois Cockerell, 1903; Coelioxys rufitarsis claripes Cockerell, 1925

Life   Insecta   Hymenoptera   Apoidea   Megachilidae   Coelioxys
Subgenus: Boreocoelioxys

Coelioxys rufitarsis fem lat comp ps
© Copyright Laurence Packer 2014 · 7
Coelioxys rufitarsis fem lat comp ps

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Coelioxys rufitarsis male lat comp ps
© Copyright Laurence Packer 2014 · 7
Coelioxys rufitarsis male lat comp ps
Coelioxys rufitarsis, female
Nancy Lee Adamson · 1
Coelioxys rufitarsis, female

Coelioxys rufitarsis, female2
Nancy Lee Adamson · 1
Coelioxys rufitarsis, female2
Coelioxys rufitarsis, female3
Nancy Lee Adamson · 1
Coelioxys rufitarsis, female3

Coelioxys rufitarsis, female4
Nancy Lee Adamson · 1
Coelioxys rufitarsis, female4
Coelioxys rufitarsis, female5
Nancy Lee Adamson · 1
Coelioxys rufitarsis, female5

Coelioxys rufitarsis, Barcode of Life Data Systems
Barcode of Life Data Systems · 1
Coelioxys rufitarsis, Barcode of Life Data Systems
Coelioxys rufitarsis, Derrick Ditchburn
Derrick Ditchburn · 1
Coelioxys rufitarsis, Derrick Ditchburn

Coelioxys rufitarsis, Derrick Ditchburnmale
Derrick Ditchburn · 1
Coelioxys rufitarsis, Derrick Ditchburnmale
Coelioxys rufitarsis, s6 acute tip with subapical notches
© USDA Bee Biology and Systematics Laboratory, Logan Utah · 1
Coelioxys rufitarsis, s6 acute tip with subapical notches

Coelioxys rufitarsis, t6 mediolateral emargination with setae
© USDA Bee Biology and Systematics Laboratory, Logan Utah · 1
Coelioxys rufitarsis, t6 mediolateral emargination with setae
Coelioxys rufitarsis, male, t2 fovea, 2
© USDA Bee Biology and Systematics Laboratory, Logan Utah · 1
Coelioxys rufitarsis, male, t2 fovea, 2

Coelioxys rufitarsis, male, t2 fovea
© USDA Bee Biology and Systematics Laboratory, Logan Utah · 1
Coelioxys rufitarsis, male, t2 fovea
Coelioxys rufitarsis, t6 mediolateral emargination with setae
© USDA Bee Biology and Systematics Laboratory, Logan Utah · 1
Coelioxys rufitarsis, t6 mediolateral emargination with setae
Overview
Reprinted with permission from: Mitchell, T.B. 1962 Bees of the Eastern United States. North Carolina Agricultural Experiment Station Technical Bulletin No. 152.


FEMALE—Length 12-18 mm.; black, tegulae more brownish-piceous, tarsal segments bright ferruginous, the tibiae and femora sometimes more or less ferruginous; eyes slightly convergent below; lateral ocelli slightly nearer margin of vertex than to each other, considerably more distant from eyes; cheeks considerably narrower than eyes, carinate posteriorly, angulate below, but this area covered with dense pubescence; clypeus nearly flat, apical margin about straight, with a fringe of rather short but dense pubescence; median length and basal width of labrum about equal; median tooth of mandible low but distinct; punctures deep and quite coarse, close but distinct across vertex, cheeks and face below ocelli coarsely rugose, becoming finely rugose below antennae and on clypeus; pubescence entirely pale on head and thorax, somewhat yellowish on face, copious around antennae and behind anterior ocellus, very short, dense and subappressed at sides of face, laterad of clypeus, thin and very short and obscure over most of clypeus, dense and white on cheeks below, becoming thin above; pubescence rather short but quite copious over pleura, more elongate and whitish on propodeum, very thin and inconspicuous on dorsum of thorax; transverse carina of tubercles rather low but distinct, more or less obscured by pubescence; posterior margin of scutellum slightly out- curved, posterior face not sharply differentiated from dorsal face; axillae acute, projecting conspicuously from sides of scutellum, but length not exceeding median length of scutellum; coxal spines reduced to short but acute tubercles which are densely pubescent posteriorly; wings hyaline basally, becoming very faintly infuscated apically, recurrent veins reaching 2nd submarginal cell subequally distant from base and apex; scutum coarsely, closely and deeply impunctate, punctures slightly separated only in center posteriorly, densely crowded and somewhat finer laterally, but distinctly separated to anterior margin; scutellum coarsely rugoso-punctate or reticulate, the axillae somewhat more finely so; pleura more reticulate than punctate, punctures coarse and closely crowded; lateral faces of propodeum somewhat smoother but very dull, posterior face dull, with closely crowded, minute punctures, dorsal area smooth but velvety, becoming narrowly substriate along upper margin; basal abdominal tergum shining, punctures deep and distinct, well separated medially, becoming close laterally, and very close and fine toward the depressed apical margin, this narrowly white fasciate, but becoming rather broadly so at extreme sides; transverse grooves of terga 2 and 3 sub- median at center, becoming somewhat more basal at extreme sides, punctures quite regular, coarse and close on basal side of groove, the apical side being largely impunctate, but with scattered punctures toward the apical margin, this depressed and densely white fasciate; terga 4 and 5 rather closely punctate toward base, sparsely so toward apical margin, the margins depressed and white fasciate; tergum 6 abruptly narrowed just beyond mid point, resulting in a pair of distinct lateral angles, a median, slightly raised, impunctate line at base, continuous with an elevated ridge toward the tip, surface deeply excavated on each side of this, the tip narrowly rounded, punctures toward the base deep, distinct, rather coarse and close, becoming densely crowded at the lateral angles, the apical part finely rugose (fig. 65); sternum 6 elongate, extending much beyond tip of tergum, narrowed apically, with a pair of distinct, lateral notches, this area much longer than width at the notches, rather narrowly rounded at apex; sterna 1-4 shining, closely, coarsely, deeply and quite regularly punctate, apical margins depressed, yellowish-hyaline, densely white fasciate; sternum 5 triangular in outline, basal portion deeply and quite closely punctate, but the punctures becoming rather abruptly fine and densely crowded just beyond base, apical half dull and densely roughened, the oblique, lateral margins densely, white fasciate.

MALE—Length 11-12 mm.; black, tegulae more brownish-testaceous, tarsi ferruginous, and tibiae usually ferruginous basally and apically; eyes rather strongly convergent below; lateral ocelli subequally distant from margin of vertex and each other, considerably more distant from eyes; cheeks slightly narrower than eyes, carinate posteriorly, quite deeply excavated below, this area shining and bare, relatively smooth, margined both anteriorly and posteriorly by dense pubescence; clypeus nearly flat, apical margin straight, with a pair of small, median tubercles; median length of labrum slightly greater than basal width; median tooth of mandible well developed; punctures coarse and deep, well separated across vertex, becoming closely crowded on cheeks, face below ocelli rather finely rugose, surface below antennae and on clypeus obscured by dense pubescence, very finely rugose beneath; pubescence entirely pale. rather creamy on face, somewhat more elongate and erect around bases of antennae, subappressed and directed upward on supraclypeal area, directed downward on clypeus, cheeks below with dense, white pubescence; pleura rather copiously covered with white pubescence, not forming definite marginal fringes, the pubescence becoming somewhat more yellowish but very thin above, elongate and copious over most of propodeum; transverse carina of tubercles erect and well developed on outer side, not extended along margin of scutum, largely obscured by pubescence; posterior margin of scutellum very slightly outcurved, posterior face not sharply differentiated from dorsal face; axillae rather elongate and acute, conspicuously diverging from sides of scutellum, and slightly exceeding median length of scutellum; coxal spines well developed, somewhat spatulate and elongate, rounded apically, densely covered with white pubescence; wings hyaline basally, becoming slightly infuscated along apical margin, recurrent veins reaching 2nd submarginal cell subequally distant from base and apex; scutum coarsely, deeply and closely punctate, punctures slightly separated only in center of disc, becoming somewhat finer and more densely crowded laterally; scutellum rather coarsely rugoso-punctate or reticulate, the axillae only slightly more finely so; pleura dull, rather finely rugoso-punctate or reticulate; lateral faces of propodeum dull, densely tessellate, posterior face dull, very minutely and closely punctate, punctures rather vague, dorsal area velvety below, becoming finely and irregularly rugose along upper margin; basal abdominal tergum shining, coarsely and deeply punctate, punctures well separated medially, but becoming very close at extreme sides, apical margin abruptly depressed, yellowish-hyaline, densely white fasciate. lateral margins of the dorsal face conspicuously white tomentose; transverse grooves of terga 2 and 3 very deep and complete sub-basal on 2 and subapical on 3, but becoming more nearly basal on each one at extreme sides, punctures on basal side coarse, deep, close and quite regular, apical margin of groove more or less impunctate, medially, becoming more coarsely and closely punctate at each extreme side, apical margins abruptly depressed, yellowish-hyaline, densely white fasciate, tergum 2 with a pair of much elongated and very narrow and deep foveae on each side, just apical to the groove; terga 4 and 5 white fasciate across base, apical margins deeply depressed and fasciate at extreme sides, punctures coarse, deep, distinct, close laterally, somewhat more widely separated medially; tergum 6 fasciate across base, surface densely and irregularly rugoso-punctate, rather finely so at sides, with a deep, median concavity, dorsal protuberances short, somewhat divergent, broadly rounded or subtruncate, the ventral spines more elongate and parallel, narrowly sub- truncate apically, lateral spines rather elongate and slender (fig. 66); sterna 1-4 shining, coarsely, deeply and closely punctate, apical margins abruptly depressed, yellowish-hyaline, densely white fasciate on 2 and 3, fasciate toward sides on 4, disc of 4 not emarginate medially but the depressed margin with a very narrow emargination; sternum 5 largely retracted, apical margin slightly incurved medially, with a rather dense, apical fringe on each side; sterna 6 and 8 entirely retracted, 8 rather narrowly subtruncate apically; penis valves of genital armature fully equal to gonocoxites in length, the gonocoxites somewhat compressed, bearing copious elongate and erect pubescence.

DISTRIBUTION—British Columbia and California to Nova Scotia, the New England states, North Carolina and Georgia; June to October.

FLOWER RECORDS—Asclepias, Baptisia, Helianthus, Melilotus, Rudbeckia, Solidago, Spiraea and Vicia. Robertson (1929) lists the following additional plant genera: Aster, Blephilia, Lobelia, Monarda, Petalostemum and Verbesina.

HOSTS—Megachile latimanus Say and M. melanophoea Smith.
The subspecific designation of rufitarsis is made necessary by the present recognition of two additional subspecies in the western states.

Identification
Theodore B Mitchell. A Subgeneric Revision of the Bees of the Genus Coelioxys of theWestern Hemisphere. Department of Entomology, North Carolina State University. 1973


Female

HEAD: 1. Ratio of median length to maximum width, in facial view - 32:49.

2. Eyes - Length, 33; lateral width, 13; ratio of upper, to maximum, to lower inter¬spaces - 28:31:23. Normally convergent below (A-3); inner margins nearly straight (B-2); orbital hairs short, 1.0 (C-2).

3. Clypeus - Median length, 12; basal width, 11; apical width, 22.Surface slight¬ly convex (A-2); apical margin straight (B-l) and dentate or tuberculate (C-2)med¬ian length equal to basal width (D-l); apical width twice the basal width (E-l).

4. Supraclypeal area - Very slightly con¬vex (A-2); median length less than that of clypeus (B-l) .

5. Parocular areas - Surface below level of antennae quite flat (A-l); maximum width about two-thirds width of clypeus at base (B-3); carina distinct, slightly separated from eye by a narrow groove, and neither elevated below not expanded above (C-l).

6. Subocellar area - Surface slightly convex (A-2) ; densely punctate or rugose (B-l), with a rosette of fine pale hairs radiating from median ocellus laterally and ventrally.


7. Antennal sockets - Distance to eyes, 6.5; to clypeus, 9; to median ocellus, 10; and to each other, 10. Sockets nearer eyes than to clypeus (A-2), to each other (B-2), and to median ocellus (C-2); subequally dis¬tant from clypeus and each other (D-l), and from median ocellus (E-l); and equidistant from median ocellus and each other (F-l).

8. Antennae - Relative lengths of scape, 8; pedicel, 3; flagellar segment one, 5; two, 5.5; six, 5; ten, 7.5; breadth of six, 4. Median flagellar segments longer than their breadth (A-3) ; apical segment slight¬ly shorter than scape (B-2).

9. Vertex - Surface nearly flat (A-2),but ocellar triangle somewhat elevated; occipi¬tal margin strongly incurved (B-2), and not carinate (C-2).


10. Lateral ocelli - Distance to eyes,7.5; to occipital margin, 5; to each other, 6.5. Nearer occipital margin than to eyes (A-2), and to each other (B-2); nearer each other than to eyes (C-2).

11. Genae - Maximum width much exceeded by width of eye in lateral view, 9:13(A-2); narrowed below (B-l) where it is slightly concave (C-l); occipital margin carinate (D-l), forming an obscure, subacute angle with the hypostomal carina.


12. Labrum - Median length equal to basal width (A-l).

13. Mandible - Median tooth nearer apex than to inner tooth (A-2); inner tooth not much larger than median tooth (B-l) ; outer margin curved apically (C-l).

14. Labial palpi - Segment 1 much shorter than segment 2, 10:15 (A-2). MESOSOMA:

15. Scutum - Median length, to breadth, 27:39. Surface evenly convex (A-l), and densely rugose (B-l); median line barely visible (C-2), obscured by the sculpture; parapsidal lines visible, slightly elevated and somewhat shining (D-l).

16. Pronotal tubercle - Carina produced or elevated and conspicuous (A-2), margin somewhat recurved, not extended toward mid¬line.


40
Subgeneric Revision of Coelioxys

17. Mesepisterna - Carina distinct, sharply dividing the anterior and lateral surfaces (A-2).

18. Scutellum - Median length, 9. Dorsal surface densely sculptured (A-l); dorso-median ridge absent (B-2); posterior margin broadly rounded in dorsal view (C-2); edge of margin abrupt or subcarinate, not protub¬erant (D-3).

19. Axillae - Dorsal surface densely sculptured (A-l); apical free process slen¬der and elongate, slightly compressed, acute in dorsal view (B-1).

20. Metanotum - Lateral extensions pos¬terior to base of hind wing strongly depres¬sed beneath carinate margin of scutellar crest (A-l).

21. Metepisterna - Unmodified.

22. Propodeum - Median length of posterior triangle considerably greater than that of median groove below, 9:6 (A-l); lateral mar¬gins of posterior face rounded and indefin¬ite (B-1), dorsal striations short, more or less parallel, but with only two or three rectangular pits toward each end (C-l); an¬terior margin of propodeal spiracle elevat¬ed and thickened, but not carinate (D-2); dorso-lateral triangles quite distinct (if area denuded), margins definitely carinate but quite low, with a dorsal groove just beneath lateral extensions of metanotum (E-3).

23. Fore wings - Length, 102; radial cell 27. Recurrent veins subequally distant from base and apex of second submarginal cell,or the second nearer the apex than first is to base of the cell (A-3).

24. Tegulae - Testaceous (A-2).

25. Front coxae - Anterior surface slight¬ly convex (A-2); tubercles low but distinct (B-5).

26. Mid coxae - Typical.

27. Hind coxae - Anterior surface broadly convex (A-l); surface dull, coarsely or densely punctate (E-l).

28. Trochanters - Front and mid pairs un¬modified; hind pair considerably longer than broad (A-l).

29. Femora - Lower surface of front pair coarsely and closely punctate, and usually well clothed with tomentum (A-l); basal concavity of mid pair deep and distinct (B-1); outer surface of hind pair completely and densely tomentose (C-2).

30. Tibiae - Outer faces of front and mid pairs densely sculptured and dull (A-l) and (B-1); outer faces of hind pair shining be¬tween distinctly separated punctures (C-2); inner hind tibial spur distinctly longer than the outer spur, 8:6 (D-2).

31. Relative lengths of tibiae and tarsi, and basitarsai breadth:


Tibiae Basi- Tarsi Basitarsai 2-5 tarsi breadth
Fore 24 12 15 2.5 Mid 24 10 17 4.0 Hind 31 16 21 5.0 Claws and concavity of strigilis unmodified. METAS0MA: 32. Relative widths of terga 1-6. Tergum one, 50; two, 49; three, 45; four,38; five, 30; six, 21. median length of six, 30.

33. Basal tergum - Dorsal surface strong¬ly convex medially (A-l), downcurved' to mar¬gin of concavity which is neither carinate (B-2) nor fasciate (C-2); ventro-lateral areas rather vaguely subcarinate laterally (D-2). 34 & 35. Terga 2 & 3 - Gradular grooves complete (A-l); postgradular areas exceeded by the pregradular areas in median length (B-3), but much broader laterally (C-l); graduli not fasciate (D-l).

36. Tergum 4 - Gradular line very shallow and obscure, even laterally (A-l), and not fasciate (B-1).

37. Tergum 5 - Gradular line very shallow and obscure, even laterally (A-l).

38. Tergum 6 - Basal and apical areas not sharply divided (A-2); apical area rel¬atively bare (B-2); lateral margins abrupt¬ly and strongly angulate (C-l), deeply sul-cate at angle (D-l), but much narrowed apically; sulcus more or less fasciate (E-l) beneath the angle; dorso-median line or ridge distinct from near base, to apex (F-l); a pair of narrow but deep, lateral excavations present (G-l), separated by the median ridge; apex narrowly rounded (H-2).

39. Sternum 1 - Surface slightly swollen medially, gently declivous to the narrow base (A-l); apical margin broadly but slightly depressed medially (B-1).

40. Sternum 2 - Gradulus distinctly evi¬dent (A-2); apical margin slightly depressed laterally but not medially (B-1).

41 & 42. Sterna 2 & 3 - Graduli indis¬tinct but indicated by a line of division between the chiefly impunctate pregradular areas and the closely punctate postgradular areas, that coincide with apical margin of the preceding sterna (A-3); apical margins slightly depressed laterally, but not medi-ally (B-1).

43. Sternum 5 - Apical margin triangular in form, apex subacute (A-l); basal width about equal to median length of the post¬gradular area.

44. Sternum 6 elongate-triangular, its ventral surface strongly convex (A-l); apex with a pair of lateral, subapical notches (B-2); margins slightly recurved (incurved toward base, outcurved or straight toward apex) (C-4); marginal fringe partially de¬veloped toward base (D-2). See fig.17,p.19.


Male

HEAD: 51. Ratio of median length to max¬imum width, in facial view - 30:47.

52. Eyes - Length, 32; lateral width, 9; ratio of upper, to maximum, to lower inter¬spaces - 28:28:22. Normally convergent be¬low (A-3); inner margins straight (B-1); orbital hairs long, 2.0 (C-4).

53. Clypeus - Median length, 12; basal width, 10; apical width, 21. Surface norm¬ally convex (A-3) ; apical margin straight (B-1), with a pair of small, submedian den¬ticles and a minute one in the center(C-2); median length slightly greater than the basal width (D-2); apical width twice the basal width (E-l).


54. Supraclypeal area - Flat (A-l); slightly shorter than clypeus (B-1).

55. Parocular areas - Surface below level of antennae quite flat (A-l); maximum width about two-Lhirds basal width of clypeus (B-1) ; carina distinct, slightly separated from eye by a narrow groove (C-l).

56. Subocellar area - Surface flat (A-l); densely sculptured (B-1).

57. Antennal sockets - Distance to eyes,5; to clypeus, 8; to median ocellus, 9; and to each other, 10. Sockets much nearer eyes than to clypeus (A-2), to each other (B-2), and to median ocellus (C-2); nearer clypeus than to each other (D-2); subequally distant from clypeus and median ocellus (E-l), and from median ocellus and each other (F-l).


58. Antennae - Relative lengths of scape, 8; pedicel, 2.5flagellar segment one, 4.5; two, 4; six, 4; eleven, 5; breadth of six, 3. Median flagellar segments slightly long¬er than their breadth (A-2); apical segment much shorter than scape (B-1).

59. Vertex - Surface nearly or quite flat (A-3); occipital margin slightly incurved (B-1); not carinate (C-2).

60. Lateral ocelli - Distance to eyes, 9; to occipital margin, 6; to each other, 5. Nearer occipital margin than to eyes (A-2); subequally distant from occipital margin and each other (B-1) ; and nearer each other than to eyes (C-2).

61. Genae - Maximum width about equal to that of eyes (A-l); constricted below (B-1); hypostomal area distinctly concave (C-2); concavity shining and bare in large part (D-l); occipital carina not protuberant below (E-2).

62. Labrum - Basal width equal to median length (A-l).

63. Mandible - Median tooth equidistant from apical and inner teeth (A-l); inner tooth not much larger than median tooth (B-1); outer margin curved apically (C-l).


64. Labial palpi - Segment 1 much shorter than segment 2, 9:13 (A-2). MES0S0MA:

65. Scutum - Median length, to breadth, 25:34. Surface evenly convex (A-l), coarsely and closely punctate (B-1) ; median line barely visible, obscured by the surface sculpture (C-l); parapsidal lines visible, slightly elevated (D-l).

66. Pronotal tubercle - Carina low and inconspicuous (A-l).

67. Mesepisterna - Vertical carina dis¬tinct, sharply dividing the anterior and lateral surfaces (A-2).


68. Scutellum - Median length, 7. Dorsal surface densely sculptured (A-l); dorso-median ridge absent (B-2) ; posterior margin broadly rounded in dorsal view (C-2); the edge rounded, the dorsal and posterior sur¬faces thus merging (D-l).

69. Axillae - Dorsal surface densely sculptured (A-l); free apical processes slender and elongate, slightly compressed, acute in dorsal view (B-1).


70. Metanotum - Lateral extensions pos¬terior to base of hind wing strongly de¬pressed beneath carinate margin of scutellar crest (A-l).

71. Metepisterna - Unmodified.

72. Propodeum - Median length of posterior triangle longer than median groove below,9:6 (A-l); lateral margins of posterior surface rounded and indefinite (B-1), the dorsal striations indistinct medially, becoming definite and parallel laterally, possibly forming quadrangular pits at each end (C-l); anterior margin of spiracle only slightly elevated and not carinate (D-l) ; dorsolat¬eral triangles weakly developed, bounding ridges low and obscure (E-l).

73. Fore wings - Length, 96; radial cell, 27. Recurrent veins subequally distant from base and apex of second submarginal cell (A-l).

74. Tegulae - Testaceous (A-2).

75. Front coxae - Anterior surface nearly flat (A-2); spines robust and erect (B-1).

76. Mid coxae - Typical.

77. Hind coxae - Anterior surface broadly convex (A-l) ; surface dull and densely punctate (B-1).


78. Trochanters - Front and mid pairs un¬modified; lower surface of hind pair convex (A-l).

79. Femora - Lower surface of front pair quite coarsely and closely punctate, and usually well clothed with pubescence (A-l); basal concavity of mid pair absent (B-3); outer surface of hind pair completely and densely tomentose (C-2).

80. Tibiae - Outer faces of front and mid pairs densely sculptured and dull (A-l) and (B-l) ; outer face of hind pair shining be¬tween distinctly separated punctures (C-2); hind tibial spurs nearly equal in length, 7:6 (D-1).

81. Relative lengths of tibiae and tarsi, and basitarsal breadth:

Tibiae Basi-tarsi Tarsi 2-5 Basitarsal breadth Fore Mid Hind 21 21 27 11 10 14 16 20 24 2.5 3.0 4.5 Claws and concavity of strigilis unmodified. METASOMA:

82. Relative widths of terga 1-6. Tergum one,44; two,42; three,39; four,36; five,29; and six,23.

83. Tergum 1 - Midline of dorsal surface strongly convex (A-l); margin of concavity neither carinate (B-2), nor fasciate (C-2); ventro-lateral areas not carinate (D-2).

84. Tergum 2 - Gradular groove complete (A-l); anterior margin of groove distinct but not carinate (B-l); groove not fasciate (C-2); postgradular area shortened medially (D-2), somewhat convex at midline (E-2),well punctured throughout (F-l), with a deep, transversely elongate fovea (G-l); apical margin deeply depressed (H-1), the rim broad and distinct (J-l), with an entire white apical fascia (K-l)» length of hairs compos¬ing the fascia about equal to width of rim.

85. Tergum 3 - Except for the shining and impunctate posterior margin of the gradulus (F-2) and the absence of the foveae (G-2), quite similar to tergum 2, with the same code numbers: (A-l); (B-l); (C-2); (D-2); (E-2); (H-1); (J-l); (K-l).

86. Tergum 4 - Gradular groove complete (A-l); anterior margin of gradulus carinate (B-l) ; gradulus fasciate (C-l); postgradular area rather sparsely punctate medially, be¬coming closer laterally (D-2); apical margin deeply depressed laterally (E-l); the rim broad and distinct (F-l).

87. Tergum 5 - Gradular groove complete (A-l); anterior margin of groove indistinct (B-2); the groove fasciate (C-l); apical margin deeply depressed laterally (D-1); the rim broad and distinct (E-l); latero-apical angles protuberant (F-l).

88. Tergum 6 - Median length of postgrad¬ular area more than half the maximum width (A-2); median excavation of dorsal surface deep, apical, separating the divergent pro¬cesses (B-l), the latter short, flattened, apices rounded (C-2); ventral surface very limited, with no area of separation between the apical margin and the bases of the ven¬tral processes (D-1), the latter slender, spinose and nearly parallel (E-l); lateral
processes triangular and acute (F-3).

89. Tergum 7 - Apical margin narrowly rounded, rim becoming quite broadly depres¬sed laterally (A-l); gradulus straight(B-l).

90. Sternum 1 - Surface slightly swollen medially, gently declivous to the narrow base (A-l); apical margin broadly but slightly depressed medially (B-l).

91 & 92. Sterna 2 & 3 - Gradulus distinctly evident (A-2); apical margin deeply and ab¬ruptly depressed over entire width (B-4).

93. Sternum 4 - Gradulus carinate (A-l); apical margin entire and simple (B-l); the rim strongly depressed and medially emarg-inate (C-3).

94. Sternum 5 - Postgradular area normally retracted, surface lightly sclerotized(A-2); gradulus rather strongly incurved toward base (B-2); apical margin not thickened (C-l); basal margin rather strongly incurv¬ed, thus constricting the pregradular area medially (D-2).

95. Sternum 6 - Gradulus indistinct, but apparently reaching basal margin (A-l); pre¬gradular areas broad medially, narrowed api-cally, not markedly produced at base (B-l); postgradular area broad medially and at base, widely separating lateral elements of the pregradular area (C-l); setose areas composed of dense, more or less oblique patches of setae on each side of midline (D-1); the apical membraneous lip rounded.

96. Sternum 7 - Obsolescent.

97. Sternum 8 - Lateral margins convergent apically (A-l); apex incised (B-3).

98. Genitalia - Gonocoxites equal to penis valves in length (A-l), compressed (B-2) and copiously long setose apically (C-l). SIZE: Female - Breadth of head, 49; scutum, 39; tergum 1, 50; median length of combined scutum and scutellum, 36; fore wing, 95; radial cell, 25.

Male - Breadth of head, 47; scutum, 34; ter¬gum 1, 44; median length of combined scutum and scutellum, 32; fore wing, 100; radial cell, 27.

SCULPTURE: Head densely punctate,coarsely so above, becoming finer below on genae, very fine on clypeus; entire mesosoma more rugose or reticulate, only the legs finely punctate in part, smooth and impunctate on some sur¬faces; punctures of metasoma coarse and close in general, details as indicated in the preceding descriptions.

COLOR: Integument chiefly black, but tegulae and legs variable, chiefly black in some populations or areas, but more often the tarsi and tibiae ferruginous to testaceous, the more basal segments usually but not always black. Fore wings hyaline at base, be¬coming lightly infuscated apically, veins brownish*to testaceous.


Names
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FamilyScientific name @ source (records)
Anacardiaceae  Rhus copallina @ UCMS_ENT (1)
Apiaceae  Daucus carota @ UCMS_ENT (1)
Apocynaceae  Asclepias sp @ UCMS_ENT (2)
Asteraceae  Achillea millefolium @ UCMS_ENT (1)

Aplopappus sp @ BBSL (1)

Aster sp @ UCMS_ENT (3)

Centaurea repens @ BBSL (1)

Chrysothamnus sp @ BBSL (2)

Cichorium [duplicate] @ UCMS_ENT (1)

Cirsium horridulum @ UCMS_ENT (1)

Cirsium neomexicanum @ BBSL (1)

Cirsium undulatum @ AMNH_BEE (2)

Cirsium vulgare @ UCMS_ENT (2)

Ericameria nauseosa @ BBSL (1)

Grindelia sp @ BBSL (3)

Grindelia squarrosa @ BBSL (4); AMNH_BEE (6)

Grindelia @ AMNH_BEE (1)

Gutierrezia sarothrae @ BBSL (2)

Helianthus annuus @ AMNH_BEE (1); BBSL (2)

Helianthus anomalus @ BBSL (2)

Helianthus maximiliani @ AMNH_BEE (1)

Helianthus tuberosus @ AMNH_BEE (2)

Helianthus @ AMNH_BEE (1)

Heliopsis scabra @ AMNH_BEE (2)

Lactuca pulchella @ AMNH_BEE (1)

Solidago tenuifolia @ UCMS_ENT (1)

Solidago @ AMNH_BEE (3)

Symphyotrichum lanceolatum @ AMNH_BEE (1)

Viguiera multiflora @ BBSL (1)
Brassicaceae  Erysimum asperum @ AMNH_BEE (5)

Raphanus raphanistrum @ UCMS_ENT (1)
Capparaceae  Cleome sp @ BBSL (1)
Fabaceae  Amorpha canescens @ AMNH_BEE (2)

Lotus corniculatus @ UCMS_ENT (1); BBSL (2)

Medicago sativa @ BBSL (6)

Melilotus alba @ BBSL (1)

Melilotus officinalis @ CUIC_ENT (4); AMNH_BEE (3)

Vicia cracca @ UCMS_ENT (1)
Grossulariaceae  Ribes sp @ BBSL (3)
Herndon, j.d.  1487 @ JRYA__OLYM (1)
Hydrophyllaceae  Phacelia sp @ BBSL (3)
Hypericaceae  Hypericum perforatum @ CUIC_ENT (1)
Lamiaceae  Caryopteris x @ CUIC_ENT (2)

Leonurus cardiaca @ UCMS_ENT (1)

Lycopus sp @ UCMS_ENT (5)

Marrubium vulgare @ BBSL (1)

Nepeta cataria @ BBSL__ERR (1)

Teucrium canadense @ UCMS_ENT (5)
Liliaceae  Allium cepa @ BBSL (1)

Allium sp @ BBSL (1)
Rosaceae  Rubus sp @ UCMS_ENT (1)
Verbenaceae  Verbena hastata @ AMNH_BEE (1)
_  Withheld @ BBSL (14)

cucurbit @ NLA (2)

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Updated: 2018-11-20 07:59:39 gmt
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